United States
              Environmental Protection
              Agency
               Environmental Research
               Laboratory
               Corvallis OR 97333
EPA/600/3-86/033
July 1986
              Research and Development
vvEPA
Ecoregions of the
Pacific  Northwest

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                                            EPA/600/3-86/033
                                            July 1986
     ECOREGIONS OF THE PACIFIC NORTHWEST
                      by

               James M. Omernik
Environmental Research Laboratory --  Corvallis
     U.S. Environmental Protection  Agency
             200 S.W. 35th Street
           Corvallis, Oregon  97333

                     and

               Alisa L. Gallant
           Northrop Services, Inc.
         1600 S.W. Western Boulevard
           Corvallis, Oregon  97333
                          0.'?,  Environmental Protection Agency
                              -n &, Library (5PL-16)
                          '•--» S. Dc-arborn Street,  Boom 1670
                          Ciucago, -Ii   60604

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                                   DISCLAIMER

     The information in this document has been funded wholly by the United
States Environmental Protection Agency.   It has been subjected to the Agency's
peer and administrative review, and it has been approved for publication as an
EPA document.  Mention of trade names or commercial  products does not consti-
tute endorsement or recommendation for use.
                                       11

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                                    ABSTRACT

     A map of ecoregions of the Pacific Northwest has been compiled to assist
managers of aquatic and terrestrial  resources to better understand the regional
patterns of the realistically attainable quality of these resources.   The
ecoregions are based on perceived patterns of a combination of causal  and
integrative factors including land use, land surface form, potential  natural
vegetation, and soils.  Descriptions and photographs are included to provide  a
synopsis of the more significant characteristics affecting and distinguishing
each ecoregion.  A synoptic approach similar to that used to define these
ecoregions is also useful  for applications of the map.   Initial  efforts to use
the framework are at the state level of management and center on identifying
attainable ranges in chemical quality, biotic assemblages, and lake trophic
condition.
                                  111

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                                     PHOTOS

Number                                                                    Page

   1   Coast Range Ecoregion, aerial  photograph at 5,000 feet 	 11

   2   Coast Range Ecoregion, aerial  photograph at 2,000 feet 	 12

   3   Coast Range Ecoregion, ground  photograph 	 12

   4   Puget Lowland Ecoregion,  aerial  photograph at 5,000 feet 	 14

   5   Puget Lowland Ecoregion,  aerial  photograph at 2,000 feet 	 15

   6   Puget Lowland Ecoregion,  ground  photograph 	 15

   7   Willamette Valley Ecoregion,  aerial  photograph at 5,000 feet .... 16

   8   Willamette Valley Ecoregion,  aerial  photograph at 2,000 feet .... 17

   9   Willamette Valley Ecoregion,  ground  photograph 	 17

  10   Cascades Ecoregion, aerial  photograph at 5,000 feet  	 19

  11   Cascades Ecoregion, aerial  photograph at 2,000 feet  	 20

  12   Cascades Ecoregion, ground  photograph  	 20

  13   Sierra Nevada Ecoregion,  aerial  photograph at 5,000 feet 	 21

  14   Sierra Nevada Ecoregion,  aerial  photograph at 2,000 feet 	 22

  15   Sierra Nevada Ecoregion,  ground  photograph 	 22

  16   Eastern Cascades Slopes and Foothills Ecoregion,  aerial photograph
       at 5,000 feet	24

  17   Eastern Cascades Slopes and Foothills Ecoregion,  aerial photograph
       at 2,000 feet	25

  18   Eastern Cascades Slopes and Foothills Ecoregion,  ground photograph . 25

  19   Columbia Basin Ecoregion, aerial  photograph at 5,000 feet  	 26

  20   Columbia Basin Ecoregion, aerial  photograph at 2,000 feet  	 27

  21   Columbia Basin Ecoregion, ground  photograph  	 27

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22   Northern Rockies Ecoregion,  aerial  photograph at 5,000 feet   ....  28

23   Northern Rockies Ecoregion,  aerial  photograph at 2,000 feet   ....  29

24   Northern Rockies Ecoregion,  ground  photograph   	  29

25   Blue Mountains Ecoregion,  aerial  photograph at  5,000  feet   	  31

26   Blue Mountains Ecoregion,  aerial  photograph at  2,000  feet   	  32

27   Blue Mountains Ecoregion,  ground  photograph   	  32
28   Snake River Basin/High Desert  Ecoregion, aerial  photograph  at  5,000
     feet	3
29   Snake River Basin/High Desert  Ecoregion, aerial  photograph  at  2,000
      feet	34

30   Snake River Basin/High Desert  Ecoregion, ground  photograph  	 34

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INTRODUCTION

     Some of the most difficult and seemingly unanswerable problems plaguing
water resource managers center on "regional  representativeness"  of data and
study results, and "attainable" ecosystem conditions or quality.  Spatial
applicability of site specific (or watershed specific) ecosystem quality
studies have long been a subject of controversy.   Processes and  relationships
that are apparent in one part of the country commonly do not appear the same in
other parts.  Interrelationships between natural  and anthropogenic factors
affecting ecosystem quality vary spatially and temporally in such a complex
fashion that mathematical  models developed to predict land use/water quality
relationships are of questionable value at best when used outside the specific
area in which they were developed.  These spatial  and temporal  interrelation-
ships are simply too complex to allow the development of effective quantitative
geographic predictive schemes until a qualitative understanding  of the regional
patterns of the appropriate interrelationships or processes has  been reached.

     Ecoregions"'" of the United States have been mapped to alleviate these
problems and to provide a geographic framework for more efficient management of
aquatic ecosystems and their components.  The ecoregion maps are intended  to
help water resource managers better understand the regional patterns of eco-
system quality and the relative importance of factors that may  be determining
this quality.  Most important, this geographic framework can establish a
logical basis for characterizing ranges of ecosystem conditions  or quality that
are realistically attainable.  Realistically attainable quality  is, of course,
relative to regional patterns of cultural as well  as physical  constraints.  For
most of the conterminous United States, particularly regions of  cultivated
cropland, it is unrealistic to expect an attainable surface water quality  or a
quality of terrestrial conditions of the level possible before  major human
settlement.  What is realistically attainable is  a quality possible given  a set
of economically, culturally, and politically acceptable protective measures
that are compatible with regional patterns of natural and anthropogenic charac-
teri sties.

     The ecoregion maps were compiled at two scales.  A national map was
prepared at a scale of 1:7,500,000, and more detailed regional  maps were
prepared at a scale of 1:2,500,000.  The regional  map of the Pacific Northwest
comprises the first of the larger scale maps and  is the first in a series  of
regional maps covering the conterminous United States.  The maps provide
general information on the spatial extent and characteristics of each eco-
region .
"I" The term ecoregions was coined by J.  M.  Crowley (1967)  and popularized by
  Robert G. Bailey (1976) to define a mapped classification of ecosystem
  regions of the United States.   Unlike many coined words or words used to
  express special  new meanings,  such as "landscape" or "stream order,"  there is
  little disagreement, misunderstanding, or difference of opinion  about the
  meaning of "ecoregion."  Ecoregions are  generally considered to  be regions of
  relative homogeneity in ecological  systems or in relationships between
  organisms and their environments.

                                       1

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BACKGROUND

     Neither the concept of ecological  regions  nor the  attempt  to  delineate
them is new.  Water resource managers and scientists  have  commonly recognized
the existence of regions.  However,  most who have  compiled similar type  maps
have based the boundaries of their regions on a single  geographic  characteris-
tic such as physiographic division or vegetation type.   Although these maps are
helpful in some areas of the country, their usefulness  for resource management
varies from one place to another.

     Part of the difficulty that has been encountered in defining  ecological
regions probably lies in the perception of "region"  and the common misconcep-
tion that, once ecological  regions have been drawn,  they will provide a  power-
ful mechanism for making quantitative assessments  about ecosystems and their
components.  John Fraser Hart's definition of a "region" and its utility is
particularly appropriate to the definition and  understanding of ecoregions.
Hart (1982) stated:

                 The concept of the  region is not  a  powerful
            technical tool, but it is merely a  useful  intellectual
            device for organizing  and presenting information.   A
            region is a more or less homogeneous area that differs
            from other areas.  To  use a more contemporary  jargon,
            within region variance is less than between region
            variance.  The best regions are those  that  are based on
            the greatest amount of interrelatedness.

     Ecological  regions that have  been patterned after  mapped information on a
single geographic characteristic such as physiography or vegetation are  gener-
ally based on understandings of relationships and/or  processes  between these
characteristics and ecosystem components, and inductive logic that these
relationships and processes will reflect useful patterns of ecosystems.
Regional patterns of aquatic ecosystems and their components (physical habitat,
hydrology, water chemistry, and biotic assemblages)  are related to a combina-
tion of characteristics (such as climate, soils, physiography,  and vegetation);
moreover, the relative importance  of each characteristic varies considerably
from one place to another.   This applies regardless  of  the resolution of
regional definition; that is, whether one is dividing the  United States  into
the broadest of regions (say four  to eight), or smaller regions of greater
homogeneity.  Ecoregions at whatever scale of definition should not be based on
single characteristics, nor are there definite  rules  for defining  them.

     There are two notable examples  of classification that have been used for
illustrating ecological regions and  that have incorporated spatial  patterns of
several factors in their design.  The first is  the Land Resources  Regions and
Major Land Resource Areas of the United States  (USDA  Soil  Conservation Service,
1981).  Originally developed by Morris Austin (1965), this map  has since
undergone two revisions.  In general, Land Resource  Regions and Major Land
Resource Areas are characterized by a particular combination of soils, climate,
water resources, vegetation type,  and land uses.  The classification was
designed by the USDA Soil Conservation Service, to aid  in  making decisions

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about national and regional agricultural concerns.  The scales at which the map
units were drawn, and the bias toward agricultural applications, limit applic-
ability of the classification for defining ecoregions.

     The second example is the map of Ecoregions of the United States (Bailey,
1976).  In compiling this map, Bailey considered climate, potential  natural
vegetation, soils, and land surface form.  However, each hierarchical level of
classification of Bailey's ecoregions is based primarily on alignments from a
particular map.  Ecoregion sections (the smallest units shown on Bailey's map)
were, for example, based on map units or groups of map units from Kuchler's map
of potential natural vegetation (Kuchler, 1970).  Ecoregion divisions (two
steps up in size from the section level  on Bailey's map) are, on the other
hand, based on climate (Trewartha, 1943).

     Our approach for defining ecoregions grew out of an effort to classify
streams for more effective water quality management and was inspired by the
philosophies of Warren (1979) as well  as Bailey (1976).  Not wishing to
reinvent the wheel, in the initial stage of this work we used Bailey's eco-
region map as the spatial framework for examining aquatic ecosystems (Hughes
and Omernik, 1981a; Omernik et aK, 1982).  While characterizing the typical-
ness of portions of these ecoregions, we developed what we believe to be a more
satisfactory scheme for defining aquatic ecoregions.   Because the approach is
based on patterns of terrestrial characteristics, the regions are not exclusive
to aquatic ecosystems, but depict terrestrial  ecosystems as well.

     The second stage of our classification work, involving examination of
spatial patterns of stream quality data in several parts of the United States,
reaffirmed the need for an alternative to present schemes for regionalizing
water resource management.  The USDA's Land Resource  Regions were too coarse
and, for most parts of the country, Major Land Resource Areas were too small.
Bailey's ecoregions proved inadequate in most areas,  primarily because of their
dependence on a single mapped characteristic (potential natural vegetation) at
the section level of classification.   Although Hydrologic Units (U.S. Geolog-
ical Survey, 1982) should not be considered for illustrating ecological
regions, they have been used extensively as a geographical  framework for water
resource management.  We found the Hydrologic Units least helpful of any
available spatial classification,  due mostly to the lack of correlation between
topographic drainage areas and the basic causal  and integrative characteristics
that affect or help define regional  patterns in aquatic ecosystem quality.

     None of the available classifications afforded a means to evaluate the
typicalness of areas within ecoregions,  within region variation, or relative
typical ness of entire watersheds.   Such  a mechanism is necessary for evaluating
existing data, for designing sampling schemes to address regional representa-
tiveness and for assessing regional  patterns of attainable aquatic ecosystem
quality.  Ecoregions are based on  patterns of homogeneity in a combination of
geographic characteristics.  Areas within an ecoregion that share all of the
characteristics predominant in the ecoregion are the  most typical portions of
that ecoregion.  The quality of ecosystems in the most typical  areas should be
less variable than in the remainder of the ecoregion  where most (but not all)
of these predominant characteristics  exist.

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     Our approach for defining aquatic ecoregions  is  based  on  the  hypothesis
that ecosystems and their components display regional  patterns that are
reflected in combinations of spatial characteristics  that caused  them to  be
what they are.  These causal  characteristics include  climate,  mineral  avail-
ability (soils and geology),  vegetation,  and physiography.   Each  of these
factors are more or less related to one another,  and  the level  of  interrelated-
ness varies spatially.  We emphasize that the level  to which these factors are
important in determining the character of aquatic  ecosystems also  varies
considerably from one place to another.  Through  analysis of a combination of
small-scale maps of some of the most important causal  and integrative character-
istics (such as land use), distinct patterns of homogeneity in ecosystems can
be perceived.

MAP DEVELOPMENT

     This approach for defining aquatic ecoregions was undertaken  in several
steps (Omernik, submitted to the Annals of the Association  of  American Geo-
graphers, 1986).  The initial  step involved gathering small-scale  maps of
factors that either cause regional variations in  ecosystems, or tend to  inte-
grate causal factors.  The four small-scale maps  found to be most  useful  were:
Land Use (Anderson, 1970); Land Surface Form (Hammond, 1970);  Potential  Natural
Vegetation (Kuchler, 1970); and soils [from varying sources; Simonson (1975)
and USDA Agricultural Experiment Station  of the Wesern States  Land-Grant
Universities and Colleges (1964) were used as component maps for  the Pacific
Northwest].

     The land use map provided a strong integrative tool  for revealing eco-
system patterns in most of the United States because  it reflects  spatial
patterns in potentials and capacities of  the land.  The potential  natural
vegetation (PNV) map was also a strong, integrative tool  because  it synthesizes
a variety of natural and imposed land features.  Hammond's  land surface  form
map synthesizes regional patterns of slope, local  relief, and  profile types
(e.g., whether, and how much,  of the more gently  sloping land  is  located  in the
lowlands or the uplands) into relatively  homogeneous  classes of land surface
form.  The classification scheme of the soils taxonomy map  (USDA  Soil  Conserva-
tion Service, 1970) was probably the most appropriate for our  purposes,  but
because of its inaccuracies [due largely  to the poor  data base and inappro-
priate cartographic techniques used in its compilation (Gersmehl,  1977)], we
relied on other small-scale regional soil  maps.

     Several other maps were consulted, generally to  verify the regional
accuracy of each of the component maps and to further support  the  patterns that
indicated ecoregions.  Those most helpful  for this purpose  were Surficial
Geology (Hunt, 1979), Physical Divisions  (Fenneman, 1946),  and Land Resource
Regions and Major Land Resource Areas of  the United States  (USDA  Soil  Conserva-
tion Service, 1981).  Maps in Climates of the United  States (Baldwin, 1973) and
the Census of Agriculture, Graphic Summary (U.S.  Department of Commerce,  Bureau
of Census, 1969, 1974, and 1978) were also used.

     The second step in developing the ecoregion  map  was to analyze these maps
together to rough out regions that exhibited a relative amount of  spatial
homogeneity in characteristics of soils,  land use, land surface form, and
potential natural vegetation, and to tabulate the  identifying  classes of  each

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(Table 1).  Ecoregion size is a function of the relative spatial  homogeneity in
an area, as compared to adjacent areas, at a scale useful  for regionalizing
resource management.  For our purposes, ecoregions should be large enough to
contain entire topographic watersheds of at least 200^ square miles,  but not so
large as to encompass contrasting relatively homogeneous regions  that, them-
selves, contain entire watersheds of at least 200 square miles.  Some regions
exhibit a patchwork, or mosaic, of conditions that are common throughout the
region -- too patchy to allow individual ecoregions to be delineated  at this
scale of analysis.  The Blue Mountains Ecoregion, for example, is characterized
by contrasts as compared to some adjacent ecoregions.  There, lower elevation
foothill areas consist of mostly sagebrush and grassland steppe widely used for
rangeland.  The higher mountainous portions of the ecoregion support  coniferous
forests, and timber harvest is a major land use.   Nearly all watersheds of 200
square miles and greater in this ecoregion contain a mosaic of these  character-
istics.

     Subsequent steps in map development inlcuded delineating ecoregion bounda-
ries and defining the most typical portions of each ecoregion. The most
typical portions of each ecoregion are those areas sharing all of the charac-
teristics that typify that ecoregion.  The remainder, or generally typical
portions of each ecoregion, share most, but not all of these same characteris-
tics.

     The final step was to color-code each ecoregion to convey a  sense of the
broader, multi-region patterns.  Regions characerized by mostly cropland were
assigned shades of brown and regions characterized by forests were assigned
shades of green or blue.  Grasslands were generally illustrated by shades of
yellow, and very arid areas, shades of pink or red.  Portions of  each aquatic
ecoregion are presented in darker color tones to denote the most  typical  areas
of the ecoregions.  The lighter-toned portions of each ecoregion  represent the
generally typical areas.

APPLICATIONS

     The primary function of the ecoregion map is to provide a geo-
graphic framework for organizing aquatic ecosystem resource information.   This
framework should allow managers, planners, and scientists to better:   (1)
compare the similarities and differences of land/water relationships; (2)
establish water quality standards that are in tune with regional  patterns of
tolerances and resiliences to human impacts; (3)  locate monitoring, demonstra-
tion, or reference sites; (4) extrapolate from existing site specific studies;
and (5) predict the effects of changes in land use and pollution  controls.
Applications will vary from one state and region to another, as do the issues
of concern such as nonpoint source pollution, eutrophication, and water with-
drawals.  Resource management interests and priorities are understandably
different in the Coast Range, than they are in the Snake River Basin/High
Desert, than they are in the Northern Rockies, and so on.   In spite of (or
  The majority of references used in this research presented information  in
  English units of measure.  We have presented our measurements accordingly.

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    perhaps because of) these differences, there is a universal  need to understand
    the spatial  patterns of realistically attainable conditions  and quality of
    aquatic ecosystems.
    
         Initial  efforts to use this ecoregion framework are underway in Ohio,
    Colorado,  Texas, Arkansas,  Oregon,  and Minnesota.  These statewide projects
    focus on aquatic systems, mainly attainable ranges of chemical  quality and
    biotic assemblages.  The applications were designed in a synoptic fashion to
    allow sampling efforts to be organized to:  fit a regional  structure for
    analysis,  comparison, and reporting; select representative  sample sites; sample
    an array of conditions to ascertain the variability within  and  between regions.
    The specific steps required to structure this approach are  typically as
    follows:
    
    1.   Boundaries of ecoregions and their most typical portions were transferred
         to state level (normally 1:500,000-scale)  maps.  The most  typical  portions
         of ecoregions are generally those areas that share all^  of  the characteris-
         tics that are predominant in each ecoregion (Table IT.   The remaining
         portions, generally typical of each ecoregion, share most, but not all, of
         these same characteristics.
    
    2.   Streams and watersheds were selected within the most typical portions of
         each ecoregion in such a way to represent the least impacted conditions
         and a realistic range of stream sizes.  Streams were characterized by the
         relative level of disturbance  in the watersheds, size,''" and the extent to
         which their watershed occupied a particular ecoregion and  most typical
         portion of that ecoregion.  The key to this process was ensuring typical-
         ness  of each stream/watershed  in terms of overall  land  surface form,
         vegetation, soil, and land use characteristics as well  as  a relative lack
         of human disturbance.   For example, one would not expect to find a typical
         stream draining 125 square miles in the Willamette Valley  to be void of
         cropland, or even with less than 75% cropland.  However, there are streams
         of that size in that ecoregion that drain watersheds having relative lack
         of population density and industrial  impact, relatively little stream
         channelization, and a relative abundance of woody riparian vegetation.
    
    3.   Mapping and statistical analyses were performed to determine:   (1) the
         Tegional patterns of attainable quality of aquatic ecosystem characteris-
         TTTcs  of interest; (2)  how these characteristics appear  to  be related to
         Jtream/watershed size; (3) how these characteristics vary  within arid
         bFtween ecoregions; and (4) what regional  properties appear to be most
         TJeHpful  for explaining these variations and enabling more  precise predic-
         tions of attainable quality.The latter (4) involves a synoptic sorting
         technique, incorporating qualitative analyses of mapped characteristics in
      The approximate area upgradient from a point on a stream is  a  more meaningful
      measure of stream size than order,  particularly for small  scale  (state,
      regional,  and national)  analyses (Hughes and Omernik,  1981b; Hughes and
      Omernik, 1983).  Where variations in runoff are greater within or between
      ecoregions, consideration of these variations must be  included.
    
                                           9
    

    -------
         the earlier stages.  Then, as significant regional  associations  become
         apparent, more detailed analyses leading to more quantitative methods
         become appropriate.
    
         Where lakes rather than streams are the  concern, similar  methods apply.
    In the state of Minnesota, for example,  water resource managers  are attempting
    to develop a better capability for predicting attainable nutrient levels  and
    eutrophication in lakes on a regional  basis.   Although the  design is  not
    complete, a summary of the initial  concept is to:
    
    1.   Sort available lake data on nutrient levels and  trophic state by eco-
         region.
    
    2.   Classify the lakes within each ecoregion by as few  characteristics as
         possible to include depth, clarity, and  size.
    
    3.   Categorize each lake by its relative amount of human disturbance, both
         point (e.g., sewage treatment plant) and diffuse or nonpoint (e.g.,  septic
         tanks in close proximity to lake shore).
    
    4.   Map color-coded values for certain  classes of lakes (e.g.,  shallow,  clear,
         small lakes) to enable comparison with spatial patterns of  factors that
         may help explain variations in value.  These factors include surface
         configuration (e.g., moraine vs.  outwash plain), vegetation type (e.g.,
         extent of coniferous, deciduous,  or mixed forest in the watershed),  and
         bedrock type.
    
    ECOREGION DESCRIPTIONS
    
         The ecoregion descriptions are presented here to provide  a  general
    synopsis of the more significant characteristics that affect aquatic  ecosystems
    and their major components.  In short, the descriptions  provide  an amplifica-
    tion of the combination of characteristics listed in  Table  1.  Each description
    presents a brief summary of regional  landscape features, such  as landforms  and
    local and general topographic relief,  followed by information  related to
    ecoregion stream and watershed drainage  characteristics. Subsequent  paragraphs
    outline major vegetation types, major soil  taxa, and  predominant land uses.  A
    final section lists significant disturbance types contributing to stream
    quality degradation.   The descriptions pertain only to the  area  within the
    regional map of the Pacific Northwest."''
    
         The accompanying photos have been taken  in the most typical  portion  of
    each ecoregion and illustrate typical  characteristics rather than extremes  or
    anomalies.  The high oblique photos have been taken at approximately  5,000  feet
    above mean terrain and illustrate general characteristics of the ecoregion.
    The low oblique aerial photos (taken about 2,000 feet above the  streams)  and
    ground photos are of characteristic,  relatively undisturbed streams and their
    riparian zones.
      Descriptions have not been provided for ecoregions  having  only  a  small
      portion of their total  area represented or having only "generally typical"
      area represented in the three-state area of the  Pacific Northwest.
                                           10
    

    -------
         Information included  in  the  ecoregion  descriptions has been obtained from
    a variety of sources  (Archie,  1981;  Anderson,  1970;  Crouse and Kindschy, 1981;
    Franklin and Dyrness, 1973; Hammond,  1970;  Hubbard et aK, 1982; Irwin and
    Hotz, 1981; Kuchler,  1970; NOAA,  1974,  1984;  Rickert et cH .,  1978; Simonson,
    1975; Snavely, 1981;  Tabor, 1981;  USDA  Agricultural  Experiment Stations of the
    Western Land-Grant Universities and  Colleges,  1964;  USDA Soil  Conservation
    Service, 1981; USDA Forest Service,  1970; Weeks,  1970).  Two sources that have
    proven particularly helpful and were heavily  relied upon in the production of
    this text are:  Natural Vegetation of Oregon  and  Washington (Franklin and
    Dyrness, 1973), and Land Resource Regions and Major Land Resource Areas of the
    United States  (USDA Soil Conservation Service,  1981).  Information pertaining
    to stream miles was determined by extrapolating from regionally representative
    7.5 and 15-minute topographic  maps.   Estimates  of watershed sizes necessary to
    support perennial stream flow  were interpolated from USGS Water Resources Data
    and 1:250,000-scale topographic maps.
    
    Coast Range
    
         This ecoregion includes  the  Pacific Coast  Range and coastal valleys and
    terraces (Photos 1-3).  Much  of the  region  is highly dissected by perennial
    streams, commonly having two  to three miles of  streams per square mile.  Local
    relief is between 1,500 to 2,000  feet,  with mountain tops generally below 4,000
    feet.  The combination of maritime weather  systems and high local topographic
    relief results in large differences  in  local  precipitation.  Thus, average
    annual  precipitation  ranges from  55-125 inches  and is heaviest in the winter
    months and lightest in the summer.   Perennial  streamflow is maintained in
    watersheds draining areas of  less  than  one  square mile; however, larger streams
    in this ecoregion drain greater than 300 square miles.   The ecoregion is approx-
               Photo 1, Highly productive, dense Douglas-fir forests, patch clearcuts, and average annual pre-
               cipitation of 55-125 inches are typical of the Coast Range Ecoregion.
                                           11
    

    -------
                                           Photo 2.
    Photos 2 and 3. Clear, high gradient streams with
    high runoff per unit area are common in the Coast
    Range Ecoregion. Average annual runoff in the Siletz
    River is over 100 inches.
                 PhotoS.
                                                          12
    

    -------
    imately 18,000 square miles in size.  Due to the geomorphic character of the
    ecoregion, there are relatively few lakes, generally occurring where coastal
    dunes impound streams.
    
         The Coast Range Ecoregion is densely forested with Douglas-fir, western
    hemlock, Sitka spruce, and western red-cedar.  Lodgepole pine is common along
    coastal dunes.  In the southern portion of the ecoregion, tree species also
    include Port-Orford cedar, California bay laurel, madrone, tanoak, and golden
    chinquapin.  Forest understories are continuous mats of shrub and herbaceous
    vegetation.  Common shrubs include:  salmonberry, rhododendron, willow, vine
    maple, and evergreen huckleberry.  Grasslands, occurring on headlands, repre-
    sent a small percentage of the region.
    
         Soils are developed mainly from sandstone, siltstone, shale, and basalt
    rock sources, and exhibit a wide range of characteristics.  Haplohumults, on
    mountain slopes, foothills, and terraces, and Dystrandepts, Dystrochepts,
    Dystropepts, and Haplumbrepts, on mountain slopes, foothills, coastal plains,
    and terraces are the most common soils in the region.  Tropaquents form in
    coastal lowlands and valleys.  Tropopsamments and Troporthods are formed on
    sand dunes and terraces.
    
         The wood products industry is the mainstay of the ecoregion; harvesting of
    timber has occurred extensively throughout the coastal mountains.  Cultivation
    of forage and grain for dairy cattle occurs locally in larger valleys and along
    the coast.  A small  percentage of cleared land is cultivated for vegetables and
    fruit.  Urban development is concentrated on land bordering water, particularly
    ocean embayments.
    
         Moderate to severe hillslope and stream bank erosion, resulting from
    removal of vegetation by logging, road construction, overgrazing, or cultiva-
    tion is common in many parts of the Coast Range.  Road construction for timber
    removal contributes to landslides and hillslope slumping.  Large woody debris
    in headwater reaches may be transported downstream during major hydro!ogic
    events, causing scouring and increasing stream sedimentation, which in turn can
    lead to loss of habitat for stream biota.  Some streams are chemically affected
    by industrial byproducts from paper and pulp mills, urban development along and
    bordering water, and easy accessibility of stream channels to livestock.
    
    Puget Lowland
    
         The Puget Lowland Ecoregion includes the open hills and tablelands of
    glacial and lacustrine deposits in the Puget Sound Valley (Photos 4-6).   The
    ecoregion is bordered by the Coast Range Ecoregion to the west, the Cascades
    Ecoregion to the east,  and the flatter Willamette Valley Ecoregion to the
    south.   The northern portion of the Puget Lowland ecoregion consists of low
    elevation (sea level  to 500 feet) flats surrounding the Puget Sound area
    interspersed with  high hills, approaching 2,000 feet in elevation.  The south-
    ern and peripheral  portions of the ecoregion contain a greater concentration of
    hills and low mountains, with peaks often exceeding 2,500 feet.  Local relief
    in the more rugged  portions varies from 800 to 1,000 feet.   Due to the rain-
    shadow effect of the mountains bordering the ecoregion to the west, average
    annual  precipitation is generally 35 to 50 inches.   Stream density is typically
    on the  order of one  to  two miles of streams per square mile.   Most streams
    
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    draining the 9,000 square miles comprising the ecoregion are  perennial.   The
    larger inclusive watersheds average 140 square miles, with  permanent stream
    flow emanating from drainages as small as two square miles.   Numerous  small
    lakes dot the landscape.
    
         Most of the land is forested and Douglas-fir is a major  subclimax timber
    species.  Stands of lodgepole pine, western white pine, and ponderosa  pine are
    common forest constituents with Douglas-fir.  Local vegetation  communities
    differing from the major forest vegetation are related to local  variations in
    climate and soil type; examples are prairie, oak woodland,  northwestern  paper
    birch, quaking aspen, Rocky Mountain juniper, and swamp and bog  communities.
    
         The majority of the soils in the northern portion of the ecoregion  are
    formed from glacial materials under the influence of coniferous  forest vegeta-
    tion.  The most common soils are Haplorthods, often associated  with  Xerumbrepts,
    Haplaquepts, Xeropsamments, and Haploxerolls.  Well drained Haplohumults and
    poorly drained Haplaquepts are most common under forest vegetation  in  the
    southern portion of the ecoregion and are derived from upland basic  igneous or
    sedimentary parent materials.  Argixerolls, developed under grassland  vegeta-
    tion, are also common in this portion of the ecoregion, particularly on  terrace
    land forms.
    
         Timber harvest is an important land use in the ecoregion.   Cleared  areas
    support crops of grains, wheat, vegetables, and occasional  irrigated cash
    crops.  Urban development is concentrated along stream and  coastal  stretches.
    
         Municipal  wastes and industrial  wastes from smelters, related metallurgical
    industries, and pulp and paper mills are among the major point  sources of
    pollutants affecting water quality in Puget Sound and many  streams  in  the
    ecoregion.  Water quality is also affected by erosion resulting  from road
    construction and timber harvest.  Urbanization along the Sound  and  its tribu-
    taries imposes direct human impact on water quality through seepage  from
     Photo 4. The numerous
     small lakes and hummocky
     and hilly appearance of
     the Puget Lowland
     Ecoregion attest to
     several epochs of glacial
     deposits. Timber harvest
     is an important land use
     in this region.
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                                                           Photos 5 and 6. Streams in the Puget Lowland
                                                           Ecoregion normally support corridors of dense
                                                           riparian vegetation. Some streams are impacted
                                                           from seepage through septic systems and altera-
                                                           tion of stream banks and stream-side forest.
    Photos.
                 Photo 6.
                                                     15
    

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    individual septic systems and alteration of stream banks  and  streamside  forest.
    An overall reduction in cropland to accommodate growing urbanization  suggests a
    shift in land use emphasis which may further alter water  quality  through
    increased demands on existing wastewater treatment facilities.
    
    Willamette Valley
    
         The majority of this ecoregion comprises the valley  floor  of the
    Willamette River in Oregon and consists of nearly level to  generally  sloping
    floodplains bordered by higher terraces (Photos 7-9).  The  northern portion  of
    the ecoregion extends into Washington, approximately  to the Lewis River.   The
    southern portion extends just south of the South Umpqua River.  The periphery
    of the ecoregion includes foothills grading into the  Coast  Range  Ecoregion in
    the west and Cascades Ecoregion in the east.  Elevation of  the  valley floor
    varies from 100 to 300 feet, and local changes in relief  are  gradual.  Eleva-
    tion of the foothills surrounding the valley increases from 1,000 feet in  the
    northern portion of the ecoregion to greater than 2,000 feet  in the central  and
    southern portions.  Annual precipitation averages 40  inches,  with the northern
    portion of the valley receiving more moisture than the southern portion.
    
         Whereas the majority of streams draining the northern  portion of the
    ecoregion are perennial, most streams draining the southern portion are  inter-
    mittent.  Nearly all of the larger perennial streams  originate  in the more
    mountainous ecoregions adjacent to the Willamette Valley  Ecoregion.   The
    largest streams having watersheds contained entirely  within the ecoregion
    commonly drain less than 100 square miles.  The relatively  few  natural lakes in
    this ecoregion are mainly river meander scars.  Numerous  small  impoundments
    exist for log ponds, irrigation, and livestock.
               Photo 7. Most of the Willamette Valley Ecoregion is in cropland. Due to the normal summer
               drought, irrigation is extensive.
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                                         PhotoS.
    Photos 8 and 9. Streams in the Willamette Valley are
    generally low gradient and meandering. Flooding is
    common in the winter months.
              Photo 9.
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         The natural  forest vegetation of the Willamette  Valley  Ecoregion  is
    represented by Oregon oakwoods (Oregon white oak  interspersed  with  Douglas-fir,
    grand fir, and bigleaf maple)  and mixed stands of cedar,  hemlock,  and  Douglas-
    fir.  Cottonwood, willow,  and  alder are dominant  riparian genera.   Though
    presently limited in area!  extent, prairie vegetation consists of  fescue,
    brome, and sedge.
    
         The principal  soils of the valley floor are  Argixerolls,  with  lesser
    amounts of HaploxerolIs, Haplaquolls, and Albaqualfs.   They  are derived primar-
    ily from alluvial and lacustrine deposits developed  under grassland vegetation.
    Poor drainage is  a common  soil  feature and drainage  tiles assist in removing
    excess water from croplands.  Along the adjacent  uplands  and low hills of  the
    valley, soils are largely  Haplohumults with some  Xerumbrepts,  Haploxeralfs,  and
    Argixerolls.  The upland soils are derived from igneous and  sedimentary parent
    materials.
    
         The ecoregion is typified by an agricultural  mosaic  of  grains, seed crops,
    fruit, mint, berries, and vegetables.  Cropland areas,  interrupted  by  pasture,
    woodland, and forest have essentially replaced prairies,  wetlands,  and many
    wooded areas in this ecoregion.  Subsequent expansion of  urbanization  and
    industry have resulted in  the  reduction of cropland,  woodland, and  forest  land.
    
         Stream quality is affected in a variety of ways  as a result of agricul-
    tural practices,  particularly  irrigation.  Extensive  stream  channelization,
    particularly in the southern half of the ecoregion,  has reduced the number of
    stream miles and  greatly altered the natural  character of the  riparian habitat.
    Numerous impoundments along streams facilitate water  withdrawal  needs  through-
    out the summer drought season, causing a reduction in stream flow.   This can
    result in elevated water temperatures, increased  growth of aquatic  vegetation,
    and barriers to fish migration, reducing the quality  and  quantity  of habitat
    for stream biota.  Runoff carries fertilizer and  pesticide compounds to
    streams.  Erosion of topsoil contributes to stream turbidity and necessitates
    increased use of  soil amendments.  Accessibility  of  stream channels to overuse
    by livestock promotes stream bank erosion and instream manure  deposits.
    
         Urban growth in the valley has increased the demand  for water, the load on
    existing water and sewage treatment plants, and the  number of  point source
    contributors of water pollutants.  Byproducts from pulp and  paper  mills and
    metallurgical industries are major point source contributors of pollutants into
    the Willamette River and its larger tributaries.
    
    Cascades
    
         The Cascades Ecoregion is comprised of the Cascades  Mountain  Range in
    Oregon and Washington and the  Olympic Mountains' of the Olympic Peninsula in
    Washington (Photos 10-12).   The Cascades Range consists of two distinct physio-
    graphic divisions:   the High Cascades, or eastern portion of the range, and  the
    geologically older, more dissected western portion of the range.  The  28,000
    square miles comprising the ecoregion are characterized by high mountains  and
    deeply dissected  valleys.   Elevations range from  near sea level  at the Columbia
    River to greater  than 10,000 feet in the peaks of the High Cascades.  Most of
    the ecoregion is  between 2,000 and 7,000 feet in  elevation,  and local  relief
    often exceeds 3,000 feet.   Average annual precipitation varies across  the
    ecoregion from 50 to 100 inches.
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         Stream density is commonly one and  a  half  to  two  miles  of  perennial
    streams per square mile.  Watersheds  as  small as one to  two  square miles
    support perennial streams, and larger watersheds that  are  contained completely
    within the ecoregion commonly exceed  500 square miles.   Numerous natural  lakes
    are scattered along the crest of  the  Cascades,  resulting from glacial  geo-
    morphic processes.  Man-made reservoirs  are  common at  lower  elevations in the
    ecoregion.
         Most of the area is
    fir, noble fir, Pacific
    hemlock and western red-
    subalpine fir, whitebark
    Forest understories are
    towards sparse in the Hi
    of shrubs such as:  vine
    berry, Oregon grape, and
    flower and a variety of
    grass, fescue, bluegrass
     densely forested.  Typical tree species are:  Douglas-
    silver fir, and western white pine, with western
    cedar providing climax forest cover.  Mountain hemlock,
     pine, and Englemann spruce grow at higher elevations.
    dense in the western portion of the ecoregion, grading
    gh Cascades.  Forest understories are commonly composed
     maple, rhododendron, oceanspray, huckleberry, black-
     sal al.  Forest floors are often matted with twin
    herbaceous species.  Alpine meadows consist of bent-
      and sedge.
         Soils of the Cascades Ecoregion
    are developed from pyroclastic
    materials such as breccias and  tuffs
    and basic igneous rock types, mainly
    pumice, andesite, and basalt.
    Dystrandepts, Haplumbrepts,
    Vitrandepts, Xerumbrepts, Cryorthods,
    Haplorthods, and Haploxerults devel-
    oped from pyroclastic parent mater-
    ials.  Argixerolls, Haplohumults,
    Haplumbrepts, and Xerumbrepts devel-
    oped from igneous materials.  Glacial
    deposits are abundant throughout the
    ecoregion; soils on glacial till are
    Cryandepts, Vitrandepts, Xerochrepts,
    Cryumbrepts, Haplumbrepts, Cryor-
    thods, Fragiorthods, and Haplor-
    thods.
    
         Timber harvest is the major
    industry in the ecoregion.  Wildlife
    habitat and recreation are other
    important land uses.  Some alpine
    meadows provide range for summer
    grazing of livestock and, though
    infrequent, mining for metals is
    important in some areas.
                                           Photo 10. High mountains and deeply dissected valleys are
                                           characteristic of the Cascades Ecoregion. Patch clearcuts are
                                           common.
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                                                                Photos 11 and 12. High annual precipitation and rapid
                                                                snow melt support perennial streams throughout the
                                                                Cascades Ecoregion. Steep valley walls are forested
                                                                with mixed coniferous stands including Douglas-fir,
                                                                pine, hemlock, and red-cedar.
    Photo11.
                 Photo 12.
                                                       20
    

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         Heavy periods of rainfall and rapid  snow melt, coupled with  removal  of
    forest vegetation and road construction for timber harvest, lead  to  increased
    slumping, landslides, and soil erosion.   This is particularly  true of  soils
    developed on pyroclastic materials.  Large woody debris  in headwater reaches
    may be transported downstream during major hydro!ogic events,  causing  scouring
    and disruption of stream habitat for biota.  Public recreation and mining
    create localized disturbances, but are not considered major contributing
    factors to stream degradation.
    
    Sierra Nevada
    
         The Sierra Nevada Ecoregion consists of portions of the Sierra, Trinity,
    Klamath, and Siskiyou Mountain ranges, but this description will  be  limited to
    the extent of this aquatic ecoregion in Oregon.  The region is characterized by
    steeply sloping, highly dissected mountains, narrow valleys with  gently sloping
    floodplains, and alluvial fans bordered by steeply sloping foothills (Photos
    13-15).  Elevation varies from 800 feet to greater than  7,000  feet,  and local
    relief is commonly 2,000 feet.  Average annual precipitation is as low as 18
    inches in some valleys and as great as 85 inches in some mountain locales.
    Most precipitation occurs during the winter months, while summer months are
    predominantly hot and dry.  The ecoregion is 3,000 square miles in size and is
    drained mainly by perennial streams.  Intermittent streams occur along head-
    water reaches and valley floors.  Lakes are infrequent and exist mainly as
    reservoirs.
    
         Mixed coniferous forests with Douglas-fir, ponderosa pine, sugar  pine,
    lodgepole pine, incense-cedar, white fir, and mountain hemlock predominate in
    the ecoregion, but mixed evergreen forests with canyon live oak, California
              Photo 13. Steeply sloping, highly dissected mountains typify the Sierra Nevada Ecoregion.
                                           21
    

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                                     Photo 14.
           Photos 14 and 15. This stretch of the Applegate
           River is typical of the rocky, steep-banked streams
           in the northern portion of the Sierra Nevada
           Ecoregion.
          Photo 15.
    22
    

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    black oak, tanoak, and madrone are common.  Typical  understory species are:
    manzanita, ceanothus,  oceanberry, rose, poison oak,  blackberry,  and Oregon
    grape, accompanied with mats of whipple vine and twin flower.   Prairie vegeta-
    tion includes wild oats, fescue, and brome.
    
         Soils in the ecoregion are derived mainly from  marine volcanic and sedi-
    mentary rocks, many of which have undergone  metamorphosis.  Upland soils are
    principally Haploxerults, generally derived  from sedimentary parent materials.
    Haploxeralfs, derived from metamorphic parent materials,  are less prevalent.
    Also present are Xerochrepts formed from granitic materials, Haploxeralfs and
    Haploxerolls on floodplains and alluvial  fans, and Chromoxererts on alluvial
    fans and bottomlands.
    
         Timber harvest is the principal industry in the ecoregion.   Forests also
    provide public recreation uses.  Livestock graze in  prairies and under open
    stands of timber.  Small grains, hay,  and pasture are grown on the sloping
    parts of valleys for feed for dairy cattle and other livestock.   Truck crops
    and orchards are important in valleys  with adequate  water supply.  Some streams
    are mined for placer deposits.
    
         Because of high potential for soil erodability, the  erosion hazard to
    hillslopes is severe if soils are disturbed  by logging,  overgrazing,  or culti-
    vation.  Landslides and stream bank erosion  are a serious problem and a major
    source of sediment in streams in this  ecoregion.  Placer  mining  causes local
    disturbance to biotic habitat along streams; historically, hydraulic  placer
    mining practices have caused major disturbance to adjacent hillslopes.
    
    Eastern Cascades Slopes and Foothills
    
         The Eastern Cascades Slopes and Foothills Ecoregion  comprises a  transition
    area between the moist, rugged Cascades to the west  and  the variously drier
    ecoregions to the east:  the Columbia  Basin  Ecoregion, the Blue  Mountains
    Ecoregion, and the Snake River Basin/High Desert Ecoregion (Photos 16-18).
    Elevation varies from near sea level (along  the Columbia  River)  to over 7,000
    feet, and local relief varies from 500 feet  to greater than 2,500 feet.  The
    ecoregion is geologically young, with  recent volcanic deposits in evidence as
    lava flows, volcanic vents, and cinder cones.  Bedrock is generally overlain by
    a mantle of pumice and ash.  Average annual  precipitation ranges from 12 to  25
    inches and occurs primarily during winter, spring, and fall  months.
    
         The density of perennial streams  varies greatly throughout  this  14,000
    square mile ecoregion.  Some areas are devoid of perennial stream flow while
    other areas have greater than one mile of perennial  flow  per square mile.
    Large streams, whose watersheds occur  completely within  the ecoregion, also
    vary greatly in size.   Their watersheds are  as large as  35 square miles in the
    northern panhandle of the ecoregion and 2,600 square miles in  the Klamath
    Basin.  Perennial streams drain watersheds as small  as two to  three square
    miles in the mountains; however, perennial flow is seldom found  in watersheds
    of less than 10 square miles in the lower flats.  Numerous springs occur
    throughout the ecoregion and natural lakes are common in  areas of poor drainage
    such as tableland and basin flats.  Reservoirs are numerous and  vary  in size
    from less than one square mile to well  over  50 square miles (e.g., Klamath
    lake).
                                           23
    

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         Forests of ponderosa pine predominate  throughout the ecoregion, and stands
    of lodgepole pine are common.  The  understory  is  generally composed of blue-
    bunch wheatgrass and Idaho fescue.  Green manzanita,  snowbrush ceanothus, and
    bitterbrush are also important understory species.   Sagebrush/wheatgrass steppe
    occurs in the foothills.  Quaking aspen  occurs  in riparian and poorly drained
    wet areas.
    
         The ecoregion is dominated by  large areas  of immature soils developed from
    volcanic ejecta, as well as  soils of more advanced  profile development derived
    from bedrock and glacial deposits.  Xeric soils of  moderate rainfall and mesic,
    frigid, or cryic temperature regimes are predominant, mainly as Inceptisols and
    Mollisols.  Cryorthods and Xerochrepts are  formed from weathered bedrock,
    glacial till, and volcanic debris.  Deep Haploxerolls are derived from glacial
    till and igneous rock.  Cryandepts  and Vitrandepts  are formed from recent and
    weathered volcanic ash and pumice.  Aquolls and Aquepts are formed in basins
    and on floodplains where poor drainage is a factor.
               Photo 16. The Eastern Cascades Slopes and Foothills Ecoregion is characterized by
               relatively gradual slopes, as compared with the adjacent Cascades Ecoregion to the west.
         Timber harvest  is  the  predominant land  use;  much of the forest is used for
    grazing of livestock.   Large  tracts  of the region are designated wilderness
    areas or national  forests,  and  recreation and wildlife habitat are important
    land uses.  A small  percentage  of land,  mainly in valleys, is cropland for
    small grain and  forage.
    
         Soils in this region are highly erodible and potential for stream bank
    erosion is high.   Disturbance from timber harvest and road construction can
    increase hill slope erosion  problems  and increase  stream sedimentation.  In
    
                                            24
    

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                                    Photo 17.
          Photos 17and 18. Forest stands of ponderosa pine
          predominate throughout the Eastern Cascades
          Slopes and Foothills Ecoregion. Timber harvest is the
          principal land use. Bridge Creek at this point drains
          approximately 15 square miles. Perennial flow is
          seldom found in streams draining watersheds smal-
          ler than three square miles in this portion of the
          ecoregion.
         Photo 18.
    25
    

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    grazed areas, livestock  trample  stream bank vegetation and upset stream  bank
    stability.  Precipitation  and  perennial  streams provide adequate water over
    most of the region; however, water  withdrawals for irrigation can deplete
    stream supplies in some  of the valley cropland areas.
    
    Columbia Basin
    
         The Columbia Basin  Ecoregion  (Photos 19-21) is typified by a high degree
    of variability.  Surrounded by a series of high mountain ranges -- the Northern
    Rocky Mountains, the Wai Iowa Mountains,  the Blue Mountains, and the Cascade
    Mountains -- the region  is characterized by deep, dry channels cut into  the
    underlying Columbia River  Basalt formation.  Elevation ranges from less  than
    200 feet (at the Columbia  River) to greater than 4,500 feet (on some mountain
    peaks), and local relief varies  from less than 100 feet to as much as 2,000
    feet.  The landscape is  composed of irregular plains, tablelands with high
    relief, and low mountains. Depending upon location and elevation, average
    annual precipitation is  from 9 to 25 inches.   Extensive loess deposits cover a
    large portion of the ecoregion.
    
         The ecoregion is  34,000 square miles in  size and is drained primarily by
    intermittent and ephemeral  streams.   Most of  the perennial streams originate in
    the adjacent mountainous ecoregions.
    However, perennial flow  can be found
    in some streams draining watersheds
    as small as four square  miles  where
    water is available from  groundwater
    aquifers.  Because of  water with-
    drawal for agriculture and evapo-
    transpiration, these streams are
    commonly ephemeral in  their middle
    and lower reaches (relative to more
    than 20 to 30 square miles).   The
    larger streams that have watersheds
    completely within the  ecoregion  drain
    from 400 to 1,800 square miles.
    Natural lakes are scattered and  few
    in number.  Of the few existing
    reservoirs, most are large and
    located on the major rivers flowing
    through the ecoregion.
    
         The region naturally  supports
    sagebrush/wheatgrass steppe and
    grasslands primarily of  wheatgrass
    with smaller amounts of  bluegrass and
    fescue.
    
           Photo 19. A very large portion of the Columbia
           Basin Ecoregion is in dryland wheat agri-
           culture. Riparian vegetation along drainages
           has largely been eliminated to maximize
           acreage for cropland. Small herbs and shrubs
           remain in drainage channels too deep to plow.
    
    
                                            26
    

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        Photo 20.
    Photos 20 and 21. Channelled scablands form a complex drainage network across portions of
    the Columbia Basin Ecoregion. Irrigated agriculture for pasture, potatoes, and peas occurs in
    the valleys where water is sufficiently available. Diversion of stream water for agriculture often
    results in intermittent streams. Wilson Creek was dry five miles downstream from this photo
    point.
       Photo 21.
                                              27
    

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         Although virtually all soils in the ecoregion have been  formed  under
    grassland or shrub/grassland vegetation, a variety of influences  from  climate
    and parent materials have affected soil formation.  Precipitation is generally
    heaviest along the margins of the ecoregion, and lightest toward  the lower,
    central portion.  Loess deposits cover the basalt formations  in southeastern
    Washington and adjacent portions of Oregon and Idaho.  Xerolls are the most
    common soils in much of the region.  Haploxerolls developed from  loess and
    Argixerolls developed from loess and colluvium predominate on loess  hills,
    mountain foothills, plateaus, and mesas.  Less frequent, are:  Albaqualfs,
    Haploxeralfs, and Xerorthents.  In the bowl-like center of the region, desertic
    soils are dominant.  Camborthids are the most common, but Haploxerolls,
    Xerorthents, Haplargids, Haplaquolls, Torripsamments, and Rockland are also
    present.
    
         Agriculture is the major land use in this ecoregion.  Much of the cropland
    is in dryland wheat, although irrigated farming (mainly for potatoes and  peas)
    is predominant locally.  Cattle grazing occurs where topographic  or  climatic
    conditions are marginal for cultivation of wheat or irrigated agriculture. A
    small acreage is in irrigated vegetables, fruit, and pasture.
    
         Low annual precipitation and water withdrawals for crop  irrigation cause
    many streams to dry up during summer months.  Livestock in and near  streams
    affect water quality both in terms of direct chemical influences  and physical
    habitat degradation.  In many cases, beds of small streams are plowed  and
    re-contoured to increase area for cultivation.  Where streams remain intact,
    riparian vegetation is often removed or thinned to sparse, small  shrub and
    herbaceous species.
    
    Northern Rockies
    
         This ecoregion is comprised of the northern portion of the Rocky
    Mountains.   Rugged, high mountains are the dominant feature  (Photos  22-24).
       Photo 22. Rugged,
       high mountains with
       sharp-crested ridges
       dominate the North-
       ern Rockies Eco-
       region. Local relief is
       commonly 3,000 feet
       or greater.
    

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    Elevations generally vary from 1,300  to 8,000  feet, with  local  relief  commonly
    about 3,000 feet.   Mountains have sharply-crested  ridges  and  steep  slopes  cut
    by steep-walled, narrow stream valleys.  Due to  the vast  differences in  eleva-
    tion and exposure,  average annual  precipitation  ranges  from 20  inches  to
    greater than 60 inches.  Accordingly,  stream density  varies greatly across  the
    41,000 square miles comprising this ecoregion.   Density ranges  from less than
    one, to as much as  three miles of perennial  stream flow per square  mile.  Large
    watersheds that are completely within the ecoregion average 1,200 square miles,
    and perennial flow  occurs in channels draining as  little  as two square miles.
    Lakes are common in higher glaciated  areas.
    
         Coniferous stands of western white pine,  lodgepole pine, western  red-
    cedar, western hemlock, western larch,  Douglas-fir, subalpine fir,  and
    Englemann spruce are common.  Ponderosa pine is  found in  some areas.   Forbs and
    grasses generally represent forest understory  vegetation.   Prairie  vegetation
    consists of wheatgrass, fescue, and needlegrass.
    
         Mountain soils are derived from  acidic  rock types  under  a  frigid  or cryic
    soil temperature regime.  Dominant soils in  the  mountainous portions of  the
    ecoregion are: Cryochrepts and Xerochrepts,  formed on mountain  slopes;
    Cryandepts, formed  on ridges with thin  layers  of volcanic ash;  and  stony
    Cryorthents, formed in areas of rock  outcrops.   Dominant  soils  in valleys  are:
    Calciorthids, Haploborolls, Argiborolls, Natrargids,  and  Cryoborolls,  formed on
    fans, floodplains,  and terraces.
    
         Timber harvest is the major economy, but  wildlife  habitat, recreation, and
    mining are also important land uses.   Open forests at lower elevations are  also
    used for livestock  grazing.  Small acreages  in valleys  are planted  for forage,
    grain, and peas.
    
         Timber harvest and related road  construction  contribute  towards hi 11 si ope
    and stream bank erosion, resulting in large  woody  debris  in stream  channels and
    increased stream sedimentation.  Significant stream disturbance results  from
    placer, shaft, and  open pit metal mining.  Mine  tailings  in and near drainages
    and disturbance or  removal of stream substrates  alter the physical  and chemical
    nature of streams,  affecting habitat quality and quantity for biota.
    
    Blue Mountains
    
         The Blue Mountains Ecoregion is  comprised of  several  mountain  ranges
    separated by fault  valleys and synclinal basins.   The Blue, Ochoco, Wai Iowa,
    Strawberry, and Aldrich Mountains are the major  ranges  (Photos  25-27).  Eleva-
    tion varies from 2,700 feet in low lying valleys to 7,000-10,000 feet  on
    mountain peaks.  Change in local  relief is often from 1,000 to  nearly  3,000
    feet.  Rainfall averages 10 to 20 inches annually  in  lower valleys  and basins,
    and greater than 40 inches in the mountains.
    
         Stream drainage density varies from one and a half to two  miles of  peren-
    nial streams per square mile in wetter areas,  to no perennial  stream  flow  in
    drier areas.  Watersheds of two to three square  miles commonly  support peren-
    nial stream flow in the higher mountains; however, larger catchment areas  are
    necessary to support perennial flow in lower terrain.  Watersheds  as  large as
    4,000 square miles  occur completely within the ecoregion.  Numerous springs are
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    scattered throughout  the ecoregion, and a few clusters  of  alpine glacial  lakes
    occur in the  higher mountains.   Reservoirs are found  on a  moderate number of
    streams.
    
         The mountainous  portions of the ecoregion support  forests  of grand fir/
    Douglas-fir,  ponderosa  pine,  and Englemann spruce/grand fir/Douglas-fir.
    Stands of larch  and lodgepole pine also occur.  Lower elevation areas and
    foothills support  large tracts  of sagebrush/wheatgrass  steppe and wheatgrass/
    bluegrass grasslands.
    
         Soils at moderate  to high  elevations have been formed under forest cover
    and are often derived  from volcanic ash.  Examples are  Vitrandepts and Fragi-
    orthods.  In  the forest/grassland transitional area,  common soils are:
    Haploxerolls, Argixerolls, and  Palexerolls.  At lower elevations, soils under
    shrubsteppe vegetation  are mainly Argixerolls, developed from loess and basic
    igneous rock  materials  or from  ancient lake-deposited sediments.   Haploxerolls
    and Haplargids are of  more limited distribution.
             Photo 25. High mountain ranges separated by faulted valleys characterize the Blue Mountains
             Ecoregion. Higher portions of the ecoregion support forests of fir, Douglas-fir, pine, and
             spruce; lower portions support tracts of shrub/grassland steppe.
         Forested areas  are  used  for timber production, grazing, wildlife habitat,
    and recreation.  Lower elevation shrub and grassland areas are  widely used for
    rangeland.  Meadows  on the  upper mountain slopes serve as summer  grazing
    grounds.  A very small percentage of the area is cropped for forage,  grain, and
    some vegetables.  Many streams  throughout the ecoregion have been mined
    intensively for metals.
    
         Streams and reservoirs supply water for irrigation, domestic use,  and
    livestock use.  Livestock in  and near streams affect stream bank  stability and
    
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      Photo 26.
    Photos 26 and 27. This portion of Desolation Creek, draining about 10 square miles, is typical
    of "least disturbed" streams in the Blue Mountains Ecoregion.
      Photo 27.
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    stream  sedimentation.   Historical  overgrazing has caused a drop in  the water
    tables  and  subsequent  downcutting  of streams in most of the lower valleys,
    accelerating  deterioration of stream bank stability and sedimentation.   Removal
    of vegetation for  timber  harvest contributes toward increased hi 11 si ope
    erosion.  A variety  of past and present mining practices have introduced
    significant disturbance to stream  quality.  Open pit and shaft mines  have had
    localized effects  where tailings were pushed downslope toward drainages.
    Placer  mining,  prevalent  in streams of all sizes, has resulted in major
    physical disruption  of stream beds and biota.
    
    Snake River Basin/High Desert
    
         The Snake  River Basin/High Desert Ecoregion consists of the basin and
    range topography of  southeastern Oregon and the smooth to deeply dissected  lava
    plains  and  plateaus  of southern Idaho (Photos 28-30).  Basin elevations  vary
    from 2,500  to 5,000  feet.   Mountain peaks often exceed 6,000 feet (and are
    greater than  9,000 feet on Steens  Mountain).  Local  relief is low in  the basins
    and plains, but 1,000  feet or greater in the mountains, and 2,000 to  3,000  foot
    differences in  elevation  are not uncommon along some deeply incised stream
    valleys.  Average  annual  precipitation for most of the region is between 8  to
    12 inches per year,  increasing up  to 25 inches on some mountain slopes.
               Photo 28. Basin and range topography characterizes the Snake River Basin/High Desert
               Ecoregion. The ecoregion is vegetated with sagebrush/wheatgrass steppe, providing exten-
               sive rangeland for cattle.
    
    
         The ecoregion is 57,500 square  miles  in size and is drained primarily by
    intermittent and ephemeral  streams.   Most  of the perennial streams in the
    ecoregion are large rivers  originating  in  adjacent,  more humid ecoregions,
    irrigation canals interconnected  with the  large rivers, and small  streams
    originating in the atypical, scattered  mountain ranges.  The small mountain
    streams become intermittent at  lower elevations due  to irrigation, seepage, and
    
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      Photo 29.
    Photos 29 and 30. Crooked Creek is one of the few perennial streams in the lower elevations
    of the Snake River Basin/High Desert Ecoregion. Most of the streams originating in the
    scattered mountain ranges become intermittent or ephemeral when they enter the vast
    basin area.
     Photo 30.
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    evaporation.  Several  lakes have been formed in internally drained depressions,
    and vary in size from less than one square mile to greater than 50 square miles
    (e.g., Lake Abert).  They are shallow and, generally,  saline.   Scattered
    springs occur throughout the region,  as do a moderate  number of reservoirs.
    
         Sagebrush/wheatgrass steppe is the dominant vegetation type throughout  the
    region, sometimes including stands of juniper.   Large  tracts of saltbush/grease-
    wood vegetation also occur.  Some playas and recent lava flows are entirely
    devoid of vegetation.   Streamside vegetation is generally the  same as the
    surrounding regional vegetation, due  to the intermittent or ephemeral nature of
    most streams.  Where perennial  flow does occur, dense  stands of sedges and
    forbs line the riparian zone.  In perennial streams with moderate annual  flow,
    woody vegetation consists of alder, willow, cottonwood,  clematis, rose, and
    mockorange.
    
         Upland soils, developed under shrub and grassland vegetation, are mostly
    Haplargids, Durargids, and Argixerolls,  A yariety of  soils are derived from
    lacustrine deposits.  Well drained lacustrine soils are  Camborthids,  Durargids,
    and Durorthids; poorly drained soils  are Haplaquolls and Haplaquepts.  In the
    northwestern portion of the ecoregion,  extensive pumice  deposits have produced
    Vitrandepts under forest vegetation,  and Camborthids and Haploxerolls under
    shrub-grassland vegetation.
    
         Most of this ecoregion is used as  rangeland, though some  areas  in basins
    or bordering large streams are irrigated for pasture and production  of
    potatoes, corn, alfalfa, sugar beets, and small grains.
    
         Where access by livestock is concentrated, loss or  reduction of  streamside
    vegetation is severe,  causing stream  bank erosion and  sedimentation.   Water
    withdrawals for irrigation often result in completely  dry channels downstream
    from diversions.
    
    SUMMARY
    
         The ecoregion map of the Pacific Northwest is based on the premise that
    areas of relative geographic homogeneity exist  and that  these  areas  can be
    perceived by a simultaneous analysis  of a combination  of causal  and  integrative
    factors including land surface form,  soil, land use, and potential  natural
    vegetation.  The map was compiled to  assist managers of  aquatic and  terrestrial
    resources to better understand regional patterns of the  attainable quality of
    these resources.  An accompanying set of descriptions  and photographs are
    included to provide further characterization of the features distinguishing
    each ecoregion.  More  synoptic analyses, at increasingly larger scales, are
    necessary in order to develop a more  quantitative understanding of the ranges
    of attainable resource quality (within  and between regions) and the  associative
    spatial  factors that permit regional  predictive capabilities.
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    LITERATURE CITED
    
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    Hughes, R. M., and J. M. Omernik.  1981a.  A proposed approach to determine
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    Weeks, Robert A.   1970.   Resources and production of major metals.   Map (scale
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