EPA-540/1-86-004
                                            Office of Emergency and
                                            Remedial Response
                                            Washington DC 20460
Off'ce of Research and Development
Office of Health and Environmental
Assessment
Environmental Criteria and
Assessment Office
Cincinnati OH 45268
                      Superfund
vvEPA
                       HEALTH EFFECTS ASSESSMENT

                       FOR POLYCHLORINATED BIPHENYLS

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                                           EPA/540/1-86-004
                                           September  1984
       HEALTH EFFECTS  ASSESSMENT
 FOR  POLYCHLORINATED BIPHENYLS  (PCBS)
    U.S. Environmental  Protection Agency
     Office of Research and  Development
Office of  Health and Environmental Assessment
Environmental Criteria  and Assessment Office
            Cincinnati, OH  45268
    U.S. Environmental  Protection Agency
  Office of  Emergency and Remedial Response
Office of Solid Waste and  Emergency Response
            Washington, DC  20460
                   U.S. Envirerrrx-'^ ^ lection Agency
                   Region '•/, U'-   •

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                                       DISCLAIMER

         This  report  has  been  funded  wholly or  In  part  by  the United  States
     Environmental  Protection  Agency under  Contract  No.  68-03-3112  to  Syracuse
     Research  Corporation.   It has been subject  to  the Agency's  peer  and adminis-
     trative review,  and H has been approved  for  publication as an EPA document.
     Mention of  trade  names  or commercial  products  does not  constitute  endorse-
     ment or recommendation for  use.
i) r« r
     nvironrnent?.! Pr©t©etion  Agency
                                            11

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                                    PREFACE
    This report  summarizes  and evaluates Information relevant  to  a prelimi-
nary   Interim   assessment   of   adverse  health   effects  associated   with
polychlorlnated   blphenyls.    All  estimates   of   acceptable   Intakes   and
carcinogenic  potency presented  1n  this document  should  be  considered  as
preliminary  and   reflect   limited  resources   allocated   to   this  project.
Pertinent  toxlcologlc  and  environmental  data  were  located  through  on-line
literature  searches  of  the  Chemical  Abstracts, TOXLINE,  CANCERLINE  and  the
CHEMFATE/DATALOG  data bases.   The basic  literature  searched  supporting  this
document is current  up  to September,  1984.   Secondary  sources of Information
have also  been relied upon  1n  the  preparation of this report  and represent
large-scale health  assessment  efforts that  entail extensive  peer  and Agency
review.  The  following  Office  of Health  and  Environmental  Assessment (OHEA)
sources have been extensively utilized:


    U.S. EPA.   1980b.   Ambient  Water Quality  Criteria for Polychlorl-
    nated  Blphenyls.   Environmental   Criteria  and  Assessment  Office,
    Cincinnati, OH.  EPA 440/5-80-068.  NTIS PB 81-117798.

    U.S. EPA.   1985.  Drinking Water  Criteria  Document for Polychlorl-
    nated  Blphenyls.   Prepared  by   the  Environmental   Criteria  and
    Assessment Office,  Cincinnati,  OH, OHEA for the Office of  Drinking
    Water,  Washington, DC.  Final Draft.

    The Intent in  these assessments 1s  to suggest  acceptable  exposure levels
whenever sufficient  data  were  available.  Values  were not  derived or larger
uncertainty factors  were employed  when  the  variable data were  limited  in
scope tending  to  generate conservative (I.e.,  protective) estimates.   Never-
theless, the Interim values  presented reflect  the relative degree of hazard
associated with exposure or  risk to the chemlcal(s) addressed.

    Whenever possible, two categories  of  values have  been estimated for  sys-
temic toxicants (toxicants for which  cancer  is  not the  endpolnt of concern).
The  first,  the AIS  or  acceptable  Intake subchronlc,  is  an  estimate of  an
exposure  level that  would   not  be expected  to cause  adverse  effects  when
exposure occurs  during  a limited time interval (I.e.,  for an  Interval  that
does not  constitute a  significant  portion  of  the lifespan).   This  type  of
exposure estimate  has  not  been  extensively used  or  rigorously defined,  as
previous  risk  assessment  efforts   have  been  primarily  directed  towards
exposures from toxicants  in  ambient air  or  water  where  lifetime exposure  is
assumed.   Animal  data  used for  AIS   estimates generally  include exposures
with durations  of 30-90 days.    Subchronlc  human  data are  rarely  available.
Reported exposures are  usually  from chronic  occupational  exposure  situations
or from reports of acute accidental  exposure.
                                      111

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    The AIC,  acceptable  Intake  chronic,  1s  similar In  concept  to  the  ADI
(acceptable  dally  Intake).   It  1s  an estimate  of an  exposure  level  that
would  not  be expected  to cause  adverse effects  when  exposure occurs  for  a
significant portion  of  the llfespan  [see U.S.  EPA  (1980b)  for a  discussion
of  this  concept].   The  AIC  1s  route specific  and  estimates   acceptable
exposure  for  a  given  route with  the  Implicit  assumption that exposure  by
other routes 1s Insignificant.

    Composite  scores  (CSs)  for  noncardnogens  have  also  been  calculated
where  data  permitted.   These  values  are used for  ranking reportable quanti-
ties; the methodology for their development  1s explained  1n U.S. EPA (1983).

    For compounds for which there  1s  sufficient evidence of carclnogenlclty,
AIS  and AIC  values  are  not  derived.  For  a  discussion  of  risk  assessment
methodology  for  carcinogens refer  to U.S.  EPA (1980a).  Since cancer  1s  a
process that  Is  not characterized by a threshold,  any  exposure  contributes
an  Increment  of  risk.   Consequently,  derivation of  AIS  and  AIC values would
be  Inappropriate.   For carcinogens,  q-j*s  have  been computed based  on oral
and Inhalation data  1f available.
                                       1v

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                                   ABSTRACT


    In order  to  place the risk  assessment 1n proper context,  the  reader  Is
referred to the  preface  of  this document.  The  preface  outlines  limitations
applicable to all documents of  this  series  as  well  as  the appropriate Inter-
pretation and use of the quantitative estimates.

    Limited human  data  are available  suggesting a relationship  between  PCB
exposure and  cancer.   Jji vitro mutagenldty evaluations  have  been  primarily
negative.  PCBs  have been  shown to be  carcinogenic 1n  rats  and  mice  when
administered orally.  Using data  for hepatocellular  carcinoma  and neoplastlc
nodules  In  female  rats,  a   carcinogenic  potency  (qi*J  of  4.34  (mg/kg/
day)"1 has  been  estimated for  oral  exposure  of  humans  to PCBs.   Data  were
Inadequate to develop a quantitative estimate for the Inhalation route.

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                               ACKNOWLEDGEMENTS


    The  Initial  draft  of  this  report  was  prepared by  Syracuse  Research
Corporation under  Contract  No.  68-03-3112  for EPA's  Environmental  Criteria
and  Assessment  Office,  Cincinnati,  OH.   Dr. Christopher  DeRosa and  Karen
Blackburn were the Technical Project Monitors  and  Helen  Ball  was the Project
Officer.  The final documents  1n this  series  were  prepared  for  the Office of
Emergency and Remedial Response,  Washington, DC.

    Scientists from  the  following  U.S. EPA offices  provided  review comments
for this document series:

         Environmental Criteria and Assessment Office, Cincinnati, OH
         Carcinogen Assessment Group
         Office of Air Quality Planning and Standards
         Office of Solid Waste
         Office of Toxic Substances
         Office of Drinking Water

Editorial review for the document series was provided by:

    Judith Olsen and Erma Durden
    Environmental Criteria and Assessment Office
    Cincinnati, OH

Technical support services for the document series was provided by:

    Bette Zwayer, Pat Daunt, Karen Mann and Jacky Bohanon
    Environmental Criteria and Assessment Office
    Cincinnati, OH
                                       v1

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                              TABLE OF CONTENTS

1.
2.


3.










4.




5.
6.






ENVIRONMENTAL CHEMISTRY AND FATE. . . .' 	
ABSORPTION FACTORS IN HUMANS AND EXPERIMENTAL ANIMALS . . .
2.1. ORAL 	
2.2. INHALATION 	
TOXICITY IN HUMANS AND EXPERIMENTAL ANIMALS 	
3.1. SUBCHRONIC 	
3.1.1. Oral 	
3.1.2. Inhalation 	
3.2. CHRONIC 	
3.2.1. Oral 	
3.3. TERATOGENICITY AND OTHER REPRODUCTIVE EFFECTS. . . .
3.3.1. Oral 	
3.3.2. Inhalation 	
3.4. TOXICANT INTERACTIONS 	
3.5. HUMAN EFFECTS 	
CARCINOGENICITY 	
4.1. HUMAN DATA 	
4.2. BIOASSAYS 	
4.3. MUTAGENICITY 	
4.4. WEIGHT OF EVIDENCE 	
REGULATORY STANDARDS AND CRITERIA 	
RISK ASSESSMENT 	
6.1. ACCEPTABLE INTAKE SUBCHRONIC (AIS) 	
6.2. ACCEPTABLE INTAKE CHRONIC (AIC) 	
6.3. CARCINOGENIC POTENCY (q]*) 	
6.3.1. Oral 	
6.3.2. Inhalation 	
Page
1
... 5
... 5
... 7
8
... 8
... 8
... 22
, . . 22
... 22
... 24
... 24
... 28
... 28
... 28
... 31
... 31
... 31
... 36
... 36
, , . 38
... 40
... 40
... 40
... 40
... 40
... 40
 7.  REFERENCES	   42
APPENDIX: Summary Table for Polychorlnated Blphenyls 	   55

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LIST OF TABLES
No.
1-1
1-2
2-1
3-1
3-2
3-3
3-4
4-1
5-1
6-1
Title
Approximate Composition of the Commercially Available
Aroclors 	
Identification Numbers and Selected Physical and Chemical
Properties of Aroclors 	
Uptake of PCBs by Rats 	
Effects of Subchronlc and Chronic Oral Exposure of PCBs
to Rats and Mice 	
Effects of Subchronlc and Chronic Oral Exposure of PCBs
to Other Species 	
Effects of Chronic and Subchronlc Exposure of PCBs to
Monkeys 	
Summary: Teratogenetlc, Fetotoxlc and Reproductive Effects
of Orally Administered PCBs 	
Summary of Carcinogenic Effects of Orally Administered PCBs. .
FDA Regulations for PCBs 	
Data Used as the Basis for the CM* 	
Paqe
2
3
6
g
12
13
25
32
39
41
     V111

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                             LIST  OF  ABBREVIATIONS

ADI                     Acceptable dally Intake
AIC                     Acceptable Intake chronic
AIS                     Acceptable Intake subchronlc
BCF                     B1oconcentrat1on factor
bw                      Body weight
CAS                     Chemical Abstract Service
CS                      Composite score
DNA                     DeoxyMbonuclelc add
PEL                     Frank-effect level
LOAEL                   Lowest-observed-adverse-effect level
LOEL                    Lowest-observed-effect level
mol.wt.                 Molecular weight
NOEL                    No-observed-effect level
PCBs                    Polychlorlnated blphenyls
PCDFs                   Polychlorlnated dlbenzofurans
ppm                     Parts per million
RNA                     R1bonucle1c add
SGOT                    Serum glutamlc oxalacetlc transamlnase
SGPT                    Serum glutamlc pyruvlc transamlnase
STEL                    Short-term exposure limit
TLV                     Threshold limit value
TWA                     Time-weighted average
UV                      Ultraviolet
                                      1x

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                     1.  ENVIRONMENTAL CHEMISTRY AND FATE
    PCBs  consist  of   a  mixture  of  chlorinated  blphenyls  that  contain  a
variable  number  of substituted  chlorine  atoms  on  the aromatic  rings.   The
commercial  PCBs  manufactured  In the  United  States  are  known as  Aroclors
followed  by a 4-d1g1t  number.   The  first  two  digits  Indicate the  type  of
mixture  (e.g.,   those  with  "12" as  the first  two  digits  are  chlorinated
blphenyls;  those with  "25" are  blends  of  75% blphenyls and  25% tMphenyls;
and  those with  "54"  are  chlorinated  trlphenyls), and  the  last  two digits
represent- the  approximate weight percent  of  chlorine  1n  the  product.   The
approximate  compositions  of  the commercially available  Aroclors and  Kane-
chlors are  shown 1n Table  1-1.   The relevant physical  and  chemical  proper-
ties and Identification numbers  of the Aroclors are presented 1n Table 1-2.
    The mobility of  different Aroclors  1n  soils as  1t  pertains  to  leaching
has been  studied by  Pal  et al.   (1980).  The  two most Important factors that
determine  the  Teachability of Aroclors  In  soils are  the  characteristics  of
the  soils  and the  nature  of  the  Aroclors.   For   example,  as  the  chlorine
content   of  an   Aroclor  Increases,   1t    may  Increase  the   adsorption
characteristics  1n   soils,  thereby   decreasing   Us  rate   of   leaching.
Similarly,  soils with  more organic  matter  content may  tend to Increase  the
adsorption  and  decrease  the  leaching  rate.   Therefore, maximum  leaching  of
Aroclors  Is  expected  In  sandy soils and with  Aroclors  having  lower  chlorine
content,  such  as  Aroclor  1221  and  Aroclor  1232.   Incorporation  of  1  ppm
dlchloroblphenyl  1n  a  sandy  loam  soil showed  that  no detectable  amounts
(<0.001 ppm) of  the compound  leached  beyond  30 cm  soil layer after 1  year  of
Incorporation (Pal  et al., 1980).
    B1oconcentrat1on factors have not been determined for  all  the Individual
Aroclors.   The potential  for  bloaccumulatlon  of Aroclors  Is  related  to  the
                                    -1-

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                                                                   TABLE 1-1
                                        Approximate  Compositions  of  the  Commercially Available Aroclors*
Aroclor
Nolecular
F omul a
C12H10
C12H,C1
C,2H8C12
C12H7C13
C12H6C14
C12H5C15
C12H4C16
C12H3C17
C12H2C18
C12HC19
Average MO).
No. of
Chlorine
Atoms
0
1
2
3
4
S
6
7
8
9
wt.
No. of
Isomers
1
3
12
24
42
46
42
24
12
3

mol. wt.
154
169
223
258
292
326
361
395
430
464

wt.
XC1
0
18.8
31.8
41.3
48.6
54.3
58.9
62.8
66
68.7

1221
11
51
32
4
2
<0.5
ND
ND
ND
NO
201
1232
<0.1
31
24
28
12
4
<0.1
ND
ND
ND
232
1242
<0.1
1
16
49
25
8
1
<0.1
ND
ND
267
1248
ND
ND
2
18
40
36
4
ND
ND
ND
300
1254
<0.1
<0.1
0.5
1.0
21
48
23
6
ND
ND
328
1260
ND
ND
ND
NO
1
12
38
41
8
ND
376
1016
<0.1
1.0
20
57
21
1
<0.1
ND
ND
ND
258
Kanechlor
300
ND
ND
17
60
23
0.6
ND
ND
ND
ND
NA
400
ND
ND
3
33
44
16
5
ND
ND
ND
NA
500
ND
ND
ND
5
27
55
13
ND
ND
ND
NA
'Source: Callahan et al.. 1979; IARC.  1978
ND . Not detected; NA . not available

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                                                                   TABLE  1-2
                               Identification Numbers  and Selected  Physical and  Chemical  Properties of Aroclors3
Identification Numbers








i
CO
Properties
Vapor pressure
im Hg at 2S'C
Mater solubility
mg/l
Log octanol/water
partition coefficient
CAS Registry
Number

1221 1232
[6.7xl(T»] [4.06xlO~»]

O.S9b at NA
24'C
[2.8] [3.2]
4.09 >4.S4
11104-28-2 11141-16-5


1016
I4xl0~«]

0.42 at
25'C
4.38
>5.58
12674-11-2


1242
4.0bxlO'«

0.24C at
25*C
4.11
>5.58
53469-21-9


1248
4.94x10'*

0.054 at
25*C
[5.75]
>6.11
12672-29-6


1254
7.71xlO"»

0.012e at
25-C
[6.03]

11097-69-1


1260
4.05xlO"»

0.0027C at
25*C
[7.14]
>6.11
11096-82-5


aA11 values are taken from Callahan et  al..  1979, except where noted; the bracketed data are estimated.
bVerschueren. 1983
cMackay and Lelnonen. 1975
NA = Not available

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number  of  chlorines,  with  the  BCF  value  Increasing with  higher  chlorine
content (Callahan  et al.,  1979).   The BCF  value  1n freshwater species may
vary  from  3000  for  the  muscle  of a brook  trout,  Salvellnus fontlnalls.  to
274,000 for  the  whole body of  a  fathead minnow, Plmephales promelas.  (U.S.
EPA, 1980b).
    The half-life of  PCBs  In  air  1s not known.  Although photolysis  of  PCBs
In  the  atmosphere  1s  a  likely  process,  1t  1s expected to  be  slow  (Pal  et
al.,  1980).   The removal  of  PCBs  from the atmosphere  by  physical  removal
processes,  such  as  wet and dry deposition,  may not be  significant  (CupHt,
1980).  The  half-life of  PCBs  as  a  result  of chemical reactions  with  OH
radicals and  O3  1n  air has been  estimated to  be >8  days  (CupHt,  1980).
    The half-life of a PCB In  aqueous  media will  depend on both  the  nature
of  the PCB  and  the  characteristics   of  the  body  of  water.    PCBs  with
Increasing chlorine  content, because of  their  lower  volatility  and blodegra-
datlon  and  higher  sorptlon characteristics,  will  have longer  half-lives.
Based on published reports, PCBs containing  less than three  chlorines  may  be
degraded 1n  aquatic media  (Callahan et al.,  1979;  Pal et  al.,  1980), al-
though no  estimation  of  half-lives  1n  natural bodies of water  1s  available.
PCBs with  four chlorines  appear to  be  somewhat degradable,  while  those  with
five or more  appear to be recalcitrant.
    The half-life of PCBs In soils  Is  expected to  follow the same pattern  as
1n  water.   Both  volatility and blodegradatlon  may  destroy PCBs  containing
one  or  two chlorine atoms; the half-lives  of  such  PCBs may be a  few days.
PCBs  containing  three or four  chlorine atoms  will  have longer  half-lives,
whereas PCBs  containing  five  or more  chlorine atoms may be  refractory  (Pal
et al., 1980).
                                    -4-

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           2.   ABSORPTION FACTORS  IN  HUMANS  AND  EXPERIMENTAL  ANIMALS
2.1.   ORAL
    One of  the problems associated with understanding  the  toxlcoklnetlcs  of
PCS products  1s  that  they are mixtures of  many different  Isomers,  each with
Us own  characteristic  kinetics of  behavior  1n the animal  body.   In  one  of
the early studies of oral absorption,  Albro and Flshbeln (1972)  administered
5,  50 or  100 mg  of 19 "pure"  PCB  Isomers and  the  totally  unchlorlnated
blphenyl  by  gavage  to  male  CD   rats  (number  not  reported).   Feces  were
collected for  4  days;  the total amount excreted 1n  the feces  and the  amount
absorbed  or  metabolized  1n  the gut  were  determined.   The  results  are
presented  1n  Table  2-1.  For  all  compounds  tested at  all dosage  levels,
absorption was >90%.  The authors recognized that  this  study  did not  Inves-
tigate the  possibility  of Intestinal  metabolism, enterohepatlc redrculatlon
or the effects of dietary status  on  retention  of PCBs,  all of  which would  be
required for full elucidation of the PCB absorption process.
    Gavage administration of 1.5  or  3.0 g Aroclor  1248/kg  bw to  adult  rhesus
monkeys resulted  1n  uptake of  >90%  of  the  total dose  {Allen  et  al.,  1974),
confirming  the data  of  Albro and  Flshbeln  (1972).   These  Investigators sub-
sequently administered  a single  18 mg  dose of  2,5,2',5'-tetra PCB/kg  bw  to
seven adult male rhesus  monkeys.   Excreta were  collected and analyzed  for  14
days.   Over  12% of  the  total  dose  was  recovered  unmodified  1n  the  feces
(Allen  et  al.,  1975).   Norback   et  al.   (1978)  administered one  dose  of
9H-labeled  2,4,5,2',4',6'-hexa  PCB  to two  young monkeys with bile  duct  and
duodenal  cannulae.   The  authors  reported  that 59.3-87%  of  the  total  dose
passed unmodified  through the  Intestinal tract within the first  week.   It
was unclear why relatively little  of this  Isomer was absorbed.
                                    -5-

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                                  TABLE 2-1
                           Uptake of PCBs by Rats3
                                      Percentage  Retained*3
Compound
Blpheny"!
2-C1
3-C1
4-C1
2,6-Cl2
2,2'-Cl2
2,4-Clp
2,3'-Cl2
2,A'-C12
3,4'-Cl2
4,4'-Cl2
2,5,2'-Cl3
2,3,5'-Cl3
3,4,3'-CL3
2,4,4'-Cl3
2,5,2',5'-Cl4
2,3,4,5-Cl4
3,4,3',4'-Cl4
2,4,5,2',5'-Cl5
2,4,5,2',4',5'-C16
5 mg/kg
99.5
98.9
98.0
97.3
NR
95.0
NR
NR
97.2
95.7
95.4
NR
NR
NR
NR
96.0
97.3
98.8
95.1
95.3
1 S.O.
0.11
0.45
1.01
1.73
NR
2.42
NR
NR
3.38
3.45
3.05
NR
NR
NR
NR
1.75
0.86
0.57
0.57
1.11
50 mg/kg
98.8
98.0
96.6
96.3
95.0
97.4
95.7
96.8
95.5
94.3
95.6
95.1
95.6
93.5
92.6
NR
NR
NR
NR
NR
i S.D.
0.25
0.30
0.10
0.40
0.96
0.20
0.40
0.51
1.20
0.86
1.12
3.00
1.05
2.85
0.86
NR
NR
NR
NR
NR
100 mg/kg
98.2
98.4
97.3
96.4
94.9
98.2
97.1
97.9
96.6
97.1
97.4
94.3
95.0
90.3
94.0
NR
NR
NR
NR
NR
± S.D.
0.30
0.15
0.40
0.42
0.75
2.10
0.35
0.28
0.78
0.63
1.10
3.18
1.12
3.58
1.50
NR
NR
NR
NR
NR
aSource: Albro and F1shbe1n, 1972
bŁach value 1s the average of three determinations on each of two rats:
                       (amount fed) - (amount excreted)
 Percentage Retained
x 100%
                                  amount fed
NR = Not reported; S.D. = standard deviation
                                    -6-

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2.2.   INHALATION
    One brief  report  concerning Inhalation  absorption  of PCBs was  located.
Benthe et al.  (1972)  exposed  male  Wlstar rats once  to  Pydranll A200 (number
of  rats,  concentration  of  Pydranll  A200 and  length  of  exposure  time  not
specified).   Animals  were killed  and examined  at  15  minutes,  2 hours,  24
hours and 2  days  post-exposure.  At the end of  15 minutes,  liver  concentra-
tions  were   >50%  of  the  maximum   concentrations  attained  after  2  hours.
During the  first  24 hours, rapid  Increase  1n brain and  fat levels of  PCBs
was noted.  At the end of 2 days,  liver  and  brain  levels  of  PCBs  had fallen,
and  levels  1n adipose  tissue had  reached  a maximum.  Indicating very  good
absorption;  however, absorption factors were  not  quantHated.
                                    -7-

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                3.  TOXICITY IN HUMANS AND EXPERIMENTAL ANIMALS
3.1.   SUBCHRONIC
3.1.1.   Oral.   Subchronlc  oral   exposure  of  experimental  animals  to  PCBs
has been Investigated 1n detail In several species, and  PCB  toxlclty In rats
appears to have  been  studied  most extensively.  The details  of  protocol  and
results of these studies  are summarized In  Tables  3-1,  3-2 and 3-3.   In  an
early  study,  Klmbrough  et al.  (1972)  compared  the  effects  of  Aroclor  1254
and Aroclor  1260 on  groups of 10  male  and  10 female Sherman  rats.   Aroclor
1254 was  Included 1n the  diet at 0,  20,  100 or  500 ppm  (mg/kg/dlet),  and
Aroclor  1260 was  Included  1n  the  diet at 0,  20,  100,  500  or  1000  ppm
(mg/kg/d1et).  The duration of exposure was  8  months.  Aroclor  1260  appeared
to  be  more  toxic to female  than  to  male rats,  since none of the male rats
fed Aroclor  1260-contalnlng diets  died,  but  1/10, 2/10  and 8/10 females  fed
diets  containing 100,  500  and 1000  ppm Aroclor  1260,  respectively,  died.
Aroclor 1254  at  dietary  levels  of 500 ppm resulted 1n the death of  two male
rats and one  female rat.
    Dietary  levels >500  ppm  Aroclor  1260  or  Aroclor  1254 seemed to  reduce
rate of weight  gain  severely  1n  treated rats, although  statistical  analysis
was not  performed to substantiate that  observation.   There was, however,  a
statistically significant  Increase 1n  relative   liver  weight (liver  weight
expressed as  percent  of  body weight) 1n all rats at dietary  levels  >20  ppm
of  either  of  the Aroclors  tested,  except  for  females exposed  to  20  ppm
Aroclor 1260  1n the  diet.  Because the  diets  with 20 ppm of  either Aroclor
1260  or  Aroclor  1254  were  set   up  at  a  different  time  from the  diets
containing other levels,  they  had their  own simultaneous  female   and  male
control groups.   These control  females  had   unusually high  liver weights  (no
                                    -8-

-------
                                                                          TABLE  3-1

                                          Effects of Subchrontc and Chronic Oral Exposure of PCBs to Rats and Nice
Species/
Strain Sex/No.
Rat/F344 H.F/191
Rat/ F/400
Sherman
Rat/Wtstar N/290
Rat/ N/6/group
Sprague-
Oawley
Rat/ N/96
Sprague-
Dawley
Source
of PCB Vehicle
Aroclor 1254 diet
Aroclor 1?60 diet
Kanechlor-300. diet
-400 or -500
Aroclor 1242 diet
Ar odors 1248, diet
1254 or 1262
Dosage
Schedule
0. 25. SO. 100
mg/kg diet for 2
years
0 or 100 mg/kg
diet for 21
months
0. 500 or 1000
mg/kg diet for
27-52 weeks
0. 5 or 25 «g/kg
diet for 2. 4 or
6 months
0 or 100 mg/kg
diet for 52 weeks
Duration
of Study
2 years
21 months
1 year
2, 4 or
6 months
65 weeks
Animal Effects
Reduced body weight. Stomach: •Intestinal"
metaplasia, dose-related; adenocarclnoma of
glandular stomach.
No effect on food Intake; reduced body weight.
Elevated tan liver modules (170/164), hepatocellu-
lar carcinoma (26/184). Other areas: hepatic
disruption.
Heavy mortality. Hepatomegaly, oval cell and bile
duct proliferation, fatty liver Infiltration.
Cholangloftbrosls at 1000 mg/kg level of all 3
Kanechlors, nodular hyperplasla. Depressed final
body weight.
Elevated hepatic mlcrosomal enzyme activity, llpld
content. Elevated urinary coproporphyrln levels.
Present after 2 months at 5 mg/kg.
Increased hepatic protein. RNA and llpld; decreased
DMA. Increased Mlcrosomal total protein and cyto-
chrome P-450. Induced N-demethylase nltroreductase.
Inhibited glucose-6-phosphatase.
Reference
Morgan
et a!..
1981
Klmbrough
et al..
1975
Ito et al.,
1974
Bruckner
et al..
1974
Allen and
Abrahamson,
1979
Rat/CD
f/300
Aroclor 1254     diet
Rat/Donryu  N/15. F/15
                Kanechlor-400    diet
0. 10. 30 or 100
mg/kg diet for up
to 20 weeks
                             total  Intake  450-
                             1500 mg  over  159-
                             560 days
20 weeks    Serum cholesterol, beta globulin Increased;  gamma     Zlnkl, 1977
            globlln decreased (dose-related).   >30 mg/kg:
            Reduced rate of gain,  hepatomegaly, cardtomegaly
            (dose-related). >10 mg/kg:   Hepatic porphyrlnlc
            fluorescence.  >10 mg/kg:   Erythema, crustiness,
            hyperkeratosts,~perlkeratosts on ears, dorsum of
            nose and feet, tall.

560 days    All treated rats:  fatty liver degeneration.          Klmura and
            Females 1200-1500 mg:   multiple adenomatous            Saba. 1973
            nodules.  All rats >700 mg:  hepatomegaly.  Lung
            and Intracranlal abscesses  suggested Impaired
            resistance to Infection.

-------
                                                                           TABLE 3-1 (cont.)
     Species/
     Strain      Sex/No.
                   Source
                   of PCB
                 Vehicle
Dosage
Schedule
Duration
of Study
                                                                             Animal Effects
Reference
     Rat/
     Sprague-
     Dawley
f/6/group
                                 Aroclor 1242
                                 diet
     Rat/
     Sherman
o
i
N/10. F/10      Aroclor 1254
In each group
                 N/10. f/10      Aroclor 1260
                 In each group
                                                  diet
                                 diet
     Rat/
     Sprague-
     Dawley
N/96
Aroclors 1248.   diet
12S4 or 1262
     Hlce/dd
H/114
Kanechlor-300.
-400. or -SOO
                                                  diet
                            0.  75 or  150 mg/kg
                            diet for  8 or
                            36  weeks
                             0.  20, 100. SOO
                             mg/kg diet for
                             8 months
                             0.  20.  100. 500.
                             1000 mg/kg diet
                             for 8 months
0 or 100 mg/kg
diet for 52 weeks
0. 100. 250. or
500 mg/kg diet
for 32 weeks
36 weeks    Both levels:  massive venous engorgement of liver     Jonsson
            with characteristic darkening; marked focal           et  al..
            necrosis and regeneration, enlarged hepatocytes;       1981
            many mitoses and multlnucleate cells, accumulation
            of pigment adjacent to veins, heaviest In Kupfer
            cells; accumulation of llpld droplets In cytoplasm.
            some with areas suggestive of llpld-cholesterol
            complexes; marked smooth ER proliferation; deposits
            of Iron; granular degeneration of mitochondria; many
            hepatocytes contained whorl-like membranous bodies.

8 months    Mortality (3/20) and reduced rate of gain at 500       Klmbrough
            mg/kg. Hepatomegaly, enlarged hepatocytes with        et  al..  1972
            foamy cytoplasm-containing Inclusions at >?0 mg/
            kg. Adenoflbrosls and pigment accumulation at
            >100 mg/kg.

8 months    Mortality 1/10, 2/10. 8/10 of females In 100. 500.
            1000 mg/kg groups. Decreased rate of gain at >500
            mg/kg. Hepatomegaly. MiF at >20 mg/kg. discolored
            livers with UV fluorescence, enlarged hepatocytes
            with foamy cytoplasm-containing Inclusions.
            Increased llpld content at >100 mg/kg. Pigment
            accumulations at 500 mg/kg. Adenoflbrosls at >100
            mg/kg.

65 weeks    Normal appetites, appearance, weight gain.            Allen
            Hb. PCV. UBC. serum protein. A/G ratios. Elevated     et  al..  1976
            serum total llplds. cholesterol. Total llpld and
            trlglycerld spiked very high peaks on Aroclor 1254
            (only) at 52 weeks. Cholesterol levels persisted
            at 65 weeks (13 weeks off exposure). Trlglycerlde
            levels fell 
-------
                                                                         TABLE  3-1  (cont.)
Species/
Strain Sex/No.
Nice/ H/25/group
BALB/C
Source Dosage
of PCB Vehicle Schedule
Aroclor 1221. diet 0. 3.7S. 37.5. or
1242 or 1254 37S mg/kg diet
for b Months
Duration
of Study Animal Effects
9 months 375 mg/kg Aroclor 1242 and all Aroclor 1254-
exposed groups: significant (p<0.01) Increase
In liver weight after 6 months. Significant
(p<0.01) decrease In liver weight of 375 mg/kg
Aroclor 1242 and 37.5 mg/kg Aroclor 1254-exposed
groups after 3-month recovery period. Aroclor 1254:
375 mg/kg: mortality and severe hepatopathology;
37.5 mg/kg: mild hepatopathology; 3.75 mg/kg:
Aroclor 1242: 375 mg/kg: moderate
hepatopathology.
Aroclor 1221: no liver lesions.
Reference
Roller. 1977
i,  Mice/Swiss  f/63
->  Albino
i
    HIce/BALB/  N/200
    CJ
    HIce/ddN     F/60
Aroclor 1254


Aroclor 1254



•PCB'
diet
diet
olive or
rice bran
oil
200 mg/kg diet
for 23 weeks

0 or 300 mg/kg
diet for 6 or 11
months

0. 0.5% V/V PCB
In olive oil, or
1600 mg/kg PCB In
bran oil for 13,
17. 22 or 26 weeks
23 weeks    Thickening and erylhemla of pinna of ear;              Bell. 1983
            changes In mlcrovasculature.

11 months   Both 11* and 6-month exposure:   Hepatomegaly,          Klmbrough
            hepatomas, liver degeneration and elevated            and Under,
            porphyrln.                                            1974

26 weeks    Both exposed groups:  slight weight loss,  reduced     Nlshliumt,
            activity; eczematous and ulccrattve skin              1970
            lesions, hepatomegaly and hepatopathology.
     A/6  B Albumin/globulin; Eft - endoplasmlc retlculum; Hb » hemoglobin;  PCV - packed cell volume

-------
                                                                              TABLE  3-?

                                              Effects of Subchronlc and Chronic Oral Exposure of PCBs to Other Species
Species/
Strain
Mink/
Pastel

Sex/
Number
H.F/105

Source of PCBs
Aroclor 1242
Aroclor 1016
Vehicle
diet
diet
Dosage Schedule
0, 5, 10. 20 or 40
mg/kg diet
0 or 20 mg/kg diet
Duration
of Study
B nonths

AnlMl Effects
Mortality of all
mg/kg diet.
Death of 3/12 (2!
•Ink on >20
iX) of females.
References
Bleavlns
et al.. 1980

I
~J
ro
          Ntnk/
          NR
F/7 or
8/group
Aroclor 1016,
1221, 1242 or
1254
diet
0 or 2 mg/kg diet
•10 Months
Necropsy:  emaciation, complete
absence of body fat. gastric
ulceratlon.  Aroclor 1?4? at 5
or 10 ng/kg diet:  complete re-
productive failure.  Aroclor
1016 at 20 ng/kg diet:  reduced
reproductive performances.

Aroclor 1254:  Interference with
reproduction.  All Aroclors tested:
no significant differences In body
weight gains, hemoglobin, PCV.
                                                                                         Aulerlch  and
                                                                                         Ringer. 1977
           NR  -  Not  reported; PCV . packed cell volume

-------
                                                                             TABLE 3-3

                                                   Effects of Chrontc and Subcnrontc Exposure of PCBs  to Monkeys
Strain Sex/No.
Rhesus F/24
M/NR
Source
of PCBs Vehicle Dosage
Aroclor 1240 diet 0. 2.5. or S.O
mg/kg diet.
-IB Months
Duration
of Study
-39.6
Months
AnlMal Effects
Hales (S.O Mg/kg level only): moderate erythema
and perlorbltal edema. Females: more severe skin
lesions (alopecia, acne); extreme weight loss.
Irregular menstrual cycle length, depressed serum
progesterone. Considerable Improvement after 1-
year recovery period.
Reference
Barsottl
and Allen,
1975
   Rhesus    F/30

             M/10
Aroclor 1248
Aroclor 1248
diet
diet
0. 2.5 or 5.0
mg/kg diet
0 or 5.0 mg/kg
diet. -16
months
-16 months;
total Intake
90 or 180 mg/
kg by females
In 6 months
CO
I
              F/8/group      Aroclor 1248     diet
              F/24
    Rhesus     NR/7
Aroclor 1016
Aroclor 124B
                            Aroclor 1248
 diet
trans-
placental
or
mother's
Milk
                trans-
                placental
                or
                mother's
                milk
0.5 or 1.0 mg/kg
diet 3 times
weekly for
-16.6 months

0.025. 0.25 or
1.0 mg/kg diet
for unreported
length of time

mothers exposed
to PCBs 6 months
before gesta-
tion, through
gestation and 3
to 4 months of
nursing

mothers removed
from exposure to
PCBs for 22-84
weeks before
conception
-16.6 months,
total Intake
~8 or 16 mg
after 7 months

NR
up to Infant
age of 24
months
                                                                  Skin lesions as above; 15X weight loss  In
                                                                  females.  Normal hematograms.  After 6 months:
                                                                  serum  total llplds reduced, shift In A/G ratio.
                                                                  elevated SGPT.  Menstrual cycles lengthened.
                                                                  Serum  progesterone and estradlol reduced.
                                                                  After  12 months, serum cholesterol and  trlgly-
                                                                  cerlde reduced.

                                                                  No  Irregularities In menstrual cycle, or serum
                                                                  estradlol, progesterone or reproduction success.
                                                                  Infants smaller, skin hyperplgmented.
No abnormalities of clinical, gross or reproduc-
tive parameters.  1.0 mg/kg:  Infants had reduced
birth weight.
Significantly Increased locomotor behavior
(hyperacttvlty).  Significantly retarded learn-
ing ability.
                                                   Mothers exposed to >2.S mg/kg:  hyperactlvtty.
                                                                                                       Barsottl.
                                                                                                       et al.. 1976
                                                                                                       Allen
                                                                                                       et al.. 1979a
                                                                                                       Barsottl,
                                                                                                       1981
Bowman and
Hetrontmus,
1981; Bowman
et al., 1981

-------
                                                                      TABLE 3-3 (cent.)
Source Duration
Strain Sex/No. of PCBs Vehicle Dosage of Study
Rhesus H/3/group 3,4.3'.4'-TCB diet 3 mg/kg reduced Up to 215
to 1 ng/kg reduced days
to 0.3 mg/kg

2,5.2',5'-TCB elevated to 1 mg/
kg
f
Animal Effects
Mortality of all three by day 21S In 3. 4.3'. 4'-
TCB-exposed groups: emaciation, skin lesions,
nail bed hyperplasla, loss of nails, thymis
atrophy, gastric lesions as described (Allen,
197S). 2.5.2'.5'-TCB: no signs of toxlctty.
no gross or hlstologlc lesions.

Reference
NcNulty
et al., 1980




          N/4-S/group   3.4.3',4'-TCB    diet
                        or
                        2.5.2'.5'-KB

                        Aroclor 1242
Rhesus    H/6
Aroclor 1242     diet
Cynomol-  F/4
gus
Cynomol•  f/7
gus
                        Aroclor 1254
P-KC-400*
Y-PCB>>
                        PY-PCBC

                        control
corn oil,
gelatin.
apple
juice

olive oil
In banana
             1 mg/kg for 38
             days or 133 days.
             then control diet

             1 mg/kg for 133
             days 5 mg/kg
             additional 2 months

             0,  3, 10. 10. 30
             or  100 mg/kg diet
                                                  -190
                                                  days
                      Up to 245
                      days
                             0,  100.  100  or
                             400 mg/kg bw/day.
                             3 days/week
5 mg/monkey/day
5 or 10 mg/
monkey/day

5 mg/monkey/day

NA

all above treat-
ments given 6
days/week
                      Up to 238
                      days
                                                                           20 weeks
                                      1  death:   necropsy findings  as  above.   Others:
                                      squamous  metaplasm of  sebaceous glands.
                                                                  No evidence of  toxlctty.
All PCB-exposed monkeys:  palpebral swelling,
erythemla; weight loss, rough hair coat, reduced
lib. leukocytosls.  Mortality of 4/6 by day 245.
Gastric lesions:  hyper trophic gastric mucosa con-
sisting of elongated hyperplastlc glands, destruc-
tion of parietal and zymogenlc cells.  Only spe-
cific region along greater curvature affected.

Lost fingernails, fetal toxlctty.  Substantially
reduced antibody production to SRBC antigen.
Death of 10 mg V-PCB-dosed monkeys by 8 weeks.
Height loss of P-KC-400 and 5 mg Y-PCB-dosed
monkeys.  V-PCB-dosed:  alopecia, acne, hyper-
pigmentation, perlorbltal edema.  All treatments:
reduced antibody productton to SRBC.  Htstopath-
ology Y-PCB:  enlarged hepatocytes with enlarged
smooth ER, focal necrosis, bile duct proliferation.
Dilated renal tubules with casts, epithelial
vacuoles.  Melbomlan cysts, skin hyperkeratosls.
Hlstopathology P-KC-400 and PY-PCB:  lesions In
liver, kidney as above, but more mild.  Perlorbl-
tal skin:  no lesions.
Becker
et al., 1979
Truelove
et al., 1982
Horl et al..
1982
aP-KC-400 -  Kanechlor-400 with PCDfs removed.
t>Y-PCB was prepared from Kanechlor 400. contained =400 ppm PCOfs.
fpY-PCB   >  Y-PCB with PCDFs removed.
A/G  » Albumin/globulin; Hb « Hemoglobin; NR » Not recorded; ER -  Endoplasmlc  retlculum NA
                                                                    Not available;  SRBC -  Sheep red blood cells

-------
explanation was  given)  which  undoubtedly accounted for the apparent  lack  of
statistical significance 1n relative  liver weights  of  the  females  exposed  to
20 ppm Aroclor 1260.
    At levels >20  ppm,  exposure to Aroclor 1254  or  1260  resulted  In  hepato-
cellular   enlargement   with   foamy  cytoplasm   that   contained  Inclusions.
Livers,  particularly  those  from  rats  exposed  to  Aroclor  1260,  were  dis-
colored;  many fluoresced  under  UV  light,  Indicating porphyrla.   Accumula-
tions  of  pigment were  noticed  In Hvcrs  from rats  exposed  to >100  ppm  of
Aroclor  1254  or 1260.   L1p1d  accumulation  and  extensive  foci of  adenofl-
brosls were also noted  1n  livers  from rats exposed  to 100  ppm Aroclor 1260.
Ultrastructurally,   an   Increase   of   smooth   endoplasmlc   retlculum,  cyto-
plasmlc  Inclusion   of   I1p1d-conta1n1ng  vacuoles  and  atypical  mitochondria
were  noted.   Structures of  concentrically arranged  membranous whorls  sur-
rounding  I1p1d-conta1n1ng  vacuoles were also  seen.   The most  striking  dif-
ference 1n  livers  from rats  exposed  to  these  two  Aroclors  was considerably
greater  adenof1bros1s   (synonyms:   cholanglof1bros1s, bile duct  prolifera-
tion,  flfaroadenoma)  In  livers  from  Aroclor   1254-exposed  rats.  Under  the
conditions of this study,  20  ppm  of  Aroclor  1254 or 1260 appeared  to produce
effects  such  as  hepatomegaly  and  hepatocytlc  enlargement  because   of  the
expanded  smooth  endoplasmlc  retlculum,   which  may   be  considered  adverse
effects.    Therefore,  20 ppm  Aroclor  1260  or  1254  was considered  a  PEL  for
this study.
    Subsequently, Bruckner  et al.  (1974) exposed  groups  of  six male  Sprague-
Dawley rats to 0, 5 or  25  ppm (mg/kg/d1et) Aroclor 1242  for 2, 4 or  6 months
to  evaluate   the  results  of  prolonged  exposure  to  relatively low  levels.
These  authors observed  no differences  1n  either body  weight gain  or  food
consumption  resulting   from   Aroclor   1242  exposure.   The  high-dose  group
                                    -15-

-------
(25  ppm)  suffered  a  slight  but significantly  reduced  hematocrH  after  2
months of exposure,  and both treatment  groups  suffered slight but  signifi-
cantly reduced hemoglobin levels.  These appeared  to  be temporary  phenomena,
since both  hematocrH  and  hemoglobin values  recovered  quickly for  the  dur-
ation of the study.
    MUrosomal enzyme  assays  Indicated  a  significant  (p<0.05) Induction  of
mixed-function oxldase activity after 2 months of  exposure  to  25  ppm Aroclor
1242  1n  the diet, and  after  4  months of  exposure to  5  ppm.   N-demethylase
activity was  significantly  (p<0.001)  Induced  by  25 ppm  Aroclor 1242 after  4
months  of  exposure.   Urinary coproporphyrln  levels  were  elevated after  2
months of exposure  to  25 ppm Aroclor 1242 In the  diet  (p<0.001) and after  6
months of exposure  to  5 ppm  (p<0.05).  Liver weight  expressed  as  percent of
body weight was  elevated only at the 25 ppm  dietary  level  (p<0.02)  and  only
after 4  months of  exposure.   Liver  llpld expressed  as mg/g wet  weight  was
elevated (p<0.02) after  only  2  months of exposure, but after  6 months,  only
livers from rats exposed to 25 ppm showed elevated (p<0.02)  I1p1d  content.
    Upon hlstopathologlcal  examination, no hepatic  lesions  were found 1n any
rat  after  2,   4  or  6 months  of  exposure.  Sudan  IV staining, however,  re-
vealed Increased  liver  I1p1d  1n  all  rats  exposed  to Aroclor 1242  for 2,  4 or
6  months.   Hlstopathologlcal  examination of  the  kidney  revealed  a  slight
vacuollzatlon  of  convoluted  tubules  only  after  exposure  to 25 ppm Aroclor
1242  1n  the diet  for  4  or  6 months.   In this study, 1t appeared that dietary
exposure of rats to  5  ppm Aroclor  1242 for  2,  4 or  6 months resulted 1n
Induction of  mlcrosomal hydroxylase activity, Increase 1n  urinary copropor-
phyrln  and  Increase  In liver  I1p1d  content.   These  changes are  probably
reversible and, over a  short  term,  do not appear  to threaten life or health,
since neither  body weight  gain  nor  food consumption was affected.   In  this
study, 5 ppm Aroclor 1242 for 2, 4 or 6 months appeared  to be a LOAEL.

                                    -16-

-------
    Zinkl (1977) exposed 300  female  CD  rats  to  diets  containing 0,  10,  30 or
100 ppm  (mg/kg/d1et)  Aroclor  1254 for 20 weeks, and  reported  that  there was
no  change  1n  hematocMt  throughout  the  trial.    Serum  cholesterol   was
elevated  within  2  weeks  1n  rats  exposed to  >30  ppm  dietary Aroclor  1254
(p<0.01).  SGOT  was elevated (p<0.05)  and  serum gamma globulin  was  reduced
(p<0.01)  In  rats  exposed to  100  ppm Aroclor 1254.   A  dose-related Increase
(p<0.05)  1n  beta  globulin was also  noted.   Rats exposed  to >30  ppm  Aroclor
experienced decreased body weight gain  (p<0.01)  and  Increased  relative  liver
weights (p<0.01), which appeared to be dose-related  (p<0.01).
    Liver lesions  were similar to  those described  previously  (Klmbrough et
al.,  1972;  Bruckner et  a!.,  1974).   Upon  gross examination,   the  livers of
100  ppm-exposed  rats  were  noticeably   enlarged  and  dark 1n  color,  often
exhibiting  an  accentuated  lobular   pattern.    Porphyrla  fluorescence  was
eventually  seen  1n  livers  from  all groups  of Aroclor   1254-exposed  rats.
Microscopically,  focal  necrosis was  observed,  accompanied by  Inflammation.
Kupfer  cells  were  noted to  accumulate  a  brownish,  Iron-positive  pigment.
Mldzonal  and  centrllobular  degeneration consisting of  vacuolated or  eoslno-
phlUc  hepatocytlc  cytoplasm  was  noted.   Lesions  1n  Hvers   from 30  ppm-
exposed  groups  were  similar  to  but less  dramatic   than  those  observed  1n
livers from 100 ppm-exposed rats.
    Most  striking were  skin lesions  on  the  ears,  which  eventually  Involved
15/60 of  the  high-dose  (100 ppm)  group, 4/60 of the  mid-dose  (30 ppm)  group
and 1/60  of  the low-dose  (10 ppm)  group  rats.   These lesions consisted of
alopecia,  hyperkeratosls,   perlkeratosls  and  deep   rete  pegs.  Keratlnous
plugging  of follicles was  noted.   Desslcated serum and other  blood  elements
                                    -17-

-------
were  observed  on  the   surface  of  the  skin.   Subcutaneous  tissues  were
slightly edematous  and  contained focal areas  of  polymorphonuclear  Infiltra-
tion  and  accumulation.   Similar  lesions  were  seen  on the  tall  and  on  the
dorsum of  the  nose.   This  was the only report  found of skin  lesions  1n  rats
associated with  PCBs.   Since  no  mention  was  made  of  liver pathology 1n  10
ppm-exposed  rats,  and   they   suffered  neither  reduced  body  weight  gain,
hepatomegaly  nor  altered  blood  chemistries,   10  ppm   Aroclor   1254   was
considered a LOAEL 1n this  study.
    Skin  lesions  on  the ears  of mice  were  reported by  Bell (1983),  who
exposed  female  Swiss Albino  mice to  a much  higher  level,  200 ppm  (mg/kg
diet) Aroclor 1254 for  23 weeks, and  observed  erythema and thickening of the
pinna of  the ear.  Hyperkeratosls and  occluslve cystic degeneration  of the
pHosebaceous  units  were  seen.   Basal  layer  cells   became  elongated  and
markedly  thickened.   Ultrastructurally, the  most striking  lesions  Involved
the  mlcrovasculature and  Included gross  enlargement   of  the  cells   of  the
basal  layer  of the  capillaries  and  small  venuoles.  The  resultant  lumenar
stenosis caused a failure of  the mlcroclrculatlon,  which  was associated  with
the skin lesions.
    In a  large  experiment,  Keller  (1977) fed  diets  containing  0,  3.75,  37.5
or  375  ppm  {mg/kg  diet) Aroclor  1221, 1242  or 1254  to  groups of  25  male
BALB/C  mice  for 6 months.   Groups of mice  fed 375 ppm Aroclor  1254 exper-
ienced high  mortality during  the  5th  and  6th months.  The mice exhibited the
hepatic  lesions discussed  above  that were  found  In  rats.   No  other groups
experienced  mortality.   Hepatomegaly  (p<0.01) was  observed In all  Aroclor
1254-exposed  groups  and  1n  the   375  ppm Aroclor  1242-exposed groups.   No
lesions  other  than  moderate hepatomegaly were  found 1n livers  from 3.75 ppm
Aroclor  1254-exposed   rats.   Liver   lesions  In  Aroclor   1242-exposed  rats
                                    -18-

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 occurred  only 1n the  375  ppm group.  None of  the  Aroclor  1221-exposed mice
 evidenced  hepatic  lesions.  Exposure  to  Aroclor-contalnlng  diets  was termi-
 nated after  6 months,  and  all  remaining mice  were given a control diet with-
 out  added PCBs  for  an additional 3-month  observation  period.   Hepatomegaly
 persisted  1n  the  Aroclor  1254-exposed  groups  and   the  375  ppm  Aroclor
 1242-exposed  group.    H1ld  liver  lesions  persisted  only  In  the 37.5  ppm
 Aroclor 1254-exposed group.
     In  this   study,  Aroclor  1254  was  clearly  the  most toxic  substance  and
 Aroclor  1221, the least  toxic substance  tested.   Since the lowest  dose of
 Aroclor  1254, 3.75  ppm,  resulted  only  1n reversible  hepatomegaly  without
 h1stolog1cally  demonstrated   liver  lesions,  3.75  ppm  may  be  considered  a
 LOAEL 1n this study.
     Other  studies  describing  subchronlc  exposure of rats  and  mice  to  PCBs
 are  detailed  1n Table 3-1.
     M1nk  are  especially sensitive  to  the  effects  of  PCBs  (see  Table 3-2).
 Bleavlns et al.  (1980)  Investigated  the  toxldty  of Aroclor  1242  and Aroclor
 1016 to pastel mink.   Aroclor  1242 was fed at dietary  levels of 0, 5, 10, 20
 or 40  ppm (mg/kg diet)  and  Aroclor  1016  was  fed at  levels of 0  or  20  ppm.
 The  duration of exposure was  8 months.
     Each  treatment  group  consisted  of 12  females  and  3 males;  the  control
 group contained 24 females and 6  males.   Aroclor  1242  at >20 ppm  resulted 1n
mortality  of  all  mink.  At  10  ppm diet, Aroclor 1242  resulted 1n mortality
 of 8/12 of the  females and 2/3 of the males.   Only one  female  fed the 5  ppm
Aroclor 1242  diet died.   Among controls, 3/24  females  died.  No  explanation
was  given  for death  of  control  group  mink.  Aroclor 1016 resulted  1n death
of 3/12  females  exposed  to  20  ppm  1n  the diet.  Necropsy procedures  con-
 sisted of  gross  examination  only and  revealed emaciation characterized  by
                                    -19-

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almost  complete  absence  of  body  fat  and  the presence  of  gastric  ulcers.
Reproduction was not possible  1n  mink  exposed to >5 ppm  Aroclor  1242,  while
20  ppm  Aroclor  1016   seriously   Interfered  with   reproduction.    Clearly,
Aroclor 1242  was  more  toxic to mink  than Aroclor  1016  1n this study.   The
finding of  gastric  ulceratlon  and severe cachexla  (emaciation) had  not  been
reported previously  1n  experimental  animals.  It  was  not clear  If  the  one
female  that died  1n  the  5 ppm  Aroclor  1242 group  exhibited the  lesions
associated  with PCB-lnduced  mortality  at higher dietary  levels.   Since  more
sensitive  criteria  of  toxldty  were  not  evaluated, It  Is  not possible  to
derive a NOEL or LOAEL  from this  study.
    In an  earlier  study,  AuleMch and  Ringer  (1977) exposed  groups  of  seven
or eight female mink to  0 or  2 ppm Aroclor  1016,  1221,  1242  or  1254 1n  the
diet  for  -10 months.   No overt  signs   of  toxldty Involving  any  of  these
Aroclors  occurred.   Body  weight   gain,  hemoglobin  and   hematocrlt  remained
normal.  Aroclor 1254 at  2 ppm 1n the  diet Interfered  with reproductive per-
formance.   For  nonreproductlve  parameters,  2 ppm of  the  Aroclors  tested
appeared to be a NOEL In this study.
    Monkeys  have  also  been  found to be very sensitive  to  PCBs  (see  Table
3-3).   Barsottl et  al.  (1976)  and Allen  et  al.  (1979a)  fed  diets containing
Aroclor 1248  for  7 months  to  rhesus  monkeys.  The  control  groups consisted
of  12 females and  6 males.   Treatment  groups were  as  follows:  nine females
received  200  g/day of  a  diet containing  2.5 ppm  (mg/kg  diet)  Aroclor  1248
(low-dose  group);  nine  females received  200 g/day  of a  diet  containing  5.0
ppm  Aroclor 1248  (high-dose group); and four males received  300  g/day  of a
diet  containing 5.0 ppm Aroclor 1248.
    Within  2 months,  Aroclor  1248-treated   females  exhibited  frank weight
loss,  which  continued  through the 6  months  of  exposure.  Females  In both
                                    -20-

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groups  began  to  show  alopecia, acne  of the  face and  neck,  and  palpebral
edema  and  erythema.   These  lesions  were  present  to  some  degree  1n  all
females by the end of 6  months.  Hlstologlcally,  keratlnlzatlon  of  hair  fol-
licles  (biopsy   specimens)  was  the  prominent  finding.   Hemograms  remained
normal, but an  Increase  In SGPT was observed.   After 4 months  of  exposure,
menstrual   cycle  length  and   duration  of  menstrual   bleeding were   both
Increased, and  other  disruptions of  the menstrual cycle occurred.   Concep-
tion  rate  1n  the high-dose group  females  was  depressed.  Animals  that  con-
ceived  In both  groups   experienced  a high  level of  early  fetal  mortality
(resorptlons)  and abortions.  By 6 months, one  of the low-dose  group females
had died.   Major gross findings  Included generalized  alopecia,  acne  and  sub-
cutaneous   edema.   Epidermal hyperkeratlnlzatlon,  hyperplasla  of  folUcular
epithelium and  localized Inflammation were  noted microscopically.   Hepatic
focal  necrosis,  enlarged hepatocytes  and  Upld  accumulation  resembled  the
lesions discussed 1n  other  species.
    By  6  months, only one  of   the males (high-dose  level) exhibited  slight
perlorbltal edema and   erythema.   Male  reproduction was  not  affected  by
Aroclor 1248 at  these dosage levels.
    It 1s  clear   from this study  that  monkeys are  more sensitive to  PCBs  then
rats and mice, and that  no  LOEL can be  derived for monkeys from these  data.
Truelove  et  al.   (1982)  dosed  four  cynomolgus  monkeys with 0,  100, 100  or
400 pg  Aroclor   1254  1n  apple juice  dally  for  238-267 days.   Since  this
experiment was directed  primarily toward studying  effects of Aroclor  1254 on
reproduction,  treatment  began  -100   days  prepartum.   Both  of   the  100  yg
Aroclor 1254-treated  monkeys  delivered full-term stillborn  male  Infants.
The  400  yg   Aroclor  1254-treated  monkey  delivered  an  apparently  normal
female  Infant,   which  died of  bronchlo-pneumonla  at   139  days   of   age;
                                    -21-

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this was presumed  to  be  evidence of Impaired 1mmunolog1cal competence.   The
400  v9   and  one  of  the  100  pg  Aroclor  1254-treated  monkeys  lost  their
fingernails  (Indicating  severe  hyperplastlc   hyperkeratosls)   after   -235
days.  No other overt  signs of  maternal  toxldty  were reported.
    Barsottl  (1981)  exposed 24  adult  female  rhesus monkeys  to diets  con-
taining  1.0.  0.25  or  0.025   ppm  (mg/kg/d1et)  Aroclor   1016,  presumably
throughout gestation and lactations.  Infants born  to  the  1  ppm Aroclor 1016
groups  were  significantly  smaller  at  birth.   No  other  manifestations  of
toxldty were reported.  For this study,  a  dietary  level of  0.25 ppm Aroclor
1016 appeared  to  be a NOEL.  It 1s possible that Aroclor  1016  1s less  toxic
to  monkeys  than the  other  Aroclors usually  tested, as was  demonstrated 1n
mink (Bleavlns  et al., 1980).
3.1.2.   Inhalation.   Pertinent  data  regarding  the  effects  of  subchronlc
Inhalation of PCBs were not located 1n the available literature.
3.2.   CHRONIC
3.2.1.   Oral.    Reports  of chronic  oral  exposure  of  laboratory  species  to
PCBs  are summarized  In Tables 3-1  and  3-2.  These  studies  Involved labora-
tory  animals exposed  to  maximally  tolerated  doses  1n  order to  create and
Investigate  the  frank effects  of  PCBs  on various  target  organs or tissues.
They  do  not  define LOAELs  or NOELs  that  would  be useful  1n  risk assessment;
therefore, this  discussion  will  be  a  brief  review of those studies that most
clearly  elucidated the toxic effects of PCBs.
     As was noted  1n the  discussion  of  subchronlc studies,  frank mortality In
rats  occurred  at  dietary  levels  >500  ppm   Aroclor  1254,   Aroclor  1260
(Klmbrough et  al., 1972) or Kanechlor-300,  -400 or  -500 (Ito et al., 1974).
Other  signs  reported  1n  rats  were  reduced appetites, reduced  rate of body
                                    -22-

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weight  gain  and reduced  final  body weight.   Dietary levels of  100 ppm  of
several Aroclors (Klmbrough  et  a!.,  1975;  Allen and  Abrahamson,  1979;  Allen
et al., 1976) or -200-300 ppm Kanechlor-400  (Klmura  and  Baba,  1973)  resulted
1n severe hepatomegaly  1n  rats;  often  livers were 3-  to  4-fold normal  size.
Livers  frequently   exhibited  multiple  elevated  tan  or  discolored   liver
nodules {Klmura and  Baba,  1973;  Klmbrough  et al., 1975),  oval  cell  prolife-
ration,  bile duct   proliferation  (cholanglof1bros1s)  {Ito et  al.,  1974),
fatty Hver degeneration (Ito et al.,  1974;  Klmura and Baba,  1973)  and  focal
necrosis and  degeneration  (Allen et al.,  1976).  Hepatocellular  carcinomas
were also found (Klmbrough et al.,  1975) (Chapter  4).   Concurrent with  these
liver  changes  1n  rats  were  elevated  serum I1p1ds   and  cholesterol  levels
(Allen et al., 1976); Increased hepatic protein,  RNA and  llpld  and decreased
DNA  content;  and  Increased  hepatic  mlcrosomal  total  protein and cytochrome
P-450 content (Allen and  Abrahamson, 1979).  Many hepatic  mlcrosomal  enzyme
activities were markedly  Induced  and  remained  that  way  for the  duration  of
exposure  (Allen and Abrahamson,   1979).    Morgan   et  al,  (1981)  reported
"Intestinal" metaplasia  and adenocardnoma In rats exposed  to  25  ppm Aroclor
1254 for 2 years.
    The only  other   laboratory  species  for  which  chronic PCB exposure  data
are  available 1s the monkey, which  Is considerably  more  sensitive  than  the
rat  to  PCBs.   As reported  1n  Section  3.1.,  liver  pathology 1n  monkeys  and
associated  serum  chemistries  (Barsottl  et  al.,  1976)  resembled   those  1n
other species.  No  reports of liver  neoplasla  In  monkeys  were  located  1n the
available literature, but  no studies were  found  where exposure  to  PCBs  ex-
ceeded 1-1/2 years.   Skin  lesions  were the  most  consistently  reported  toxic
manifestation of PCBs 1n monkeys.   Dietary  levels as  low as  2.5  ppm Aroclor
                                    -23-

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1248 produced  erythema  and  peMorbltal  edema,  which progressed to alopecia,
acne, facial edema and hyperplgmentatlon  (Barsottl and Allen, 1975).
3.3.   TERATOGENICITY AND OTHER  REPRODUCTIVE  EFFECTS
3.3.1.    Oral.   Several  authors  have  Investigated  the  teratogenldty  and
fetotoxldty  associated  with  various  mixtures of PCBs.   Little  evidence of
teratogenldty was found; most  reports  concerned  fetotoxldty  of PCBs.  The
more relevant studies are summarized 1n Table 3-4.
    Collectively,  these  studies reveal  several  aspects of  the  toxldty of
PCBs.  First,  manifestations  of fetotoxldty 1n  mice (Orberg and Klhlstrom,
1973),  rats (Under et al., 1974) and monkeys  (Barsottl, 1981) were observed
1n  the absence of  overt signs of maternal toxldty.  The ability of PCBs or
their  metabolites to  cross  the  placenta  and   cause   abortions  or  fetal
mortality has been demonstrated 1n rats  (Baker et al.,  1977; Spencer,  1982),
rabbits  (VUleneuve  et  al.,  1971) and  mice  (Orberg   and  Klhlstrom,  1973,
Marks  et  al., 1981).   Reduced  Utter  size  was  observed In rats  from  dams
exposed to  diets  containing 100-500 ppm  Aroclor  1254 from  3-4 weeks of age
to  termination of a 1-2 generation study  (Under et  al.,  1974).
    Infants from  monkeys  exposed  to >2.5  ppm  Aroclor 1248  1n  the diet  from
-6  months  before  mating  were  born with hyperplgmentatlon, a mild  sign of
PCB-1nduced  toxldty  1n monkeys  (Allen  and  Barsottl,  1976;  Allen  et  al.,
1979b).  Reduced birth weights  were observed 1n Infants  from monkeys exposed
to  1.0  ppm Aroclor 1016  1n  the diet  (Barsottl,  1981).   Nursing resulted In
loss of facial hair,  edema  of the  eyelids, gastric  hyperplasla and vomiting,
liver  degeneration and  other  general  signs of PCB-lnduced  toxldty, Indica-
ting the  ability  of  monkeys  to excrete  substantial  quantities  of PCBs and
                                    -24-

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                                                                          TABLE 3-4

                                   Sumary:  Teratogenetic, Fetotoxic and Reproductive Effects of Orally Administered PCBs
Species/
Strain        Source of PCBs
                   Dosage Level  and
                  Duration of  Treatment
                                   Maternal Response
                                     Progeny Response
                                            Reference
Rabbit
Aroclor 1254
Rat/Wlstar    Aroclor 1254
Rat/Sherman   Aroclor 1254
Rat/
Holtzman
Nice/CD]
              Aroclor 1260
              Aroclor 1254
Aroclor 1254
3.3'.4.4',5.5'
HCB
0. 1.0. 10.0. 12.5.  25.0,
50 Mg/kg/day on days 1-2B
of pregnancy; 25 mg/kg/day
on days 7-2B of pregnancy
                  0.  6.25.  12.5.  25.0. 50.
                  100 mg/kg/day on days 6-15
                  of  gestation

                  0.  1.  5.  20,  100. 500 mg/kg
                  diet fro* 3-4 weeks of age
                  to  termination  of 1- or 2-
                  generatlon study
0, 5. 20. 100,  500 mg/kg
diet fro* 3-4 weeks of  age
to termination  of  1- or 2-
generatlon study

0,10.50.100 mg/kg/day dosed
on days 7-15 of gestation

0.25.50,100,200.300.600.900
mg/kg diet/day  on  days  6-15
of pregnancy

0,0.1.1.0.2.0.4.0.8.12
mg/kg/day, on days 6-15
of gestation
dose >25 mg/kg:  maternal
deathT weight loss >10.0
mg/kg:  hepatomegaly 25
mg/kg days 7-2B:  reduced
rate of gain.

none reported
                                 none  reported
                                                   none reported
                                                   none reported
>600 mg/kg:  partial
anorexia and weight loss
>8 mg/kg/day:  reduced
rate of gain, lethargy.
vaginal bleeding
dose >12.5 mg/kg/day:  fetal death.
resorptlons, abortions
                                                                 none reported
                                F]a 500 mg/kg:  reduced litter size;
                                100 mg/kg:  reduced Utter size;
                                Fit 10° *9/kg:  reduced survival at
                                weaning; 20. 100 mg/kg:  reduced
                                litter size;
                                F?a 100 mg/kg:  reduced Utter size and
                                reduced survival at weaning;
                                20 mg/kg:  reduced Utter size;
                                Fjb 20. 100 mg/kg:  reduced Utter size.

                                Fjj 500 mg/kg:  reduced Utter size
                                and reduced survival at weaning.
                                FII> 500 mg/kg:  reduced Utter size.
                                   100 mg/kg/day:  reduced Utter size.
>30 mg/kg:  fetal death at delivery
>100 mg/kg:  reduced Utter weight
               fetal mortality.resorp-

               Increased Incidence of
>4 mg/kg/day:
lions;
>2 mg/kg/day:
cleft palate;
>4 mg/kg/day:  Increased Incidence of
hydronephrosls;
>1 mg/kg/day:  Increased Incidence of
cream colored liver;
>1 my/kg/day:  Increased Incidence of
undersized renal papillae.
                                            Vllleneuve et
                                            al..  1971
                                            Llnder  et  al..
                                            1974
                                            Spencer,
                                            1982
Harks et al.,
1981

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                                                                       TABLE  3-4 (cent.)
Species/
Strain
Rat/Sprague-
Dawley
Rat/Ulstar
HIce/NNRl
MIce/NR
, Rabbit
cr
' Mink
Source of PCBs
Aroclor 1221
1242 or 1260
Aroclor 1254
Clophen A60
2.2»,4.4»,
5.51 hexa-
chloroblphenyl
Aroclor 1221.
Aroclor 1254
Aroclor 1242
Dosage Level and
Duration of Treatment
0 or 30 mg/kg bw, days 14-
20 of pregnancy
0 or 70 mg/t drinking water
(6.4 Mg/kg bw); 9 wks
0 or 0.025 mg/day; 72-76 days
0 or 0.5 mg/day; 7 or 13
days of pregnancy
0,1.0 or 10 mg/kg bw;
first 28 days of pregnancy
0-40 Mg/kg diet; B months
Maternal Response Progeny Response
none none
mortality at 7 wks fetal resorptton
lengthened estrus cycle fetal resorptlon
hepatomegaly none
none none
>10 mg/kg: significantly NA
Increasing mortality
>5 mg/kg: complete re- NA
Reference
Gellert and
Wilson. 1979
Baker et al..
1977
Or berg and
Klhlstrom. 1973
Nattsson
et al.. 1981
Vllleneuve
et al.. 1971
Bleavlns
et al.. 1980
              Aroclor 1016      20 mg/kg diet
Nonkey/       Aroclor 1254      0.100 or 400 mg/kg bw/day;
cynomolgus                      continuous starting at 60
                                days of pregnancy

Nonkey/       Aroclor 124B      0.2.5 or 5.0 mg/kg diet;
rhesus                          18 months starting 6 months
                                pregestatlonal
              Aroclor 1248      0.2.5 or 5.0 mg/kg diet;
                                off treatment for  1 year

Nonkey/       Aroclor 1016      0.0.025. 0.25 or  1.0 mg/
rhesus                          kg diet for unspecified
                                length of time
                                                                  productive  failure

                                                                  25JC mortality,  reduced
                                                                  fertility

                                                                  fingernail  loss;  Immuno-
                                                                  loglc  Incompetence
                                                                  facial  edema,  cachexla,
                                                                  hair  loss,  hyperplg-
                                                                  mentatton
                                                                  persistent  hyperplgmen-
                                                                  tatlon

                                                                  none
Increased mortality, decreased
body weight by 4 weeks

fetal mortality; Immunologic
Incompetence
fetal death. Infant facial and eye-
lid edema, loss of facial hair.
facial hyperplgmentatlon. gastric
hyperplasla and vomiting, lymphotd
degeneration, hypocellular bone
marrow, fatty liver

fetal death; reduced neonatal weight
hyperplgmentatlon

1.0 mg/kg diet:  Infants had reduced
birth weight
                                                                                                                                              Truelove
                                                                                                                                              et  al.,  1982
                                                                                                                                             Allen and
                                                                                                                                             BarsotM. 1976;
                                                                                                                                             Allen et al..
                                                                                                                                             1979b;
                                                                                                                                             Allen et al..
                                                                                                                                             1980
                                                                                                                                             Barsottl. 1981
NA . Not applicable

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 their  toxic metabolites  by .lactation  (Allen  and  Barsottl,  1976;  Allen  et
 a!.,  1979b,  1980).   This phenomenon had been  suggested  earlier  by Under  et
 al.   (1974),  who  noticed decreased  survival  at weaning  1n rats  from  dams
 exposed continuously to dietary concentrations >100 ppm Aroclor 1254.
    Only  one  report associating  teratogenldty with  oral  exposure  to  PCBs
 has been  found.   Marks  et  al.  (1981) treated  13-63  pregnant  CD,  mice/group
 with  0,  0.1,  1.0, 2.0,  4.0,  8  or 12 mg 3,3',4,4',5.5'-hexa CB/kg bw/day  by
 gavage  for  days  6-15 of  gestation.   Although no deaths occurred  1n exposed
 dams,  lethargy  and  vaginal  bleeding  were  observed  1n dams  exposed to  >8
 mg/kg/day.  At dosages  >4 mg/kg/day, the average number of  live fetuses/dam
 was reduced.  A reduction  1n  the  number  of  Implants and an Increase 1n fetal
 resorptlons  were   observed 1n  mice  treated  with >8  mg/kg/day.   Possibly
 because of  the greater  care used In  examining  fetuses  1n  this study, terata
 were  found.   Significant  (p<0.001)  Incidences  of  cleft palate occurred  1n
 groups  dosed  at   >2  mg/kg/day.    The occurrence of  cleft  palate  was  2/247,
 4/83, 5/63, 20/117, 69/154, 118/176  and  82/108  1n  fetuses  from dams, respec-
 tively,  1n  the   groups  described  above.    Hydronephrosls   also  followed  a
 significantly (p<0.001)  dose-related trend  In  fetuses  from dams  exposed  to
 >4 mg/kg/day.  Cream-colored  liver  nodes were  significantly  (p<0.001) assoc-
 iated with  dosage levels >1  mg/kg/day.  The authors mentioned  briefly  that
 they  had  also   found   3,3',4,4'-tetra  CB   to  be  teratogenlc,   but  that
 3,3',4,4',5,5'-hexa CB  was the more potent  teratogen.
    In  these  studies,   as  1n  the  toxldty  studies  reviewed  In  Chapter  3,
monkeys appeared  to  be  the species most  sensitive to fetotoxlc  effects  of
 PCBs.    Barsottl   (1981)  reported  reduced   birth  weights  1n  Infants  from
monkeys treated with  1.0 ppm Aroclor  1016  1n the diet.  The  observation  of
 fetotoxlc  effects  at  levels lower  than  those  reported to cause  toxldty  1n
                                    -27-

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animals  after   birth  reduces  considerably  the confidence  placed  In  risk
figures derived solely from subchronlc and chronic  toxldty  studies  (Section
6.1.1.).  The fact  that  Aroclor  1016 Is  probably the least  toxic of  the  PCB
products  1n  the   environment  further  erodes  confidence  1n  risk   figures
derived from studies using Aroclor  1016.
3.3.2.   Inhalation.  Pertinent  data regarding the  fetotox1c1ty  or  terato-
genldty associated with Inhalation exposure to PCBs could be  located  In  the
available literature.
3.4.   TOXICANT INTERACTIONS
    Many of  the Interactive effects between PCBs  and  other xenoblotlcs  can
be  predicted  from an understanding  of  the mlcrosomal  enzyme-Inducing  capa-
bilities  of  PCBs.    For   example,   PCB  pretreatment  of  rats  resulted  In
Increased rate  of metabolism and excretion of  pentobarbltal, and 1n  reduced
pentobarb1tal-1nduced  sleeping times (Chu et  al.,  1977;  VUleneuve et  a!.,
1972).  Hepatotox1c1ty Induced by vinylldene fluoride (Conolly et al.,  1979)
and halothane anesthesia  (S1pes  et  al.,  1978)  was  Increased 1n rats  by  pre-
treatment with  PCBs.    Cadmium,  however,  appeared  to  have  an  antagonistic
effect  on  the  biochemical  changes  that result   from  PCB-1nduced  hepatic
enzymes  (Suzuki,   1980;  Horlo  et  al.,  1981).   The exact  nature  of  this
protective  effect  has  not been  elucidated,   but  the  effect  appears  to  be
additive (Suzuki,  1980).
3.5.   HUMAN EFFECTS
    Several   reports  of   human  effects  resulting from  occupational  or  acci-
dental exposure are available.   Melgs  et al.   (1954) reported  seven  cases  of
chloracne among 14 workers  Intermittently exposed  to  Aroclor vapors.   The
mean  length of exposure  was  14.3  months  for  affected  and  11.4 months  for
                                    -28-

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unaffected workers, but no  significant correlation  between  duration  of  expo-
sure and  effects  occurred.   Apparently,  a one-time measurement  revealing  an
Aroclor concentration of 0.1 mg/m3 was made 19 months  after  exposure  began.
    F1schbe1n et al. (1979) reported  a cross  sectional  clinical  field survey
of  326  workers 1n  two capacitor-manufacturing  plants  who  were exposed  to
Aroclors  1254,  1242,  1016  and  1221.   A1r levels  (8-hour  TWA)  of   PCBs  In
these  plants  were  0-11.0  mg/m3.  Dermatologlc  symptoms  reported 1n 76/168
(45%) male and 87/168  (55%) female workers were  rash,  pruritus,  acne, hyper-
pigmentation, and thickening and  discoloration of  the  fingernails.   Respira-
tory complaints  were  made  by  3.4-13.8%  of  these workers,  and 48.2%  com-
plained of eye Irritation (Warshaw et al.,  1979).   Gastrointestinal  symptoms
(18%) and neurological  symptoms  (39% of males and  56%  of  females) were also
reported  (F1schbe1n et  al.,   1979).   The authors  performed extensive  bio-
chemical and hematologlcal  studies on  these workers and concluded that  there
was no  correlation  between  any symptoms and  duration of  employment  but that
a positive association  between plasma PC8 level  and serum  SCOT  level (Indi-
cating  extent of liver  damage)  existed.
    Ouw et al.  (1976)  compared dermatologlc  and hepatic  function parameters
of 34 electrical  Industry workers exposed to electrical grade  ("no" Impuri-
ties) Aroclor  1242  with those  of 30  control  volunteers.   Control volunteers
had no  history of  occupational exposure to PCBs,  and PCBs  were  not  detected
1n  their  blood.   Exposed  workers had  hepatic  function  tests   that  yielded
scattered abnormal  values:  one worker had chloracne and five had eczematous
rashes   on hands and legs.   These effects were associated with air  levels  of
<1 mg Aroclor 1242/m3.
    Accidental 1ngest1on of PCBs  occurred 1n  >1000 persons  1n Japan 1n 1968
as a result  of contamination  of  rice oil with Kanechlor-400, a  PCB product
                                    -29-

-------
used  as  a  heat  transfer  agent.   The  average  total  PCB  consumption  by
affected Individuals was estimated to be  -2 g, with  0.5  g  being  the  smallest
total amount  consumed by  a  group of  325 people (Kuratsune  et  al.,  1972).
The  contaminated  rice oil  also  contained  ~5 ppm  PCDFs,  presumably  formed
from  continual  reheating  of  PCBs.   PCDFs  are thought  to be several  times
more  toxic than PCBs.   The dermal and respiratory symptoms reported  1n  this
case  of  "Yusho"  poisoning were similar  to  those reported  1n  occupatlonally
exposed  workers  but  were considerably  more  severe.   Additionally,  palmar
sweating  and muscular  weakness were  common  complaints  (Kuratsune  et  al.,
1969).   Yamashlta  (1977) reported premature  delivery, reduced  birth  weight
and  height,   hyperplgmentatlon of   skin  and  mucous  membranes,  precocious
dental  eruption  and  decreased  calcification  of  the  cranium  1n  Infants  of
Yusho women.   In  addition to  these  findings,  Kuratsune et al.  (1969)  also
reported delivery of 3 stillborn and  10 live Infants from 11 Yusho women.
    By  1979,  31 Yusho patients had died, 11  (35.4%)  of  these from malignant
neoplasms  (Urabe  et al.,  1979).   These  11  malignancies  Included 2  stomach
cancers,  1  stomach  and  liver  cancer  (possibility  of  metastasis not  dis-
cussed),  2 liver  cancers 1n  clrrhotlc livers, 2  lung  cancers,  1 lung tumor,
1 breast  tumor and 2 malignant lymphomas.
    In  1979,  -2000  persons  1n  central  western  Taiwan  developed  symptoms
practically  Identical to  those  observed 1n  Yusho patients  (Chen  et  al.,
1981).   These persons had also consumed  rice oil  contaminated  by PCBs  used
as  a  heat transfer  agent  In the  manufacturing process.   Samples of contami-
nated rice oil were  obtained  and were found  to  contain  53-405  ppm  PCB;  no
estimates  of  total PCB Intake or duration of exposure were reported.
                                    -30-

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                              4.   CARCINOGENICITY
4.1.   HUMAN DATA
    There  are  few  data  regarding  the  cardnogenlclty  of  PCBs  1n  humans.
Urabe et  al.  (1979)  reported that 11/33 deaths  (35.454)  among Yusho patients
who  had  died  by  1979  resulted  from  malignancies.   These  mallg-  nancies
Involved  various  body  sites,  and  expected  Incidences  based on  unexposed
populations were not available; therefore,  the  significance  of these  data 1s
uncertain.  Bahn et  al.  (1976,  1977)  reported  2  cases  of malignant melanoma
among 31  workers "heavily exposed" and  1 case of  malignant melanoma among 41
workers "less  heavily  exposed"  to Aroclor  1254.  IARC  (1978)  estimated that
only  0.04 cases  would  be expected  1n  an  unexposed population of  this size
and  concluded  that  this  was  suggestive  evidence  that  PCBs   are  human
carcinogens.   Davldorf  and  Knupp (1979)  examined the  Incidence  of  ocular
melanoma  In  several  counties  of   Ohio  during  1967-1977,  attempting  to
associate  higher  Incidences with areas  known to have  higher  concentrations
of  PCBs  because of  Industrial contamination.   No significant  associations
were found.
4.2.   BIOASSAYS
    Protocol and  results  of the  major cardnogenlclty  bloassays  are  sum-
marized  In Table  4-1.   Early  cardnogenlclty  bloassays Involving  dietary
administration of PCBs  to  rats and mice either Involved  too  few animals or
had   too   short   a   duration  to  generate   statistically  or  biologically
significant data.   Klmura  and  Baba   (1973)  fed  diets  containing  levels  of
Kanechlor-400 that varied from  38.5-616 ppm to 10  male  and  10 female Oonryu
rats for  22-80 weeks.  The  study  was  Inconclusive because of early  mortality
from  a  number  of unrelated causes.   Hyperplastlc  liver  nodules,  which  the
authors  suggested  were  precancerous  lesions,  occurred  In  all female  rats
                                    -31-

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                        TABLE 4-1



Smmary  of  Carcinogenic Effects of Orally Administered PGBs
Hepatopalhology: No. Affected/No. Necropsled
Species/ PCB
Strain Sex Source
Rats/ H control
Donryu
F control
H Kanechlor-400
Ł, f Kanechlor-400
1 Nice/ N control
dd
Kanechlor-SOO
Kanechlor-SOO
Kanechlor-SOO
Kanechlor-400
Kanechlor-400
Kanechlor-400
Kanechlor-300
Kanechlor-300
Kanechlor-300
Dietary Level
(ppm)
0
0
38.5
38.5
0
500
250
100
500
250
100
500
250
100
Number
Treated
5
5
10
10
6
12
12
12
12
12
12
12
12
12
Exposure
•days)
NA
NA
159-530
244-560
224
224
224
224
224
224
224
224
224
224
Neoplasttc
Nodules
0/10
0/10
0/10
6/10
0/6
0/12
0/12
0/12
0/12
0/12
0/12
0/12
0/12
0/12
Adenof Ibrosts
0/10
0/10
0/10
0/10
NR
NR
NR
NR
NR
NR
NR
NR
NR
NR
Hepatoma
NR
NR
NR
NR
NR
NR
NR
HR
NR
NR
NR
NR
NR
NR
Hepatocellular Reference
Carcinoma
0/10 Klmura and
Baba. 1973
0/10
0/10
0/10
0/6 Ito et al.,
1973
5/12
0/12
0/12
0/12
0/12
0/12
0/12
0/12
0/12

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                                                                       TABLE 4-1  (cent.)
co
i
Hepatopathology: No. Affected/No. Necropsled
Species/
Strain
Rats/
Ulstar











Nice/
CJ
Balb

Rats/
Sherman
PCB
Sex Source
N control
Kanechlor-SOO

Kanechlor-SOO

Kanechlor-SOO
Kanechlor-400
Kanechlor-400
Kanechlor-400
Kanechlor-300
Kanechlor-300
Kanechlor-300
N control
Aroclor 1254
Aroclor 1254
F control
Aroclor 1260
Dietary Level Number
(ppn) Treated
0
1000

SOO

100
1000
SOO
100
1000
SOO
100
0
300
300
0
100
IB
290
rats
total
In
1 con-
trol. 9
treatment
groups




100
SO
50
200
200
Exposure
(days)
378
378

378

378
378
0/8
378
378
378
378
330
330
180
630
630
Neoplastlc
Nodules
0/18
5/13

5/16

3/25
3/10
0/8
2/16
0/15
0/19
1/22
NR
NR
NR
0/173
144/184
Adenof Ibrosis
0/18
4/13

0/16

0/25
2/10
NR
0/16
2/15
0/19
0/22
0/58
22/22
0/24
NR
NR
Hepatoma Hepatocellular Reference
Carcinoma
NR
NR

NR

NR
NR
0/8
NR
NR
NR
NR
0/58
9/22
1/24
NR
NR
0/18 Ho et a!.,
1974
0/13

0/16

0/25
0/10

0/16
0/15
0/19
0/22
NR Klmbrough
and Under.
NR 1974
NR
1/173 Klmbrough
et al.. 1975
26/184

-------
                                                                         TABLE 4-1 (cont.)
 i
09
Hepatopathology: No. Affected/No. Necropsled
Species/
Strain
Rats/
Fischer
344






Sex
M
M
N
M
F
F
F
F
PCB
Source
control
Aroclor 12S4
Aroclor 1254
Aroclor 1254
control
Aroclor 1254
Aroclor 1254
Aroclor 1254
Dietary Level Number
(ppn) Treated
0
25
50
100
0
25
50
100
74
24
24
24
24
24
24
24
Exposure
(days)
735
735
735
735
735
728-735
728-735
735
Neoplastlc
Nodules
MR
NR
NR
NR
NR
NR
NR
NR
Adenof Ibrosls
0/24
0/24
0/24
1/24
0/23
0/24
1/22
2/24
Hepatoma
NR
NR
NR
NR
NR
NR
NR
NR
Hepatocellular Reference
Carcinoma
0/24 NCI. 1978
0/24
1/24
2/24
0/23
0/24
0/22
0/24
           NA-Not  applicable; NR-not  reported

-------
that  survived  long enough  to  consume >1200 mg  Kanechlor-400.   Hyperplastlc
nodules were not found 1n male rats.
    Ito  et  al.  (1973)  produced hepatocellular  carcinomas  1n  5/12 male  dd
mice  exposed  to 500  ppm Kanechlor-500 for  32  weeks.   Lower dietary  levels
(250,  100  ppm) of  Kanechlor-500  and all  three  levels of Kanechlor-400  and
Kanechlor-300  failed  to  cause hepatocellular carcinoma.  These  authors  sur-
mised  that  degree  of  chlorlnatlon  was Important 1n determining  the  carclno-
genlcHy of PCB  products.   A  subsequent  study by Ito  et  al.  (1974)  produced
considerable mortality but  no cancer In  Wlstar  rats  exposed to 100,  500  or
1000 ppm Kanechlor-300, -400 or -500  1n the diet  for  378 days.
    Exposure of 200 female  Sherman rats  to 100  ppm Aroclor  1260 (similar  to
the PCBs  contained 1n Kanechlor-500)  1n  the diet for 630  days resulted  1n
144 livers  containing  neoplastlc nodules  from the 184  rats  that survived  to
termination.  Hepatocellular carcinomas were  found  In  26 of the 184  treated
survivors.  No neoplastlc .nodules  and only  one hepatocellular carcinoma  were
found In the 173 control  group survivors  (Klmbrough et  al.,  1975).
    The NCI (1978)  exposed  groups  of 24 male and 24 female  Fischer  344  rats
to dietary  levels of 0,  25, 50  or  100 ppm Aroclor  1254 for  728-735 days.   No
Hver tumors were found  1n  any  female rats.   Among the male  groups,  1/24 and
2/24  rats  1n   the 50  ppm and   100  ppm  groups,   respectively,   developed
hepatocellular   carcinomas.   The  Incidence   did   not  seem  to   follow   a
dose-related pattern and was  not  significant by either  the  Cochran-ArmHage
or Fisher exact  tests.  The NCI (1978)  concluded  that Aroclor   1254 was  not
carcinogenic under  the conditions of  this  bloassay.
                                    -35-

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4.3.   MUTAGENICITY
    Schoeny et  al.  (1979)  Investigated the  mutagenlclty  of Aroclor  1254  to
Salmonella typhlmurlum In the Ames assay.  Negative  results  were  obtained  1n
S.  typhlmurlum  strains  TA1535, TA1539,  TA98 and  TA100  without  hepatic  S-9
activation, and with  S-9 activation from  rats  pretreated with Aroclor  1254
as a hepatic  enzyme  Inducer.   Schoeny  (1982) also obtained  negative  results
1n  these  same four  strains  with  one  monolsomer,  two separate  tetralsomers
and one hexalsomer of  PCBs.   Activation with hepatic S-9  fractions with and
without several kinds  of Induction  failed  to elicit positive  results.   The
only report of positive  response  In  the Ames  assay Involved 4-chlorob1phenyl
and,  to  a  lesser extent,  Aroclor  1221,  tested  1n S.  typhlmurlum  strain
TA1538  (Wyndham et al..  1976).   These  authors  used rabbit  liver mlcrosomal
preparation  as  the  metabolic  activator;   no  reason  for   this  choice  was
mentioned.
    PCB products  failed  to  produce  positive  results In the  dominant  lethal
assay  1n  rats  (Green  et al.,  1975a)  or  chromosomal changes 1n  D.  melano-
gaster  (Nllsson and  Ramel,  1974)  or 1n  sperm and bone marrow  cells  of  rats
(Green et al., 1975b; Garthoff et  al.,  1977).
4.4.   WEIGHT OF EVIDENCE
    IARC  (1978) has  developed and used a method  of  qualitative risk  assess-
ment of potential carcinogens,  based upon  sufficiency of  evidence from human
studies and  animal  bloassays.  The  degrees  of evidence  for carc1nogen1c1ty
1n  human  studies  were categorized  as  sufficient  evidence,  limited  evidence
or  Inadequate evidence.   Based upon the studies  of PCB-lnduced  cancers  1n
humans,  the  evidence for a  carcinogenic  role  of  PCBs  1n  humans  was judged
Inadequate.   As discussed  In Section  4.2.,  Kanechlor-500 has  been  shown  to
be  carcinogenic  1n  mice (Ito  et  al.,  1973)  and Aroclor 1260  has been shown
                                    -36-

-------
to be carcinogenic to rats (Klmbrough et al.,  1975).   There  was  judged  to  be
sufficient evidence  to  conclude  that  PCBs  are  carcinogenic  1n  laboratory
rodents.   Applying  the  criteria for  evaluating the  overall  weight of  evi-
dence for  cardnogenlcHy to  humans  proposed  by  the Carcinogen  Assessment
Group of  the U.S.  EPA  (Federal  Register, 1984), PCBs  are  most appropriately
classlfed as  Group B2 - Probable Human Carcinogens.
                                    -37-

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                     5.   REGULATORY STANDARDS AND CRITERIA
    Under  Section  6{e)  of  the  Toxic Substances  Control   Act  (TSCA)  (P.I.
94-469), the  manufacture,  sale  and  distribution  of  PCS  products have  been
restricted.   PCB  products  were  further  restricted  to  sealed  systems as  of
1977, and manufacture and distribution were banned  In  1979.
    The U.S.  EPA (1980b) has set ambient water  quality  criteria  for  PCBs for
the  protection  of humans  from  Increased  lifetime  risk of cancer of  10~5,
10~6  and  10~7  at 0.79,  0.079  and   0.0079  mg/i.   Because these  compounds
have a large  BCF, these criteria apply regardless of  whether  exposure occurs
through  consumption  of  2 I of  water  and  6.5 g of  fish/day or through  con-
sumption of  fish  alone.  The  FDA  has set  temporary  tolerances  for  PCBs  In
food and related products as  reported 1n Table 5-1 (Code of  Federal  Regula-
tions, 1981.)
    Occupational exposure limits recommended by the ACGIH (1980)  for  Aroclor
1254  are a  TLV of  0.5 and  a  STEL  of  1.0  mg/m3.   For Aroclor  1242,  the
recommended TLV  Is  1 and the  STEL  Is 2 mg/m3.   The  NIOSH (1977)  criterion
1s 1.0 vg/m3 for an  exposure of 10  hours/day,  40 hours/week.
                                    -38-

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                                  TABLE  5-1
                           FDA  Regulations  for PCBs*
             Commodity                                   Temporary Tolerances
                                                               (ppm)
M1lk (fat basis)                                                1.5
Manufactured dairy products (fat basis)                         1.5
Poultry (fat basis)                                             3.0
Eggs                                                            0.3
Finished animal feeds                                           0.2
Animal feed components of animal origin                         2.0
Edible portion of fish and shellfish                            5.0
Infant and junior foods                                         0.2
Paper food packaging material                                    0.0

*Source:  Code of Federal Regulations, 1981
                                    -39-

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                             6.  RISK ASSESSMENT
6.1.   ACCEPTABLE INTAKE SUBCHRONIC (AIS)
    PCBs have  been  shown to be  carcinogenic  1n laboratory animals and  data
are sufficient for estimation of carcinogenic potency.   It  Is  Inappropriate,
therefore,  to derive an AIS for  PCBs.
6.2.   ACCEPTABLE INTAKE CHRONIC (AIC)
    PCBs have  been  shown to be  carcinogenic  1n laboratory animals and  data
are sufficient for estimation of carcinogenic potency.   It  1s  Inappropriate,
therefore,  to derive an AIC for  PCBs.
6.3.   CARCINOGENIC  POTENCY (q  *)
6.3.1.   Oral.   Data  from  female  rats  1n  the study  by  Klmbrough  et  al.
(1975) were  chosen  by the U.S.  EPA Carcinogen  Assessment  Group  to quantify
the carcinogenic  risk  from  exposure  to  PCBs  (U.S.  EPA,  1980b).   The  TWA
dietary  level was  determined  to  be  88.4  ppm,   equivalent   to  4.42  mg/kg
bw/day, assuming  an  adult rat  eats food equal  to  5% of  Us body weight/day
and weighs  0.4  kg.   For  statistical  analysis,  the  Incidences  of  hepato-
cellular carcinoma   (26/184  1n  treated  rats   and 1/173   In  controls)  and
neoplastlc   nodules  (144/184 1n  treated  rats   and  0/173  1n  controls)  were
combined to  produce total tumor  Incidences  of 170/184  1n treated rats  and
1/173  1n controls,  respectively.   Using  these  data,  the data 1n Table  6-1
and the  linearized  multistage  model  developed  by  Crump and  adopted by  the
U.S.   EPA    (1980a),   a  q^   for  human   exposure   to   PCBs   of   4.3396
(mg/kg/day)"1 was calculated.
6.3.2.   Inhalation.   No  studies   of  carclnogenlcHy  of  PCBs   related   to
Inhalation   exposure  have been   located  1n  the  available literature,  and  no
carcinogenic risk data for this  route can  be  calculated.
                                    -40-

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                                   TABLE  6-1
                      Data Used as the Basis for the q-j*
     Criteria
         Details
Species
Strain
Sex
Body weight (assumed)
Length of exposure
Length of experiment
Tumor site
Tumor type

PCB product tested
Dose
(mg/kg/day)
0
4.42
rat
Sherman
female
0.4 kg
645 days
730 days
liver
combined hepatocellular carcinomas
and neoplastlc nodules
Aroclor 1260
Incidence
(no. responding/no, tested)
1/173
170/184
                                    -41-

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                                7.   REFERENCES

ACGIH  (American  Conference  of  Governmental Industrial  Hyg1en1sts).   1980.
Documentation of the Threshold Limit Values, 4th ed.   (Includes  Supplemental
Documentation,  1981).   Cincinnati,  OH.   486  p.

Albro, P.W.  and L.  F1shbe1n.   1972.   Intestinal  absorption of  polychlorl-
nated blphenyls In rats.  Bull. Environ. Contam. Toxlcol.  8: 26.   (CHed  1n
U.S. EPA, 1985)

Allen, J.R. and L.J.  Abrahamson.   1979.  Responses of rats  exposed  to  poly-
chlorinated blphenyls  for  52 weeks.   II.  Compositional  and enzymlc  changes
In the liver.  Arch. Environ. Contam.  Toxlcol.  8:  191.   (Cited  In  U.S.  EPA,
1985)

Allen, J.R.  and D.A.  Barsottl.   1976.   The effects  of transplacental and
mammary movement of the PCBs on Infant  rhesus monkeys.  Toxicology.   6:  331.

Allen, J.R.,  D.H.  Norback  and  I.e.  Hsu.   1974.    Tissue  modifications  In
monkeys  as  related to  absorption  distribution  and excretion of  polychloM-
nated  blphenyls.   Arch. Environ. Contam.  Toxlcol.  2:  86.   (CHed  1n  U.S.
EPA, 1985)

Allen, J.R.,  L.A.  Gartens, L.J.  Abrahamson and  R.J.  Marlar.   1975.  Re-
sponses  of  rats   and  nonhuman  primates  to  2,5,2'.5'-tetrachlorob1phenyl.
Environ.  Res.  9:  265.  (CHed 1n U.S.  EPA,  1985)
                                    -42-

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Allen,  J.R.,  L.A.  Carsten  and L.J.  Abrahamson.   1976.   Responses  of  rats
exposed to polychlorlnated  blphenyls  for  52 weeks.   I. Comparison of  tissue
levels of  PCB  and  biological  changes.  Arch.  Environ.  Contam. Toxlcol.   4:
409.  (Cited 1n U.S.  EPA,  1985)

Allen,  J.R.,  W.A.  Hargraves,  M.T.  Hs1a,  et al.   1979a.   Comparative  toxi-
cology  of chlorinated  compounds  on  mammalian  species.   Pharmacol.   Ther.
7(3): 513-147.  (Cited 1n  U.S.  EPA,  1985)

Allen,  J.R.,  O.A.  Barsottl,   L.K.  Lambrecht and  J.P.  Van  Miller.    1979b.
Reproductive effects  of  halogenated  aromatic  hydrocarbons on nonhuman  pri-
mates.  Ann NY Acad.  Scl.   320: 419-424.   (Cited  1n  U.S.  EPA,  1985)

Allen, J.R.,  O.A. Barsottl, L.A. Carstens.   1980.  Residual effects  of  poly-
chlorinated blphenyls  on  adult  nonhuman  primates and  their  offspring.   J.
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AuleMch,   R.J.  and  R.K.   Ringer.   1977.   Current status  of  PCB toxUHy  to
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Bahn,  A.K.,   I.  Rosenwalke,  N.  Herrmann,   P.  Grover,   J.  Stellman  and  K.
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                                    -43-

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Bahn, A.K..  P.  Grover,  I.  Rosenwalke,  K.  O'Leary and  3.  Stellman.   1977.
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Baker, F.D., B. Bush,  D.F.  Tumasonls  and F.C. Lo.  1977.   Toxlclty and  per-
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Barsottl,  D.A.   1981.   Gross,  clinical  and  reproductive  effects  of  poly-
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Barsottl, D.A. and  J.R.  Allen.   1975.  Effects of polychlorlnated blphenyls
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Barsottl, D.A., R.J. Marlar and  J.R.  Allen.   1976.  Reproductive dysfunction
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Becker,   G.M.,  W.P.   HcNulty  and M.  Bell.   1979.   Polychlorlnated blphenyl-
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Bell, H.   1983.   Ultrastructural features of the murlne  cutaneous mlcrovas-
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                                    -44-

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Benthe, H.F.,  J.  Knop and A.  Schmoldt.   1972.   Absorption and distribution
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Bowman, R.E. and M.P. He1ron1mus.  1981.  HypoactWUy In  adolescent monkeys
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Bowman, R.E.,  M.P. He1ron1mus  and  D.A.  Barsottl.   1981.   Locomotor  hyper-
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Bruckner,   J.V.,  K.L.  Khanna  and  H.H.   Cornish.   1974.    Polychlorblphenyl-
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Callahan.   M.A.,  M.W.  Sllmak,  N.W.  Gabel,  et  al.   1979.   Water-Related
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                                    -45-

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Chen, P.H.,  K.T.  Chang and  Y.O.  Lu.   1981.   Polychlorlnated blphenyls and
polychloMnated dibenzofurans  1n  the  toxic  rice-bran  oil  that  caused PCB
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Chu,  C.K.,  V.J.  Stella,  J.V.  Bruckner  and W.O.  Jiang.   1977.   Effects  of
long-term exposure  to  environmental levels  of  polychlorlnated blphenyls  on
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Code of  Federal Regulations.  1981.  Tolerance  for  polychlorlnated  blphenyls
(PCBs) (para.  51,725).   21:  109.30.

Conolly,  R.B.,  S. Szabo and  R.J.  Jaeger.  1979.  V1nyl1dene  fluoride:  Acute
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CupHt,   L.T.  1980.   Fate  of  Toxic  and  Hazardous Materials  In  the A1r
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Davldorf,  F.H.  and  J.A.   Knupp.    1979.   Epidemiology  of  ocular  melanoma.
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                                    -46-

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Flschbeln, A.,  M.S.  Wolff, R. L1l1s.  J.  Thornton and  I.J.  Sellkoff.   1979.
Clinical finding among  PCB-exposed capacitor manufacturing workers.   Ann.  NY
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Garthoff,  L.H.,  L.  Friedman,  T.M.  Farber,  et al.   1977.   Biochemical  and
cytogenetlc effects  1n  rats  caused by  short-term Ingestlon  of Aroclor  1254
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EPA, 1985)

Gellert, R.J.  and  C. Wilson.  1979.   Reproductive  function  In rats  exposed
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437.  (Cited In U.S.  EPA, 1985)

Green, S.,  F.M.  Sauro  and L.  Friedman.   1975a.   Lack  of dominant  lethality
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Green, S.,  J.V.  Carr,   K.A.  Palmer  and E.J. Oswald.   1975b.  Lack of  cyto-
genetlc effects In bone marrow and spermatogonlal cells  1n rats  treated with
polychlorlnated blphenyls  Aroclors  1242  and 1254.   Bull.   Environ.  Contam.
Toxlcol.  13:  14-22.   (Cited  1n U.S.  EPA,  1985)

Hor1, S.,  H.  Obana,  T. Kashlmoto, et  al.   1982.   Effect of polychlorlnated
blphenyls  and polychlorlnated  quaterphenyls  1n  cynomolgus  monkey  (Hacaca
fasdcularlsl.  Toxicology.  24(2):  123-139.   (Cited  1n  U.S.  EPA,  1985)
                                    -47-

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Klmbrough,  R.D., R.E.  Under  and  T.B.  Galnes.  1972.  Morphological changes
in  livers  of  rats  fed  polychlorlnated  blphenyls.  Arch.  Environ. Health.
25: 354.  (CHed 1n  U.S.  EPA,  1985)

Klmbrough,  R.D., R.A. Squire,  R.E. Under, J.D. Strandberg, R.J. Hontall and
V.W. Burse.  1975.   Induction  of  liver  tumors 1n Sherman  strain female  rats
by  polychlorlnated   blphenyl   Aroclor  1260.   J.  Natl.  Cancer  Inst.   55:
1453-1459.   (CHed 1n U.S. EPA, 1985)

Klmura,  N.T.   and  T.  Baba.   1973.   NeoplastU  changes  1n  the  rat  liver
Induced  by  polychlorlnated blphenyls.   Gann.   64:  105-108.  (CHed 1n  U.S.
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Koller,  L.D.   1977.   Enhanced  polychlorlnated blphenyl  lesions  In  Moloney
leukemia virus-Infected  mice.  Clln.  Toxlcol.   11(1):  107-116.    (CHed  1n
U.S. EPA, 1985)

Kuratsune,  M.,  et al.  1969.   An epldemlologlc  study  on  "Yusho" or  chloro-
blphenyls  poisoning.   Fukuoka  Acta  Med.   60:  513.    (Jap.)   (Cited  1n
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Kuratsune,  M.,  T. Yoshlmura,  J.  Matsuzaka and A.  Yamaguchl.  1972.   Epldeml-
ologlc  study  on Yusho, a  poisoning caused  by 1ngest1on of Mce oil contami-
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                                     -49-

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 Under,  R.E.,  T.B.  Galnes and R.O.  Klmbrough.   1974.   The effect of PCB  on
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 1178-1180.

 Marks  T.A.,  G.L.  Klmmel and R.E.  Staples.   1981.  Influence of  symmetrical
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 Mattsson, R.,  A. Mattsson,  J.E.  Klhlstroem and K. L1ngahl-K1essl1ng.   1981.
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.fetuses  1n  pregnant  mice.   Arch. Environ.  Contam. Toxlcol.  10(3): 281-288.
 (CUed 1n U.S. EPA,  1985)

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                                     -50-

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Morgan,  R.W.,  J.M.  Hard  and  P.E.  Hartman.   1981.   Aroclor  1254-lnduced
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Nllsson,  B.  and  C.  Ramel.   1974.  Genetic  tests  on DrosophUa melanogaster
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NIOSH  (National   Institute  for  Occupational   Safety   and   Health).    1977.
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N1sh1zum1, M.  1970.   Light and electron microscope study of  chloroblphenyl
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Norback,  D.H., E.   Mack,  K.A.  Blomqulst  and  J.R.  Allen.    1978.  Metabolic
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                                    -51-

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Orberg. J. and J.E. Klhlstrom.  1973.  Effects of long-term feeding of poly-
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Ouw, H.K.,  G.R.  Simpson  and D.S.  S1yal1.   1976.   Use and health effects of
Aroclor  1242,  a  polychlorlnated blphenyl  1n  an  electrical  Industry.  Arch.
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Schoeny,  R.S., C.C. Smith and  J.C. Loper.   1979.  NonmutagenUHy for  Salmo-
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S1pes,  I.G.,  G.E.  McLaln,  Jr.,  T.L. Podolsky  and B.R.  Brown,  Jr.  1978.
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Spencer,   F.   1982.   An  assessment  of  the reproductive  toxic  potential of
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                                    -52-

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Suzuki,  T.   1980.   Additive effects  of dietary  cadmium and  PCB 1n  rats.
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U.S.  EPA.   1980a.   Guidelines  and Methodology  Used  1n  the  Preparation  of
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U.S.  EPA.   1980b.   Ambient  Water Quality  Criteria  for  PolychloMnated  B1-
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U.S.  EPA.   1983.   Methodology  and Guidelines for Reportable Quantity  Deter-
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U.S.  EPA.   1985.   External  Review Draft of  Drinking Water Criteria Document
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Assessment  Office,  Cincinnati,  OH.,  OHEA for  the  Office of Drinking  Water,
Washington,  DC.

                                    -53-

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           Verschueren,  K.   1983.   Handbook of Environmental Data on Organic Chemicals,

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           (Cited  In  U.S. EPA,  1985)



           VUleneuve,  D.C.,  D.L.  Grant   and  W.E.J.  Phillips.   1972.   Modification of

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           Warshaw,  R.,  A.  F1schbe1n,  J.  Thornton, A.  Miller and  I. J. Sellkoff.  1979.

           Decrease  1n vital  capacity  1n  PCB-exposed  workers  1n a capacitor  manufac-

           turing  facility.  Ann.  N.Y. Acad. Sd.  320: 277.  (Cited 1n U.S.  EPA, 1985)



           Wyndham,  C.,  0.  Devenlsh  and  S.  Safe.   1976.  The In  vitro  metabolism,

           macromolecular  binding  and bacterial  mutagenldty  of  4-ch1orob1pheny1,  a

           model  PCB  substrate.   Res.  Commun.  Chem.  Pathol.  Pharmacol.   15:  563-570.

           (Cited  1n  U.S. EPA,  1985)


           YamashHa,  F.   1977.  Clinical features of polychloroblphenyls  (PCB)-lnduced

           fetopathy.  Paediatrician.  6:  20-27.   (Cited 1n  U.S.  EPA, 1985)
 *
*
*» •         Z1nkl,  J.G.   1977.   Skin and  Hver   lesions  1n rats  fed  a polychlorlnated

           blphenyl  mixture.  Arch. Environ.  Contam.  Toxlcol.   5(2):  217-228.   (Cited

           1n U.S. EPA,  1985)
                                              -54.          U.S. Envirorrr ••:' •  r, •'. ,;t;on Agency
                                                            Region V, Li,.,
                                                            230 Soulii E.\- . '    :  ,  ;'32t
                                                            Chicago,  Illinois
 *
V •

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                                                           APPENDIX
                                                    Summary Table for PCBs
en
i
Carcinogenic
Potency
Inhalation
Oral
Species
NA
rat
Experimental
Dose/Exposure
NA
4.42 mg/kg/day
Effect
NA
hepatic
tumors
Unit Risk or
qi*
NO
4.34
(mg/kg/day)-1
Reference
NA
Klmbrough et al., 1975;
U.S. EPA, 1980b
         NA = Not applicable; ND = not derivable

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