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                                 903979005
[    U.S. ENVIRONMENTAL  PROTECTION AGENCY
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MIDDLE ATLANTIC REGION-III  6th and Walnut Streets, Philadelphia, Pennsylvania 19106

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EPA 903/9-79-005
                                         BIOCHEMICAL  STUDIES



                                               OF THE



                                    POTOMAC  ESTUARY—SUMMER 1978

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EPA 903/9-79-005
                          BIOCHEMICAL STUDIES

                                OF THE

                     POTOMAC ESTUARY—SUMMER 1978
                               May 1979
                          Joseph Lee Slayton
                          E.  Ramona Trovato
                        Annapolis Field  Office
                              Region III
                 U.S.  Environmental  Protection Agency

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                                Table of Contents
                                                                         Page
      Tabulation of Figures  	       ii
      Tabulation of Tables	       iii
  I.  Introduction 	        1
 II.  Conclusions  	        4
III.  Procedures	        6
 IV.  C30D and NOD Kinetics 1n The Potomac Estuary	        8
  V.  Oxygen Demand of Algal  Respiration and Algal  Decay 	       19
 VI.  Phytoplankton Elemental  Analysis/Methods of TKN  	       25
       Digestion of Algal  Samples
VII.  Potomac Long-Term BOD Survey Data	       28
      References	       35

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      Figures
                                                                      Page
 No.
  1.   Study Area	      3
  2.   General  BOD Curve:   Y  *  L0(l-10'kt)	      8
3-4.   River Samples-Oxygen Depletion  Curves   	   10-11
  5.   Plot of  NOD2Q  vs  (TKN  x  4.57)	     15
  6.   STP Effluent Samples-Oxygen  Depletion  Curves  	     17
7-9.   Oxygen Depletion  Curves  of Algal  Respiration  and  Decay  ...   20-22

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     Tables
                                                                          Page
1.  Station Locations 	    2
2.  Thomas Graphical  Determinations of RIQ. L0, and r for River CBOD's  .   12
3.  Thomas Graphical  Determinations of k]g> L0> and r for River NOD's .  .   13
4.  Thomas Graphical  Determinations of k-|Q, L0, and r for STP CBOD's  .  .   16
5.  First Order Correlation Coefficients for STP NOD's  	   18
6.  Phytoplankton Oxygen Depletion  	   23
7.  BODs Requirements for Algal  Decay and Respiration 	   24
8.  Phytoplankton Elemental Analysis  	   26
9.  Results from Three TKN Digestion Methods  	   27

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  I.  Introduction

      During the summer of 1978 an intensive survey of the middle
reach of the Potomac River was undertaken by the  A.P.O. (Table 1,
Figure 1).   As part of this work biochemical  assays were  performed to:
      (1)  determine the carbonaceous  and nitrogenous  oxygen  demand
           rate constants for river and STP effluent samples;
      (2)  establish the relative contributions to the BOD5 of algal
           respiration and the oxygen  utilized in algal decay; and
      (3)  characterize the elemental  composition of the  phytoplankton
           present and establish the relative digestion efficiencies
           of several  methods of algal  TKN determinations.
      The mention of trade names  or commercial  products  in  this  report
is for illustration purposes and  does  not constitute  endorsement or
recommendation by the U.S. Environmental  Protection Agency.

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     Table 1.  Station Locations
Station Number
Station Name
RMI
Buoy Reference
P-8
P-4
1
1-A
2
3
4
5
5-A
6
7
8
8-A
9
10
10-B
11
12
13
14
15
15-A
16
Station Number
S-l
S-2
S-3
S-4
S-5
S-6
S-7
S-8
Chain Bridge
Wndy Run
Key Bridge
Memorial Bridge
14th Street Bridge
Ha ins Point
Bellevue
Woodrow Wilson Bridge
Rosier Bluff
Broad Creek
Ft. Washington
Dogue Creek
Gunston Cove
Chapman Point
Indian Head
Deep Point
Possum Point
Sandy Point
Smith Point
Maryland Point
Nanjemoy Creek
Mathias Point
Rt. 301 Bridge
Treatment Plant Name
Piscataway STP
Arlington STP
Blue Plains STP East &
Alexandria STP
Westgate STP
Hunting Creek STP
Dogue Creek STP
Pohick Creek STP
0.0
1.9
3.4
4.9
5.9
7.6
10.0
12.1
13.6
15.2
18.4
22.3
24.3
26.9
30.6
34.0
38.0
42.5
45.8
52.4
58.6
62.8
67.4



West










C "1"
FLR-23' Bell

C "87"
N "86"
FL "77"
FL "67"
R "64"
FL "59"
N "54"

R "44"
N "40"
N "30"
G "21"
N "10"
C "3"











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Figure 1.   Study Area
                                                Potomac  Estuary

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II.   Conclusions

(1)    The carbonaceous  oxygen  demand  of  the  Potomac  River  samples
      followed  first order  kinetics with an  average  deoxygenation
      constant  ke = 0.12  day "^  and standard deviation  = 0.03 day""*
           = 0.051  day'1).
(2)    The growth kinetics  of river nitrification  were  more  erratic
      but in general  were  first order with an average  ke  =  0.10  day"
      and standard deviation of 0.06.

(3)    The CBOD5 on the average was 58% of the BODs for river samples
      and therefore estimates of CBOD5 from BODs  values are prone to
      error unless a nitrification inhibitor is employed.

(4)    The CBOD of the Potomac STP effluent samples followed first .
      order kinetics with  an average ke =0.16 day'1  and  standard
      deviation of 0.05.
(5)    The NOD for the STP  effluent samples had a  significant lag
      time resulting in poor correlation coefficients for first
      order fit.  This lag time was  probably an artifact of the
      APHA dilution method, since nitrification in the receiving
      waters was immediate.

(6)   The NOD2Q observed for river samples did not significantly
      differ from the product  of  4.57 and the TKM concentration
      (4.57 x TKN).

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(7)    In concentrated algal  samples the average algal  contribution
      to the BOD5 was 0.027  mg BOD5/yg chlorophyll  a_.   The predominant
      species present was the filamentous blue green algae Pseudanabaena,

(8)    Phytoplankton decay represented 70% of the algal BODs and algal
      respiration accounted  for the remaining 30% of the five day
      oxygen depletion.

(9)    The average composition of the phytoplankton present in the
      study area was (mg/vg):
      Org C/Chlor a. = .021 ;  P04/Chlor a. » .002; TKN/Chlor a_ =  .005

(10)   Relative to manual  digestion the Technicon continuous digestor
      and Technicon block digestor recovered respectively an average
      of 58% and 83% of the algal TKN.

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III.  Procedures
  Biochemical Oxygen Demand:  The BOD test is outlined in Standard
     Methods APHA, 14th edition1.  All dissolved oxygen measurements
     were made with a YSI BOD probe #5720 and a YSI model 57 meter.
     The BOD of river water was determined on unaltered samples.  STP
     effluent samples were altered by:  the addition of 1 ml of stale
     settled sewage (seed); sufficient sodium sulfite (NazSOs) to
     dechlorinate the samples; and dilution with APHA dilution water.

  Nitrification:  Formula 2533 nitrification inhibitor (Hach Chemical
     Co.) was dispensed directly into the BOD bottles.  Two bottles
     were filled with each sample—one received the inhibitor and
     represented CBOD and the uninhibited bottle expressed total BOD.
     The NOD was determined by difference2.

  A1 gal BOD Measurements;  The algae in 4 to 10 liters of sample were
     concentrated by continuous centrifugation (Sharpies Continuous
     Centrifuge Model T-l at 12,000 rpm and 1.5-2 liters/min).  The
     pellet was resuspended in 500 ml of collected supernatant.  The
     resultant suspension was diluted in a 300 ml BOD bottle as follows:

         a.  50 ml suspension +  250 ml supernatant
         b.  50 ml suspension (freeze dried) + 250 ml supernatant
         c.  50 ml deionized water + 250 ml supernatant

         a1. 50 ml suspension +  249ml supernatant + 1 ml seed/bottle
         b1. 50 ml suspension (freeze dried) + 249ml supernatant +
             1 ml seed/bottle
         c1. 50 ml deionized water +  249 ml supernatant + 1 ml  seed/bottle

     The sample composite on September 6 consisted of approximately
     2 gallons each from stations:  8; 8A; 9; 10; and 10B.

     The composite of September  14 consisted of about 1/2 gallon each
     from stations:  8; 8A; 9; and 10.  Twenty ml volumes were  used
     instead of the 50 ml volumes indicated above for this  composite.

  Freeze Drying:  Samples were freeze-dried in a Virtis model 10-100
     Unitrap freeze-drier.  The  suspension was spread as a  thin sheet
     and slowly frozen  to avoid  foaming and to shorten drying time.
     Samples required 4 to  6 hours to reach the manufacturer's  specified
     end point.

     The freeze-dried samples were washed  into BOD bottles  with
     supernatant from centrifugation.

  Elemental  Analysis:

     1.  Sample  Preparation:  Samples were stored on  ice and  returned  to
         the laboratory where 4  to  8  liters were  immediately  concentrated
         using a Sharpies T-l Continuous  Centrifuge  at  12,000  rpm  and
         1.5-2.0 liters/min.  Microscopic  examination  revealed  no

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    apparent morphological  damage to the predominant phytoplankton
    species present.  The pellet was resuspended in 250 ml  of
    clear supernatant, which had been collected during centrifugation,
    Aliquots of the suspension and the supernatant were chemically
    analyzed.  The supernatant values were used for blank corrections,

2.  Chlorophyll a:  The photosynthetic pigment from 5-20 ml of
    algal suspension was retained on a 0.45y Millipore filter and
    extracted into 90% acetone with grinding.  The extracted
    solution was centrifuged and measured spectrophotometrically3.

3.  Total Organic Carbon (TOG):  10 ml of algal suspension  was
    diluted to 100 ml  in a  volumetric flask using deionized water.
    A blank was run using 10 ml of supernatant river water
    diluted to 100 ml  in deionized water.  The samples and
    calibration standards were then acidified by the addition of
    1 ml of 6% phosphoric acid to 25 ml and purged free of
    inorganic carbon with oxygen.  The total organic carbon
    was then determined on  a Beckman 915 TOC analyzer**.

4.  Total Phosphate:  5 ml  of sample and blank were diluted to
    25 ml with deionized water.  The sample and blank were
    placed in aluminum foil covered pyrex test tubes to which
    ammonium persulfate and sulfuric acid were added and auto-
    el aved at 15 psi for 30 minutes.  The digests were then
    analyzed for total phosphate by the Technicon automated
    ascorbic acid reduction method .

5.  Algal Nitrogen;  5 ml of sample and blank were diluted  to
    25 ml with deionized water.  The prepared solutions were
    then analyzed for TKN by the following methods:

        A.  Helix:  Samples and blanks were digested by a Technicon
            Continuous Digestor (Helix) and analyzed by the
            automated colorimetric phenolate method4.

        B.  Manual :  Samples and blanks were manually digested
            with 10 ml aliquots placed in reflux tubes and  8.0 ml
            of H2S04/K2S04  digestion solution added.  The tubes
            were placed over flame until boiling and reflux
            stopped.  The contents of the tubes were washed
            into a graduated cylinder with deionized water  and
            brought to 50 ml.  The resultant digests were analyzed
            using a Technicon Continuous Digestor (Helix) and
            the Technicon automated colorimetric phenolate  method1*.

        C.  Block:  Samples and blanks were analyzed by a Technicon
            Block Digestor  BD-40 and analyzed by the salicylate/
            nitroprusside method5.

    The blank carried throughout these methods was used to  correct
    for non-algal nitrogen.

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o
o
CO
IV.  CBOD and NOD Kinetics in the Potomac Estuary

     Biochemical  oxygen demand (BOD)  is  a bioassay in  which  the
oxygen utilization of a complex and changing population  of micro-
organisms is measured as they respire in a changing mixture  of
nutrients.  That portion of the BOD due  to the respiration of organic
matter by heterotrophic organisms is  termed the carbonaceous oxygen
demand and that portion resulting from autotrophic nitrification
is termed nitrogenous oxygen demand.    Nitrification is  the  conversion
of ammonium to nitrate by biological  respiration.  These BOD
components were delineated using an inhibitor to nitrification.   The
inhibitor, formula 2533 of the Hach Chemical Company,  has been shown
                                              2J&.7
to effectively stop the growth of Nitrosomonas   .  The  product consists
of 2-chloro-6 (trichloromethyl) pyridine, known as nitrapyrin, plated
onto an inorganic salt.  The salt serves as a carrier because it  is
soluble in water.  The organic component is not biodegradable, even
after 30 days of BOD incubation, and therefore does not  contribute
to the measured carbonaceous oxygen demand2.
     The shape of the oxygen depletion curves (Figures 2, 3, and  4)
were such that the slope of the curves decreased with increased time
of incubation.
Figure 2:  General BOD  Curve
                                   Curve  Equation:  y = L0(l-10"kt)
                           t  =  elapsed time  of incubation  in the dark at 20°C
                           y  =  BOD; mg/1  oxygen  consumed after time t
                          L0  =  ultimate  BOD;  the  oxygen used in the total
                               degradation  of the  substrate
                           k  =  deoxygenation  constant; a constant which
                               reflects  the  rate  at which  a  substance is
                               oxidized--a  function of temperature, biota
                               and  the nature of the substrate.
                                   Time

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The rate of reaction associated with oxidation-respiration (Ay/At)
was initially rapid corresponding to an initial relatively large
substrate concentration.  This rate decreased with time as the
oxidizable substrate was depleted.  Other nutrients are provided
in excess and do not effect the rate of oxygen consumption in the
standard BOD test.  The quantity and nature of the organic material
in the sample will limit oxygen consumption and determine the rate
of depletion.  This type of reaction, in which the rate is proportional
to the amount of the reactant remaining at any time is referred to
as a "first order" reaction.  In general, the first order reaction
pattern was observed for both the carbonaceous oxygen demand and the
nitrogenous oxygen demand BOD components of Potomac River samples.

     Long-term BOD incubation data were used to give the best available
estimate of k-jQ and L0 using the Thomas Graphical Determination8-'9''10 in
                     1 /^
which a plot of (t/y) '   vs. t yielded a linear relation where
k-jQ = 2.61 x (slope/intercept) and Lo = (2.3 x (intercept)3 x k-|o)  •
The correlation coefficient of the linearized data was taken as a
measure of goodness of fit to first order reaction kinetics.

     The CBOD results for river samples were compiled in Table 2.
The average (n=23) k-|Q value for river CBOD's was 0.051 day"  or
ke = 0.12 day"'' with an average correlation coefficient = 0.98 and
standard deviation = 0.03 (base e).  The value of ke obtained in a 1977
Potomac study6 was 0.14 day"  ,  with  n  =  43  and a  standard
deviation of 0.02.  The ratio of CBODs to BOD5 was found to be 0.58 in
the 1978 study.

     The NOD of the river samples (Table 3) followed first order kinetics
with a correlation coefficient of 0.85 (n=22) and an average ke of 0.10
day""'.  The standard deviation of ke was 0.06.

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Figure 3:   River Samples-Oxygen Depletion Curves
                     Woodrow Wilson Bridge Station 5
                            Sept. 11, 1978
                                                      = .054
                            8      10     12
                              Time (Days)
                        Ft. Washington Station 7
                            Sept. 11, 1978
16
18    20
                             8      10    12    14    16
                              Time (Days)
       IB
                    Total
                     BOD
                                                                         NOD
                                                                        CBOD
                                                                            »•*
                                                                         BOD
                                                                        CBOD,
                                                                         NOD.

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 Figure 4 :   River Samples-Oxygen  Depletion  Curves
 en
 E
   7 -
   6


   5
£ 4 -
 OJ o
 c 3

 c
 0)
 X
o
                          Indian  Head  Station  10
                              August 28,  1978
                              8    10    12
                                Time  (Days)
16    18    20
^ 8

•S

16
4J
O)
 c
 
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Table 2;  Thomas Graphical Determinations of k10, L0, and r for River CBOD's
Date
Aug.







Aug.







Sept.







Sept.







- Sta
14
5
7
8A
10
n
14
16
28
5
7
8A
10
n
14
16
n
5
7
8A
10
n
14
16
25
5
7
8A
10
n
14
16
r

.931
.951
.966
.958
.991
.984
.985

.993
.996
.992
.994
1.000
.990
.996

.994
.990
.987
.989
.940
.981
.997

.999
.996
(.931)
(.231)
(-.231)
(.126)
(.557)
(day'1 )

.045
.059
.038
.057
.067
.062
.089

.046
.040
.039
.033
.029
.027
.056

.059
.054
.044
.044
.041
.054
.069

.079
.049
(.020)
Lag



r:  (correlation coefficient)
n  = 23
Average = .98
Std. deviation = .02 (base  10)
                                  4.5
                                  5.7
                                  6.5
                                  5.2
                                  6.7
                                  3.4
                                  5.8
                                  5.0
                                  5.9
                                  7.9
                                  6.7
                                  5.1
                                  3.5
                                  5.5
                                  5.4
                                  7.2

                                (15.7)
                                            1.8
                                            2.1
                                            2.4
                                            1.7
                                            1.9
                                            0.9
                                            2.8
                                            2.5
                                            2.7
                                            3.1
                                            2.7
                                            1.9
                                            1.6
                                            3.0
                                            3.2
                                            3.1

                                           (3.2)
 3.9
 4.7
 5.4
 4.1
 5.0
 2.4
 5.4
 4.7
 5.4
 6.8
 5.9
 4.3
 3.2
 5.3
 5.3
 6.5

(9.5)
43
43
71
51
60
38
93
4.2
4.9
3.4
2.8
3.2
2.4
3.0
.39
.61
.70
.69
.49
.41
6.4
4.4
4.4
3.9
3.6
7.9
                                                calc.  = Calculated value based upon
                                                        Thomas Graphical determina^io
                      or  ke = .12 day
n  = 23
Average = .051 day"1       c
Std. deviation = .015 day"1 (base  10)

CBOD5/POD5:
n  = 19
Average = .58
Std. deviation = .15
                                      -1

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 *""   Table 3:   Thomas  Graphical  Determinations of k
10'
                                                       -0'
        and r for River NOD's

Date
Aug.








- Sta
14
5
7
8A
10
n
14
16

r

.957
.780
.939
.600
.949
.802
-.441

(da>)

.077
.032
.037
.019
.037
.024
Lag

"-0
(nig/1 )

1.7
4.7
5.5
5.3
3.0
3.6

Calc.*
MOD5
(mg/1 )

1.0
1.4
1.9
1.0
1.0
.8

Calc.
NOP2o
(mg/1 }

1.7
3.6
4.5
3.0
2.4
2.4

Potential**
MOD
(rag/1 )

2.5
2.9
2.8
1.9
2.3
1.3
( .9)
     Aug.  28
5
7
8A
10
11
14
16
.600
.99.5
.978
.996
.989
.876
.877
.017
.067
.039
.037
.048
.049
.030
13.8
5.2
2.9
3.1
3.1
1.9
0.8
2.4
2.8
1.0
1.1-
1.3
1.5
0.2
7.4
5.0
2.4
2.5
2.7
1.6
0.6
     Sept.  11
    Sept. 25
5
7
8A
10
n
14
16
.974
.216
-.276
.658
.727
-.735
.995
.104
Lag
Lag
.022
.023
Lag
.088
    r:   (correlation coefficient)
    n  = 22
    Average =  .86
    Std. deviation  =  .14  (base 10)
                                          6.7
                                          4.0
                                          5.2

                                          1.1
  4.7
   .9
  1.2

  0.7
6.7
2.5
3.4

1.1
                                                                            7.2
                                                                            5.1
                                                                            2.5
                                                                            2.4
                                                                            2.3
                                                                            1.5
                                                                            1.4
5
7
8A
10
11
14
16
.877
.994
.628
.755
.937
-.619
-.381
.049
.098
.028
.023
.039
Lag
Lag
9.1
2.6
4.8
5.0
4.7


3.9
1.7
1.3
1.2
1.7


8.1
2.5
3. A
3.3
3.9


7.0
2.9
2.9
3.1
2.3
(1.4)
(1.4)
 8.3
(5.0)
(4.3)
 3.4
 3.7
(3.5)
 3.3
  * calc.  = calculated
 ** Potential  MOD = 4.57 x TKN
       = 22
    Average =  .045 day""1  or  ke  =  .104  day'1
""*   Std. deviation =  .026 day'1 (base  10)

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The NOD results agreed with previous Potomac demand studies8  in which
the average NOD ke was 0.14 day""1 with a standard deviation of 0.05.

     The larger standard deviation observed for the NOD reflects
both the more fragile nature of nitrification11 and the method by
which it was determined—uninhibited depletion minus inhibited depletion.
The NOD20 was found not to be significantly different from the
potential NOD expressed as 4.57 x TKN (Figure 5).  The critical value
of the paired t-test at a 95% confidence level was 2.08 and the
calculated value was 0.37.  The 4.57 constant is the stoichiometric
conversion factor for the milligrams of oxygen consumed by the  oxidation
of ammonia to nitrate.

     The CBOD kinetics observed for the sewage treatment plant  effluents
were first order with an average ke of  0.16 day"^  (n = 36 and standard
deviation of 0.05).  The average correlation coefficient was 0.985
(Table 4, Figure 6).
     The NOD kinetics observed for the  sewage treatment plants  were
characterized by a lag period  (Figure 6) which resulted in poor
correlation to  first order reaction kinetics  (Table 5).  This lag
time was probably an artifact  of the APHA  dilution method, since
nitrification in the receiving waters was  immediate.  Because the
Potomac waste treatment effluents are characterized by high ammonia
levels6, the initial lack of nitrification is  probably the result  of
 an  insignificant  number of nitrifying  bacteria  in  the  samples  and/or
in the seed  innoculum.  The long term BOD  oxygen depletion data is
included  in Section  VII.

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       Figure 5:  NOD20 (Inhibitor) vs NOD (TKN x 4.57)  River  Water  Samples
    n -


    10-


     9-


     8-
P   ^
in
*i-
x— 6^
*£  en
5-
4-
3-
2-
     1 -
             	v = 19778
             	. = 1978
                                                   1978

                                                 NOD9n vs (TKN) X 4.57
                                                    ^H = 22
                                                     Correlation coefficient = .87
                                                     Least Squares:  Slope = .886;
                                                     y-intercept = .455

                                                 Paired t test
                                                     Degrees of freedom = 21
                                                     t found = .374
                                                     t critical (a = .050;
                                                       a/2 = .025) = 2.080
                   234     56789     10    11
                             NOD20  (Inhibitor)

                                   (mg/1)

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Table 4:  Thomas Graphical Determinations of k-|0, L0, and r for STP CBOD's
Date - Sta
Aug. 14
     S-l
     S-2
     S-3 E
     S-3 W
     S-4
     S-5
     S-6
     S-7
     S-8

Aug. 28
     S-l
     S-2
     S-3 E
     S-3 W
     S-4
     S-5
     S-6
     S-7
     S-8

Sept. 11
     S-l
     S-2
     S-3 E
     S-3 W
     S-4
     S-5
     S-6
     S-7
     S-8

Sept. 25
     S-l
     S-2
     S-3 E
     S-3 W
     S-4
     S-5
     S-6
     S-7
     S-8
                    Name

              Piscataway
              Arlington
              Blue Plains
              Blue Plains
              Alexandria
              Westgate
              Hunting Creek
              Dogue Creek
              Pohick Creek
              Piscataway
              Arlington
              Blue Plains
              Blue Plains
              Alexandria
              Westgate
              Hunting Creek
              Dogue Creek
              Pohick Creek
              Piscataway
              Arlington
              Blue Plains East
              Blue Plains West
              Alexandria
              Westgate
              Hunting Creek
              Dogue Creek
              Pohick Creek
              Piscataway
              Arlington
              Blue Plains
              Blue Plains
              Alexandria
              Westgate
              Hunting Creek
              Dogue Creek
              Pohick Creek





East
West


k




r
1.000
.997
.999
.997
.999
.995
1.000
.993
1.000

Mo
(day'l)
.060
.032
.081
.054
.080
.053
.050
.064
.024

Lo
(mg/1)
12.8
17.3
109.4
21.1
45.9
18.3
29.3
24.7
31.4
Calc.*
CBOD5
(mg/1)
6.4
5.3
66.3
9.7
27.7
8.3
12.9
12.9
7.44
Calc.
CBOD20
(mg/1 )
12.0
13.2
106.7
19.3
44.8
16.7
26.4
23.4
20.8


East
West



-------
         Figure 5:   STP Effluent Samples  -  Oxygen  Depletion  Curves
      50 -
 ?   40 -
 C
 o
 >
 X
' O
30


20


10
                              Piscataway STP  Station  1
                                   August 14,  1978
                                                           r  =  -.744
                                                                   -N-
                                                                              Total
                                                                                BOD
                                             NOD
                       r =  1.0   k10  -  .060
                                  8    10    12    14
                                     Time  (Days)
                                           •CSOD
                                                    16    18    20    22
 c
 o
 cu
 d)
 a
 C51
 >>
 X
 o
90


80


70


60


50


40


30


20


10


 0
                              Westgate STP Station 5
                                September 11, 1978
                                                                                Tota"
                                                                                 BOD
                                                                                 NOD
                                                                                CBOD
i
8
                                        10
                                        12    14
 i
16
18    20
 i
22

-------
Table 5:  First Order Correlation Coefficients for STP NOD's
Sta.           Name
S-l      Piscataway
S-2      Arlington
S-3      Blue Plains East
S-3      Blue Plains West
S-4      Alexandria
S-5      Westgate
S-6      Hunting Creek
S-7      Dogue Creek
S-8      Pohick Creek

r = correlation coefficient
Aug 14
r*
-.744
.060
-.574
-.335
-.597
-.591
-.538
.957
-.722
Aug 28
r
-.863
-.995
-.886
-.892
-.905
-.902
-.582
-.993
-.982
Sept 11
r
-.629
.351
-.642
.972
-.994
-.778
-.594
-.778
-.709
Sept 25
r
-.210
.987
-.816
-.833
-.872
-.619
-.816
-.829
-.619

-------
  V.  Oxygen Demand of Algal Respiration and Algal  Decay

      Potomac BODs samples containing algae historically6'12 expressed
significantly high oxygen demand.  The oxygen demand of such samples
was the result of:  algal respiration; the decay of phytoplankton; and
the carbonaceous and nitrogenous demand of other non-algal  sample
constituents.  To resolve the BOD fractions of the sample,  it was
assumed that algae represented the only significant particulate
contribution to the BOD of the sample.  The non-algal BOD of the
sample was assumed to be associated with the soluble organic and
ammonium/nitrite fractions of the sample.  The non-algal or background BOD was
measured in the supernatant which had been obtained from the
centrifugation of the algae containing samples.  It was further assumed
that the BOD of freeze-dried algae corrected for seed addition and
the BOD of the dilution water (river water supernatant) represented
the biochemical  oxygen demand of algal decay.  Freeze-drying has been
shown to effectively kill phytoplankton without significantly altering
their physical structure13 thus providing a method of separating algal
respiration and algal decay measurements in a BOD analysis.
      The results of these experiments are presented in Figures 7,8,and 9
and Tables 6 and 7.  Algal decay was found to be the major contribution
to algal 8005 with an average mg algal BODg per yg chlorophyll a_ of
0.019.  Algal respiration represented about 30% of the algal BODc
contribution with an average of £.008 mg algal BOD^ per yg chlorophyll a_.
The predominent species present in the Potomac during this  study was the

-------
     Figure 7:  Oxygen  Depletion  Curves of Algal  Respiration and Decay
                                September  14,  1978
   7 4
   6 -
   5 -
I 4
 OJ
 QJ
 c
 »
 X
o
   3 -
   1 -
           River water supernatant used as dilution water
           O  Algal suspension
                                                                              Freeze,
                                                                              Dried
X   Algal  suspension,
     freeze-dried
    River  water
    supernatant
    blk
                                                               066 day'1   L0 = 2.0
                                   r = .909  kin
                                       11        14
                            Time of Incubation (days)
                                                                                 25™
   7 -
   5 -
-  4 -
OJ
   3 -
c
O)
en

52
   1  -
           River water supernatant used as dilution water

           1  ml seed/300 ml  BOD bottle
                                                                             Freeze*
                                                                             Dried
                                                             049 day'1   L0  =  2.3
                                       11        14
                           Time of  Incubation  (days)
                                                                                  w
                                                                                 2C
                                                                                 '•N»

-------
       Figure 8:  Oxygen Depletion Curves of Algal Respiration and Decay
                                  September 6, 1978
                                          1  ml  Seed/BOD Bottle
   20


   18


   16


   14


°  12


   10
 en
 e
 -*->
 (U
 O)
 en
 X
 O
     8 -
6


4
             O  Algal  suspension

             X  Algal  suspension, freeze-dried

             •  River  water supernatant blk
!   I  1
                       i
                       5
                I   1   f  1
                          8
                                           I
                                          12
                                                               =  .989
                                                                                                           Freeze
                                                                                                           Dried
                                                                                                         Supernatant
                                                                                                             Blk
 i
19
                                               Time  of Incubation  (days)

                           I   E   1  I  1   I  I  I   1   !  I   I   !   i   1
i
33
                                                                                        i   i  I  I   I   I

-------
                    ,Ji DlrJt1d.  jUrvl-  if /i^JI Rl.rlrafc. i an! . ica>l
                                    September 6, 1978
                                                                        II   I   i   If  I  i   I   J   f
20



18



16



14
0)
r—
c
(U
   10 -
    8 -
    6


    4


    2


    0
             O Algal suspension

             X Algal suspension,  freeze-dried

             • River water supernatant  blk
                                                                 =  .987
                                                               T
                                          12                   19
                                         Time of Incubation  (days)
                                                                                                         Freeze
                                                                                                         Dried
                                                                                                    33

-------
Table 6:  Phytoplankton Oxygen Depletion
Date/Sample
Days of Incubation
Sept. 6, 1978
Algal Suspension
Algal Suspension
Freeze-Dried
River Water
Supernatant Blk
Seeded Algal
Suspension
Seeded Algal
Suspension
Freeze-Dried
Seeded" River Water
Sept. 14, 1978
Algal Suspension
Algal Suspension
Freeze-Dried
River Water
Supernatant Blk
Seeded Algal
Suspension
Seeded Algal
Suspension
Freeze-Dried
Seeded River Water
5
9.8
6.4
3.0
10.0
9.3
2.8
5
2.4
2.2
1.4
2.6
2.0
1.1
8
12.0
8.5
3.4
12.6
10.8
3.3
11
5.0
3.8
1.2
5.0
3.5
1.4
12
13.8
9.8
3.6
14.4
12.0
3.6
14
5.1
3.6
1.7
5.3
3.2
1.8
19
16.6
11.4
9.3
17.2
14.3
4.4
25
6.7
5.0
1.9
7.0
4.8
2.1
33
19.1
13.1
5.1
19.8
16.1
5.2








-------
Table 7:  BODs Requirements for Algal Decay and Respiration
Decay





Date

Sept. 6
Sept. H
Sept. 6
Sept. H

//
I I BOD5 -
1 ufreeze-
\ dried
algal
suspension)
mg/1
6.4
2.2
9.3
2.0


Background^
BOD5 /



mg/1
3.0
1.4
2.8
1.1

\
DilutionU
factor /
/



6.0
15.0
6.0
15.0


chloro a_




yg/l
1386
810
1386
810
average
Respiration

/

\

Date

Sept. 6
Sept. 14
Sept. 6
Sept. 14

/.
f( BOD5 -
1 algal
t, Vsuspension
\

mg/1
9.8
2.4
10.0
2.6


BODs \X
(freeze-
dried /
algal
suspension)
mg/1
6.4
2.2
9.3
2.0

\
Dilution U
factor /
/



6.0
15.0
6.0
15.0


chloro a_




yg/l
1386
810
1386
810
average
5-Day
Algal Decay
=

mg 62 depletion
yg cm or a_


.0147
.0148
.0281
.0167
.019
5 -Day
Algal Respirati
=

mg 02 depletion
yg cnior a_


.0147
.0037
.0030
.0111
.008
«


•M

""


— '
•w.
K.<
W^
wm
-
fei(
1
W>^


«•

Ml
^.
-
HUH
mg
IBI

-------
filamentous blue-green algae Psuedanabaena. Figures 7,8,and9 also
revealed that seeding of the samples with 1  ml  per bottle of stale
settled sewage1 had little effect upon the amount and rate of oxygen
depletion.  This suggested that the supernatant contained sufficient
microorganisms for algal decay.
  VI. Phytoplankton Elemental Analysis and Methods of TKN Digestion
      of Algal Samples
     The algae bloom of Psuedanabaena occurred in mid to late September
with a chlorophyll a_ peak of 159 yg/1 on September 27.  The elemental
composition of the phytoplankton is compiled in Table 8.  The average
elemental ratios to chlorophyll a_were:  .021 mg C/yg chlorophyll a_;
.0054 mg N/pg chlorophyll a_; and .0020 mg P04/yg chlorophyll a_.  It
should be emphasized that the results are based on the overall
phytoplankton standing corp.  The nitrogen values reported for elemental
analysis were obtained by the automated colorimetric phenolate procedure
employing the continuous  (helix) digestor with preliminary manual
digestion.  Neither the Technicon block digestor nor the Technicon  continuous
digestor alone provided satisfactory digestion of algal TKN.  The data
from side-by-side algal digestions  are compiled  in  Table 9.   The
average recovery relative to preliminary manual digestion for the
Technicon continuous digestor and block digestor were -58%  and 83%
respectively.  This suggested that  42% of algal nitrogen was refractory
to the Technicon continuous  digestor.  This  agreed  with  a  50% TKN  recovery
estimate  suggested  in  a  previous study.lk

-------
i  i      i  i   ii   t   j   11   11   i
Table 8;  phytoplankton Elemental  Analysis
If  II   II   1  i   II   I  i  I   i   11   II   li
Date


Sept. 7
Sept. 11
Sept. 26
Sept. 28

Station
5-A
8-A
9
10
10-B
8-A
9 .
10-B
11
8-A

9

10

10-B

11

7
9
10
10-B
11
rage
. deviation
mg TOC
yg chloro a^
.017
.019
.015
.027
.027
.024

.023

.019

.026

.018

.020

.021

.018
.018
.019
.020
.022
.021
.004
mg TOC
mg TSS
.147
.147
.093
.171
.124
.130

.130

.096

.119

.127

.134

.112






.13
.022
mg TKN
yg chloro a^
.0057
.0052
.0052
.0054
.0065
.0054

.0052

.0058

.0086

.0060

.0060

.0060

.0042
.0043
.0035
.0039
.0044
.0054
.0012
mg TKN
mg TSS
.050
.040
.033
.035
.030
.029

.028

.029

.039

.042

.040

.035






.036
.007
mg P04
yg chloro a_
.0021
.0020
.0021
.0020
.0029
.0023
.0024
.0023
.0023
.0021
.0021
.0026
.0028
.0017
.0016
.0018
.0018
.0020
.0022
.0012
.0012
.0010
.0010
.0014
.0020
.0005
mg P04
mg TSS
.018
.015
.013
.013
.013
.012
.013
.012
.012
.011
.010
.012
.013
.012
.012
.012
.012
.011
.012





.013
.002
mg TSS
yg chloro a_
.115
.130
.158
.156
.220

.185


.201

.218

.142

.148

.189






.169
.035

-------
Table 9:  Results From Three TKN Digestion Methods


Date
Sept.








Sept.



Sept.



Sept.













Station
7 5-A
8-A

9

10

10-B

11 8-A
9
10-B
n
n s
8-A
9
10
26 8-A

9

10

10-B

n

average
std. deviation
Manual
mg/1
TKN
14.52
15.14
15.14
14.89
15.89
15.89
15.89
14.52
15.14
29.27
28.28


23.32
29.05


21.73

25.17

34.66

31.95

26.74



Block
mg/1
TKN
11.10
14.50
13.03
14.47
14.09
13.63
14.06
13.09
14.36
19.49
20.00


-__
—


16.58
17.74
20.63
19.46
30.88
28.02
24.00
26.30
20.60
20.32


Helix
mg/1
TKN
9.15
9.52
9.27
9.52
9.27
9.21
8.81
8.24
8.06
12.92
-12.61


11.83
11.83


13.65

16.86

22.36

22.84

18.53




Helix
Manual
.63
.63
.61
.64
.58
.58
.55
.57
.53
.44
.45


.51
.41


.63

.67

.65

.71

.69

.58
.09

Block
Manual
.76
.95
.86
.97
.89
.86
.88
.90
.95
.67
.71


_..
—


.76
.82
.82
.77
.89
.81
.75
.82
.77
.76
.83
.08

Helix
Block
.82
.66
.71
.66
.66
.68
.63
.63
.56
.66
.63


_-_
___


.82
.77
.82
.87
.72
.80
.95
.87
.90
.91



-------
VII.  Potomac River Long-Terra  BOD  Survey Data - Summer 1978
 Date 8/14/78

 Station
    5
                        Days  of  Incubation
                         10           15
   8-A
   10
   n
   14
   16
T*
C*
N*

T
C
N

T
C
N

T
C
N

T
C
N

T
C
T
C
N
2.4
1.3
1.1

2.7
1.3
1.4

4.3
2.3
2.0

3.9
2.9
1.0

4.6
3.5
1.1

3.5
2.6
0.9

1.8
1.6
0.2
3.0
1.4
1.6

4.4
1.3
3.1

6.3
2.8
3.5

5.3
3.1
2.2

5.8
4.0
1.8

4.7
2.9
1.8

2.0
1.6
0.4
3.4
2.1
1.3
4.9
1.7
3.2

8.0
3.9
4.1

6.8
4.0
2.8

7.0
4.7
2.3

5.6
3.7
1.9

2.4
2.0
0.4
21

3.8
2.2
1.6

5.3
1.9
3.4

8.7
4.4
4.3

7.2
4.4
2.8

7.3
5.0
2.3

6.2
3.8
2.4

2.9
1.8
1.1
 Date 8/28/78

 Station
    5
T
C
N
            Days of Incubation
   7                13               20

   4.3              9.3             10.8
   2.4              3.2              3.9
   1.9              6.1              6.9
   8-A
T              6.2
C              2.7
N              3.5

T              4.4
C              3.1
N              1.3

*T-BOD  (mg/1)
*C-CBOD (mg/1)
*N-NOD  (mg/1)
                    8.0
                    3.8
                    4.2

                    6.4
                    4.3
                    2.1
                        9.4
                        4.7
                        4.7

                        7.7
                        5.4
                        2.3

-------
VII.   Potomac  Rfver  Long-Term BOD Survey Data * Summer 1978 (con't)
 Date 8/28/78 (con't)
                 Days of Incubatfon
             7           13
 Station
   10
   11
   14
   16
 Date  9/11/78

 Station
    5
   8-A
   10
   11
   14
   16
T
C
N

T
C
N

T
C
N

T
C
N
3.6
2.2
1.4

4.2
2.5
1.7

1.4
1.2
0.2

3.8
3.5
0.3
5.2
3.2
2.0

6.1
3.9
2.2

2.7
1.8
0.9

4.9
4.5
0.4
20

6.6
4.1
2.5

7.6
4.9
2.7

3.9
2.4
1.5

5.8
5.2
0.6
                        Hays  of Incubation
                               10        14
                                        21
T
C
N
T
C
N
T
c
N
T
C
N
T
C
N
T
C
N
T
C
N
3.7
1.7
2.0
3.3
1.9
1.4
2.1
2.1
—
2.5
1.9
0.6

___
—
1.2
1.2
0
2.2
2.1
0.1
8.9
2.9
6.0
4.9
3.1
1.8
4.8
3.6
1.2
4.4
2.9
1.5
3.9
2.0
1.9
2.0
1.7
0.3
3.5
3.5
0
9,8
3.5
6.3
6.0
3.9
2.1
7.7
4.6
3.1
6.6
4.2
2.4
6.3
3.2
3.1
2.8
2.3
0.5
4.3
4.2
0.1
11.0
4.0
7.0
6.7
4.5
2.2
9.1
6.2
2.9
7.8
5.0
2.8
7.1
4.1
3.0
3.8
2.7
1.1
5.0
4.6
0.4
12.2
4.6
7.6
7.6
5.4
2.2
9.9
6.7
3.2
8.9
5.9
3.0
8.0
4.0
4.0
4.5
3.2
1.3
5.8
5.0
0.8

-------
VII.  Potomac River Long-Term BOD  Survey  Data  - Summer 1978 (con't)
 Date 9/25/78

 Station
    5
   8-A
   10
   n
   14
   16
T
C
N

T
C
N

T
C
N

T
C
T
C
N

T
C
N

T
C
N
         Days of Incubation
   3              7             14

  6.1            8.5           11.0
  2.3            3.8            4.8
  3.8            4.7            6.2

  2.7            6.2            8.4
  2.1            3.8            5.7
  0.6            2.4            2.7

  2.5            7.1           10.5
  2.1            4.1            7.6
  0.4            3.0            2.9

  2.5            7.6           11.0
  2.0            6.2            9.1
  0.5            1.4            1.9

  2.3            5.7           11.2
  1.5            3.8            8.6
  0.7            1.9            2.6

  0.8            2.0            4.5
  0.7            1.1            2.9
  0.1            0..9            1.6

  1.1            1.6            2.7
  0.6            0.7            1.7
  0.5            0.9            1.0
 Date 8/14/78

 Station
   S-l
T
C
N
20.1
 7.2
12.9
                         Days of Incubation
                          10            15
 38.7
  9.6
 29.1
           41.6
           10.8
           30.8
                21

               43.5
               11.4
               32.1
   S-2
   S-3 (E)
T
C
N

T
C
N
21.0
 6.0
15.0

81.0
75.0
 6.0
 22.8
  9.0
 13.8
157
 88
.5
.5
           41,
           11,
           29,
                                          69.0
174
 96.0
 78.0
               55.5
               13.2
               42.3
181,
 96,
                             85.5

-------
VII.  Potomac STP Long-Term  BOP Survey Data - Summer 1978 (con't)
 Date 8/14/78 (con't)
 Station
   S-3 (W)
   S-4
   S-5
   S-6
   S-7
   S-8
 Date 8/28/78

 Station
   S-l
 S-2
 S-3  (E)
 S-3  (W)
T
C
N

T
C
N

T
C
N

T
C
N

T
C
N

T
r
N
T
C
N

T
C
N

T
C
N

T
C
N
              Days of Incubation
  6            10            15            21

21.6          60.0          73.8          77.4
10.8          15.0          18.0          18.3
10.8          45.0          55.8          59.1

36.0          72.0          87.0          92.3
31.5          36.8          40.5          40.3
 4.5          35.2          46.5          52.0

14.1          41.7          59.4          72.6
 9.6          12.8          14.4          16.5
 4.5          28.9          45.0          56.1

18.6          39.9          51.3          55.8
14.7          20.0          23.6          25.8
 3.9          19.9          27.7          30.0

30.6          44.4          43.5          46.8
15.2          18.0          20.7          22.5
15.4          26.4          22.8          24.3

10.2          38.7          56.4          75.5
 8.7          13.1          17.4          21.2
 1.5          25.6          39.0          54.3

          Days of Incubation
   7              13              20

  9.6            43.7           71.7
  7.8             9.8           10.5
  1.8            33.9           61.2

 12.3            22.8           46.8
  7.8             8.4            8.6
  4.5            14.4           38.2

 28.5            79.5          148.5
 27.0            36.0           36.8
  1.5            43.5          111.7

 24.0            67.5          117.8
 21.0            27.0           28.5
  3.0            40.5           89.3

-------
VII.  Potomac STP Long-Term  BOD Survey Data - Summer 1978 (con't)
 Date 8/28/78 (con't)
 Station
   S-4
   S-5
   S-6
   S-7
   S-8
 Date 9/11/78

 Station
   S-l
   S-2
   S-3  (E)
   S-3
   S-4
                    Days of Incubation
                          13
                                                        20
T
C
N

T
C
N

T
C
N

T
C
N

T
C
N
T
C
N

T
C
N

T
C
N

T
C
N

T
C
N
    42.0
    33.0
     9.0

     9.5
     8.9
     0.6
    19,
    13,
     6,

    25,
    15,
    10,

    n,
    10,
                            0.9
11.4
 7.8
 3.6

28.8
 6.0
22.8

13.5
13.5
13.5
12.0
 1.5

 1.8
15.5
 1.5















Days
6
39.0
10.2
28.8
50.4
8.4
42.0
20.3
20.3
0
22.5
18.0
4.5
27.0
24.0
3.0
87.0
39.8
47.2
22.8
10.4
12.4
42.0
17.7
24.3
41.4
20.1
21.3
22.4
16.1
6.3
132.0
42.8
89.2
47.7
11.7
3F.O
47.9
20.1
27.8
53.6
21.6
36.0
52.4
20.4
32.0
of Incubation
10
52.8
11.4
41.4
68.4
8.4
60.0
34.5
22.5
12.0
49.5
21.0
28.5
46.5
27.0
19.5
14
62.4
13.2
49.2
70.8
8.4
62.4
69.0
24.0
45.0
78.0
22.0
56.0
76.5
27.0
49.5
 21

63.0
15.0
48.0

87.0
10.4
76.6
79,
28,
51.0

90.0
24.0
66.0

99.0
31.0
68.0

-------
VII.   Potomac  STP  Long-Terra  BOD Survey  Data  - Summer 1978  (con't)
 Date 9/11/78

 Station
   S-5
   S-6
   S-7
   S-8
 Date  a/25/78

 Station
   S-l
   S-2
   S-3  (E)
   S-3  (W)
   S-4
   S-5
                    Days of Incubation
                   6        10        14
                                     21
T
C
N
T
C
N
T
C
N
T
C
N
9.0
9.0
0
9.9
9.9
0
9.6
9.0
0.6
7.8
7.8
0
14.4
13.2
1.2
15.0
15.0
0
14.4
13.2
1.2
12.0
10.2
1.8
44.4
16.2
28.2
32.4
17.4
15.0
31.8
16.2
15.6
42.6
14.4
28.2
76.2
16.8
59.4
51.6
18.0
33.6
55.8
18.6
37.2
69.0
16.8
52.2
91.2
18.6
72.6
55.2
21.0
34.2
64.2
22.8
41.4
79.8
21.6
58.2
T
C
N

T
C
N

T
C
N

T
C
N

T
C
N

T
C
N
 7.8
 5.4
 2.4

22.8
 5.4
17.4
31,
19,
12.0

63.0
27.0
36.0

30.0
24.0
 6.0

 9.0
 9.0
  0
Days of Incubation
       7            14

     40.2          49
     13.8          14
     26.4          34.8

     60.0          91.8
     12.6          13.8
     47.4          78.0
     69,
     31,
     37.5

    123.0
     45.0
     78.0
     52,
     31,
     21.0

     15.6
     11.4
      4.2
   .2
   .4
108
 37,
 70,

163.
 60,
103,

m,
 37,
 73.5

 59.4
 13.8
 45.6

-------
VII.  Potomac STP Long-Term BOD Survey Data - Summer 1978 (con't)
 Date 9/25/78 (con't)                Days of Incubation
                              3             7            14
 Station
   S-7
   S-8
T
C
N
T
C
N
11.4
11.4
0
14.4
9.6
4.8
21.0
16.2
4.8
60.0
11.4
48.6
42.0
20.4
21.6
94.8
15.6
79.2

-------
I I
             References
1.  "Standard Methods for The Examination  of Water  and  Wastewater,"
    14th ed.f APHA, 1975.

2.  Slayton, J.L. and Trovato, E.R.,  "Simplified  N.O.D.  Determination,"
    34th Annual  Purdue Industrial  Waste Conference, Purdue University 1979,

3.  Strickland,  J.D.H. and Parsons, T.R.,  "A Manual of  Sea Water
    Analysis," Bulletin 125, Fisheries  Research Board of Canada,
    Ottowa, 1960, p.  185.

4.  Environmental Protection Agency,  Methods for  Chemical Analysis
    of Water and Wastes. 1974.

5.  Gales, M.E., "Evaluation of The Technicon Block Digester  System
    for Total Kjeldahl Nitrogen and Total  Phosphorus,"   EP4-600/4-78-015,
    Feb. 1978,  Environmental Monitoring Series.  E.P.A.  Cincinnati,
    Ohio.

6.  Young, J.C., "Chemical Methods for  Nitrification Control,"
    24th Industrial Waste Conference, Part II Purdue University,
    pp. 1090-1102, 1967.

7.  Young, J.C., "Chemical Methods for  Nitrification Control,"
    J.W.P.C.F..  45, 4, pp. 637-646 (April  1973).

8.  Slayton, J.L. and Trovato, E.R.,  "Carbonaceous  and  Nitrogenous
    Demand Studies of The Potomac  Estuary, AFO Region III, Environmental
    Protection Agency, 1977.

9.  Thomas, H.A., "Graphical Determination of B.O.D. Curve Constants,"
    Water and Sewage Works, p. 123-124, (March 1950).

10. Moore, W.E.  and Thomas, H.A.,  "Simplified Methods for Analysis of
    B.O.D. Data," Sewage and Industrial Works. 22,  p. 1343-1355,  1950.

11. Finstein, M.S., et al, "Distribution of Autotrophic  Nitrifying
    Bacteria in  a Polluted Stream," The State Univ., New Brunswick,
    N.J., Water  Resources Res. Inst.  W7406834, Feb. 1974.

12. Clark, L.J.  and Roesch, S.E.,  "Assessment of  1977 Water Quality
    Conditions In The Upper Potomac Estuary, E.P.A. 903/9-78-008,
    July 1978.

13. Fitzgerald,  G.P., "The Effect  of  Algae on B.O.D. Measurements,"
    J.W.P.C.F.,  Dec.  1964, pp. 1524-1542.

14. Slayton, J.L. and Trovato, E.R.,  "Algal  Nutrient Studies  of the
    Potomac Estuary", AFO Region III, Environmental  Protection
    Agency, 1977.
                                           35

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