Environmental Requirements and Pollution Tolerance of Common Freshwater Chironomidae


                                                    EPA-600/4-77-024
                                                    April 1977
ENVIRONMENTAL REQUIREMENTS AND POLLUTION TOLERANCE

           OF COMMON FRESHWATER CHIRONOMIDAE
                            by

                 William M. Beck,  Jr.
           Laboratory of Aquatic Entomology
                Florida A & M  University
              Tallahassee,  Florida 32307
                    Grant No. R802754
                     Project Officer

                 Donald J.  Klemm, Ph.D.
                Aquatic Biology  Section
                Biological Methods Branch
     Environmental Monitoring and Support Laboratory
                  Cincinnati,  Ohio  45268
     ENVIRONMENTAL MONITORING AND SUPPORT LABORATORY
           OFFICE OF RESEARCH AND DEVELOPMENT
          U.S. ENVIRONMENTAL PROTECTION AGENCY
                 CINCINNATI, OHIO  45268

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U.S."Environmental

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                                 FOREWORD

     Environmental measurements are required to determine the quality of
ambient water, the character of effluents, and the effects of pollutants
on aquatic life.  The Environmental Monitoring and Support Laboratory-
Cincinnati conducts research to develop, evaluate, and promulgate methods
to:

    * Measure the presence and concentration of physical, chemical, and
      radiological pollutants in water, wastewater, bottom sediments, and
      solid waste.
    * Concentrate, recover, and identify enteric viruses, bacteria, and
      other microorganisms in water.
    * Measure the effects of pollution on freshwater, estuarine, and marine
      organisms, including the phytoplankton, zooplankton, periphyton,
      macrophyton, macroinvertebrates, and fish.
    #Automate the measurement of the physical, chemical, and biological
    .  quality of water.
    # Conduct an Agency-wide quality assurance program to assure standardi-
      zation and quality control of systems for monitoring water and waste-
      water .

     The effectiveness of measures taken to protect the biological integrity
of the Nation's surface waters is dependent upon our knowledge of the
environmental requirements and pollution tolerance of aquatic organisms
and our understanding of the complex relationships that prevail in aquatic
ecosystems.  This study focuses on one of the most important groups of
insects in aquatic ecosystems—the chironomids (commonly called midges
or bloodworms).  The chironomids are one of the most useful biological
indicators of water quality because of the wide range in their environmental
requirements and habitat preferences, and their abundance, diversity,
cosmopolitan distribution, and distinct morphological differences.  Also,
they form a significant portion of food for many groups of organisms,
especially fish.

     This report is the second in a series of reports in preparation on the
environmental requirements and pollution tolerance of aquatic organisms.
Water quality profiles have been developed to serve as companions to the
EPA biological methods manual and identification manuals to assist biologists
in interpreting field data collected during studies of the effects of pol-
lutants on indigenous communities of aquatic organisms.

                                Dwight G. Ballinger
                                Director
                                Environmental Monitoring and
                                Support Laboratory - Cincinnati
                                     iii

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                                                 ««P  '€ *
                               ABSTRACT
                                  " ! "     , ° .-    • • ,   '-•*.'*.••*    ""  ,       '.

     Data on the environmental requirements and pollution tolerance of
230 taxa of freshwater chironomids were cpmpiled from 33 references.
This compilation'was prepared to assist biologists in evaluating data
from macroinvertebrate samples collected for the assessment of water
quality.  The following parameters were considered:  developmental stage,
pH, salinity, nutrients, degfadable dissolved organics, oxygen, tempera-
ture, turbidity, current, general habitat, specific habitat, seasonal
distribution.                    .           „                            !

     -This report was submitted in fulfillment of Grant No. R-802754 by
Florida A & M University under the sponsorship of the U. S. Environmental
Protection Agency.  Work was completed in November 1975.
                                                                          1;'
                                    lv

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                                 CONTENTS





Foreword                                                                ill




Abstract                                            .                     iv




Acknowledgment                                                           vi




    I    Recommendations                                             ,     1




    II   Introduction                                                     2




    III  Ecological parameters                                            4




    IV   Data compilation                                                10




    V    Chironomids included in this compilation                       JL1




    VI   Selected references for chironomid ecology profiles             21




    "VII  Ecological profiles for 230 taxa of freshwater Chironomidae     24

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                             ACKNOWLEDGMENTS

     It would be virtually impossible to acknowledge fully all those who
have in any way contributed to this project.  Those contributing directly
have my sincere thanks.  Among those are Ms. K. Patricia Fan, Ms. Kehinde
Akinyemi, Ms. Carolee Farmer, Ms. Linda Perez, and Ms. Lorna Radford.
The cooperation of the Aquatic Biology Section, Biological Methods Branch,
Environmental Monitoring and Support Laboratory was gratefully accepted.
                                   vL

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                                SECTION I

                              RECOMMENDATIONS

    This compilation should be used in conjunction with literature for
chironomid identification to help assess water quality.  As additional
information becomes available about a chironomid species, it should be
added to the summary sheet.  It is hoped that this compilation will
become more valuable to the user when used as an active resource.

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                                 SECTION II

                               INTRODUCTION

     The objective of this project is to compile, standardize, and
centralize data on the environmental requirements and pollution tolerance
of the larval forms of the common species of chironomids.  Many of these
species are very widespread both ecologically and geographically, while
others are very narrowly distributed in both regards.

     During recent years, chironomids (DipterarChironomidae) have been
used with increasing frequency to assess the condition of both inland
and coastal waters.  This group is particularly well-suited for this
function for several reasons.

     Many chironomid larvae can be identified to the species level
relatively easily.  Probably no other group of aquatic macroinvertebrates
has had the intensity of study of the immature stages by as many people
as have the chironomids.

     Chironomids are among the most ubiquitous of aquatic invertebrates.
They are found in a wide variety of aquatic habitats, including lakes,
ponds, bogs, marshes, swamps, rivers, creeks, temporary pools, pitcher
plants, bromeliads, tree-holes, and soil and leaf mold.  Even man-made
habitats such as sewage treatment plants, water treatment plants, fish
pools, irrigation diteches, and bird baths have been exploited by chirono-
mids.  In this variety of aquatic habitats, chironomids occupy an even
greater number of different microhabitats and are often dominant.

     Finally, each chironomid taxon is thought to occupy a different
niche in the aquatic ecoystem, responding individually to different
chemical, physical, and biological parameters.  Some species have broad
tolerance ranges and are abundant in a wide variety of habitats and
ecological conditions.  Other species have very limited tolerance ranges
and are restricted to specific habitats favorable to their existence.
By careful analysis of a chironomid community and a knowledge of com-
munity members' ecological "preferences" and tolerances, especially in
conjunction with the other macroinvertebrates, a great deal can be
ascertained about the habitat and environment.

     As various investigators have studied chironomid communities or
macroinvertebrate communities, bits of information concerning the aute-
cology of species have become available.  This information is distributed
throughout many journals and innumerable unpublished reports of various
levels of governmental agencies.

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     In this form, the information is of limited use to those who would
apply it.  One difficulty of the present compilation has been to determine
the reliability of identifications, especially to species, in reports,
published or unpublished, available from a great variety of environmental
agencies.  In other words, it became necessary to determine who was respon-
sible for identification or verification of the organisms in question.  It
speaks well for the quality of environmental work that many of these un-
published reports are among the finest sources of ecological data.

     In the present report, by far the greatest source of information has
been the writer's personal collections and records covering 28 years of
aquatic environmental work.  In addition, the writer has specimens and
records from environmental agencies and universities for which he has done
identifications.

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                                SECTION III

                           ECOLOGICAL PARAMETERS

     Chironomids are affected by many different parameters in the environ-
ment, and it is probably a combination of factors that limit their distri-
bution.  Several investigators have designated "spectral ranges" for indi-
vidual chemical and physical parameters.  Those selected here were recom-
mended by the Aquatic Biology Section, Environmental Monitoring and
Support Laboratory, U.S. Environmental Protection Agency, Cincinnati,
for use with the macroinvertebrates.

                                DEFINITIONS

1.  STAGE;  Stage or stages to which source of data refer:  eggs, larvae,
            pupae, or adults.

2.  pH;  Acidobiontic - occurring at pH below 7, preferring pH below 5.5.

         Acidophilous - occurring at pH around 7, but usually below 7.

         Neutral - best development around pH of 7.

         Alkaliphilous - occurring at a pH around 7, but usually above 7.

         Alkalibiontic - occurring at pH over 7, preferring pH over 8.5.

         Indifferent - common in both acidic and alkaline waters, not pH
                       dependent.

3.  SALINITY;

        Polyhalobous - occurring in salt concentrations of over 40,000
                       mg/1.

        Euhalobous - marine forms occurring in salt concentrations of
                     30,000 mg/1.

        Mesohalobous - brackish water .forms occurring in salt concentra-
                       tions of 500 to 30,000 mg/1.

        Oligohalobous - fresh water forms occurring in salt concentra-
                        tions of less than 500 mg/1.

        Euryhalinous - occurring under a broad range of salt concentra-
                       tions, often encompassing two or more spectra.

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4.  NUTRIENT;

        Eutrophic - characteristic of water with high nutrient concentra-
                    tions.

        Mesotrophic - characteristic of water with moderate nutrient con-
                      centrations.

        Oligotrophic - characteristics of water with low nutrient concentra-
                       tions.

        Dystrophic - characteristics of waters rich in humic materials.

5.  DEGRADABLE DISSOLVED ORGANICS:

        Saprophilic - occurring usually in polluted waters accompanied by
                      brief periods of dissolved oxygen concentrations
                      below 5.0 mg/1 , pH changes of up to 2 units, and/
                      or temperatures exceeding 25°C; but also present in
                      limited numbers in clean water habitats.

        Facultative - wide range  of tolerance to organic pollution; common
                      in both polluted and clean waters.

        Saproxenous - characteristic of clean water habitats but also
                      tolerant of organic enrichment if the dissolved
                      oxygen concentration remains above 5.0 mg/1, and
                      pH and water temperature are not adversely altered.

        Saprophobic - restricted  to waters that have not been exposed  to
                      pollutants.

 6.   OXYGEN;

        Euoxyphilous -  characteristic of water having high 0^ concentration
                        (saturated).

        Mesoxyphilous - characteristic  of water having moderate  02  concen-
                         tration.

        Oligoxyphilous  - characteristic of water  having  low 02 concentra-
                          tion.

        Anoxyphilous  -  facultative anaerobic.

 7.   TEMPERATURE;

         Euthermal - warm-water forms,  usually occurring  at  temperatures
                     greater than 30°C.

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        Mesothermal  -  temperate-water  forms, usually  occurring between
                       15°  and  SO^C.

        Oligothermal - cold-water  forms, usually  occurring between 0° and
                        15°C.

        Stenothermal - occurring over  a temperature range of no greater
                        than 5°C.

        Metathermal  -  occurring over a temperature range of 5° to 15°C.

        Eurythermal  -  occurring over a temperature range of 15°C or greater.

8.  TURBIDITY;

        Eulichotophilous - characteristic of low  turbidities, clear water.

        Mesolichtophilous - characteristic of waters, generally clear,
                            occasionally clouded.

        Polylichtophilous - occurring  at wide range of turbidities.

        Oligolichtophilous - characteristic of high turbidities, murky
                             waters.

9.  CURRENT;

        Limnobiontic - characteristic  only of standing waters.

        Limnophilous - characteristic  of standing water but may be found
                       in running water.

        Indifferent - common in both standing and flowing waters.

        Rheophilous - characteristic of running water but may be
                      found in standing water.

        Rheobiontic - characteristic only of running water.

10.  GENERAL HABITAT;

        Marine -. characteristic of oceans and seas.

        Estuary - characteristic of estuaries and brackish water habitats.

        Lake - characteristic of large inland bodies of standing water.

        Pond - characteristic of small bodies of lentic water.

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10.  GENERAL HABITAT;  (Cont'd)

        River - characteristic of larger flowing waters.

        Stream - characteristic of smaller flowing^waters.

        Spring - characteristic of springs and seepages.

        Other - characteristic of intermittent streams and temporary
                pools and other aquatic habitats (when using this category
                specify type of habitat in notes).

11.  SPECIFIC HABITAT;

        Epibenthic - occurring on, but not penetrating, the substrate
                     and submerged objects.

        Embenthic - occurring in  (penetrating) the substrate and submerged
                    obj ects.

        Epipelic - occurring on (or in) mud and silt.

        Episabulic - occurring on (or in) sand.

        Epilithic - occurring on  (or under) rocks.

        Epixylous - occurring on  (or burrowing into) wood.

        Epizooic - occurring on (or within) animals.

        Epiphytic - occurring on  (or within) plants.

        Attached - normally sessil.

        Unattached - normally free living and capable of  locomotion.

12.  SEASONAL DISTRIBUTION OF MATURE FORMS  (Emergence):

        Winter - sexually mature  form present during winter.

        Spring - sexually mature  form present during spring.

        Summer - sexually mature  form present during summer.

        Autumn - sexually mature  form present during autumn.

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13.  FEEDING BEHAVIOR;

        Predator - feeds on other animal forms.

        Herbivore - feeds primarily on plants.

        Omnivore - feeds on both plants and animals.

        Scavenger - feeds on non-living plant and animal matter.

14.  GEOGRAPHIC DISTRIBUTION - Figure 1.  U.S. Environmental Protection
                               Agency regions.

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                                SECTION IV

                             DATA COMPILATION

     In most instances, it has been necessary to take limited liberties in
the interpretation of authors' comments on chironomid ecology.  Some
authors did not designate spectral categories for some of the parameters.
Often inferences were made and enough information was given that enabled a
designation to be made in the compilation.  An investigator may have
reported individual pH values, but gave no indication of a range of pH
applied to a given species.

     In the column headed "Consensus & Notes", emphasis has been placed
on the latter.  It is felt that both at present and in the future it would
be of greater value for the reader to make his own judgment as to a con-
sensus.  Frequently, there are only two references for a species, one of
which may list that species as a rheobiont and the other as rheophil.  In
these instances, a consensus at present would result in a classification of
"indifferent", which is hardly justifiable.

     This compilation is not intended to be used a rigid definitive assess-
ment of the autecology of the species included.  It should be remembered
that the consensus for any parameter may represent the findings of a
wide variety of investigators.  These individuals may have used different
methods of specimen collection and analysis of ecological parameters.

     Keeping these considerations in mind, the compilation should prove
useful to those employing chironomids in the assessment of water quality.
As more information becomes available on any species, it may be added.
Species not included in this project may be added on blanks provided at
the end of the taxa ecological profiles section.
                                    10
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                                  SECTION V




              CHIRONOMIDS INCLUDED IN THIS COMPILATION





Ablabesmyia aequifasciata Dendy & Sublette




Ablabesmyia americana Fittkau




Ablabesmyia annulata (Say)




Ablabesmyia aspera (Roback)




Ablabesmyia basalis (Walley)




Ablabesmyia cinctipes (Johannsen)




Ablabesmyia hauberi Beck & Beck




Ablabesmyia illinoensis (Malloch)




Ablabesmyia mallochi (Walley)




Ablabesmyia monilis (Linnaeus)




Ablabesmyia ornata Beck & Beck




Ablabesmyia parajanta Roback




Ablabesmyia peleensis (Walley)




Ablabesmyia philosphagnos Beck &  Beck




Ablabesmyia rhamphe Sublette




Ablabesmyia tarella Roback




Arctopelopia flavifrons (Johannsen)




Boreochlus persimilis Johannsen




Brillia  flavifrons  (Johannsen)




Brillia  par (Coquillett)
                                      11
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Brillia parva Johannsen




Calopsectra  confusa  (Malloch)




Calopsectra  dendyi  (Sublette)




Calopsectra  neoflavella  (Malloch)




Calopsectra  xantha  (Sublette)




Cardiocladius obscurus (Johannsen)




Cardiocladius platypus (Coquillett)




Chironomus atreila  (Townes)




Chironomus attenuatus Walker




Chironomus carus  (Townes)




Chironomus chelonia  (Townes)




Chironomus crassicaudatus  (Malloch)




Chironomus plumosus  (Linnaeus)




Chironomus riparius  (Meigen)




Chironomus staegeri Lundbeck




Chironomus stigmaterus Say




Chironomus tentans Fabricius




Chironomus tuxis Curran




Cladotanytarsus Kieffer




Cladotanytarsus viridiventris (Malloch)




Clinotanypus Kieffer




Clinotanypus pinguis (Loew)




Clunio marshalli Stone & Wirth




Coelotanypus concinnus (Coquillett)




Coelotanypus scapularis  (Loew)




Coelotanypus tricolor (Loew)
                                     12
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 Conchapelopia Fittkau

 Corynoneura taris  Roback

 Cricotopus  sp.  I (Beck)*

 Cricotopus  belkini Bendy & Sublette

 Cricotopus  bicinctus  (Meigen)

 Cricotopus  exilis  Johannsen

 Cricotopus  fugax (Johannsen)

 Cricotopus  politus (Coquillett)

 Cricotopus  remus Sublette

 Cricotopus  sylvestris (Fabricius)

 Cryptochironomus blarina (Townes)

 Cryptochironomus curtilamellatus Malloch

 Cryptochironomus fulvus (Johannsen)

 Cryptocladopelma Lenz

 Cryptotendipes Lenz

 Cryptotendipes casuarius (Townes)

 Cryptotendipes darbyi Sublette

 Cryptotendipes emorsus (Townes)

 Demeij erea  atrimanus  (Coquillett)

 Demeijerea  brachialis (Coquillett)

 Demicryptochironomus  vulneratus  (Zetterstedt)

 Diamesa nivoriunda (Fitch)

 Diamesa spinecies  Saether

 Dicrotendipes californicus  (Johannsen)

 Dicrotendipes fumidus Johannsen

*This is a very distinctive species thus far unreared and unnamed.  It
 shows promise as a useful addition to this compilation when identified.

                                     13
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 Dicrotendipes incurvus (Sublette)




 Dicrotendipes leucoscelis (Townes)




 Dicrotendipes lobus (E.G. Beck)




 Dicrotendipes modestus (Say)




 Dicrotendipes neomodestus Malloch




 Dicrotendipes nervosus (Staeger)




 Einfeldia austeni Beck & Beck




 Einfeldia brunneipennis Johannsen




 Einfeldia natchitocheae (Sublette)




 Endochironomus nigricans (Johannsen)




 Epoicocladius flavens  (Malloch)




 Eukiefferiella sp.  Florida




 Eukiefferiella coerulescens  group




 Glyptotendipes barbipes (Staeger)




 Glyptotendipes lobiferus (Say)




 Glyptotendipes meridionalis  Dendy & Sublette




 Glyptotendipes paripes (Edwards)




 Goeldichironomus  holoprasinus  (Goeldi)




 Guttipelopia  currani Beck &  Beck




 Harnischia amachaerus  (Townes)




 Harnischia boydi  (Beck)




 Harnischia edwardsi (Kruseman)



Harnischia galeator (Townes)




Harnischia viridulus (Linnaeus)




Heterotrissocladius Sparck




Kiefferulus dux (Johannsen)




Krenosmittia Thienemann



                                     14
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Labrundinia beck! Roback




Labrundinia floridana Beck & Beck




Labrundinia j ohannseni Beck & Beck




Labrundinia neopilosella Beck & Beck




Labrundinia pilosella (Loew)




Labrundinia virescens Beck & Beck




Larsia decolorata (Malloch)




Lauterborniella agrayloides (Kieffer)




Lauterborniella varipennis (Coquillett)




Leptochironomus nigrovittatus (Malloch)




Lobodiamesa Pagast




Macropelopia decedens (Walker)




Macropelopia hirtipennis (Loew)




Metriocnemus abdomino-flavatus Picado




Metriocnemus hamatus Johannsen




Metriocnemus knabi (Coquillett)




Micropsectra Kieffer




Micropsectra dives (Johannsen)




Micropsectra dubia (Malloch)




Micropsectra nigripila (Johannsen)




Micropesectra polita (Malloch)




Microtendipes pedellus (De Greer)




Monopelopia boliekae Beck & Beck




Monopelopia tillandsia Beck & Beck




Nanocladius alternantherae Bendy & Sublette




Natarsia fastuosa (Johannsen)
                                     15
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 Nilodorum deveineyae (Beck)




 Nilotanypus  americanus  Beck & Beck




 Nimbocera Reiss




 Odontomesa fulva (Kieffer)




 Omisus  pica  Townes




 Orthocladius annectens  Saether




 Orthocladius obumbratus Johannsen




 Pagastiella  orophila (Edwards)




 Parachironomus alatus (Beck)




 Parachironomus carinatus (Townes)




 Parachironomus directus (Bendy &  Sublette)




 "Parachironomus" demeij era  (Kruseman)




 Parachironomus hirtalatus Beck &  Beck




 Parachironomus monochromus  (Wulp)




 Parachironomus pectinatellae  (Bendy  &  Sublette)




 Parachironomus scheideri Beck & Beck




 Parachironomus tenuicaudatus  (Malloch)




 Paracladopelma loganae  Beck & Beck




Paracladopelma undine (Townes)




Paralauterborniella elachista  (Townes)




Paralauterborniella nigrohalteralis  (Malloch)




Paralauterborniella subcincta  (Townes)




Paramerina anomala Beck & Beck




Paramerina smithae (Sublette)




Parametriocnemus lundbeckii (Johannsen)




Paratendipes albimanus  (Meigen)
                                     16
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Paratendipes subaequalis (Malloch)




Paratendipes thermophilus Townes




Parochlus kiefferi (Garrett)




Parorthocladius Thienemann




Pedionomus beckae Sublette




Pentaneura comosa Sublette




Pentaneura inconspicua (Malloch)




Phaenopsectra Kieffer




Phaenopsectra profusa (Townes)




Polypedilum apicatum Townes




Polypedilum aviceps Townes




Polypedilum braseniae (Leathers)




Polypedilum convictum (Walker)




Polypedilum digitifer Townes




PolypedHum fallax (Johannsen)




Polypedilum halterale (Coquillett)




Polypedilum illinoense (Malloch)




Polypedilum labeculosum (Mitchell)




Polypedilum nigritum Townes




Polypedilum obtusum Townes




Polypedilum Ontario (Walley)




Polypedilum scalaenum (Schrank)




Polypedilum simulans Townes




Polypedilum trigonum Townes




Polypedilum tritum (Walker)




Potthastia Kieffer
                                     17
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Procladius  adumbratus  (Johannsen)




Procladius  bellus  (Loew)




Procladius  culiciformis  (Linnaeus)




Procladius  denticulatus  Sublette




Procladius  riparius  (Mailoch)




Prodiamesa  bathyphila  (Kieffer)




Prodiamesa  olivacea  (Meigen)




Psectrocladius elatus  Roback




Psectrotanypus dyari (Coquillett)




Psectrotanypus venustus  (Coquillett)




Psectrocladius vernalis  (Mailoch)




Pseudochironomus aix Townes




Pseudochironomus chen  Townes




Pseudochironomus fulviventris  (Johannsen)




Pseudochironomus richardsoni Mailoch




Pseudodiamesa Goetghebuer




Rheocricotopus robacki (Beck)




Rheotanytarsus exiguus Johannsen




Sergentia coracina (Zetterstedt)




Smittia aterrima (Meigen)




Stempellina Bause




Stenochironomus hilaris (Walker)




Stenochironomus macateei (Mailoch)




Stictochironomus devinctus (Say)




Stictochironomus varius Townes




Sympotthastia Pagast
                                      18
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 Tanypus  carinatus  Sublette

 Tanypus  clavatus Beck

 Tanypus  grodhausi  Sublette

 Tanypus  parastellatus Sublette

 Tanypus  neopunctipennis  Sublette

 Tanypus  punctipennis  Meigen

 Tanypus  stellatus  (Coquillett)

 Tanytarsus  buckleyi Sublette

 Tanytarsus  confusus Malloch

 Tanytarsus  dendyl  Sublette

 Tanytarsus  neoflavellus  Malloch

 Tanytarsus  quadratus  Sublette

 Tanytarsus  recens  Sublette

 Tanytarsus  xanthus Sublette

 Telmatogeton japonicus Tokunaga

 Thalassomyia bureni Wirth

 Thienemannimyia barberi  (Coquillett)

 Thienemannimyia senata (Walley)

 Thienemanlella xena Roback

 Tribelos sp.A  (Beck)*

 Tribelos fuscicornis  (Malloch)

 Tribelos j ucundus  (Walker)

 Tribelos quadripunctatus (Malloch)

 Trissocladius Kieffer
*This is a very distinctive species thus far unreared and unnamed.  It
 shows promise as a useful addition to this compilation when identified.

                                      19
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Xenochironomus rogersi (Beck & Beck)




Xenochironomus scopula Townes



Xeno chironomus taenionotus Say




Xenochironomus xenolabis Kieffer




Zavrelia Kieffer




Zavrelimyia carneosa Fittkau
                                       20
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                                SECTION VI

            SELECTED REFERENCES FOR CHIRONOMID ECOLOGY PROFILES

(23)*  Anderson, J.F., and S.W. Hitchcock.  1968.  Biology of Chironomus
            atrella in a tidal cove.  Ann. Entomol. Soc. Am. 61(6):1597-
            1603.

(14)   Anonymous.  1967.  A biological survey of the Suwannee River Basin.
            U.S. Dept. of the Interior, Federal Water Pollution Control
            Administration, Southeast Region, Atlanta, Georgia,  mimeo;
            17 p.

(17)   Anonymous.  1972.  Flint River Basin Study.  Georgia Water Quality
            Control Board, Atlanta, Georgia,  mimeo, 210 p.

(1)    Beck, Jr., W.M.  Personal collection and files.

(7)    Beck, Jr., W.M. , and E.G. Beck.  1966.  Chironomidae (Diptera) of
            Florida I. Pentaneurini (Tanypodinae).  Bull. Florida State
            Mus. 10(8):305-379.

(33)   Beck, E.G., and W.M. Beck, Jr.  1969.  Chironomidae (Diptera) of
            Florida III. The Harnischia complex (Chironominae).  Bull.
            Florida State Mus, 13(5):277-313.

(6)    Curry, L.L.  1954.  Notes on the ecology of the midge fauna
            (Diptera:Tendipedidae) of Hunt Creek, Montmorency County,
            Michigan.  Ecology 35(4):541:550.

(9)    Curry, L.L.  1965.  A survey of environmental requirements for the
            midge (Diptera:Tendipedidae).   In; Biological Problems in
            Water Pollution, U.S. Dept. of Health, Education, and Welfare,
            Cincinnati, Ohio.  p. 127-141.

(22)   Darby, R.E.  1962.  Midges associated with California rice fields,
            with special reference to their ecology (Diptera:Chironomidae),
            Hilgardia 32(1):1-206.

(28)   Dendy, J.S.  1971.  Phenology of midges in experimental ponds.
            Can. Entomol. 102(3):376-380.
*Number assigned to the "sources" in the Water Quality Profile forms.
                                    21
-------
(26)



(27)




(10)


(21)



(ID



(13)
(25)



(19)




(18)


(5)


(24)
Fagan, E.B., and W.R. Enns.  1966.  The distribution and biology
     of aquatic midges in Missouri lagoons.   (Diptera:Chironomidae).
     Proc. Entomol. Soc. Wash. 68(4):277-289.

Frommer, S.I. , and J.E. Sublette.  1971.  The Chironomidae
     (Diptera) of the Philip L. Boyd Deep Canyon Desert Research
     Center, Riverside Co., California.  Can. Entomol. 103(3):414-
     423.

Gaufin, A.R., and C.M. Tarzwell.  1955.  Environmental changes in a
     polluted stream during winter.  Am. Midi. Nat. 54(1):78-88.

Grodhaus, G.  1967.  Identification of chironomid midges commonly
     associated with waste stabilization lagoons in California.
     Vector Views 14(1):1-12.

lovino, A.J., and F.D. Miner.  1970.  Seasonal abundance and
     emergence of Chironomidae of Beaver Reservoir, Arkansas
     (Insecta:Diptera).  J. Kans. Entomol. Soc. 43:197-216.

Johannsen, O.A.  1937.  Aquatic Diptera Part III. Chironomidae:
     Subfamilies Tanypodinae, Diamesinae, and Orthocladiinae.
     Cornell University Agricultural Experiment Station Memoir
     204:1-84.

Johannsen, O.A.  1937.  Aquatid Diptera Part IV. Chironomidae:
     Subfamily Chironominae.  Cornell University Agricultural
     Experiment Station Memoir 210:1-56.

Kimerle, R.A., and W.R. Enns.  1968.  Aquatic insects associated
     with midwestern waste stabilization lagoons.  J. Water
     Poll. Contr. Fed. 40(2):R31-R41.

Kimerle, R.A., and N. H. Anderson.  1971.  Production and bioener-
     getic role of the midge Glyptotendipes barbipes (Staeger) in
     a waste stabilization lagoon.  Limnol. Oceanogr. 16(4):646-
     659.

Lindeman, R.L.  1942.  Seasonal distribution of midge larvae in
     a senescent lake.  Am. Midi. Nat. 27:428-444.
Morrissey, T.  1950.  Tanypodinae of Iowa  (Diptera).
     Midi. Nat. 43(1):88-91.
III.   Am.
Neel, J.K.  1968.  Seasonal succession of benthic algae and their
     macroinvertebrate residents in a head-water limestone stream.
     J. Water Poll. Contr. Fed. 40(2)Part 2:R10-R30.
                                    22
-------
(8)    Paine, Jr., G.H., and A.R. Gaufin.  1956.  Aquatic diptera as indi-
            cators of pollution in a midwestern stream.  Ohio J. Sci. 56
            (5):291-304.

(2)    Roback, S.S.  1953.  Savannah River Tendipedid larvae (Diptera:
            Tendipedidae (=Chironomidae).  Proc. Acad. Nat. Sci. Phila.
            105:91-132.

(3)    Roback, S.S.  1955.  The Tendipedid fauna of a Massachusetts
            cold spring (Diptera:Tendipedidae).   Not. Nat. Acad. Nat.
            Sci. Phila. 270:1-8.

(4)    Roback, S.S.  1957.  The immature Tendipedids of the Philadelphia
            area.  Monogr. Acad. Nat. Sci. Phila. 9:1-152.

(20)   Sadler, W.O.  1934.  Biology of the midge Chironomus Tentans
            fabricius, and methods for its propagation.  Cornell
            University Agricultural Experiment Station Memoir 173:1-25.

(16)   Stahl, J.B.  1966.   Characteristics of a North American Sergentia
            Lake.  Gewass. u. Abwass. 41/42:95-122.

(32)   Stone, A., and W.W. Wirth.  1947.  On the marine midges of the
            genus Clunio Haliday (Diptera, Tendipedidae).   Proc. Entomol.
            Soc. Wash. 49:201-224.

(15)   Sublette, J.E.  1957.  The immature stages and female of Polypedi-
            lum (Polypedilum) obtusum Townes (Tendipedidae:Diptera).
            Ent. News 68(2):37-40.

(12)    Sublette, J.E.  1964.  Chironomidae (Diptera) of Louisiana I.
            Systematics and immature stages of some lentic chironomids
            of West-Central Louisiana.  Tulane Stud. Zool. 2(4):109-150.

(31)    Wirth, W.W.  1952.  Notes on marine midges from the Eastern
            United States  (Diptera, Tendipedidae (=Chironomidae).   Bull.
            Mar. Sci. Gulf Caribb. 2(1):307-312.

(30)    Wurtz, C.B.,  and S.S. Roback.  1955.   The invertebrate fauna
            of some gulf coast rivers.  Proc.  Acad. Nat. Sci.  Phila.
            107:167-206.

(29)    Usinger, R.L., and W.R.  Kellen.   1955.   The role of insects in
            sewage disposal beds.  Hilgardia 23(10):263-321.
                                    23
-------
Taxon:  Ablabesmyia aequifaaciata  Bendy & Siiblette
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Acidophilous
Neutral
Alkaliphilous
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Indifferent
Polyhalobous
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Olicolialobous
Euryhalinous
Eutrophic
Hesotrophic ,
Olieotroohic
Dvstrophic
Sapronhllic
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Limnobiontic
Liranophilous
Indifferent
Kheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Eoibenclilc
Embenthic
Epipelic
Eoissbulic
Epilithic
Epixvlous
Epizooic
Eoiphvtic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
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Taxon:  Ablabeamyia amerieana  Fittkau

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Taxon:   Ablabesmyia annulata (Say)
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Alkaliphilous
AJkalibiontic
Indifferent
'olyhalobous
Euhalobous
lesohalobous
Dligohalobous
^uryhalinous
^utrophic
Mesotroohic
Olieotrophic
Dystrophlc
Saprophilic
Facultative
Sap roxenous
Saorophobic
Euoxyphilous
Hesoxvnhilous
Otteoxvohilous
Anoxvphilons
Ptlthcrmnl
^feaot-hprmal
Oligothermal
Stenothermal
Hetathermal
Eurythermal
Eulichotophilous
Hesolichtophilous
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Limnobiontic
Limnophilous
Indifferent
Rheophtlous
Rheobiontic
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Estuary
Lake
Pond
River
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Efnbenthic
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Spring
Summer
Autumn
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Region II
Region III
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                               26
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Taxon!   Ablabesmyia  aspera  (Roback)
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Neutral
Alkaliphilous
Alkalibiontic
Indifferent .
Polyhalobous
Euhalobous
lesohalobous
Olieohalobous
iuryhalinous
Eutrophic
•fesotrophic
OliEOtrophic
Dys trophic
Saprophilic
Facultative
Saproxenous
Saprophobic
iuoxvohilous
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Jliaoxvohilous
Anoxvphflnus
Cuthennal
tesnrhRrmal
Oligothermal
Steno thermal
letathermal
Eurythermal
Eulichotophilous
iesolichtophilous
Polylichtophilous
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Limnobiontic
Limnophilous
ndif ferent
Rheophilous
Rheobiontic
Marine
Es tuary
Lake
Pond
River
Stream
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Epibenthic
Embenthic
Enipelic
Eoisabulic
Epilithic
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Unattached
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Summer
Autumn
Predator
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Region II
Region III
Region IV
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Region VI
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                             27
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Taxon:	Ablabesmyia basalis  (Walley)
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Polyhalobous
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Euryhalinous
Eutrophic
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DliEotrophic
Dys trophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Etioxvohilous
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DliEoxvohilous
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Stenothermal
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Eurythermal
Eulichotophilous
Mesolichtophilous
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Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
liver
Stream
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Kmbenthic
Epioelic
Enisabulic
Enilithic
Epizooic
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                                29
-------
laxon:   Ablabesmyia hauberi  Beck & Beck
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Mkaliphilous
UkalibionCic
indifferent
Polyhalobous
Suhalobous
^lesohalobous
Dlieohalobous
Buryhalinous
Eutrophic
^esotrophic
Dlizotrophlc
Dystronhic
Sapronhilic
Facultative
Saproxenous
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Dllgothermal
Stenochermal
Mecatherraal
Eurythermal
Eulicho tophilous
Kesolich tophi lous
Polylicheophilous
OTLigdlichtophilous
Limnobiontlc
Limnophilous
Indifferent
Rheophilous
Uieobiontic
larine
Estuary
Lake
Pond
River
Stream
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Embenthic
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Epilithic
Eplxvlous
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Region II
Region III
Region IV
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Region VI
Region VII
Region VIII
Region IX
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                               30
-------
Taxon!  Ablabeamyia illinoensis   (Malloch)
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Taxon!   Ablabesmyia mallochi   (Walley)
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^arvae
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feutral
Mkaliphilous
Mkalibioncic
[ndifforent
Polvhalobous
luhalobous
•tesohalobous
^lieohalobous
Euryhalinous
EuCrophic
tesocrophic
31iEotrophic
DysCrophic
Saptophilic
Faculcacive
Saproxenous
Saorophobic
Euoxvphilous
^fesoxvnhitous
alicoxvohllous
IVnoxvpM 1 oiis
Etithermnl
fpRorhormfll
Dligothermal
Scenothermal
Mecacherraal
iurytherraal
Eullchotophilous
Mesolichtophilous
Polylichcopliilous '
Oligolichcophilous
LimnobionCic
Limnophilous
Indiffetenc
Iheophilous
UieobionCic
Marine
SsCuary
Lake
Pond
River
Scream
Spring
Other
Epihsnthic
Embenthic
Eoioel.ic
Episabulic
Epilithic
Enixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
qprjn_
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IX
Region X
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Taxon: 'Ablabesmyia monilis   (Linnaeus)
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3upae
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icidobiontic
Vci'dophilous
Neutral
Ukaliphilous
V3kalibiontic
Endifferent
3olyhalobous
^uhalobous
lesohalobous
)liKohalobous
luryhalinous
Sutrophic
lesotrophic
3liROtrophic
^ystrophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Euoxvohilous
lesoxvohilous
Dlieoxvohilous
\noxvphi1 otis
Eu thermal
'•fesorhermat
Jligothermal
Stenothermal
^letathermal
Eury thermal
Eulicho tophilous
4esoiichtophilous
Polylicheophilous ,
Dligolichtophilous
CimnobiontTc
Limnophilous
Indifferent
Rheophilous
Uieobiontic
larine
Estuary
Lake
Pond
River
Stream
Soring
Other
Epibenthic
Embenthic
Epipelic
Eoisabulic
Enilithic
Epixvlous
Epizooic
Epiphytic
Attached

Winter




Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Reeion VII
Region VIII

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Concensus
&
Motes






































































-------
Taxon;    Ablabesmyia ornata  Beck & Beck
-------
Taxon:   Ablabesmyia parajanta  Roback

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Geographical
Distribution
Rgs
arvae
upae
dults
cidobiontic
cidophilous
eutral
Ikaliphilous
Ikallbiontic
ndifferent
olyhalobous
uhalobous
lesohalobous
lisohalobous
.uryhalinous
^utrophic
lesotrophic
)lieotrophic
Jystrophic
Japrophilic
^acultative
saproxenous
Saproohobic
luoxvphilous
lesoxvphilous
Dlisoxvohilous



)ligcthermal
Steno thermal
letathermal
Jurythermal
Eulichotophilous
lesollchtophilous
'olylichtophilous
Dligolichtophilous
Limnobiontic
Limnophilous
IndifEerent
Rheophilous
iheobiontic
larine
Estuary
Lake
Pond
River
Stream
Spring
Other
Eoibenthic
Embenthic
Epipelic
Episabulic
Eoilithic
Enixvlous
Epizooic
Epiphytic
Attached

Winter
Spring
Summer

Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Reeion V
Region VI
Region VII
Region VIII
Reeion IX
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                                35
-------
Taxon:    Ablabesmyia peleensis   (Walley)
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Habitat
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Habitat
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-------
Taxon;    Ablabesmyla phllosphagnos  Beck & Beck


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31igothermal
Stenothermal
leCa thermal
!urythermal
iulichotophilous
> esolichtophilous
?olylichtophilous
)ligolichtophilous
jimnobiontic
jimnophilous
Indifferent
Hheophilous
Rheobiontic
Marine
Estuarv
Lake
Pond
River


Other
Epibenthic
Embenthic

Episabulic
Enilithic


Epiphvtic

irnat-tacbed
Winter '




Herbivore
Omni vo re 	
Region I
Reeion IT

Region IV
Region V
Reeion VI
Region VII
Reeion VIII
Region IX 	 	

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                                57
-------
Taxon:    Ablabesmyia rhamphe   Sublette

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Larvae
Pupae
Adults
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Acldophllous
Neutral
Alkaliphilous
Aakalibioncic
Indifferent
Polyhalobous
Euhalobous
Mesohaloboiis
Oligohalobous
Survhalinous
Eutrophic
Mesotrophlc
Olleotrophic
}ystrophic
Saprophilic
Facultative
Saproxenous
Saorophobic
luoxvphilous
lesoxvphilous
Jliuoxvphilous
Inoxvphiloiis
iUthermal
•lesochprmal
Oligothermal
Stenothermal
fetathermal
Eury thermal
Eulichotophilous
lesolichtophilous
Polylichtopiiilous
Oligolichtophilous
Limnobiontic
Limnophilous
ndifferent
heophilous
Tieoblontio
Estuary
Lake
Pond
River
Stream
Sprina
Dther '
Enibenthtc
Embenthic
Entoellc
Eptsabulic
Epilithic
Bnixvlous
apizooic
Boiohvtic
Attached
Jnattached
•Mnter
Soring
Summar
\utumn
lEredator 1
Herbivore
>mnivore
Scavenger
Region II
Reeton III
Reuton IV
^eeion V
?eaion VI
Region VII
leeion VIII
teeion IX
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                             38
-------
Taxon:    Ablabesmvia tarella Roback
ystrophic .
aprophilic
'acultative
aproxenous
Jaorophobic
^uoxvphilous

)lipoxvnhilous
\n^vvphiloiic;


}ligothermal
3 teno thermal
let a thermal
Surythermal
Sulicho tophilous
lesolichtophilous
'olylichtophilous
Dligolichtophilous
jimnobiontic
.imnophilous
[ndifferent
theophilous
IheoblonCic
nrine
stuarv
.ake
'ond
River
Scream
Snrine
Qt;her
Epibenthic
Embenthic
Epioelic
Eoisabulic
Eoilithlc

Epizooic
Eoinhvtic
Attached
Unattached
; Winter



Predator
Herbivore
Omnivore
Scavenser
Region I
Region II
Region III
Region IV
•Region V
Region VI
Reeioa VII •
Region VIII
Region IX
_ .Region X 	


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                                39
-------
Taxon:  Arctopelopia flavifrons  (Johannsen)

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Habitat
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Habitat
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Geographical
Distribution



Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Euhalobous
Mesohalobous
Oligohalobous
!urvhalinous
Eutrophic
Mesotrophic
OliEOtroohic
]ys trophic
Saprophilic
•acultative
Saproxenous
Saorophobic
luoxvphilous
fesoxvphllous
llieoxvphllouB
inoxvphilniis
iuthermal
tesofhRrmal
Oligothermal
Stenothermal
letathermal
Eurythermal
Eulichotophilous
[esolichtopliilous
olylichtophilous
ligolichtophilous
imnobiontic
imnophilous

heophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Enibenthlc
Embenthic
Entnelic
Episabultc
Enillthic
Epixvlous
Eolzooic
Epiphytic
Attached
Jnttttached
Mnter
Sprinj
Summer
Autumn
Predator
ilerbivore
Omnivo re
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
legion VIII"
^eelon IX
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-------
Taxon:    Boreochlus persimilis  Johannsen.
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Distribution
SRS
arvae
upae
dults
cidobiontic
cidophilous
eutral
Ikaliphilous
Ikalibiontic
ndlfferent
olvhalobous
uha].obous
esohalobous
lisohalobous
uryhalinous
utrophic
lesotrophic
liEOtroohic
)vs trophic
>aprophilic
'acultative
Sap roxenous

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5tenothermal
letathermal
lurythermal
Sulichotophilous
esolichtophilous
'olylichtophilous
Jligollchtophilous
jimnobiontlc
jimnophilous
Indifferent
Uieophilous
Uieobioncic
arine
Estuary
Lake
Pond
River
Stream
Spring
Other
Epibenthic
Embenthic
Epipelic
Episabulic
Epilithic
Epixvlous
Epizooic
Epiohvtic
Attached
Unattached
Winter
Spring


Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IX
Region X 	
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Concensus
&
Notes












































































-------
Taxon:  Brillia flavifrons  (Johannsen)

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Distribution
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Eggs
Larvae
Pupae
Adults
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Acidophilous
Neutral
Alfcaliphilous
Alkalibiontic '
[ndifferent
'olyhalobous
^uhalobous
Mesohalobous
Oligohalobous
^uryhalinous
iutrophic
tesotrophic
Oli^otrop_hic
Dystrophic
Saprophilic
'acultative
Saproxenous
Saprophobic
iuoxvphilous
(esoxvphilous
>lieoxvphilous
inoxvpliiloiis
•uthermal
fesocliprmal
Oligothermal
Stenothermal
letathermal
Eurythermal
Eulicliotophilous
Icsolichtophilous
olylichtopliilous
Oligol ichtophilous
mnobiontic
imnophilous
ndif ferent
heophilous
EUieobiontic
Marine
Escuary
Lake
Pond
River
Stream
Sprinz
Hther
Eoibenthic
Embenthic
EnlDelic
Episobulic
Epilicbic
Epixvlous
Epizooic
Epinhvtic
Attached
Unattached
[«nter
Snrinir
Summer
Allttimn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Recion IV
Rocion V
Reeion VI
Reelon VII
Reeion VIII
Reeion IX
Reeion X
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Concensus
&
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                             Ifi
-------
Taxon!    Brillia  par  (CocmilletO
-------
Taxon: Brlllia parva  Johannsen

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Habitat:
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Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
AJkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
OliKohalobous
luryhallnous
Mesotroohic
Olieotrophic
)ys trophic
Facultative
Saproxenous
Saprophobic
iuoxvphilous
•tesoxvphilous
lllKoxvphllous
moxvpMlntis
•fesnMiormal
Oligotherraal
Stenothermal
[etsthermal
Eurycherraal
esolichtophilous
olylichtopliilous
ligolichtophilous

ndlCferent
neophilous
Rheobiontic
Estuary
Lake
Pond
River
Stream
Sprine
3ther
Eplbenthic
Embenthic
Eptpelic
Bpisabulic
Epilithic
ipixvlous
Spizooic
ipiphvtic
\ttached
Jnattached
•Hnter
Spring
Summer
kueumn
{erbivore
Omnivore
Scavenger
Jecion II
Reeion III
leeion IV
legion -V
teeion VI
teeion VII
ieeion VIII
teeion IX
leeion X
3
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Concensus
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-------
Taxon:  Calopseetra confusa  (Malloch)
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Habitat
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Geographical
Distribution
ECS.
arvae
upae
dults
cidobiontic
cidophilous
eutral
Ikaliphilous
Ikalibiontic
ndifferent
olyhalobous
uhalobous
esohalobous
licohalobous
uryhalinous
utrophic
esotrophic
lieotrophic
)y s trophic
aprophilic
"acultative
Japroxenous
»aprophoblc
juoxvphilous
lesoxvohilous
)liffoxvphilous
iPfiXvphllotis


31igo thermal
3 teno thermal
etathermal
Surythermal
Julichotophilous
lesolichtophilous
'olylichtophilous
Dligolichtophilous
jimnobiontic
jimnophilous
[ndifferent
iheophilous
Uieobiontic
arine
Estuary
Lake
Pond
River
Stream
Sorine
Other
Enibenthic
Embenthic
Enioelic
Eoisabulic
Eoilithic

Epizooic
Epiohvtic
Attached
Unattached
Winter




Herbivore
Omni vo re
Scavenger
Region I
Region II
Reeion III
Region IV
Region V
Region VI
Reeion VII
Region VIII
Region IX
_. .Region X 	
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Concensus
&
Notes













































































-------
Taxon:    Calopsectra deudyi  QSubleCte)

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Habitat:
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Larvae
Pupae
Adults
Acidopliilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
Olieohalobaus
Survhalinous
lesotrophic
Oliaocrophic
Jystrophic
Saproiihilic
Facultative
Saproxenous
Saprophobic
Euoxvphilous
•tesoxvphilous
)licoxvDhilous
LnOXVDhlloilB
;utherm,il
•tesot-liermal
Oligothermal
Stenothermal
tetathermal
Eurythermal
!esolichtophilous
olylichtophilous
ligol ichtophilous
•imnophilous
ndif ferent
heophiious
Elheobiantlc
Marina
Estuary
Lake
Pond
Rivsr
Stream
Sprinz
Dther
Epibenthtc F
Smbenthic
Sninellc
SpisabiUic
inillthic
Soixvlous
5pizooic
ininhvtic
Vttached
Jnattached
JInter
Snrinp
Summer
Aubumn
Predator
lerbivore
Omnivore
Scavenger
URion II
?CKton III
Reelon IV
legion V
tesion VI
(onion VII
(eeton VIII
(eeion IX
!eeion X
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Concensus
&
Notes

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-------
Taxon:    Calopsectra neoflavella  (Malloch)
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13
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Habitat
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Distribution B

Sgs
arvae
upae
dults
cidobiontic
cidophilous
eutral
Ikaliphilous
Ikalibiontic
ndifferent
uhalobous
esohalobous
lieohalobous
uryhalinous
utrophic
esotrophic
liHotroohic
ystrophic
aprophilic
acultative
aproxenous
aoroohobic
uoxvphilous
tesoxvohilous
)li2oxvph.i?°us
tooxvphilnus


)ligothermal
itenothermal
[eta thermal
Zury thermal
lulicho tophilous
tesolichtophilous
'olylichtophilous
Higolichtophilous
j-imnobiontic
jimnophilous
Indifferent
•theophilouB
Uieobiontic
nrine
Sstuarv
Lake
Pond
River
Stream
prine
Other 	
Eplbenthic


Episabulic
Rpilithic

Kplzooic

Attached
Unattached
Winter '

Summer 	
Predafcor
Herbivore
Omnivore 	 ^_
Region I
Region II
Reelon III
Region IV
Reeion V
Reeion VI
Region VII
Region VIII
Region IX 	

28

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Concensus
&
Notes







































































-------
Taxon;_   Calopsectra xantjia   (Sublette)

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uhalobous
tesohalobous
licohalobous
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Stenothermal
letathermal
Surythermal
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lesolichtophilous
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Estuary
Lake
Pond
River
Stream
Spring
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Epibenthic
Embenthic
EpiDe3.ic
Episabulic
Enilithic
Epixvlous
Epizooic
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Upjittached
Winter


Autumn

Herbivore
Omnivore
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Region I
Region II
Region III
Region IV
Region V
ReEion VI
Region VII
Region VIII
Reelon IX
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-------
Taxon!     Cardioeladius  platypus   (Coqulllett)

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Euhalobous
Mesohalobous
)licohalobous
iurvhalinous
iutrophic
{esotrophic
Olieotrophic
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Sapronhilic
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tenothermal
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urythermal
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Lake
Pond
River
Stream
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Epipelic
Episabulic
Eoilithic
Epixvlous
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Unattached
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Spring
Summer
Autumn-
Predator
Herbivore
Omnivore
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Region I
Reqion II
Region III
Region IV
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Concensus
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                             50
-------
Taxon: Chironomus atrella.  (Townes)

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Taxon: Chironomus attenuatus  Walker
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Polyhalobous
tahalobous
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Euryhalinous
Eutrophic
tesocrophic
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Dystrophic
Saprophilic
FaculCaCive
Saproxenous
Saprophobic
Euoxvphilous
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Stenothermal
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Eurychermal
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^lesolichcophilous
Polylichcophilous
Oligolichcophilous
Limnoblontic
Limnophilous
Indifferenc
Rheophilous
Rheobiontic
Marine
iscuary
Lake
Pond
River
Scream
Spring
oeher
Eoibenchic
Embenchic
Eptnelic
Episabulic
Enilithic

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Attached
Unattached
Winter
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Summer
Auttima
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII-
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Region IX
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                               52
-------
Chironomus carus  (Townes)
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Distribution
KRS
arvae
upae
dults
cidobiontic
cidophilous
eutral
Ikaliphilous
Ikalibiontic
ndifferent
olyhalobous
uhalobous
lesohalobous
lisohalobous
^uryhalinous
jUtrophic
lesotrophic
)liKOtroohic
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Saprophilic
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letatherraal
Surytherraal
iulichotophilous
^esolichtophilous
'olylichtophilous
Dligolichtophilous
jimnobiontic
jimnophilous
[ndifferent
iheophilous
yieobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring

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Embenthic
Epioelic
Episabulic
Eoilithic
Eoixvlous
Epizooic
Epiphytic
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Unattached
Winter


Autumn

Herbivore
Omnivore
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Region I
Region II
Region III
Region IV
Region V
Region VI
Region' VII
Region VIII
Region IX
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                       53
-------
Taxon! Chironomus chelonia  (Townes)
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Distribution
ERRS
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
£uhalobous
•fesohalobous
Olicohalobous
Euryhalinous
lutroohic
fesotrciphic
Oligotrophic
Dys trophic
Saprophilic
Facultative
Snproxenous
Saprophobic •
Euoxvphilous
Mesoxvphilous
OlieoxvPhllous
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Stenothermal
Metathermal
Eurythermal
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Polylichtophilous
Qligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Kheophilous
Rheobiontic
Slarine
Estuary
Lake
Pond
River
Stream
Sprinz
Other
Enibenthic
Embenthic
Enipelic
Episabulic
Enilithic
Epixvlous
Epizooic
Epiphvtic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
ReRion I
Recion II
ReRion III
P.eRion IV
ReRion V
ReRion VI
Reeion VII
Region VIII
Reeion IX
Reeion X
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-------
Taxon:    Chironomus  crassieaudatus  (Malloeh)
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Taxon: Chironomus plumosus   (Linnaeus)
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Ukaliphilous
Ukalibiontic
Indifferent
Polyhalobous
Euhalobous
^fesohalobous
Dliffohalobous
Burvhalinous
SutrODhic
^lesoCrophic
Dlizotrophic
Dystrophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Suoxvohilous
Hesoxvphilous
Olicoxvphilous
Anoxvphilons
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Stenothermal
MeCa thermal
Eurythermal
Eulichotophilous
Mcsolichtophilous
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Dligolichtophilous
LimnobionCic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
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Epibenthic
Embentbie
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Eoilithic
Epixvlous
Epizooic
Epiphytic
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Unattached
Winter
Spring
Summer
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Herbivore
Omnivore
Scavenger
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Region II
Region III
Region IV
Region V
Region VI
Region VII-
Region VIII
Region IX
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                                56
-------
Taxon: Chironomus rigarius  (Meigen)
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Geographical
Distribution
Eggs
jarvae
?upae
Adults
Acidobiontic
Acidophilous
Neutral
Ukaliphilous
Mfcalibiontic
Indifferent
'olyhalobous
Duhalobou's
lesohalobous
Olieohalobous
Buryhalinous
Eutrophic
tesotrophic
Olieotrophic
Dystrfiphic
Saprophilic
Tacultati've
Saproxenous
Saprophobic
Euoxvphilous
Mesoxvphilous
Olieoxvphilous
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^uthermal
^eBoMiormal
Oligothermal
Stenotherma.l
Metathermal
Eurythermal
Eulichotophilous
^tesolichtophilous
Polylichtophilous
Dligolichtophilous
Limnobiontic
Liranophilous
Indifferent
Kheophilous
Elheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Epibenthic
Embenthic
Epipelic
Eoisabulic
Epilithic
Epixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V .
Region VI
Region VII •
Region VIII
Reifion IX
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                              57
-------
Taxon: Chironorous staegeri  Lundbeck
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Distribution
Eggs
jarvae
>upae
Wults
icidobiontic
^cidophilous
Neutral
(Ukaliphilous
Mkalibiontic
Indifferent
'olyhalobous
Euhalobous
lesohalobous
Qligohalobous
Suryhalinous
Jutrophic
Mesotrophic
Dligotrophic
Dys trophic
Sapronhilic
Facultative
Saproxenous
Saprophobic
Euoxvphilous
Mesoxyphilous
Olisoxvohilous
Anoxvphilons
Euthermal
^esother-nal
Oligothermal
Stenothermal
Metathermal
Eurythermal
Eulichotophilous
Mesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Snrine
Other
Eoibenthic
Embenthic
Eoipellc
Episabulic
Epilithic
Epixvloiis "'
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn*
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Omnivore
Scavenger
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Region II
Region III
Region IV
Region V
Reeion VI
Region VII
Region VIII
Region IX
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&
Notes

















































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                               59
-------
Taxon: Chironomus tentans  Fabriclus
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Distribution
'•KRS
jarvae
'upae
\dults
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leutral
Mkallphllous
Mkallblontlc
Indifferent
'olvhalobous
^uhalobous
^esohalobous
311eohalobous
Euryhalinous
Eutrophic
^esotroohlc
Dlieotroohic
Dystrophic
Saprophlllc
Facultative
Saproxenous
Saprophobic
'uoxvphllous
Mesoxvphilous
Ollsoxvphilous

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^feqor-hprmal
Oligothermal
Stenothermal
Metathermal
Eurythermal
Bullcho tophllous
Mesollchtophllous
Polylichtophllous
aligolichcophilous
Llmnoblontic
Llmnophllous
Indifferent
Rheophilous
RhcobionCic
Marine
Estuary
Lake
Pond
River
Stream
SprinK_
Other
Epibenthic
Embenthic
Epipelic
Episabulic
Eoilichic
Epixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Reeion II
Reeion III
Reeibn IV
Region V
Resion VI
Reeion VII
Rceion VIII
Region IX
Region X
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Concensus
&
Notes

















































waste-stabilization lagoons



























                               60
-------
Taxon! Chironomua tuxis  Curran
01
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Habitat
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Geographical
Distribution
Eggs
.larvae
Pupae
Adults - . -
Acldobiontic
Acidophilous
Ueutral
Alkaliphilous
Ukalibiontic
Indifferent
Polyhalofaous
Cuhalobous
^lesohalobous
Dlieohalobous
Euryhalinous
Eutrophic
*iesotrophic
Olieotroohic
Dys trophic •
Saprophilic
Facultative
Saoroxenous
Saoroohobic
Euoxvphilous
Mesoxvohilous
Olieoxvphilous
ftnoxvphil ous
Euthermal
^fesothermal
Oligothermal
Steno thermal
Metathermal
Eurytliermal
Eulichotophilous
Mesolichtophilous'
Polylichtophilous
Oligolichtophilous
Limnobiontic
Litnnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Es tuary
Lake
Pond
River
Stream
SorinE
Other
Eoibenthic
Embenchic
Eoinelic
Eoisabulic
Eoilithic
Eoixvlous
Epizooic
Eoiohvtic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Reeion VI
Region VII •
Region VIII
Region IX
Reeion X 	
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Concensus
&
Notes
































































                               61
-------
Taxon:    Cladotanytarsus
                             Kieffer


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Distribution
c
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Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphiloua
Alkalibiontic
Indifferent
?olyhalobous
Euhalobous
lesohalobous
Jlieohalobous
Survhalinous
LUtrophic
Hesotrophic,
Olieotrophic
Dystrophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Juoxvphilous
^fesoxvphilous.
>lieOXVDhilOUS
\noxvphi1otts
Ellthermal
fesothprmal
Dligothermul
Stenochermal
Mecnchertr.nl
Eurythermal
Sullchotophilous
Icsolichtophilous
Polylichtophilous
Dligolichtophllous
Limnobiontic
Llmnophilous
Indifferent
Kheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Eptbenthic
Embenthic
Enipe].ic
Eoissbulic
Enilithic
Enixvlous
Eoizooic
Eninhvtic
Attached
Unattached
HJnter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
ScavenRer
Region I
ReKion It
Reeion III
Region IV
Reeion V
ReKion VI
Reeion VII
Rezion VIII
Reeion IX
Reeion X
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Concensus
.&
Notes

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                              62
-------
Taxon: Cladotanytarsus vlrldiventris (MalloctQ
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-------
Taxon:   Clinotanypus  Kieffer

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Geographical
Distribution
1
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Eggs
Larvae
Pupae
Adults
Acidobiontic
Vcidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohnlobous
Oligohalobous
Euryhalinous
Eutrophic
lesotrophic
Oligotrophic
Dystrophic
Saprophilic
Facultative
Saproxenous
Saorophobic
iuoxvohilous
lesoxvphilous
(linoxvnhilous
Anoxvph-tloiiB
•tlthermal
leBorhermal
Oligothermal
Scenoehermal
letathermal
Eurytherraal
Eulichotophilous
lesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
ndifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sorine
Other
Enibenchic
Embenthic
Eplpelic
Eoisabultc
Eoilithic
Eoixvlous
Epizooic
Epinhvtic
Attached
Unattached
Winter
Spring
Summer
Autumn
Prpdator
Herbivore
Omnivore
Scavenger
ReRion I
Reeion II
Region III
P.ecion IV
Region V
Region VI
Region VII
Region VIII
Region IX
Region X
1

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Concensus
&
Notes










































marsh




-------
Taxon: Clinotanypus pinguis   fl-oew)
to
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Habitat
to
u
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fn OT
Geographical
Distribution
gRS
arvae
upae
dults
icidobiontic
^cidophilous
eutral
dkaliphilous
Llkalibiontic
ndifEerent
'olyhalobous
luhalobous
lesohalobous
)lleohalobous
^uryhalino'us
'utrophic
leso trophic
31ieotrophic
)ys trophic
Saprophilic
?acultative
Saproxenous
Saproohobic
Suoxvphilous
•lesoxvphilous
Dlicoxvohilous



Dligothermal
Stenothermal
letatharmal
Jury thermal
Eulichotophilous
'•lesolichtophilous
Jolylichtophilous
DligoJ ichtophilous
Limnobiontic
jimnophilous
Indifferent
^heophilous
Iheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Eoibenthic

Enipelic
Eoisabulic
Epilithic
Epixvlous
Epizooic
Epiohvtic
Attached
Unattached
Winter

Summer
Autumn
Predator
Herbivore
Omni vo re '

Region I
Region II
Region III
Region IV
Region V
Region VI
Reeion VII
Region VIII

Reeion X 	
?.R

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Concensus
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Notes


























,














































-



                                 65
-------
Taxon:   Clunlo marshalll  Stone & Wirth

<
J
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1 Salinity
g
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Taxon!    Coelotanypus coneinnus   (Coqulllett)
0)
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Distribution
ggs
arvae
upae
idulcs
Lcidobiontic
Icidophilous
eutral
Ukaliphilous
Ukalibiontic
Cndifferent
'olyhalobous
'uhalobous
lesohalobous
)lieohalobous
:uryhalinous
^utrophic
lesotrophic
)lieotrophic
)ystrophic
Saprophilic
facultative
Saproxenous
Saprophobic
Suoxyphilous
lesoxvphilous
}lieoxvDhilous

F.nthermal
lesot-hermal
Dligothermal
Stenothermal
teta thermal
!ury thermal
Bulichotophilous
Hesolichtophilous
Polylichtophilous
)ligolichtophilous
Limnobiontic
jimnophilous
Indifferent
Rheophilous
Jheobiontic
larine
Estuary
Lake
Pond
River
Stream
Spring'
Other
Eoibenthic
Embenthic
EpipeJ.ic
Episabulic
Epilithic
Epixvlous
Eoizooic
Epiphytic
Attached
Unattached
Winter



Predator
Herbivore
Omnivore
Scavenger
Region I
Reeion II
Reeion III
Region IV
Region V
Region VI
Region VII
Reeion VIII
Reeion IX
Reeion X
1

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Concensus
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                                67
-------
Taxou;	Coelotanypus  scapularls  (Loew)

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Habitat
ee
1
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Geographical
Distribution


Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral .
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
Olieohalobous
Survhalinous
Eutrophic
•lesotronhic
Olieotroohic
Dystrophic
Saprophilic
Facultative
Saproxenous
suoxvphilous
iesoxvohilous
Jlicoxvphilous
inoxvphlloiiB
EuChermal 1
fesothermal
Oligo thermal
Steno thermal
letathermal
EurytHerraal
Eulichotophilous
lesolichtophilous
olylichtopliilous
ligolichtophilous
limnophilous
ndifferent
heophilous
EUieobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Dther
Epibenthic
r,mbenthic
Eplpelic
Eoisabulic
Enilithic
Epixvlous
Epizooic
Epiplivtic
Attached
Unattached
«nter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Recion II
Reeion III
Reeion IV
Reeion V
Reaion VI
Region VII-
*etion VIII
Reeion IX
Reeion X
1
~X~







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X


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X
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Concensus
&
	 	 Notes









































-------
Taxon:    Coelotanypus tricolor  (Lowe)

•a nj
-------
Taxon;    Conchapelopia  Fittkau

a)
e
t-
VI
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Isalinicy
c
4
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Habitat
Specific
Habitat
ff
I
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T 0
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L Geographical
Distribution
c
EKfis
Larvae
Pupae
Adults
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Euhalobous
Mesohaldbous
Oligohalobous
Suryhalinous
Eutrophic
Mesotrophic
OliEOtrophic
Jystrophic
Saproohilic
Facultative
Saproxencus
Saprophobic
iUOXVDhilOUS
fesoxvnhilous
(llBOxvohilous
Anoxvphflous
Euthermnl
fesorfiBmwl
Oligothermal
Stenothermal
letathermal
Eurythermal
Eulichotophilous
lesolichtophilous
Polylichtophilous
Oligolichtophilous
.imnophilous
ndifferent
heophilous
Rheobiontic
Estuary
Lake
Fond
River
Stream
Snrine
Other
Eoibenthtc
Erobenthic
Entpelic
Episabulic
Epllichlc
Epixvlous
Eoizooic
Eoinhvtic
Attached
Unattached
Winter
Snr-tnp
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Reeion I
Res-ion II
Reelon III
Region IV
Region V
Reaion VI
Recion VII
Reeion VIII
Reeion IX
Reeion X
1

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X
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Concensus
&
Notes











































                             70
-------
Taxon:   Corynoneura taris  Roback
olylichtophilous
Diigolichtophilous
jimnobiontic
jimnophilous
CndifEerent
Iheophilous
iheobiontic
Marine
Es tuary
Lake
Pond


ipring_ 	
<3ther 	
Enifaenthic


Eoisabulic
Eoilithic


Epinhvtic
Attached 	
Unattached 	
Winter




Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Reeion V
Reeion VI
Region VII
Region VIII
Reglon..TX 	

1

X



X







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X
X

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Concensus
&
Notes











































































                               71
-------
          Taxon;    Crieotopus sp.  I   (Beck)*

fl>
c
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w



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&


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o n
a



KEBS
Larvae
PupaQ
Adults
Acidobiontic
Acidophilous
Neutral
Alkallphilous
A]kallbiontlc
Indifferent:
Polyhalobous
Euhalobous
Mesohalobous
Olieohalobous
Etaryhalinous
Eu trophic
Mesotronhic
Olieotrophic
Dystrophic
Saprophilic
Facultative
Saproxenous
Sanrophobic
Juoxvphilous
tesoxvphilous
JliBoxvnhllous
Anoxvphilons
•uthermal
teBOI-hnrmal
Oligo thermal
Stenothermal
letathernal
Eurytherraal
Eulichotophilous
[esolichtophilous
Polylichtophilous
Oligolichcophilous
Limnobiontic
Limnophilous
Indifferent
Kheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Epibenthtc
Embenchic
Eptpelic
Episabulic
Epilithic
Enixvlous
Epizooic
Epiphvtic
Attached
[Tnattaclied
Winter
Spring
Summer
Autumn-
Herbivore
Omnivore
Scavenger
Region I
teEion II
?eelon III
P.eElon IV
teeion V
?eelon VI
leeion VII
(eaion VIII
teeion IX
?eeion X
1

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Concensus
&
Notes


—










































































*This is a very distinctive species 'thus far unreared and unnamed.
promise as a useful addition to this compilation when identified.

                                      72
It shows
-------
Taxon:  Cricotopus belkini   Bendy S Sublette


AC
4J
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Distribution
0)
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fa
Eggs
Larvae
Pupae
Adults
Acidobtontic
Acidophilous
Neutral
Alkaliphilous
A]kalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
Oligohalobous
Suryhalinous
Eutrophic
fesotrophic
Oligotrophic
Dys trophic »
Sapronhilic
Facultative
Saproxenous
Saprophobic
Juoxvphilous
Mesoxvphilous
Olieoxvohilous
Anoxvphilous .
Suthermal
Mesot-hnrmal
Oligothermal
Stenothermal
Hetathermal
Eurythermal
Eulichotophilous
Hesolichtophilous
Polylichtophilous
OligoJ ichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Enibenthic
Embenthic
Epipe3.ic
Eoisabulic
Eoilithic
Epixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII •
Region VIII
Reeion IX
Region X
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                            '75
-------
Taxon;    Crlcotopua bleinetus  (Melgen)


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Larvae
Pupae
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Acidophilous
Neutral
Alkaliphilous
Alfcalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
Olicohalobotis
Euryhalinous
Butrophic
fesotrophic
Jlieotrophic
Dystrophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Suoxvphilous
Mesoxvphilous
)lieoxvphilous
Anoxvphlloiis
SnthermflT
Mpsoth^rmal
Dligothermal
Stenothermal
He ta thermal
Eurytherraal
EulichoCophilous
Mesolichtophilous
PolylichCophilous
Dllgolichtophilous
Limnobiontic
Liranophilous
Indifferent
Rheophilous
EUieobiontic
Marine
Estuary
Lake
Fond
River
Stream
Spring
Other
Enibenthic
Embenchic
Eplpelic
Enisabulic
Enilithic
Enixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Reeion III
Recion IV
Keeion V
Reston VI
Reeion VII
Reeion VIII
Reeion IX
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-------
Taxon:  Cricotopus exilis  Johannsen
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Taxon;   Cricotopua fugax  (Johannsen)


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Eggs
Larvae
Pupae
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Polyhalobous
Euhalobous
Hesohalobous
Oligohalobous
£uryhalinous
Eutrophic
Hesotroohic
Olieotrophic
Dystrophic
Saprophilic
Facultative
Saproxenous
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fesoxvphilous
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Eurythermal
Eulichotophilous
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Embenthic •
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Attached
Unattached
Winter
Sorfnfl
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Residn I
Region II
Region III
P.egion IV
Region V
Reeion VI
Region VII
Region VIII
Reeion IX
Reeion X
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•
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                              76
-------
Taxon;    Cricotopus politus   (Coqulllett)
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-RRS
^arvae
upae
Adults
teidobiontic
teidophilous
leutral
akaliphilous
Ukalibiontic
Undif ferent
'olyhalobous
3uhalobous
lesohalobous
3uryhalinous
jUtrophic
lesotrophic
Dligotrophic
3vstronhic
Saprophilic
Facultative
Saproxenous
Saorophobic
Euoxvphilous
'•fesoxvphilous
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Euthermal
fesothprmal
Dligothermal
Steno thermal
teta thermal
Eury thermal
Eulichotophilous
Mesolich tophi lous
Polylichtopliilous
Oligolichtophiious
Limn ob ion tic
Limnophilous
Indifferent
Rheophilous
Jheobiontic
Marine
istuary
Lake
Pond
River
Stream
Spring
Other
Eoibenthic
Embenthic
Epioelic
Eoisabulic
Eoilithic
Epixvlous
Epizooic
Epiphytic
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Unattached
Winter

Summer

Predator
Herbivore
Omnivore
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Region I
Region II
Reeion III
Region IV
Region V
Resion VI
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Region VIII
Region IX
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                                77
-------
Taxon:	Criootopus remus  Sublette

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Acidophilous
Neutral
Alkaliphilous
A3kalibiontic
Indifferent
Polyhalobous
Euhalobous
llesohalobous
OliRohalobous
luryhalinous
Eutroohic
•lesotroohic
OliEOtroohic
)ys trophic
Saprophilic
Facultative
Saproxenous
Saoroohobic
•fesoxvDhilous
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Lnoxvphlloiis
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resol-hprmsl
Oligothermal
Steno thermal
^eta thermal
Eurythermal
Euiichotophilous
7es^olichtophiious
olylichtophilous
ligolichtophilous
-imnophilous
ndifferent
heophilous
leobiontic
Marine
Estuary
Lake
Pond
Wver
Scream
Sorine
Ither
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voioelic
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vnilithic
ioixvlous
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ioiohvcic
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Summer
kuttimn
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lerbivore
Omnivore
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lesion II
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teeion VIII
Lesion IX
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                            7B
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Taxon:  Cricotoous svlvestris.  (Fabricius)
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Mesohalobous
Oligohalobous
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Hesotronhic
Ollgotrophic
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Stenothermal
letathermal
Eurythermal
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ndifferent
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leobiontic
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Estuary
Lake
Pond
River
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Enlzooic
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Unattached
Winter
Soring
Summer
Autumn
Predator
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Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
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-------
Taxon:    Cryptochironomus  curtilamellatus  Malloch
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Distribution
Eggs
jarvae
?upae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
[ndifferent
Polyhalobous
Suhalobous
lesohalobous
Dligohalobous
Euryhalinous
Sutrophic
Mesotrophic
Oligotrophic
)ystrophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Euoxvphilous
Mesoxyphilous
Olicoxvohilous
Anoxvphilous
Euthermal
Mesotrhermal
Oligo thermal
Stenothermal
Hetathermal
Eurythermal
Eulichotophilous
Mesolich tophi lous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheojihilous
Rheobiontic
Marine
Estuary .
Lake
Pond
River
Stream
Sprine
Other
Enibenthic
Embenthic
Epioelic
Episabulic
Enilithic
Epixvlous
Epizooic
Epiohvtic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III •
Region IV
Region V
Region VI
Region VII •
Region VIII
Reeion IX
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                              81
-------
Taxon:	Cryptochironomus fulvus   (Johannsen)
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Distribution
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>upae
Wults
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^cidophilous
teutral
Ukaliphilous
Mkalibiontic
indifferent
Polyhalobous
tahalobous
^fesohalobous
Dlicohalobotis
Eurvhallnous
^utrophic
^lesotrophic '
^liROtrophic
Dys trophic
Saprophilic
Facultative
Saproxenous
Saprophobic
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^noxvphll ous
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Stenothermal
Metathermal
Eurythermal
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klesolichtophilous
Polylichtoptiilous
Qligollchtophilous
Liranobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
tarine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Epibenthlc
Embenthic
Enipelic
Eptsabulic
Epilithic
Epixvlous
Epizooic
Epiphytic
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Unattached
Ulnter
Sprlnt>
Summer
Autumn
Predator
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Omnivore
ScavenRer
Region I
Rezion II
Region III
Region IV
Region V
Region VI
Reeion VII
Region VIII
Region IX
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-------
Taxon: Cryptocladopelma   Lenz
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Habitat
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Geographical
Distribution
Eggs
Larvae
Pupae
Adults
Acidoblontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
Olinohalobous
Euryhalinous
Eutrophlc
tfesotrophic
Oligotroohic
Dys trophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Euoxvohilous
tesoxvphilous
ttieoxvphilous
Anoxvphilous
Suthermal
•feso thermal
Oligothermal
Steno thermal
Hetathermal
Eurychermal
Eulichotophilous
Mesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent:
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Eoibenthic
Embenthic
Epioelic
Episabulic
Epilithic
Epixvlous
Epizooic
Eoiohvtic
A'ttached
Unattached '
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
.Region IX
Region X
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Concensus
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                              83
-------
Taxon:
          Cryptotendipes  Lenz
s
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Taxon:    Cryptotendipes casuarius   (Townes)
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Habitac
Specific
Habitat
60
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Geographical
Distribution
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
A3kalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
Olicohalobous
Euryhalinous
Eutroohic
besotrophic
Olieotroohic
Ltys trophic •
Saprophilic
Facultative
Saproxenous
Saoroohobic .
iuoxvphilous
•lesoxvnhilous
Olisoxvohilous
Anoxvphilous
Suthermal
'tesot-hprmal
Oligothermal
Stenothermal
Metathermal
Eury thermal
Eulichotophiious ,
^ esolichcophilous
Polylichtophilous
Dligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
SprinE
Dther
Eoibenthic
Embenthic
Eoioelic
Eoisabulic
EDilithiC
Epixvlous
Epizooic
Eoinhvtic
Attached
Unattached
Winter
Sorine
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
.Region III
Region IV
Region V
Region VI
Region VII •
Region VIII
Region IX
Region X
33

X



X







X

X


X



X

x



X



X

X





X




X
X
X



X







X
X
X

X


X




x













































































































































































































































































































































































































































































































































































































































































































































Concensus
s
Notes J















































                             85
-------
Taxon;   Cryptotendioes darbyi,  Sublette
s
14
1

u
(9
u
V*


~
-•»


5
5
ft
tn
u

a
s
3
•3
S
. S

IN
0



1
1

•
•0
S
1
H

%
z
%
s
General
Habitat
Specific
Habitat
tli
u
I
33
•ss
S.S
Geographical
Distribution
Eggs
jarvae
>upae
Adults
(Vcidobiontic
Acldophilous
(eutral
MkalAphilous
Ukalibiontic
Indifferent
'olyhalobous
tahalobous
^tesohalobous
Dligohalobous
Euryhalinous
Eutroohic
^lesotroohic
Oligotroohic
)ystrophic
Saprophilic
Facultative
Snproxenous
Sarroohobic
Euoxvohilous
"•lesoxvphilous
OlieoxvpKilous
\noxvptinonq
Ruthermal
tesnhhpmml
Dligo thermal
Scenothermal
letathermal
Euryclietmol
iUlichotophilous
lesolichtophilous
Polylichtoplillous
31igollchtophilous
Uimnobiontic
Limnophilous
Indifferent
Kheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sorinz
Other
Eoibenthlc
Embenthic
Eotoelic
Eoisabulic
Eoilithic
Eoixvlous
Eoizooic
Eoinhvtic
Attached
Unattached
HJnter
Spring
Summer
Autumri
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Resion IX
Reeion X
22

X




X






X

X





X



X


X


X


X


X







X





X







X


X













X








































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes
















































-------
Taxon:	Cryptptendipes emorsus  (Townes)
QJ
U
M
3
&
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EC
£


^
4J
5
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a
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rQ
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§

U
General
Habitat
Specific
Habitat
ti-
M
00 O
c *H
•H >
•o td
0) £
m o
FEI pq
Geographical
Distribution
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
iluhalobous
lesohnlobous
OllRolialobous
Suryhalinous
Sutrophic
•lesotroohic
OliRotrojDhic
lys trophic
Saprophillc
Facultative
Saproxenous
Saprophobic
Euoxvohilous
Mesoxvnhilous
Olieoxvphilous
Anoxvphilous
Eufchermal
leBotheyinal
Oligothermal
Stenoehermal
Mecntliermnl
Eurytharmai
Eulicho tophlloue •
Mesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
istuary
Lake
Pond
River
Stream
Spring
Other
Enibenthic
Kmbenthic
EnipeliQ
Episabulic
Eoilithic
Epixvlous
ICpizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
P*egion IV
Region V
Region VI
Region VII
Region VIII
Region IX
Region X
28

X





X





X.

X





X


X



X



X

X


X







X




X








X

X
X








X













































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes






















































-------
Taxon!  Cemeljerea atrimanua  (Coquillett)
3
O
W
«
g
•1



s
• ?


>
il
5
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en
^
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a
s
0
B
S
£

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0



§
H


•B
3
1
H

S
i
s
3
General
Habitat
Specific
Habitat
60
H
23
3§
U X
££
Geographical
Distribution
BSES
[.arvae
'upae
Wults
\cidobiontic
teidophilous
feutraL
Mkaliphilous
Mkalibiontic
Indifferent
Polyhalobous
Euhalobous
tesohalobous
Dligohalobous
Euryhalinous
Eutrophic
^lesotrophic
31ieotrophic
Dystrophic
Saprophilic
Facultative
Saproxenous
Ssprophobic
Euoxvphilous
(esoxvnhilous
Dli^oxvphiloua
^pQxyph•^ OHK
Kuthermal
fpsnt-tiprmal
Dligo thermal
Steno thermal
'leta thermal
Eurythermal
Eulichotophilous
[csoliciitophilous
Polylichtophilous
Jligolichtophilous
Limnobiontic
Limnophilous
Indifferent
ttheophilous
Rlicobioncic
•larine
istuary
Lake
Pond
River
Stream
Spring
Other
Enibenthic
Embenthic
F.plpelic
Eoisabulic
Eptlithic
Epixvlous
Eptzooic
Eoiphvtic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Reeion I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Reeion IX
Reeion X
28

X





X





X







X


X



x



X

X


X







X




x








X


X








X

































.





















































































































































































































































































J





















































































































































































































































































































































































































Concensus
&
Notes


























- - •







































-------
Taxon:   Demeilerea brachialls  (Coquillett)
0)
; g
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01
§
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1
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0
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&
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g>5
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•o rt
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h 0
Geographical
Distribution
-BBS
uarvae
»upae
Wults
\cidobiontic
\cidophilous
Neutral
Mkaliphilous
Mkalibiontic
Indifferent
polyhalobous
iuhalobous
'lesohnlobous
Hiaolialobous
3uryhalinous
iutronhic
lesotrophic
Jlieotrophic
Dystrophic
Saoronhilic
Facultative
Saproxenous
Saoronhobic
Euoxvohilous
fesoxvphilous
DliBoxvohilous
\noxvphi1 oits
EuthermAl
lesttthormal
Dligo thermal
Stenothermal
[etathermal
Eurythermal
Eulicho tophilous
^lesolichtophilous
Polylichtophilous
Dligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Kheophilous
Rheobiontic
Marine
Estuary
Lake •••'
'ond
River
Stream
Sorine
Other
Enibenthic
Embenthic
Eninelic
Eoisabulic
Epilithic
Epixvlous
Epizooic
Epinhvtic .
Attached
Unattached
Winter
•Spring
Jium^er

Predator
Herbivore
Omni vo re
Scavenger •
Region I
Region II
Reeion III
Region IV
Reeion V
Reeion VI
Reeion VII
Reeion VIII
Reeion IX
_ Region X 	
28

X





X





X







X


X



X



X

X


X







X




X








X


X
X







X













































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes










































































-------
Taxon t___ Demicryptochironomus  vulneratus  (Zetterstedt)

-------
Taxon:    Diamesa nivoriunda.   (Fitch)


61
U



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d


>

5
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AJ
ti
5
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u
c
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fi

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>H
S
H

S
0)
s
<->
General
Habitat
Specific
Habitat
ftC
I.
Feeding
Behavior
Geographical
Distribution
-------
Taxon:    Diamesa spinecies   Saether
«
CJ
s
o
CO

oc
u




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u
5
iH
en
s
g
h
s
	 o
c
s
s

-------
Taxon:    Dicrotendlpes  californicus  (Johannsen.)
tu
CJ
)-l
i
CO
0)
oc
u



t


^
U
5
•3
en
JJ
^
S
• s
a
G
S
0

^
0



|
1


V
3
S
s

S
-------
Taxon:	 Dicrotendlpes fumidus  Johannsen
01
a
i

ef
jj



s



3
5
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£
£
|
s
S
£
s
S
o

A1
0


i
H
^



s
s
H

u
0
«
o
General
Habitat
Specific
Habitat
ec
M
eo o
C *ri
ss
s-g
FEI m
Geographical
Distribution
SEES
^arvae
?upae
Adults
&cidobiontlc
ftcidophilous
Neutral
Mkallphilous
Mkallbiontic
EndiEferent
'olyhalobous
iuhalobous
^[esobnlobou's
DliKohalobous
Eurvhallnous
Eutrophlc
rtesotrophic
OlieotroDhic
Dys trophic
Sapronhilic
facultative
Snproxenous
Saprophobic
Euoxvphllous
Mesoxvphllous
Olipoxvphllous
^oxvphilnits
RuHiermnl
^lesot-Tiprmal
Oligothermal
Steno thermal
Metattiermal
Eurythermal
Culichotophilous
Mesoiichtoptiilous
Poly lich toph ilous
Oligol ichtophilous >
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
•larine
Estuary
Lake
Pond
River
Stream
Sorine
Other
Enibenthic
Embenthic
Enioelic
Episabulic
Eolllthlc
Tipixylous
Eoizooic
Epiphvtic
Attached
Unattached
Winter
Spring
ftumijier
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Reeion II
Recion III
Reeion IV
Region V
Reeion VI
Region VII
Region VIII
Region IX
Region X
11

X











X


X








X-



X

X





X






X





X








X


X










X











































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes













































































-------
Taxon:
           Dicrotendipes incurvus   (Sublette)
0)
i

1
u
w


a


4J
5

CO
£
1
i!
s
u
c
«
14
o

£?
°



§
0}


13
s
M
H

u

B
o
General
Habitat
Specific
Habitat
M
M
M O
K -H
•H >
•a rt
-------
Taxon:    Dicrotendipes leucoscelis  (Townes)
0)
o
I
W

1
u




a



4J
I
3
w
AJ
I
M
.P
li
c
s
s

N




§
-------
Taxon:
           Dicrotendipes lobus   (E.G. Beck)
a
o
M
3
O
w

£
•U



33
a



j->
5
r-i
VI
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1
M
3.
1
•g
S
g

ci

.


§
1


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1
H

£
01
y
3
General
Habitat
Specific
Habitat
AC
14
0}
1
II
T3 (0
0) A
&&
Geographical
Distribution
gss
arvae
upae
dults
icidobiontic
tcidophilous
eutral
Llkaliphilous
llkalibiontic
ndifferent
olyhalobous
.uhalobous
lesohalobous
liEohalobous
luryhalinous
lutrophic
lesoCroohic
JliKOtroohic
)ys trophic
Saproohilic
^acultative
Saproxenous
Saprophobic
jlioxvoHilous
lesoxvphilous
Dlieoxvphilous
Vnoxvphilotis
jitthermal
•Ie*5nthermal
)ligothermal
Stenochermal
letathermal
5ury thermal
Sulicho tophilous
lesolich tophi lous
Polylichtophilous
Dligolichtophilous
Ljimnobio'ntic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Eoibenthic
Embenthic
Epipelic
Episabulic
Epilithic
Epizooic
Epiphytic
Att-aphed
Unattached
Winter
s prtng
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IX
Reeion X 	
1

X



X







X

X


X



X

X



X



X
X







X





X


X







X

X









X




































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes












































































                                97
-------
Taxon:    Dicrotendipes modestus  (Say)


tt




a
0



5
5

in
S
0)
H
5
a
c
3


C4




1
£


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2
S
H

tJ

§
0
General
Habitat
Specific
Habitat
M
Jil
-o ta
0) X
0) 
-------
Taxon:   TH<;rnt:gnt»pag neoTimdestus	Malloch
0)
o
}J
3
o
w

u
u
M



s

	
4J
a
^
VI
4J

"S3
S^
Geographical
Distribution

ggs
arvae
upae
dults
cidobiontic
cidophilous
eutral
Ikaliphilous
Ikalibiontic
olyhalobous
uhalobous
esohalobous
ligohalobous
uryhalinous
u trophic
leso trophic
ligotroohic
)vstrophic
>aprophilic
"acul'tative
japroxenous
Saprophobic
"uoxvphilous


\nQxvphi1oiis


Higo thermal
5 teno thermal
letathermal
•Xirythermal
iulichotophilous
esolichtophilous
'olylichtophilous
Dligolichtophilous
..imnobiontic
jimnophilous
[ndifferent
iheophilous
Jheobiontic
arine
Estuary
Lake '
Pond


Spring
Other


Epipe3.ic
Episabulic
Epilithic


Epiphvtic
Attached
Unattached
Winter




Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV

Region VE

Region VIII
Region IX 	

1

X










X


X








X



X

X





X






X






X







X


X










?


































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes




























'















































                                   99
-------
Taxon;    Dierotendlpes nervosus  (Staeger)

s
R
•u
w
5
.,,
•H
C
«
en
S
Tl
4J
U
C
3
°

•sj
££
L Geographical
Distribution
j
L
Eggs
Larvae
Pupae
Adults
Acidoblontic
Acidophilous
Neutral
Alkaliphllous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
leSohalobous
Olisohalobotts
^uryhalinous
^utrophic
fesotrophic
Oligotrophic
)ys trophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Juoxvphllous
(esoxvnhilous
llisoxvohilous
Anoxvph lions
!uthennal
teBortiprmal
Oligothermal
Steno thermal
letathermal
Eurytherraal
Eulichotophilous
lesolichtophilous
Polylichtophilous
Oligolichtophilous
tironobiontic
Limnophilous
ndifferent
Rheophtlous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
3ther
Enibenthic
Embenthic
Rpipelic
Bpisabulic
Eoilithic
Enixvlous
Eplzooic
Eoiphvtic
Attached
Unattached
Winter
Soring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Reeion IX
Resion X
1

X







X



X

X


X


X


X







X

X





X



X

X
X


X








X
X
X
X
X


X




X
X





q

X





X





X


















X








X '


X





X








X

X
X
X









X




17

X






X




X










X



X






X

X






X






X







X


X










X




11

f











X











X



X

X





X







X




X








X


X













X

n

r




X






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X





X



X


X


X


X


X







X




X








X
X
X
X
X







X






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X





x





X

X





X


X



X



X

X


X




































































































































































































































































































































































.Concensus
&
Notes















































                             100
-------
Taxon:
          Einfeldia austeni  Beck & Beck
0)
o
s
o
(O

ec
JJ




CL



4J
5

w
JJ
§
M
3
o
c
3
g

e-i
O



§
IH


tJ
•e
H

u
(U
y
o
General
Habitat
Specific
Habitat
C£
1
w
CO O
C -H
•H >
*a nj
s-s
fn M
Geographical
Distribution
'sss
jarvae
Jupae
Vdults
^cidobiontic
^cidophilous
•eutral
Vlkaliphilous
Ukalibiontic
Lndifferent
aolyhalobous
^uhalobous
lesohalobous
)liEohalobous
'uryhalinous
^utrophic
lesotroohic
Dlieotroohic
)ys trophic
Sapronhilic1
Facultative
aaproxenous
Saorophobic
Euoxyphilous
-lesoxvohilous
3lieoxvDhilous
\noxvphilous
Eu thermal

3ligothermal
Stenothermal
letathermal
Surythermai
Eulicho tophilous
vlesolichtophilous
Polylich tophilous
Dligolich tophilous
Limnobiontic
^imnophilous
IndifEerent
ttheophilous
Rheobiontic
•larine
Estuary
Lake
Pond
River
Stream
Snrine
Other
Epibenthic
Embenthic
Kpioelic
Eoisabullc
Eoilithic

Epizooic
Epinhvtic
Attached
Unattached
Winter
Spring


Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Reeion V
Reeion VI
Reeion VII
Reeion VIII
Region IX
_Reaion_JC 	
1

X





X




X


X




x-



X



X



X





X




X







X







X


X




x



•X

y











































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes









































































                              101
-------
Taxon;  Einfeldia brunneipennis  Johannsea

6)
tt
ca
u
CO
33
ft




5
i-t
CO

-------
Taxon:    Einfeldia natchitocheae   (Sublette)
QJ
O
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2
O
en

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Taxou;    Endoehironomus  nigricans   (Johannsen)

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-------
Taxon:    Epoicocladius flayens  (Malloch)
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Geographical
Distribution
RRs
arvae
upae
dults
cidobiontic
cidophilous
eutral
IkaLiphilous
Ikalibiontic
ndif ferent
olyhalobous
uhalobous
esohalobous
ligohalobous
uryhalinous
utrophic
lesotrophic
Hieotroohic
)ystrophic
Japrophilic
"acultative
Japroxenous
Japro phobic
^uoxvphilous

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Stenothermal
letaCherraal
Jury thermal
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iesolichtophilous
Jolylichtophilous
Jligolichtophilous-
^mnobiontic
jimnophilous
[ndif ferent
*heophilous
•Uieobiontic
arine
Es tuary
Lake
Pond
River
Stream
Spring
Other
Epibenthlc
Embenth,ic
Enipelic
Epi^abulic
Epilithic

Epizooic
Epiphvtic
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Unattached
Winter




Herbivore
Omni vo re
Scavenger
Region I
Region II

Region IV
Region V
Region Vt
Region VII
Reeion VIII

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Concensus
&
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This species lives attached
to burrowing mayflies.














5


                                105
-------
Taxon;  Eukiefferiella sp. Florida

i
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c
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Habitat
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eographical
istribution
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Eggs
Larvae
Pupae
Adults
Acidoblontic
Acidophilous
Neutral
Alkaliphilous
Alkalibioncic
Indifferent
Polyhalobous
Euhalotuous
Kesohalobous
Olieohalobous
lurvhalinous
lesotrophic
Olieotroohic
)ys trophic
Saprophilic
?acultative
Saproxenous
Saorophobic
luoxvohtlous
lesoxvnhilous
llicoxvohilous
inoxvphllons
!utherma1
lesot-hnrmal
Oligothermal
Stenothermal
tetathermal
Eurythermal
Eulichotophilous
lesolichtophilous
Polylichtophilous
Oligolichtophilous
.imnophilous
ndif ferent
Rheophtlous
heobiontic
Marine
Estuary
Lake
Pond
River
Stream
Snrine
Other
Enibenthic
Embenthic
Enioelic-
Eoisabulic
Enllithic
Eoixvlous
Eoizooic
Eoinhvtic
Attached
Unattached
Sorine
Summer
Aucrnnn
Predator
Herbivore
Omnivore
Scavenger
Region II
Seeton III
*6R Ion IV
Region V
teuton VI
Reeion VII
legion VIII
\eeion IX
leeion X
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                            106
-------
Taxon; Eukiefferiella coerulescens group




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Concensus
&
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                                107
-------
Taxon; Glyptotendipes barbipes  (Staeger)

J
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Habitat
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Habicac
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Larvae
Pupae
Adults
Acidophilous
Neutral
Alkaliphilous
Alkallbioncic
Indifferent
Polyhalobous
Euhalobous
tfesohnlobous
Olicohalobous
5urvhalinous
Eutrophic
Mesotrophic
Oligotrophic
Jystrophlc
Saprophilic
Facultative
Saproxenous
Saprophobic
Euoxvphilous
tesoxvohilous
lllEoxvnhilous
mnxvphlloiis
.uthermql
Tesot-hprmal
Oligothermal
Stenothermal
ictathermal
Eurythermal
Eulichotophllous
lesolichtophilous
olylichtophilous
ligolichtophilous
.imnophilous
ndifEerent
heophilous
RheobiAntic
Marine
Estuary
Lake
Pond
Uver
Scream
Sprine
Jther
iPibenthic
inbenthic
-.ntnelic
loisabullc
Snllithic
inixvlous
!olzooic
ininhvtic
Attached
fnattached
Hnter
Snrtn^
Summer
Autumn
'redator
lerbivore
)mnivore
Scavenger
RORton 1
Reeion II
ieelon III
SeElon IV
teelon V ,
teeion VI
Region VII
(eeion VIII
ieeion IX
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Concensus
&
Notes













































-------
Taxon:
           Glvptotendipes loblferus   (Say)
10
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Habitat
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Habitat
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p^
Geographical
Distribution
Sggs
jarvae
>upae
Adults
teidobiontic
Icidophilous
eutral
akaliphilous
klkallbioneic
ndifferent
'olyhalobous
luhalobous
lesohalobous
)liEohalobous
luryhalinous
3utrophic
fesotrophic
)liKOtrophic
)ys trophic
Saprophilic
Facultative
Sap roxenous
5aprophobic
Suoxvphilous
^esoxvphilous
31ieoxvphilous
\noxvphil OHS
Euthermal
^feBot-bermal
31igothermal
Stenothermal
'•let a thermal
Eurythermal
Eulichotophilous
^lesolichtophilous
Polylichtophilous
Jligolichtophilous
Linuiobiontic
Limnophilous
Indifferent
Rheophilous
Uieobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Enibenthic
Embenthic
Epipe] ic
Epissbulic
Eoilithic
Eoizooic
Epiphytic
Attached
Unattached
liinter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region- I
Reeion II
Region III
Region IV
Region V
Reeion VI
Region VII
Region VIII
Reg-ion IX
Region X
1

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Concensus
&
Notes



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                                109
-------
Taxon: Glyptotendipes meridionalis  Dendv & Sublette

0)
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u
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General
Babltat
Specific
Habitat
bo
u
1
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5|
•Bn n
Geographical
Distribution
c
£
C
w
EBKS
Larvae
Pupae
Adults
Acidoblontic
Acldophilous
Neutral
Alkaliphilous
Alkallbiontic
Indifferent
Polyhalobous
Cuhalobous
Kesohalobous
OliKohalobous
Euryhalinous
Eutrophic
Hesotrophic
OliEOtrophic
Dystrophic
Saproohilic
Facultative
Saproxenous
Saorophobic
Suoxvphilous
lesoxvphilous
)lieoxvphilous
Anoxvphilous
Cuthermal
Mesot-hpi-mal
Oligothermal
Stenothermal
Me ta thermal
Eurytherraal
Eulicho tophilous
^ esolichtophilous
Polylichcopliilous
Jligolichtophilous
Llmnobiontic
Limnophilous
Indifferent
Kheophilous
Elheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Enibenthic
Embenthic
EoipeHc
Eolsabullc
Epilithic
Eoixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnlvore
Scavenger
Region I
Region II
Recion III
Renlon IV
Region V
Reeion VI
Reeion VII
Reeion VIII
Region IX
Reeion X
12

X







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Concensus
&
Notes











































in clay.



                             110
-------
Taxon:    Glyptotendipes paripes  (Edwards)
o)
0
s
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CO

61
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Habitat
Specific
Habitat
00
n
01
1
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SfS
•H >
is rt
s-s
fm m
Geographical
Distribution
SRS
arvae
upae
dults
cidobiontic
cidophilous
eutral
Ikdliphilous
Ikalibiontic
ndif ferent
olyhalobous
uhalobous
6sohalobous ,
ligohalobous
uryhalinous
M trophic
leso trophic
)lisotrophic
Jystrophic
Japrophilic
''acultative
saproxenous
Saprophobic
"uoxvphilous
lesoxvphilous
51ieoxvphilous
\noxvphi1oiis

•lefiot-hernifll
Dligothermal
Stenothermal
[etathermal
!urythermal
EuLichotophilous
^esolichtophilous
'olylichtophilous
31igolichtophllous
Litiuiobiontic
jimnophilous
Indifferent
iheophilous
Iheobiontic
•larine
Estuary
Lake
Pond
River
Stream
Spring
Other "
Epibenthic
Embenthic
Epipelic
Eoisabulic
Epilithic
Epixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter




Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IX
Region X 	
1

X







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X

X
X

X

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,















































Concensus
&'
Notes












































































                                III
-------
Taxon:    Goeldichironomus holoprasinus  (Goeldi)
01
s
5
CO

tx
4J



EC
o



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c
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CO
AJ
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is
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u

s
0
General
Habitat
Specific
Habitat
u
u •
a
<9
g.3
3§
4) J=
£S
Geographical
Distribution
ERRS
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
AJkalibiontic
Indifferent
Polyhalobous
Suhalobous
•fesohalobous
Oligohalobous
Suryhalinous
Sutrophic
fesotrophic
Oligotrophic
Dys trophic
Saprophilic
Facultative
Saproxenous
Saprophobic
;uoxvphtlous
'fesoxvphilous
OliEOxvohilous
Anoxvphilous
Elithermal
HeBOChprmal
Oligo thermal
Stenothermal
Metathermal
Eury thermal
Eulichotophilous
Hesolichtophilous
Polylichtophilous
Oligolichtophilous
£4imnobiontic
Limnophitous
Indifferent
Rheophilous
RheobionCic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Enibenthic
Embenthic
Eplpelic
Episabulic
Epilithic
Epixvlous
Eplzooic
Epiphytic-
Attached
Unattached
Hfnter
Spring
Summer
Atitiimn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Res-ion V
Region VI
Region VII
Region VIII
Reeion IX
Region X
1

X







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X

X


X

X




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X
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25

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X



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X

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26

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X

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X


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Concensus
&
Notes










































Tree holes .




                              112
-------
Taxon:   Guttipelopia currani  Beck & Beck
01
u
M
1

u
u



SB
a



4-1
c


•a td
Q] J-
SS
Geographical
Distribution
KRS
arvae
upae
dults
cidobiontic
Lcidophilous
eutral
Ikaliphilous
Ikalibiontic
ndifferent
olyhalobous
uhalobous
iesohalobous
ligohalobous
Airyhalinous
]utrophic
[esotrophic .
Jligotrophic
)ystrophic
Japrophilic
^acultative
>aproxenous
Saprophobic
]uoxvph ilous
lesoxvphilous
Higoxvphilous
Vnoxvphilous
""uthermal
lesot-hprmal
)ligo thermal
Stenothermal
•letathermal
Zurythermal
Zulichotophilous
lesolichtophilous
'olylichtophilous
Dligolichtophilous
Limnobiontic
Limnophilous
tndifferent
^heophilous
Iheobiontic
Marine
istuarv
Lake
Pond
River
Stream
Sprine
Other
Epibenthic
Embenthic
Epipel.ic
Episabulic
Epilithic
Enixvlous
Eoizooic
Eoinhveic
Attached

Winter

Summer


Herbivore
Omnivore
Scavenger
Region I
Region II
Resion III
Region IV
Region V
Region VI
Region VII •
Reeion VIII
Region IX
Reeion_X 	
1

X



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X


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X







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X




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X
X

X

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X
X
X
X
X

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X





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X

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Concensus
&
Notes

















































Swamp .



























-------
Taxon:    Harnischia amachaerus  (Townes)

01
u
a
u
w
5

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U
f
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Habitat
Specific
Habicat
tc
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11
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tM n
Geographical
Distribution
(U
u
t.
c
en
ERBS
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
OliRohalobous
Buryhalinous
Eutrophic
Hesotronhic
Olieotroohic
Dystrophic
Saprophilic
Facultative
Saproxenous
Saproohobic
Suoxvphilous
•lesoxvnhilous
ninoxvphilous
Anoxvphilons
!uthermat
^fesoChprmHl
Oligo thermal
Stenothermal
Metathermal
Eury thermal
Eulicho tophilous
t esolichtophilous
Polylichtophilous
Dligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Hheobloncic
Marine
Estuary
Lake
Pond
River
Stream
Snrina
Other
Epibenthic
Embenthic
Enlpelic
Enisabulie
Epilithic '
Epixvlous
Enizooic
Eninhvtic
Attached
Unattached
Winter
Sortn?
Cummer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Recion II
Region III
P.egion IV
Reeion V
ReRion VI
Reeion VII '
Reuion VIII
Region IX
Reeion X
1

X







X



X

X


X


X


X







X
X




X





X
X
X
X









X

x-
X
X
X
X

X





X






4

X



X







X
















X








X





X

X














X







X































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes















































-------
Taxon:    Harnischia boydi  (Beck)
0)
o
M
3
O
en

•a rt
SJi
(U
fo FQ
Geographical
Distribution
ERSS
Larvae
'upae
\dults
\cidobiontic
\cidophilous
Neutral
\lkaliphilous .
Mkallbiontic
Indifferent
'olyhalobous
Cuhalobous
esohalobous
DliRohalobous
Euryhalinous
Eu trophic
tesotraphic
Dlieotroohic
Dys trophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Euoxvohilous
fesoxvphilous
")lieoxvohilous
'Vnoxvphil ous
Blithe ^mal
^feBot-hprmal
Dligothermal
Stenbthermal
^letathermal
Eurythermal
Euiichotophiious
Mesolichtophilous
Polylichtophilous
aTigoIichtophilous
Lininobioritic
Limnophilous
Indifferent:
Iheophilous
Rheobiontic
larine
Estuary
Lake
Pond
River
Stream
Spring
Other
Enibenthic
Embenthic
Epipelic
Eoisabulic
Enilithic
Epixvlous
Epizooic
Eoinhvtic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
.Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII-
Reeion VIII
Region IX
Reeion X
1

X



X







X

X


X


X


X







X

X



X





X
X

X









X

X
X
X
X
X

X





X













































































































































































































































































































































































































































































































































































































































































































































Concensus
& •
Notes













































































                              IB
-------
Taxon!  Harnisehia  edwardsi   (Kruseman)
«
jj
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tn
0
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u




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General
Habitat
Specific
Habitat
M
S
S
ea o
5-3
T3 (3
s-s
FM n
Geographical
Distribution
iKKS
^arvae
>upae
\dults
\cidobiontic
\cidophllous .
Neutral
Mkaliphilous
Mkalibiontic
[ndif Cerent
Polyhalobous
Euhalobous
lesohalobous
31iKohalobous
Euryhalinous
Eu trophic
tesotrophic
Dlieotroohic
Dvstronhic
Saoronhilic
Facultative
Saproxennus
Saprophobic
Euoxvphilous
^fesoxvphilous
Dlieoxvphilous
Anpvvphlloiis
Ktif-hprmal
^pRot-hf rmal
Dllgotherraal
Stenothermal
'•tetathermal
Eurythermal
Eulicho tophilous
tesolichtophilous
Polylichtophilous
Dligolich tophilous
Limnobiontic
Liranophllous
Indifferent
Kheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprint!
Qtjh.er
Epibenthic
Bmbenthic
Epinelic
Eoisabulic
Epilithic
Epixvlous
Kpizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnlvore
Scavenger
Region I
Region II
Region III
Heeion IV
Region V
Region VI
Region VII
Region VIII
Ren ton IX
Region X
27

X





X





X



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X

X



X


X









X





X














X













X

28

X





X





X

X





X


X



X



X

X


X







X




x








X


X








x






























































































































































































































































































































































































































































































































































































































































Concensus
&
Notes

-








































































                               116
-------
Taxon:    Harnischia  galeator  (Townes)
0)
a
1

91
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a



5
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u
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0
a
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^
2
S
H

S

13 «
S^
^a
Geographical
Distribution
Eggs
Larvae
Pupae
Wults
icidobiontic
Acidophilous
Neutral
Ukaliphilous
Ukalibiontic
Indifferent
Polyhalobous
Suhalobbus
lesohalobous
Dligohalobous
Euryhalinous
Eutrophic
'•lesotrophic
Jligotrophic
Dys trophic
Saprophilic
racultative
Saproxenous
Saprophobic
Suoxyphilous
tesoxvphilous
Dlieoxvphilous
\noxvphllonfi
Eiithermal
^fesofhprmal
Dligo thermal
Stenothermal
tetathermal
Eurythermal
Eulichotophilous
lesolichtophilous
Polylichtophilous
31igolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Snrine
Other
Epibenthic
Embenthic
Enipelic
Episabulic
Enilithic
Epixvlous
Epizooic
Epiphvtic
At tached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Reeion VI
Region VII
Region VIII
Region IX
Region X
1

X







X



X

X


X


X


X







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X



X





X
X

X


X








X
X
X
X
X



X



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12

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X



X






X

X






X












X

X


X
X









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11

X











X


X








X



X

X





X






X












X

X


X










X




23

X





X





X

X





X


X



X



X

X


X







X




X








X

X









X
































































































































































































































































































































































































































































































Concensus
&
Notes




























































                               117
-------
Taxon:  Harnischia viridulus  (Linnaeus)
OJ
o
u
a
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w
0
1
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General
Habitat
Specific
Habitat
£f
-------
Taxon:
          Heterotriasocladius   Sparck
<11
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o
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Habitat
Specific
Habitat
' b£
Vj
33
3 §
s-s
h m
Geographical
Distribution
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
?olyhalobous
luhalobous
•lesohalobous
Oligohalobous
Suryhalinous
Sutroohic
Mescjtrophic
OlieotroDhic
3ys trophic
Saprophilic
Facultative
Saproxenous
Saoroohoblc
Euoxvohilous
•lesoxvphilous
Oligoxvohilous
Anoxvphilous
Euthermal
Mesnt-hprmal
Oligothermal
Stenothermal
Metathermal
Eurs'thermal
Eulicho tophilous
Mesolichtophilous
Polylichtophilous
Oligolichtophilous
LiLmnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobioncic
Marine
Estuary
Lake
Pond
River
Stream
Sorine
Other
Eoibenthic
Embenthic
Epioelic
Eoisabulic
Enilithic
Epixvlous
Epizooic
Epiohvtic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Reeion V
Region VI
Region VII •
Region VIII
Reeion IX
Region X
1

X



X







X


X

X



X

X







x

X






X




X
X







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X
X
X
X


X





X













































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes





















































                              119
-------
Taxon:    Kiefferulus  dux  (Johannsen)
0)
1
CO

a
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0


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5
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CO
£
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5
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1
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General
Habitat
Specific
Habitat
to
1
60 O
5-5
13 &
§•§
fe m
Geographical
Distribution
se.es
.arvae
'upae
\dulcs
teidobiontic
\cidophilous
•leutral
Mkaliphilous
Mkalibiontic
Indifferent
Polvhalobous
Suhalobous
^esohalobous
DliRohalobous
Euryhalinous
Eutroohic
lesotronhlc •
Jlieotrophic
Dystrophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Buoxvphilous
^[esoxvphilous
Dlicoxvohilous

Riit-hprmal
^esothprmal
Dligothermal
Steno thermal
kleta thermal
Eurythermal
Eulichotophilous
ttesolichtophilous
Polylichtophilous
Jligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Kheophilous
Rheobionclc
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Epibenthlc
EmbenChic
Enioellc
Eplsabulic
Eoilithic
Epixvlous
I^ptzooic
Epiphytic
Attached
Unattached
Winter
.*lpriny .
Summer
A",t1im.n
Predator
Herbivore
Omnivore
Scavenger
Reeion I
Region II
Reeion III
Reeion IV
Reeion V
Reeion VI
Ree-ion VII
Reeion VIII
Reeion IX
JlegioiLjC 	
1

X



X







X

X


X

X




X






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X


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X
X




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X





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14

X







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X




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X






11

X











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X

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X







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X







X


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X




8

X





X





X


X




X


X



X



X

X






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X



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Concensus
&
Notes




































































                               ]20
-------
Taxon:   Krenosmittia •  Thienemann


DC
4->



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a
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Habitat
Specific
. Habitat
00
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II
11
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Geographical
Distribution
0)
CJ
to
g
«
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
?olyhalobous
Suhalobous
Mesohalobous
Oligohalobous
Suryhalinous
Sutrophic
•lesotrophic
Oligotrophic
3ys trophic
Saprophilic
Facultative
Saproxenous
Saorophobic
Juoxvphilous
•lesoxvnhilous
Higoxvphilous
AnoxvpMlous
Suthermal •
tesorhermal
Oligothermal
Stenothermal
Metathermal
Eurythermal
Eulicho tophilous
Mesolichtophilous
Polylichtophilous
Oligolich tophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
^hedfaiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Epibenthic
Embenthic
Epinelic
Episabulic
Enilithic
Eoixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Reeion HI
Region IV
Region V
Region VI
Region VII •
Region VIII
Reeion IX
Reeion X
1

X



X







X


X

X



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X

X





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Concensus
&
'Notes




















































                             121
-------
Taxon:    Labrundinla beckl  Roback

a
1
4J




5


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-------
Taxon:
          Labrundinia floridana  Beck & Beck

•o a
0) .C
-------
Taxou:    Labrundinia  johannseni  Beck & Beck


tt
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Habitat
Specific
Habitat
ec
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Geographical
Distribution
01
o
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1
EgES
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polvhalobous
£uhalobous
Kcsohalobous
Olicohalobous
£uryhalinous
Eutrophic
•lesotrophic
Olicotrophic
Systrophic
Saprophilic
facultative
Saproxenous
Saprophobic
Suoxvphtlous
•fesoxvphilous
Olicoxvohilous
Anoxvphilons
RuChernial
Me^ot-hprmal
Ollgothermal
Stenothermal
Metathermal
Eurythermal
Eulicho cophilous
Kesollchtophilous
Polylichtophilous
Ollgolichtophilous
Linnobionclc
Liranophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Enibenthtc
Embenthic
Epinelic
Episabulic
Eptlithi'c
Epixvlous
Eoizooic
Eplphvtic
Attached
Unattached
Winter
Sprlnp
Stimmpr
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
P,eeion IV
Region V
Region VI
Reeion VII
Region VIII
Rceion IX
Reeion X
1

X







X



X

X
X

X


X


X







X

X




X




X


X









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X
X
X
X

X





X






7

X



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X

X


X



X

X




X

X

X



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X











X

X
X




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Concensus
&
Notes




























































                              124
-------
Taxon: Labrundinia neopllosella  Beck & Beck
01
o
p
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Habitat
Specific
Habitat
M
jj
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g>5
5 §
-------
Taxon: Labrundinia pilosella   (Loew)


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General
Habitat:
Specific
Habitat
ec
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Geographical
Distribution
-------
Taxon:
           Labrundinia virescens  Beck & Beck

T3 TO
-------
Taxon:    Larsia decolorata^  (Malloch)
,
5
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Habitat
Specific
Habitat
ec
^4
1
II
•o a
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U V
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Geographical
Distribution
ERRS
Larvae
'upae
(Wults
&cidobiontic
^cidophilous
leutral
Mkaliphilous
UfcalibionCic
Indifferent
'olyhalobous
Suhalobous
^esohalobous
31iKohalobous
£urylralinous
Eutrophic
lesotrophic
31ieot:roDhic
Dystrophic
Sapronhilic
Facultative
Saproxenous
Saprophobic
Euoxvphllous
fesoxvphilous
OliEOxvphilous
^nowphi^o^ls
K"Kh^rmai
^esol-hormal
Dligothermal
Stenothermal
^leCatherraal
Eurychermal
Eulicho tophilous
Icsolichtophilous
Polylichtopliilous
Jllgolichtophilous
Limnobiontic
Limnophilous
Indifferent
EUieophilous
Elheobiontic
Marine
Estuary
Lake
Pond
River
Stream .
Spring
Other
Eoibenthic
Embentliic
Eplpellc
Boisabulic
Epillthic
Epixvlous
Eoizooic
Epiphvtic
Attached
Unnttached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Reeion-III
Region IV
ReEion V
Recion VI
Reeion VII
Region VIII
Region IX
Re B ion X
11

X











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Concensus
&
Notes































































                              128
-------
Taxon:
          Lauterborniella agrayloidas  (Kieffer)
OJ
o
3
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w
a
ca
u


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a

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3
w
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13
3
-------
Taxon:    Lauterbornlella varipennis  (Coquillett)

o
cc
s
en
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-------
Taxon:   Leptochironomus  nigrovlttatus   (Malloch)

01
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«
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
Olisohalobous
Euryhalinous
Eutrophic
Hesotrophic
Olieotrophic
Elystrophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Suoxvphilous
klesoxvphilous
Olicoxvphilous
Anoxvphilous
2u thermal
Msso thermal
Oligothermal
Stenothermal
Metathermal
Eurythermal
Eulichotophilous
Mesolichtophilous
Polyliclitophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
SprinE
Other
Epibenthic
Embenthic
Enipelic
Episabulic
Enilithic
Eoixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
•Region II
Region III
Region IV
Region V
Reeion VI
Region VII
Region VIII
Reeion IX
Reeion X
1

X







X



X

X
X




X


X



X


X


X



X





X

X



X








X

X
X
X






X
X













































































































































































































































































































































































































































































































































































































































































































































• Concensus
&
Notes














































































                             131
-------
Taxon:
           Lobodiamesa  ?agast
a
c
M
at
u
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General
Habitat
Specific
Habitat
£
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&
33
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o cl
t>« »
Geographical
Distribution
ERRS
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polvhalobous
Euhalobous
Mesohalobous
OliRohalobous
Euryhalinous
Eutrophic
Hesotroi>hic
Olieotrophic
Dys trophic
Saprophilic
Facultative
Saproxencus
Saprophobic
Euoxvphilous
•fesoxvphilous
Oltcoxvnhilous
Anoxvphilous
Ellthermnl
^esofhormal
Oligo thermal
Stenochermal
Hetathermal
Eury thermal
Eulicho tophilous
Mesolichtophilous
Polylichtophilous
Oligolich tophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
EUieobiontic
Harine
Estuary
Lake
Pond
River
Stream
Snrine
Other
Epibenthlc
Embenthic
Enioe].ic
Episabulic
Epllithic
Eoixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter-
Spring
Summer
Autumn
Prpdator
Herbivore
Omnivore
Scavenger
Region I
Recion II
Reeton III
Reelon IV
Reeion V
Resion VI
Reeion VII
Region VIII
Reeion IX
Rezion X
1

X



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X


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X

X






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X




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X






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X



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Concensus
&
Notes















































                             132
-------
Taxon:  Macropelooia decedens  (Walker)
CD
u
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c
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Habitat
Specific
Habitat
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w
00 O
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•H >
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0} jZ
0) OJ
N W
Geographical
Distribution
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
Oligohalobous
Suryhalinous
Eutrophic
lesotrophic
OliEOtrophic
Dystrophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Euoxvphilous
•fesoxvphilous
)lieoxvphilous
Anoxvphilous
!u thermal
lesot-hprmal
Oligothermal
Stenothermal
letathermal
Eury thermal
Eulicho tophilous
lesolichtophilous
olylichtophilous
ligolich tophilous
imnobiontic
imnophilous
ndif ferent
heophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Epibenthic
Embenthic
Epipelic
Episabulic
Epilithic
Eoixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII •'
Reeion VIII
Reeion IX
Reeion X
3

X


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X



X




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X


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» Concensus
&
Notes















































                             133
-------
Taxon! Maeropelopia hirtipennis  (Loew)

flj
cc
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V.
X
d
>,
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C
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ffl
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e
-------
Taxon:    Metrioenemus abdoaino-flavatus  Picado
a
a
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c
en
03
«
10
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ft
>
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Hablcat
Specific
Habitat
60
M
-------
Taxon: Metrlocnemus  hamatus  Johannsen
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Taxon;   Metriocnemus knabi  i
u
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s-s
fo M
Geographical
Distribution
a
j
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
Olieohalobous
Euryhalinous
EutroDhic
rtesotrophic
OliEOtroahic
Oystrophic
Saprophilic
Facultative
Saproxenous
Saproohobic
Euoxvohilous
fesoxvphilous
Hicoxvohilous
Anoxvphiloiis
Suthermal
lesochermal
Oligothermal
Stenothermal
t etathermal
Eur^'thermal
Eulicho tophilous
Hesolich tophi lous
Polylichtophilous
Dligolich tophilous
Limnobiontic
Limnophilous
Indifferent

Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Epibenthic
Embenthic
Eoipelic
Episabulic
Enilithic
Epixvlous
Epizooic
Epiphvtic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IX
Reeion X
1

X



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x


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X





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X







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Concensus
&
Notes










































Inhabits water in the
purpurea .




                             137
-------
Micropsectra  Kieffer
s
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Habitat
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Geographical
Distribution
~(ZES
^arvae
3upae
Wults
kcidobiontic
^cidophilous
Neutral
Mkaliphilous
Mkalibiontic
Indifferent
Polyhalobous
Euhalobous
[esohalobous
DliKohalobous
Euryhalinous
^utrophic
[esotrophic
31iBOtroDhic
Dvs trophic
Saprophilic
Facultative
Saproxenous
Saoroohobic
Euoxvphilous
^lesoxvohilous
Dlieoxvnhilous
f\poxyphi1otts
R^t;bermal
fesof-hermal
Oligothermal
Stenothermal
letathermal
Lurythermal
Eulichotbphilous
Mesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
^imnophilous
Indifferent
Rheophilous
Jheobiontic
larine
Estuary
Lake
Pond
River
Stream
Sorine
Other
Eoibenthic
Embenthic
Enioelic
Eoisabulic
Enilithic
Eoixvlous
Eptzooic
EoiDhvtic
Attached
Unattached
Winter

ilUI^n^r

Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
• Reeion III
Region IV1
Region V
Region VI
Region VII
Region VIII
Region IX
Region X 	
1

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Concensus
&
Notes












































































                   138
-------
Taxon:  Micropsectra dives   (Johannsen)

•O (fl
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FE< n
Geographical
Distribution

EES
arvae
upae
dults
cidobiontic
cidophilous
eutral
Ikaliphilous
Ikalibiontic
ndif ferent
olyhalobbus
uhalobous1
esobalobous
liEOhalobous
uryhalindus
utrophic
lesotrophic
lieotrorhic
Jvstrophic
>aprophilic
'acultative
>aproxenous
JaoroDhobic
juoxvphilous
lesoxvohilous
)lieoxvphilous
VnovvphiloiiB


)ligothermal
Stenotherraal
"letathermal
Surj'thermal
Sulichotophilous
lesolichtophilous
'olylichtophilous
31igolichtophilous
jiranobiontic
jimnophilous
[ndifferent
Iheophilous
Iheobiontic
•larine
Estuary
Lake
Pond
River
Stream
Sprine
Other
EoibenChic
Embenthic
Enioelic
Episabulic
EDilithic
Epixvlous
Eoizooic
Epiphytic
Attached

Winter




Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Reeion VI
Reeion VII
Reeiori VIII
Reeion IX
Resion X

3

X



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Concensus
&
Notes














































































                               139
-------
Taxon:  Micropseetra dubia  (Malloch)
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Habitat
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Habitat
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Feeding
Behaviot
Geographical
Distribution
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkallbiontic
Indifferent
Polyhalobous
Euhalobous
tesohalobous
Oligohalobous
Suryhalinous
Eutroohic
Hesotrophlc
Olieotroohic
Dys trophic
Saprophilic
Facultative
Saproxenous
Saproohobic
Euoxvohilous
Mesoxvphilous
Olieoxvphilous
Anoxvphil oils
Ktithermal
McRothnrmal
Oligothermal
Stenothermal
Mecachermal
Eurythermal
Eulichotophilous
Mesolichtophilous
Polylichtophilous
Oligolichtophilous
Liranobiontic
LimnophilOus
Indifferent
Rheophilous
Rheobiontic
Marine
Estuarv
Lake
Pond
River
Stream
Sorine
Other
Enibenthtc
Embenthic
Eoipelic
Episabulic
Enilithic
Epixvlous
Epizoolc
Epiphvtic
Attached
Upat^ached
Winter
Spring
Puiltn^r
Anttimn
Predator
Herbivore
Omnivore
ScavenKer
Region I
Region II
Reelon III
Reeion IV
Reeion V
Reeion VI
Reeion VII
Reeion VIII
Reeion IX
Reeion X
28

X





X





X

X





X


X



X



X

X


X







X




X








X

X









X













































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes















































-------
Taxon:	Micropsectra nigripila  (Joharmsen)

"O ct)
gj 4J
-------
Taxon;	Hicropsectra pollta   (Malloch)
01
a
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5




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General
Habitat
Specific
Habitat
efi
M
I
II
•o rt
0) ^
ss
Geographical
Distribution
-ggs
^arvae
'upae
\dults
\cidobiontic
Vcidophilous
teutral
Mkaliphilous
Ukalibiontic
Indifferent
Polyhalobous
Suhalobous
fesohalobous
31lEohalobous
Suryhalinous
Sutrophic
fesotrophic
!)lieotrophic
Dvs trophic
Sapronhilic
Facultative
Saproxenous
Saorophobic
Euoxvphilous
^tesoxvphilous
DUgoxvphilous
\noxvph-------
Taxon;    Mierotendipes pedellus   (De Geer)

•a «i
-------
Taxon!    Monopelopia boliekae  Beck & Beck

0)
tf
0)
u
CO
X
Q.
Salinity
9
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Habitat
Specific
Habitat
6C
U
I
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•a 5
at 42
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Geographical
Distribution
i
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
Oligohalobous
:.uryhalinous
lutrophic
•lesotrophic
)lieotroohic
>ys trophic
Saprophilic
Facultative
Saproxenous
Saprophobtc
luoxvohilous
tesoxvnhilous
(lieoxvphilous
Anoxvphllons
5uthertn.il
leROt-hprmal
Oligothermal
Stenotliermal
letathermal
Eurythermal
Eulicho tophilous
Icsolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
ndifferent
Rheophilous
Rheobiontic
Marine
Es tuary
Lake
Pond
River
Stream
Sprine
Other
Epibenthlc
Embenthic
Enioelic
Eoisabulic
Epilithic
Eoixvlous
Epizooic
Epinhvtic
Attached
Unattached
Winter
Sorinif
Summer
Autumn
Prsdafnr
Herbivore
Omnivore
Scavenger
Reeion I
Reeion II
Reeion III
PvEEion IV
Region V
Reeton VI
Region VII
Region VIII
RCEton IX
Reeion X
7

X




X






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X



X

X



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x



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X
X
X


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X



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X
X
X
X


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Concensus
&
Notes















































                           1HH
-------
Taxon:   Monopelopia tillandsia  Beck & Beck

•a «i
-------
Taxon: Nanocladius alternantherae  Dendy & Sublette

5
33
s-s
b m
Geographical
Distribution


J&ggS
Larvae
Pupae
Adults
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Euhalobous
^esbhalobous
Olleohalobous
mryhalinous
Eutroohic
•tesotroohic
Olieotroohic
Jystroohic
Saprophilic .
••acultative
Saproxenous
Saproohobic
iuoxvphllous
fesoxvohilous
)liKOxvohilous
Anoxvphilous
iuChermal
•tesothnr-mal
Oligothermal
Stenothermal
letathermal
Eurytherraal
Eulichotophilous
lesolicntophilous
Polylichtophilous
Oligolichtophilous
.imnophilous
ndifferent
heophilous
leobiont-ic
Marine
Estuary
Lake
Pond
River
Stream
SprinE
Other
Epibenthic
Etnbenthic
Epipelic
Eoisabulic
Epilithic
Epixvlous
Eoizooic
Eoiphvtic
Attached
Unattached
;«nter
Sorlnu
Summer
Autumn
Predator
ilerbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
teeion VII
Region VIII
lesion IX
Region X
1
X






X


X


X







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Ji.










X



X






X

X
X
X
X
X

X







X




28
X




X




X

X





X


X



X



X

X









X




X








c


X







































































































































































































































































































































































































































































































































































































































Concensus
&
	 Notes











































-------
Taxon: Natarsia fastuosa  (Johannsen)
-------
Taxon!    Hilodorum deveineyae   (Beck)

I
i
to
s
X
e
a
CO
|
j-
u
c
c
lieoxvphilous
Anoxvphllons
•.llCtiennal
fesohhprmal
Oligotjiermal
Stenothermal
letachermal
Eurythermal
Eullchotophilous
lesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophllous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Epibenthic
Embenthic
Eotpelic
Episabulic
Epilithic
Eoixvlous
Epizooic
Epinlwtic
Attached
Unattached
IHnter
Spring
Summpr
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
KCKion III
Region IV
Reeion V
Reeion VI
Reeion VII
Renion VIII
Reeion IX
Reeion X
1

X







X


X








X


X






X


X




X



X







X







X
X


X



X



X













































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes
















,






























                            148
-------
Nilotanypus americanus  Beck & Beck
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General
Habitat
Specific
Habitat
60
u
tto o
•s-g
•o rt
0) J=
ta 
-------
Taxon:
           Nimbocera  Reiss


a
u



S



2
s
l-l
w

01
M
3
U
1
U
s

-------
Taxon:   Odontomesa  fulva   (Kleffer)


a
u
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33
•i.5
2Ti
O (li
S3


UJ
a
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VI
RRS
arvae
tipac
dults
cidobiontic
cidophilous
eutral
Ikaliphilous
IkalibionCic
ndif ferent
olyhalobous
uhalobous
lesohalobous
)liRohalobous
:uryhallnous
!utrophic
leso trophic
)li.EOtroph£c
)ys trophic
Japrophilic
""acultative
Saoroxenous
Saoroohobic
2uoxvphilous
fesoxyphilous
Jlicoxvnhilous



31igothermal
Stenothermal
• etathermal
Surythermal
Sulichotophilous
lesolichtophilous
?olylichtopliilous
Uigolichtophilous
jiranobiontic
jimnophilous
Indifferent
Hheophilous
ilheobiontic
arine
Estuary
Lake
Pond
River
Stream
Spring
Other
Epibenthic
Embenthic
Epipelic
Eoisabulic
Eoilithic
Eoixvlous
Rpizooic
Epiphvtic
fa Cached
Unattached
Winter


AMtumn

Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII

_ .Reaion._X 	


X







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X





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X







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x






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X





X





X
















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Concensus
&
Notes














/































































                                151
-------
Taxon:
           Omisus  pica  Townes

<
ffi
D
C Salinity
c
1
^
'f
K
tn
I
f
O
,:
H

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2
3
General
Habitat
Specific
Habitat
to
H
i
S3
•a a
s-s
£ n
f-l C
35
SS
B.M
« *H
MU
0 m
S-H
a


Eccs
Larvae
Pupae
Adults
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
£uhalobous
Mesohalobous
Olicohalobous
Survhalinous
lesotrophic
Olieotrophic
lystrophic
Facultative
Saproxenous
Saorophobic
iuoxvphllous
fesoxvnhllous
IHeoxvohilous
.noxvph-nniis
tesorhormsil
Oligothermal
tenothermal
letathermal
tarythermal
esollchtophilous
olylichtophilous
lligolichtophilous
.imnophilous
nditjerent
leophilous
tieobiontic
Marine
Estuary
..ake
?ond
Jiver
Stream
Sprlne
)ther
ipibenthic
imbenchic
!i)tnelic
inisabulic
ipilithic
iplxvlous
ioizoolc
ipiphvtic
Attached
Jnat cached
»nter
Spring
Summsr
lutumn
Jlredator 1
terbivore
Omnivore
Scavenger
legion II
(eelon III
Eegion IV
teeion V
teEton VI
Reeion VII •
Reeion VIII
leelon IX
[eeion X
1



TT






X

—

X

X
X





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X

X



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Concensus
&
	 	 Notes


































                           152
-------
Taxon;     Orthocladius anneetens  Saether
V
I


a
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W



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u
5
r-l
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= >J
3

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S
§
H
O
General
Habitat
Specific
Habitat
M
!
60 O
(3 -H
^H >
•a nj
ystrophic
aprophilic
^acultative
aproxenous
Japrophobic
^uoxvohilous
lesoxvphilous
Uieoxvohilous



Dligo thermal
stenothermal
letatherraal
Eury thermal
Sulichotophilous
lesolichtophilous
Polylichtophilous
Dligolichtophilous
Limnobiontic
Limnophilous
[ndifferent
Rheophilous
iheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine

Epibenthic
Embenthic
Koine lie
Episabulic
Epilithic

Epizooic
Epiphvtic
/\t Cached
Unattached
Winter
Sprinp
Summer

Predator
Herbivore
Omnivore
Scaveneer
Region I
Region II
Reaion III
Region IV
Region V
Reeion VI
Re E ion VII
Region VIII
Region IX
Region X 	


^



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X

X


X


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X







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X









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X
X
X

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Concensus
&
Notes



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• -








                               153
-------
Taxon! Orthocladius obumbratus  Johannsan

0)
01
a
u
Ul
1

c
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1
7
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u
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s


General
Habitat
Specific
Hablcat
tc
u
01
s
Si
•85
&&
Geographical
Distribution

Eggs
Larvae
Pupae
Adults
Acldobiontic
Acidophilous
Neutral
Alkalipjiilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
Oligohalobous
^uryhalinous
Eutrophic
•lesotroDhic
JliKotroohic
)ys trophic
Saprophilic
"acultative
Saproxenous
Saoroohobic
luoxvohilous
fesoxvnhilous
llieoxvohilous
inoxvphllot.s
leso thermal
Oligothermal
Stenothermal
letatherrcal
Eurythermal
EesolichtophiloUs
Polylichtophilous
Oligolichtophilous
ranobiontic
Limnophtlous
ndif fercnt
heophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Eoibenthic
Embenchic
Bnlpelic
Episabulic
Enilithic
Eoixvlous
Boizooic
Eoinhvtic
Attached
Unattached
rTJnter
Sortnf
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Reeion II
Region III
Region IV
Region V
Region VI
Region VII •
Iggion VIII
Seeion IX
Reeion X
3

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Concensus
&
Notes













































-------
Taxon:
           Pagastiella orophila   (Edwards)
qj
8
1

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Habitat
opecific
Habi,tat
«
J*
§•3
•H >
XJ nj
a) A
a a
& ffl
Geographical
Distribution
-Sgs
^arvae
3upae
Adults
\cidobiontic
\cidophilous
Neutral
\lkaliphilous
Ukalibiontic
Indifferent
Polyhalobous
Suhalobous
•lesohalobous
31iKohalobous
Euryhalinous
Sutrophic
lesotrophic
Jligotro^liic
Dystrophi-c
Sapronhilic
•"acultative
Saproxenous
Saprophobic
Euoxvphilous
tesoxvphilous
31ieoxvphilous
\noxvphllous
Fluthermal
lesofhermal
Dligothermal
Steno thermal
Metathermal
Eurytherraal
Eulichotophilous
'•tesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobioncic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Sstuary
Lake
Pond
River
Stream
Spring
Other
Epibenthic
Embenthic
Epipel.ic
Eoisabulic
Epilithic
Epixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore ,
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IX
Region X
1

X



X







X

X
X

X


X


X







X

X




X




X

X
X


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X
X
X
X
X



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X

X













































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes







































































                              155
-------
alatus  (Beck)
0)
J
c
CO
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t-4
M
B
Hi
M
4J
f
a
c
3
o

-------
Taxon:
           Parachironomus  carinatus   (Townes)
a
1
•,
4J
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e
rH
W
JJ
g
fc
U
O
c
3
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-------
Taxon:	Parachironomus directus   (Dendy  &  Sufalette)
01
CJ
f.
1

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5
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General
Habitac
Specific
Habitat
to
H
el
&
to o
c *•-<
31
8-S
(M w
Geographical
Distribution
ERRS
larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
AJkalibiontic
Indifferent
'olyhalobous
Euhalobous
lesohalobous
)liaohalobous
Eurvhalinous
Eutrophic
Hesotrophic
)lieotrophic
)ystrophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Euoxvnhilous
^tesoxvphilous
Dlieoxvohilous
AnoxvphtlonR
Rtlthermal
ff»Bof hormal
Dligothermal
Stenothermal
[etathermal
Eurythermal
Eulichotophilous
lesolichtopliilous
Polylichtopliilous
^llgolTchtophilous
Limnobiontic
[jimnophilous
Indifferent
Kheopnilous
Rheobiontic
Marine
Bstuarv
Lake
Fond
R-iver
Stream
Sprine
Other
Epibenthtc
Kmbenthtc
Entoellc
Episabulic
Eoilithic

Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI ,
Region VII
Region VIII
Region IX
Recion X
2?

X





X





X

X





X


X



X



X

X


X







X




X








X

X
X
X







X













































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes



















































                              158
-------
Taxon: "Parachironomus" deraeijerea  (Kruseman)
0)
a
J
a
«
4J



a
P.


u
5
•3
W
S
1
ii
0
c
M


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1



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M
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4J
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General
Habitat
Specific
Habitat
tc
M
CO O
C iH
*H >
TJ td
01 ,C
0) Q]
Fn W
Geographical
Distribution
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Zuhalobous
lesohalobous
Dliaohalobous
Suryhalinous
2u trophic
Mesotrophic
Olieotroohic
Dys trophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Euoxvohilous
Mesoxvphilous
Oligoxvphilous
Anoxvphilons
F.u thermal
MGBo^hp•^ma^
Oligo thermal
Stenothermal
Metatherraal
Eurythermal .
Eulicho tophilous
Mesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Kheophilous
Rheobiontic
•larine
Estuary
Lake
Pond
River
Stream
Spring
Other
Eoibenthic
Embenthic
Epinelic
Episabulic
Epilithic
Epixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III .
Region IV
Region V
Region VI
Region VII
Reeion VIII
Region IX
Region X
i

X







X



X


X

X



X
X




X





X






X




X
X



X

X





X
X
X
X
X
X
X


X


X
X












































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes























































                              159
-------
Taxon:
           Paraehironomus  hirtalatus  Beck & Beck


•o ta
0) .C
si et
ft* CQ
Geographical
Distribution
0)
o
J.
5
ERRS
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
Olieohalobotis
Euryhalinous
Eutrophic
fesotronhic
OliEOtroohic
Dys trophic
Saproohilic
Facultative
Saproxenous
Saoroohobic
Euoxvphilous
lesoxvDhilous
Jliuoxvnhllous
Anoxvphllous
"llthermal
lesochcrmal
Oligo thermal
Stenothermal
t etathermal
Eurythermal
Eulichotophilous
Mesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Liranophilous
Indifferent
Rheophllous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Enibenthtc
Embenehic
Eoioellc
Enisabulic
Enilithic
Eoixvlous
Enizooic
Eoinhvtic
Attached
Unattached
Winter
Soring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Recion IV
Region V
Reaton VI
Region VII
Region VIII
Reeion IX
Region X
1

X







X



X

X


X


X


X.







y

X




X




X
X
X
X









X

X
X
X
X
X


X



X
X
X







































































































































































































































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Concensus
&
Notes















































                            160
-------
Taxon:
           Parachirgnomus monochromus   (Wulp)
s
s
o
C/l
0)
s
t-i


a
&


>,
s
5
•3
w
s
5
i!
s
3
c
S
s


15
' £
MS
53
•8.S
CU 01
P* «
Geographical
Distribution
Eggs
Larvae
Pupae
Wults
4cidobiontic
kcidophilous
Neutral
Ukaliphilous
\lkalibiontic
Indifferent
Polyhalobous
Euhalobous
^lesohalobous
Dlieohalobous
Euryhalinous
Eutrophic
'•lesotrophic
DliEOtrophic
)ys trophic
Saprophilic
•"acultative
Saproxcnous
Saprophobic
Euoxvphilous
^esoxvphilous
Dlieoxvphilous
\noxvphi1ons
Enthermal
^esot->iRi-mal
31igo thermal
Stenothermal
4etathe.rmal
Eury thermal
Eulichotophilous
4esolichtophilous
Polylichtophilous
Dligolichtophilous
Limnobiontic
jimnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Epibenthic
Embenthic
Sninelic
Episabulic
Epilithic
Enixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
Sprin_
Bummer
Autumn
Predator
Herbivore
Omnivore
ScavenRer
Region I
Region II
Region III
Region IV
Region V
Reeion VI
Region VII
Region VIII
Region IX

1

X







X



X

X
X

X

X



X







X

x



X





X
X
X
X


X








X
X
X
X
X


X




X
X





12

X







X



X










X



X






X

X






X












X

X

X
X










X




11

X











X


X








X



X

X





X






X












X

X
X
X
X
X









X




22

X




X






X

X





X



X


X


X


X


X







X











X

X


X













X

28

X





X





X

X





X


X


..
X



X

•x


X







X




X








X
X
X
X








X

















































































































































































































































































































































































































Concensus
&
Notes





























































                              161
-------
Taxon:    Parachlronomus  pectinatellae   (Dendy & SubleCte)
01
u
£
c
w

8
4J
«



a



U
c
l-t
w
JJ
01
£
3
U
c
u
o

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H



•H
W
H

u

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General
Habltac
Specific
Habitat
ec
jj
6)
&
60 O
•S3
•o 5
ei .e
CJ 0>
t< n
Geographical
Distribution
EBBS
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
luhalobous
•tesohalobous
Olieohalobous
Suryhalinous
Sutrophic
klesotrophic
Olieotroohic
•lystrophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Luoxvphilous
Hesoxvphilous
Oliffoxvphilous
Anoxvpliilous
Etithermfll
Meqot-Ti^rmaT
Oligo thermal
Stenothermal
Metathermal
Eurytherraal
Eulichotophilous
Mesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Litnnophilous
Indifferent
Rheophilous
RheobionCic
Marine
Estuary
Lake '
Pond
River
Stream
Snrine
Other
Enibenthic
Embenthic
Epipelic
Episabulic
Epilithic
Eplxvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Itarbivore
Omnivore
Scavenger
Region I
Reeion II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IX
Region X
1

X







X



X

X
X

X


X


X






X


X





X



X

X
X








X

X


X
X
X






X
X
X












































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes












































This species burrows in the
gelatinous matrix of
Pectinatella magnifica.







                              162
-------
Taxon:
           Parachironomus  scheideri  Beck & Beck
«
u
h
3
0
cn
«
!
u




0.


.»
s
5

M
£
§
^
5
u
C
s
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CS
0
.


1
H



£
£
H

JJ

£
O
General
Habitat
Specific
Habitat
(i
(J
g"5
•H >
•a ta
Sx
0)
[C4 m
Geographical
Distribution
Eggs
Larvae
Pupae
\dults
\cidobiontic
\cidophilous
Neutral
Ukaliphilous
Mkalibiontic
Indifferent
Polyhalobous
[uhalobous
lesohalobous
Dligohalobous
Euryhalinous
Eutrophic
^lesotrophic
Dlieotrophic
Dystrophic
Saprophili'c
Facultative
Saproxenous
Saprophobic
Euoxvphilous
vIesoxvDhilous
Dligoxvphilous
taoxvphilous
Euthermal
^esot-hormal
Dligothermal
Stenothermal
^fetatherinal
Eurythermal
Eulichotophilous
^lesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
[iheobiontic
iarine
Estuary
Lake
Pond
River
Stream
Sprina
Other
Eoibenthic
Embenthic
Epioe].ic
Episabulic
Enilithic
Eoixvlous
Epizooic
Epinhvtic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Reeion IX
Reeion X
1

X



X







X

X


X

X



X






X

X




X





X
X

X









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Concensus
&
Notes



























































                              163
-------
Taxon:
           Parachironomus tenuicaudatus  (Malloch)
s
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Habitat
Specific
Habitat
ec
14
ei
&
60 O
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SS
Geographical
Distribution
ERRS
Larvae
Pupae
Adults
AcidobionClc
Acidophilous
Neutral
Alkaliphllous
Mkalibiontic
Indifferent
'olyh'alobous
Zuhalobous
^esohalobous
31it>ohalobous
:tiryhalinous
lUtrophic
^tesotrophic
DliBOtrophic .
)ystrophic
Saprophilic
facultative
Saproxenous
Saoroohobic
Euoxvphtlous
Hesoxyphllous
Dlicoxvphilous
^oxvphtl otis
Etlthermal
^Bsotliprmat
31igo thermal
Stenothermal
Metnchermal
Eurythermal
Eulichotophilous
[esolichtophildus
Polylichtophilous
Dligolichtophilous
LiAinobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sorine
Other
Epibenthic
Embenthic
Eolpelic
Episabulic
Epilithic
Goizooic
Eoinhytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
ReEion I
Rezion II
Recton III
Rocion IV
Region V
Realon VI
Reeion VII •
Reeion VIII
Reeion IX
Reeion X
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Concensus
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-------
Taxon:
          Paracladopelma loganae  Beck & Beck

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s«s
Geographical
Distribution
u
o
t
c
CO
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiondic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
Olieohalobous
Eurvhalinous
Eutrophic
ylesotrophic
Oligoerophic
Rys trophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Suoxvotiilous
•tesoxvohilous
3HeoxvDhilous
Anoxvphilons
Euthermal
Mesot-hormal
Oligo thermal
Stenothermal
Metathermal
Eurythermal
Eulicho tophilous
Hesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream .
Sorine
Other
Enibenthic
Embenthic
EoiDelic
Eoisabulic
Eoilithic
Eoixylous
Eoizooic
Epirhvtic
Attached
Unattached
Winter
Snrintf
Summer
Autumn
Predator
Herbivore
Omni vo re
Scavenger
Region I
Region II
Region III
Recion IV • ••
Region V
Reyjiori VI
Region VII •
Region VIII
Reeion IX
Reeion X
1

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Concensus
&
Notes















































                           ,165
-------
Taxon: Paracladopelma undine  (Tqwnes)

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Taxon:    Paralauterborniella elachista   (Townes)
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Distribution
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
"olyhalobous
Suhalobous
lesohalobous
Oligohalobous
Suryhalinous
Sutrophic
•lesotrophic
Oligotro'phic
)ystrophic
Saprophilic-
Facultative
Saproxenous
Saprophobic
Euoxvphilous
Mesoxyphilous
OliEoxvphilous
Anoxvohilous
Euthermal
4esofhormal
Oligo thermal
Stenothermal
Metathermal
Eurythermal
Eulicho tophilous
Mesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Epibenthic
Embenthic
Enipelic
Eoisabulic
Epilithic
Epixvlous
Eoizooic
Epiphytic
Attached
Unattached •
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivo re
Scavenger'
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IX
Region X
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Notes













































































                             167
-------
Taxon:	Paralauterbprniella nigrohalteralis   (Malloch)
-------
laxon:   Paralauterborniella subeincta  (Townes)
,
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Taxon:
         •P«ramerina anomala  Beck & Beck
1
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Habitat
Specific
Habitat
CJS
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I
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60 O
C *H
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•0 a
-------
Taxon:   Paramerina smithae  (Sublette)

•o «
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0) 01
tt. «
Geographical
Distribution
zss
^arvae
upae
kdults
IcidobionCic
^.cidophilous
eutral
akaliphilous
Ukalibiontic
tndifferent
'olyhalobous
Zuhalobous
lesohalobous
lllcohalobous
Zuryhalinous
^utrophic
tesotroohic
Jlieotroohic
)vstrophic
aaprophilic
Facultative
Saproxenous
Saoroohobic
Euoxvphilous
^esoxvohilous
Dlieoxvphilous

Ru thermal

Dligothermal
Steno thermal
'letathermal
!ur3T thermal
Eulichotophilous
^lesolichtophilous
Polylichtopliilous
Oligolichtophilous
Limnobiontic
Limnophilous
CndifEerent
Rheophilous
Iheobioncic
•larine
Estuary
Lake
Pond
River
Stream
Spring
Other
Enibenthic
Embenthic
Epioelic
Eoisabulic
Enilithic
Eoixvlous
Eoizooic
Epiphytic
Attached
Unattached
Winter

Summer

Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Reeion ,V
Reeion VI
Region VII
Region VTII
Region IX
Region X 	
27

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Concensus
&
Notes













































































                               171
-------
Taxon:    Parametrloenemus lundbec'kil (Johannsen)

0)
8
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Habitat
Specific
Habitat
u
M
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s
60 O
C *H
35
s-s
Pn «
Geographical
Distribution
a
c
CO
ERRS
Larvae
Pupae
Adults
Acidobiontlc
Acidophllous
Neutral
Alkaliphilous
Alkalibiontic
.ndifferent
Polyhalobous
Euhalobous
Mesohalobous
OliEohalobous
taryhalinous
Eutrophic
fesotrophic
Olieotrophic
)ys trophic
Saprophilic
Facultative
Saproxenous
Saproohobic
iuoxvphilous
tesoxvohilous
ilieoxvphilous
Anoxvphnous
•llChermal
esothermal
Dligothermal
tenothermal
ietathermal
Eury thermal
Eulichotophilous
lesoiichtophilous
olylichtophilous
ligolichtophilous
imnobiontic
iranophilous
ndi££erent
heophilous*
heobiontic
Marine
Estuary
Lake
Pond
River
Stream
Snrine
Other
Eptbenthic
EmbenChic
Eoloelic
Eoisabulic
Entlithic
Epixvlous
Eplzooic
Eolnhvtic
Attached
Unattached
Winter
Spring
Summer
Autumn
("redator
Herbivore
Omnivore
Scavenger
ReRion I
ReRion II
ReKion III
ReRion IV
Region V
ReEion VI
Reeion VII
Reeion VIII
Reeion IX
Reeion X
1

X



X







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• Concensus
&
Notes

















































                              172
-------
Taxon: Paratendipes albimanus  (Melgen)
0)
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s
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Habitat
Specific
Habitat
f
I
P
iy "§
Geographical
Distribution
-KRS
jarvae
'upae
Vdults
icidobiontic
Vcidophilous
feutral
Ukaliphilous
akalibiontic
[ndifferent
'olyhalobous
^uhalobous
lesohalobous
Dlieohalobo'us
:uryhalinous
Zutrophic
"lesotrophic
)liEOtroohic
Dvstrophic
Saprophilic
Facultative
Saproxenous
Baprophobic
Euoxvphilous
^esoxvohilous
Jlieoxvohilous
\noxvphilous
Euthermal
^esot:hGnna1
Dligothermal
Stenothermai
letathermal
Eurythermal
Eulicho tophilous
M.esolichtophilous
Polylichtophilous
Migolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Iheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Eoibenthic
Embenthic
Epioelic
Episabulic
EDilithic
Epixvlous
Epizooic
Epinhytic
Attached

Winter .

Summer

Predator
Herbivore
Omni vo re . .
Scavenger
Region I
Region HI
Reeion III
Reeion IV
Region V
ReEion VI
Reeion VII •
Reeion VIII
Region IX
Reeion X 	
6

X











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11

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X





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X


X





X

X



X



X

X






X





X


X








X












X














































































































































































































































































































































































































































































































































































Concensus
&
Notes









































































                               173
-------
Taxon:
           Paratendipes' subaequalis   (Malloeh)
lit;ohalobous
Euryhalinous
^utrophic
tesotrophic '
)liKOtrophtc
)ys trophic
Saprophillc
Facultative
Saproxenous
Saproohobic
Euoxvphilous
^tesoxvphilous
^ligOKYPhiloM8
^oxvphllous
RutTiermfll
^lesothprmal
Dligo thermal
Stenothermal
^[etathermal
Eurythermal
Eulichotophilous
'[esolichtoptiilous
Polylichtophiious
Dligolichtdphilous
Limnbbiontic
Llmnophilous
Indifferent
Rheophilous
Rheobiontic
larine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Enibenthtc
Embenthic
Enipelic
Eoisabulic
Eoilithic
Epixvlous
Eoizooic
Eoiphvtic
Attached
Unattached
Winter
Sprtny
Sqnim&r
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
ReEion II
Reeion III
ReEion IV
Reeion V
ReKton VI
ReEion VII
Reeion VIII
Reeion IX
Reeion X
1

X







X



X

X
X

X


X


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X

X




X




X
X
X
X


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X
X
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X
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X
X

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Concensus
&
Notes





















































-------
Taxon:	Paratendipes thermophilus  Town.es
-------
Taxon:   ?aroehlus  kiefferi   (Garrett)
(
ti
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Habitat
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Geographical
Distribution
EBBS
Larvae
Pupae
\dultfe
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mcsobalobous
)liRohalobous
luryhalinous
Eutrophic
Kesotrophic
Olieotroohic
)ys trophic
Saprophilic
facultative
Saproxenous
Saprophobic
iuoxvphilous
tesoxvDhilous
llieoxvnhilous
Anoxvphllons
luthermal
iesothormal
Oligo thermal
Stenothermal
[etathermal
Eurythermal
Eulichotophilous
Icsolichtophilous
Polyliclitophilous
Oligolichtophilous
Limnobiontic
Liranophilous
ndif ferent
Kheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Enibenthlc
Embenthlc
Enloellc
Episabulic
IvRilithic
Epixvlous
Eoizooic
Eoiohvtic
Attached
Unattached
[«nter
Sor-lny
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
ReRion I
Region II
Reelon III
RenLon IV
Region V
ReEion VI
Region VII
teeion VIII
Recton IX
Recion X
27

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X













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;

















































































































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Concensus
&
Notes















































                            176
-------
Taxon:
           Parorthocladius   Thienemann
OJ
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en
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Habitat
ec
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Geographical
Distribution

EES
arvae
upae
dults
cidobiontic
cidophilous
eutral
Ikaliphilous
Ikalibiontic
ndifferene
olyhalobous
uhalobous
lesohalobous
liEohalobous
uryhalinous
.utrophic
lesoCrophic
JliEOtroohic
)ystrophic
Japrophilic
"acultative
Japroxenous
Saprophobic
"uoxvohilous
•lesoxvDhilous
Dlisoxvphilous

"!u thermal

Dligothermal
Stenothermal
letathermal
iurythermal
Eulichotophiious
^lesollchtophilous
>olylichtophilous
Dligolichtophilous
jimnobiontic
jimnophilous
[ndifferent
Uieophilous
Iheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Epibenthic



Epilithic



Attached 	
Unattached 	
Win car
Spring



Herbivore
Omnivore
Scavenger
Region I
Region II

Region IV
Hep; ion V
Reeion VI
Region VII •
Reeion VIII
Region IX 	

1

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X



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Concensus
&
Notes










































































                               177
-------
Taxon:
          Pedlonomus beckae  Sublette


w
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Habitat
Specific
Habitat
a
%
&
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Taxoni   Pentaneura comosa  Sublette
01
1
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Habitat
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Habitat
ti
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g'.S
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o <3
Fn P3
Geographical
Distribution
RRS
arvae
upae
adults
Lcidobiontic
Lcidophilous
eutral
ilkaliphilous
akalibiontic
ndifferent
Jolyhalobous
.uhalobous
lesohalobous
)lisohalobous
luryhalinous
lutrophic
lesotrophic
)lieotroohic
)ystcophic
Saprophilic
racultative
Saproxenous
saprophobic
Zuoxvnhilous
•lesoxvphilous
Dlisoxvphilous



Dligothermal
Stenothermal
^etathermal
Jurychermal
SuLichotophilous
Mesolich tophi lous
'olylichtophilous
Higolichtophiious
jimnobiontic
jimnophilous
Indifferent
^heophilous
•Uieobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Epibenthic
Embenthic
Epioellc
Episabulic
Enilithic
Epixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter

Summer
Autumn

Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Reeion VII
Region VIII
Reeion IX
_ ReeifiiLX 	
7.7

X





X





X



X




X

X



X


X









X





X












X















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Concensus
&
Notes












































































                               179
-------
Taxon:
           Pentaneura inconspicua   (Malloeh)

§1
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Habitat
Specific
Habitat
£f
i
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tJ 5
S4=
0
b w
Geographical
Distribution
i
c
Egss
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
AJkalibiontlc
Indifferent
Polyhalobous
Euhalobous
Hesohalobous
Ollcohalobous
luryhalinous
Eutrophlc
•lesotrobhic
Oliijotrophic
Jystrophic
Saprophilic
Facultative
Saproxenous
Saproohobic
luoxvphilous
lesoxvphilous
)ligoxvnhilous
Anoxvphllons
!u thermal
fesofhnrmat
Oligothermal •
Sceno thermal
letathermal
Eurythermal
Eulicho tophilous
lesolich tophi lous
Polylichtophilous
Dligolichtophilous
LimnobionCic
Llmnophilous
Indifferent
Rheophilous
Elheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Eptbenthic
Embenthic
Rplpelic
Episabulic
Eoilithic
Enixvlous
Epizooic
Epiphytic
Attached
Unattached
Htnter
Sprint
Summer
Autumn
Predator
Herbivore
Omnivdre
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Renion IX
Region X
i

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17

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Concensus
&
Notes















































                            180
-------
Taxon     Ehaenopsectra  Kieffer
aj
o
$
o
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£
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a



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General
Habitat
Specific
Habitat
tti
u
60 O
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a) A
Geographical
Distribution
ERS
arvae
ttpne
dults
cidobiontic
Lcidopbilous
eutral
Ikaliphilous
Ikalibiontic
ndifferent
olyhalobous
uhalobous
lesohalobous
)liEohalobous
luryhalinous
^utrophic
leso trophic
)liEOtrophic
3ys trophic
saprophilic
^acultative
Saproxenous
saoroohobic
luoxvohilous
lesoxvphilous
}liEOxvohilous

"InChermat
•feso thermal
31igo thermal
Btenothermal
[etathermal
iurythermal
Eulichotophilous
lesolichtophilous
Polylichtophilous
Jligolichtophilous
Limnbbiontic
Llmnophilous
Indifferent
Rheophilous
theobiontic
•larine
Estuary
Lake
Pond
River
Stream
SprinE
Other
Epibenthi'c
Embenthic
Eplpelic
Episabulic
Enilithic
Epixvlous
Epizooic
Eoirhytic •
Attached

I-Mnter
Spring
Summer

Predator
Herbivore
Omnivore
Scavenser
Region I
Region II
Reslon III
Reeion IV
Resion V
ReEion VI
Reeion VII
Reeion VIII •
Reeion IX
_ Re£ion_X 	
1

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X

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X
X

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Concensus
&
Notes












































































                               181
-------
Taxon;    Phaenopsectra profusa   (Townes)

V
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General
Habitat
Specific
Habitat
fee
u
i
60 O
3-g
•83
SS
Geographical
Distribution
a
u
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
AJkalibiontic
Indifferent
Polyhalobous
Euhalobous
Hesohalobous
Oligohalobous
Eurvhalinous
Butrophic
fesotrophic
Olieotrophic
Dystrophic
Saprqphilic
Facultative
Saproxenous
Saprophobic
Suoxvohilous
•fesoxvnhilous
nieoxvphilous
Anoxvphllmis
!uthermal
fcsnrhermal
Oligo thermal
Steno thermal
Metathermal
Eury thermal
Eulicho tophilous
Mesollchtophilous
Polylichtophilous
Dligolichtophilous
Limnobiontic
Limnophllous •
Indifferent
Kheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Enibenthic
Embenthlc
Eolpelic
Episabulic •
Eoilithic
Eoixvlous
Eoizooic
Epinhvtic
Attached
Unattached
Winter
Sprlnp
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Reeion IV
Reeion V
Reelon VI
Reeion VII
Reeion VHI
Roc Ion IX
Reeion X
22

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X






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X





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X


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X


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Concensus
&
Notes















































                             182
-------
Taxon:    Polypediluta apicatum  Towues
s
M
3
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cn

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General
Habitat
Specific
Habitat
be
p
I
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II
Geographical
Distribution

RES
arvae
upae
dults
cidobiontic
cidophilous
eutral
Ikaliphilous
Ikalibiontic
ndifferent
olyhalobous
uhalobous
esohalobous
lieohalobous
uryhalinous
utrophic
esotroohic
ligotrophic
ys trophic
aprophilic
^acultative

aprophobic
'uoxvohilous
lesoxvnhilous
)lieoxvphilous
Vnoxvphil mis


Dligothertnal
Stenothermal
etathertnal
Surythermal
Eulichotophilous
esolichtophilous
'olylichtophilous
Dligolichtophilous
jimnobiontic
jimnophilous
Indifferent
ttieophilous
Uieobiontic
Marine
Estuary
Lake
Pond
River
Stream
prinR 	


Eoinelic
Eoisabulic
Enilithic
Epixvlous


Attached 	
Unattached 	
Winter
Spring


Predator
Herbivore
Omnivore

Re'sion I
Resion II
Reeion III
Reeion IV
Region V

Reeion VII
Reeion VIII
j^^pion IX
Rppion X

27

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Concensus
&
Notes










































































                               183
-------
Taxon;    Polvpedilum aviceps   Townes

01
w
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Habitat
Specific
Habitat
tc
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•85
as
Geographical
Distribution
a)
s
j
Eggs
Larvae
Pupae
Adults
Acidobtontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhnlobous
Euhalobous
Mesohnlobous
Olieohalobous
luryhalinbus
Sutroohlc
lesotroohic
Oligotrophic
Dystrophic
Saprophilic
Facultative
Saproxenous
Saproohobic
Euoxvphilous
lesoxvohilous
)llB6xvnhllous
ftnoxvpM Ions
•llthermBl
fesot-Iinrmal
Dllgothernal
Stenothermal
'letathermal
Eurytheoial
Eulicho tophllous
(esolichtdphilous
folylichcopli Uous
Dligolichtophilous
Limnobiontic
[.imnophllous
Indifferent
-------
Taxott:    Polypedilum braseniae   (Leathers)
-------
Taxon;	Pglypedi.lt
                                (Walker)
01
g
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Habitat
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Habitat
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01. JS
01 0)
F*4 M
Geographical
Distribution
IRKS
larvae
Pupae
Adults
Acidobiontic
Acidophilous
teutral
AlkaliEhilous
Alkalibiontic
Indifferent
Polvhalobous
luhalobous
'lesohalobous
Olieohalobous
Euryhalinous
Sutrophic
Hesotrophic
Olieotroohic
Jystrophic •
Saprophilic
Facultative
Saproxenous '
Saproohobic
Euoxvphilous
Hesoxvphllous
Oligoxvphilous
Anoxvphtlons
Futhermal
Sfesothprmal
Oligothetmal
Steno thermal
Metathermal
Eurythermal
Eulichotophilous
Mesolicntophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Es tuary
Lake
Pond
River
Stream
Sorinz
Qt^her
Enibenthic
Embenthic
Kploellc
Eoisabulic
Eoilithlc
Epixvlous
Epizooic
Eoiphvtic
Attached
Unattached
Winter
Sprang
Summer
Auttimn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Reaion III
Recion IV
Reeion V
Reeton VI
Reelon VII
Reeion VIII
Reelon IX
Reeion X
17

X







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X


X

X





X



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X


X





X




X










X

x


X








X













































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes





































-




















                              186
-------
Taxon;  Polypedilum digitifer  Townes
OJ
o
p
3
O
w
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ff
4J

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s
5

to
4J

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3
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S
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0



|
-------
Taxon:
           Polypedilum fallax  (Johannsen)

o
w
a
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u
5
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General
Habitat
Specific
" Habitat
ff
i
II
11
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Geographical
Distribution
01
u
t-
j
Eggs
Larvae
Pupae
Adults
Acldobiontic
Acidophilous
Neutral
Alkallphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
OliBohalobous
Euryhalinous
Eutrophic
Mesotrophic
Oligotrophic
Dystrophic _
Saprophllic
Facultative
Saproxenous
Saproohobic
Euoxvphtlous
Mesoxvphilous
Olieoxvnhilous
Annxvphil otts
EltthGTTnal
Mesot-hermal
Oligo thermal
Stenothermal
Metathermal
Eurythermal
Eul icho tophilous
Mesolichtophilous
Polylichtophilous
Oligolich tophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprlne
Other
Enlbenthic
Embenthic .
EnipeJ.ic
Enisabulic
Enilithic
Epixvlous
Epliooic
Epinhvtic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
P.eRlon IV
Reeion V
Rceion VI
Reeion VII
ReEion VIII
Reeion IX
Reeion X
1

X







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X
X

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X


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X
X
X
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Concensus
&
Notes












































Found in submerged decompos-
ing wood and leaf drift.



                             188
-------
Taxon!  Polvnedilum halterale  (Coquillett)


•o (d
OJ J3
0) 
-------
Taxon!    Polypedilum illinoeosa   (Malloch)
' 
-------
Taxon;.  Polypedilum labeculosum   (Mitchell)


ec
4J




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5
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01
1
o
en
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
AJkalibiontic
Indifferent
'olyhalobous
^uhalobous
Mesohalobous
Dlisohalobous
Suryhalinous
Sutrophic
lesotrophic
Oligotrophic
Dystrophic
Saprophilic
Facultative
Saproxenous
Saproohobic
Euoxvphilous
Mesoxvphilous
OliKOxvohilous
Anoxvphilous
Euthermal
^eBofhnrmal
Oligothermal
Steno thermal
Hetathermal
Eurytherraal
Eulichotophilous
Mesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheopttilous
Rheobiontic
•larine
Sstuary
Lake
Pond
River
Stream
Spring
Other
Enibenthic
Embenthic
Epioelic
Episabulic
Enilithic

Eoizooic
EoinhvCic
Attached
Unaitached
Winter
Sprinp
Ajummer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IX
Region X
27

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Concensus
&
Notes














































































                              191
-------
Taxon:
           Polypedilum nigritum  Townes
0)
o
a
1
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0


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Habitat
4U
3S
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0) 03
CU «
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60
CO O
p
a v
tu to
Geographical
Distribution
'•RRS
Larvae
>upae
\dults
\cidobiontic
\cidophilous
Neutral
Mkaliphilous
Mkalibiontic
Indifferent
'olyhalobous
Euhalobous
^esohalobous
Dligohalobous
Suryhalinous
Eutrophic
lesotrophic
Dligotrophic
Dystrophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Euoxvphilous
fesoxvohilous
Dlieoxvphilous
Anoxvphilons
Ktlthermal
^pBOPhormal
aligothermal
Stenothermal
>[etachermal
Eurythermal
Eulichotophilous
Icsolichcophilous
Polyliclitophilous
aligolichtophilous
Cimnobiontic
Uimnophilous
Indifferent
Hheophilous
Uieobiontic
Marine
Estuary
,ake
Pond
River
Stream
Spring
Other
Epibenthic
Embenthic
Enioelic
Episabulic
Enilithic
Eoixvlous
Epizooic
Epinhvtic
Attached
Unattached
Winter


Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IX
. Region X 	
11

X











X


X








X



X

X





X






X





X








X


X










X











































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes








































































                              192
-------
Taxon;  Pglypedilum obtusum  Townes
4
1

M
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S

j-i
•5



5

3
0
q
3
o

CM




a
01




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H

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a)
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u
General
Habitat
u
•H 4J
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O -H
01 A
P. cd
n EC
U}
60 O
f!
01 
-------
Taxon:   Polypedllum Ontario   (Walley)
OJ
a
1

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General
Habitat
Specific
Habitat
ec
P
	 IT
§*
la
gs
Geographical
Discrlbution
2RS
.arvae
upae
dults
cidobionClc
cidoDhilous
eutral
Ikaliohilous
"Ikallbioncic
olyhalobous
uhalobous
Icsohalobous
Jlieohalobous
luryhallnoua
lu trophic
teso troohic
)lleotrophic
)vstrophic
Saprophilic
racultative
Saproxenous

Euoxvphilous
lesoxvohilous
31igoxvpbilous
AnnvvnM1ntt<:


Oligo thermal
Stenothermal
4cta thermal
•ur>'thermol
•ulichotophilous
^lesolichtophilous
'olylichcophllous
)ligolichtophilous
jimnobioncic
jimnophllous
CndifEerent
-------
Taxon:    Polypedilum scalaenum   (Schrank)
V
a
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en

tt
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tn


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Habitat
Specific
Habitat
ec
M
00 O
(3 *H
ss
QJ .n
fiS
Geographical
Distribution
Eggs
Larvae
Pupae
Adults
Acidofaiontic
Acidophilous
Neutral
Ukaliphilous
Alkalibiontic
Indifferent
'olyhalobous
iuhalobous
tesohalobous
Oligohalobous
Survhalinous
Sutrophic
Mesotrophic
Olieotrophic
Dys trophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Euoxvohilous
Mesoxyphilous
Oligoxvphilous
Anoxvphilous
Euthermal
Mesof-hfarmal
Oligothermal '
Stenothermal
Metathermal
Eurythermal
Eulictio topliilous
Mesolichtophiloiis
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous '
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Eoibenthic
Embenthic
Epipelic
Episabulic
Enilithic
Epixvlous
Eoizooic
Epiphvtic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IX
Region X
11

X











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X








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X





X






X






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27

X





X





X



X




X

X



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X













X

























































































































































































































































































































































































































































































































































































































































Concensus
&
Notes













































































                             195
-------
Taxon !__._Polypadilum simulans  Townes
1
o
VI

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5
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General
Habitat
Specific
Habitat
ec
n
I
uS
51!
•0 C3
-------
Taxon:  Polypedilum trigonum  Townes

01
tl
tl
V.
ffi
o
Salinity
C
-------
Taxon:  Bolypedllum tritum  CWalker)
01
u
h
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u




a



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5
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1
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3
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s
g
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General
Habitat
Specific
Habitat
tg
I
"H"
g"5
S§
0) &
S.&
Geographical
Distribution
RES
arvae
upae
dults
cidobiontic
cidophilous
eutral
Ikaliphilous
Ikalibiontic
ndifferent
olyhalobous
uhalobous
esohalobous
lieohalobous
uryhalinous
lutrophic
fesotrophic
Jligotrophic
Jystrophlc
Japrophilic
"acultative
Japroxenous
Jaorophobic
"uoxvphilous
tesoxvnhilo'us
lllcoxvohilous
Vnovvphiloiis

IpRof-hprmsl
)ligo thermal
5 teno thermal
let a thermal
Surythermal
Sulichotophilous
esollchtophilous
'olylichtophilous
31igol ichtophilous
.imnobioncic
.Iranophilous
^different
Uteophilous
UieobionCic
Marine
Istuary
Lake
Pond
River
Stream
Sorine
pj:her
Eoibenthic
Embenthic
Eplnelic
Eolsabulic
Enilithlc


F.ninhvClc
Att-ached
Unattached
Winter


Anftimn

Herbivore
Omnivore
Scavenger
Re R ion I
Region II
Region III
Re e ion IV
Region V
Region VI
Reeion VII
Region VIII
Rccion IX
Region X 	
11

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X








X



x

X





X






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X











































































































































































































































































































































































































































































































































































































































































































































Concensus
&
Notes













































































                               198
-------
Taxon:
           Potthastia  Kleffer
01
1
o
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a
3
ta

1


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§
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2
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4-1
§
3

General
Habitat
Specific
Habitat
60
p-
1
w
00 O
sv
•S5
01 0)
P4 W
Geographical
Distribution
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
lesohalobous
>ligohalobous
!urvhalinous
Eutrophic
•tesotrophic
Oligotrophic
Dystrophic
Saprophilic
facultative
Saproxenous
Saproohobic
Juoxvohilous
fesoxyphilous
Dlieoxvohilous
^oxvphll OUR
Edthermal
lesot-hermal
Oligo thermal
Stenothermal
Metathermal
Eurythermal
Eulichotophilous
Hesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Epibenthtc
Embenthic
Eoipelic
Episabulic
Epilithic
Eoixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IX
Reeion X
1

X







X



X.

X
X

X



X

X






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X






X




X
X






X




X
X
X



X





X






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X







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X







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X




X





X




X










X

X
X



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Concensus
&
Notes



























































                             199
-------
Taxon:
           Procladius adumbratus  (Johannsen)
01
1
W
«
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S




1


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Habitat
Specific
Habitat
Jf
i
g5
-H >
•O 0)
0) ,C
u ai
(14 M
Geographical
Distribution ,
IRBS
^arvae
>upae
\dults
icidobiontic
\cidophilous
Neutral
Mkaliphllous
Mkalibiontic
tn.dif£erent
'blvhalobous
^uhalobous
•tesohnlobou;;
31ieohalobous
Suryhallnous'
eutroohic
[esotroohic
^lieotroohic
Jvstrophic
Saoronhilic
Facultative
5a*proxenous
SaoroDhobic
iuoxvphilous
lesoxvohilous
31lBoxvphllous
^noxvph^ous
Euthermal
fpsot-hormal
Dligothermal
Stenothermal
^etathermal
Eurythermal
Sulicho tophilous
IcsolichCophilous
Polylichtopiiilous
31igolich tophilous
Limnobiontic
Limnophilous
Indifferent
Kheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sorine
Other
Epibenthic
Hmbenthic
F.nioelic
Eoisabulic
Enilithic
Eoixvlous
Eolzooic
Epinhvtic
Attached
Unattached
Winter
Spring
Riimmpr
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Reelon VIII
Region IX
Region _X 	
28

X





X





X

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X


X



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X

X


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X










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Concensus
&
Notes













































































                              200
-------
laxon: Procladlus bellus  
-------
Taxon; Proeladius culiciformis  (Linnaeus)
a
o
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Habitat U
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Uabitat
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11
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rHQt-i-f hut-ion B

P£S
arvae
upae 	
cidobiontic
cidophilous
eutral
Ikaliphilous
Ikalibiontic
ndifferent
olvhalobous 	 	
uhalobous
esohalobous
ligohaloboua 	
uryhalinous
utrophic 	
esotrophic
ligotrophic 	
ystrophie
aprophilic 	
acultacive
aoroxenous 	
aorophobie 	
luoxvpht^oiis
fesoxvphilous
llleoxvphllous 	
TitMienMl 	
**»«nt-hprmfll
Jligothermal
5 teno thermal
(etathermal
Sulichotophilous
lesolichtophilous
'olylichtophilous
Dllgolichtophilous
jimnobiontic
limnophilous
[ndifferent
ttieophilous
arine
Estuary
,ake
Pond ____ 	
River 	
pjrine 	 . 	
Epibenchic
Embentbig
EptpeUc
Epjgabulic
EpiHthlc 	
Epixvlous 	
Eolzooic 	
Epinhvtie 	
/\fl-»rhi.H
Mnter 	
Sprinr

Predator 	
Herbivore
Omnivore 	
ReRion I
Region II 	
Region III " 	
Region IV
Region V
Hpftton VI
Region VII 	
Region VIII 	
Region IX 	

7

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-
-1 	 1 	 	 -
                                202
-------
Taxon;   Procladius  denticulatus   Sublette

1
u
w
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a
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General
Habitat
Specific
Habitat
60
JJ
60 O
C -H
•H >
ng «
0) "S
Geographical
Distribution
-------
Taxon:    Procladlus rlparius   (Malloeh)


«
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1
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w
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arvae
upae
dults
cidoblonCic
tcidophilous
Icutral
akallphilous
^Ikalibiontic
ndlfferent
*olvhalobous
luhalobous
Icsohnlobous
)l±Eohalobous
luryhalinous
'utrophic
lesocrophic
)liEOtroohic
Jystrophic
japtophilic
facultative
Saproxenous

Suoxvphilous
Tesoxvphtlous
Dlieoxvphilous
\nowphiloiis


Dligothermal
Stenothermal
[etathertnal
iiirythermal
iulichocophilous
lesoLichtophilous
^lylichtophilous
Uigollchtophilous
jimnobiontic
jlmnophilous
Cndifferenc
theophilous
itheobiontic
larine
Estuary
Lake

River
Stream
Snrine
pther
F.nibenChic
Embenthic
Enioe3,ic
Eoisabulic
Eoilithic
Epixvlous
Epizooic
Epiphvtic

Unattached'
Winter

Bummer


Herbivore
Omni vo re
Scavenger
Re R ion I
Region II
Region III
Recion IV
Re B ion V
Rettion VI
Reeion VII
Reeion VIII
Reeion IX
_ JtegJLon X 	
78

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Concensus
&
Notes


























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                               204
-------
Taxon!  Prodiamesa bathvohila   (Kieffer)

>
u
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C
rt


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Habitat
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Habitat
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Distribution
c
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
.ndif ferent
Polyhalobous
Euhalobous
Mesohalobous
Oligohalobous
Survhalinous
Eutrophic
[esotrophic
Oligotrophic
Jystrophic
Saprophilic
l"acultatlve
Saproxenous
Saorophobic
Euoxvphilous
.esoxvphilous
ligoxvphilous
noxvphilous
^thermal
esotrhermal
ligothermal
teno thermal
letathermal
ury thermal
ulichotophilous
esolichtophilous
olylichtophilous
ligolichtophilous
imnobiontxc
imnophilous
ndifferent
heophilous
Sheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Epibenthic
Embenthic
Epipelic
Eoisabulic
Epilithic
Epixvlous
Epizooic
Epiphytic
Attached
Unattached
i-finter
Soring
Summer
Autumn
Predator
Herbivore
Cfmnivore
Scavenger
ReRion I
Region II
Region III
Region IV
Region V
Region VI
Region VII •
Reeion VIII
Region IX
Region X
6

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Concensus
&
Notes










































•




                            205
-------
Taxon:
           Prodiamesj^ ollvacea  (Meigen)
s
u
1

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Psectrocladius elatus   Roback
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u
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Habitat
Specific
Habitat
tic
M
60 O
C -H'
•H >
13 rt
0) .C
a) upae
Wults
Acidobiontic
Acidophilous
Meutral
Alkalipjiilous
Alkalibiontic
Indifferent
'olyhalobous
Suhalobous
lesohalobous
Olieohalobous
Euryhalinous
Sutrophic
Hesotrophic
Olisotrophic
)ystrophic
Saprophilic
facultative
Saprbxenous
Saoroohobic
Euoxvphilous
Mesoxvphilous
Olieoxvohilous
Anoxvphil otis
Euthermal
Mesothermal
Oligothermal
Stenothermal
Metathermal
Eury thermal
Eulichotophilous
Hesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Epibenthic
Embenthic
Enioelic
Episabulic
ERilithic
Epixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IX
Region X
1

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X

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Concensus
&
Notes











































































                   207
-------
Taxon: Psectrotanypus dyarl  (Coquillett)


s
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laxon:    Psectrotanypus venustus   (Coquillett:)


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Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkalipjiilous
Alkalibiontic
Indifferent
3olyhalobous
Suhalobous
•lesohalobous
OliEohalobous
Euryhalinous
i)u trophic
•fesotrophic
Oliaotrophic
)ys trophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Euoxvphilous
•lesoxvohilous
Olieoxvohilous
Annxvphiloiis
Euthermal
Mesothormal
Oligothermal
Stenothermal
Me ta thermal
Eurythermal
Eulichotophilous
Mesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Enibentliic
Erabenthic
Eninel.ic
Episabulic
Enilithic
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Reeion VII
Region VIII
Region IX
Reeion X
27

X





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X

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Concensus
&
Notes













































































                              209
-------
Taxon:
           Psectrocladius vernalis   (Malloch)
-------
Taxon: Pseudochironomus aix  Townes
s
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Habitat
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Geographical
Distribution
-ggs
_iarvae
'upae
Adults
icidobiontic
^cidophilous
Neutral
Ukaliphilous
Ukalibiontic
indifferent
"olyhalobous
Suhalobous
•lesohalobous
31iRohalobous
Suryhalinous
Eutrophic
tesotrophic
Dlieotrophic
Dystronhic
Saprophilic
Facultative
Saproxenous
Saorophobic
Euoxvphilous
^esoxvphilous
Dligoxvpb,ilous
^noxvphi^otls
Ruthermal
^[Gsofhprmal
Oligo thermal
Steno thermal
^letathermal
Eurythermal
Eulichotophilous
Mesolichtophilous
Polylichtophilous
Oli golichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Epibenthic
Embenthic
Eoipelic
Episabulic
Enilithic
Epixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IX
Reeion X . . ... _
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Concensus
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Notes












































































                             211
-------
Taxon;   Pseudochironomus  chen   Townes
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Discrlbuclon
ERRS
Larvae
?upae
Adults
AcidobionCic
Acidophilous
Neutral
Alkaliphllous
Ukalibiontic
IndifferenC
Polyhalobous
Euhalobous
•tcsohnlobous
)lieohalobous
Eurvhalinous
Eutrophic
Kesocrophic
OlieoCrophic
Dyscrophic
Saprophilic
Faculcacive
Saproxenous
Saprophobic
Euoxvphllous '
Mesoxvphilous
Oli^oxvphilous
Anoxvphl 1 oits
Eufchermal
MesoHiprmaT
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SCenochermal
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Eurychermal
Eulichotophilous
Hesolichcophilous
PolylichCophilous
Oligolichcophilous
LimnobionCic
Limnophilous
IndifEerenC
Rhcophilous
RheobionCic
Marine
Escuarv
Lake
Pond
River
Scream
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Eolbenchlc
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Eplsabulic
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Epixvlous
Epizooic
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Unnfctached
Winter
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Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
ReRlon III
P,eElon IV
Reeion V
Reaion VI
Reelon VII
Realon VIII
Renttin TX
Reslon X
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Concensus
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                              212
-------
Taxon:     Pseudochironomus fulviventris  (Johannsen)
-------
Taxon;
          Pseudochironomus richardsoni  Malloch


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Pupae
Adults
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Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Folyhalobous
Euhalobous
Mesohalobous
Olisohalobous
luryhalinous
Eutrophic
•tesotrophic
DliKOtroohic
Oystrophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Euoxvphilous
Kesoxvphilous
Olieoxvphilous
Anoxvphllons
Euthermal
Mesothnrmal
Oligothermal
Stenothermal
Metathermal
Eurytherraal
Eulichotophilous
Mesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Kheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
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Epibenthic
Erabenthic
Eolpelic
Episabulic
Eptlithic
Epixvlous
Epizooic
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Attached
Unattached
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Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Reeion III
Region IV
Region V
Region VI
Reeion VII
Region VIII
Reeion IX
Reeion X
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Concensus
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                             214
-------
Pseudodiamesa  Goetghebuer
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Sggs
^arvae
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Mkaliphilous
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Polyhalobous
Euhalobous
lesohalobous
Dlisohalobous
Euryhalinous
Eutrophic
fesotrophic
31iEOtroohic
Dystrophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Euoxvohilous
^fesoxvphilous
DliEOXVDhilous
Euthermal
fesofhermal
Dligothermal
Stenothermal
rtetathermal
Eurythermal
Eulichotophilous
Hesoiichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
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Enibenthic
Embenthic
Eoioelic
Eoisabulic
Eoilithic
Epixvlous
Eoizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IK
Region X
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Concensus
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Notes












































































                   215
-------
Taxon:
           Rheocricotopus robacki   (Beck)

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Larvae
Ptipne
Adults
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Acidophllous
Neutral
Alkallphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
OliRohalobous
Suryhalinous
Eutrophic
Mesotrophic
Olieotrophic
3ystrophic
Saprophilic
facultative
Saproxenous
Saprophobic
Euoxvphllous
fesoxvphilous
JHeoxvDhilous
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fesothermal
Oligothermal
Stenothermal
[etathermal
Eurythermal
Eulichotophilous
lesolichtophilous
Polylichtophilous
Dllgolichtophilous
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Limnophilous
Indifferent
flheophilous
Elheobiontic
Marine
Estuary
Lake
Fond
River
Stream
Spring
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Embenthic
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Episabulic
Eoilithic
Eoixvlous
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Unattached
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Autumn
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Omnivore
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Region I
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Reelon III
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Region VI
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ReRion VIII
Reeion IX
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                            216
-------
Taxon:
           Rheotanytarsus exiguus   Johannsen
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Distribution
Eggs
Larvae
Pupae
Adults
Acldobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
)ligohalobous
Euryhalinous
Eutrophic
•lesotrophic
Oligotrophic
Dystrophic
Saprophilic
facultative
Saproxenous
Saprophobic
Suoxyphilous
Mesoxvphilous
Oligoxvohilous
Anoxvphilous
Euthermal
Mesof-hprmal
Oligo thermal
Stenothermal •
Metathermal
Eury thermal
Eulichotophilous
Mesolichtophilous
Polylich tophi lous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sorine
Other
Epibenthic
Embenthic
Epinelic
Enisabulic
Enilithic
Epixvlous
Epizooic
EDinhvtic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III •
Region IV
Region V
Region VI
Region VII •
Region VIII
Reeion IX
Reeion X
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                             217
-------
laxon:   Sergentia coracina  (Zetterstedt)
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Taxon:   Smittia aterrima   (Meigen)
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'upae
kdults
cidobiontic
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Ukaliphilous
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'olvhalobous
3uhalobous
Eesohalobous
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:uryhalinous
:utrophic
[esotrophic
)lieotrophic
5vs trophic
aaprophilic
racultative
Saproxenous
Saprophobic
Suoxvph ilous
lesoxvphilous
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•Ip^nrtr-rmal
Dligochermal
Scenothermal
tetachermal
iurythermal
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^esoLichtophiious
Polylichtophilous
Dligolichcophilous
Lironobioncic
Limnophilous
Indifferent
theophilous
Uieobioncic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Epibenthic
Embenthlc
Enlpel.ic
Enlsabullc
Epilithic
Enixvlous
Eplzooic
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Unattached
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Reaion II
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Reeion VIII
Reeion IX
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                              220
-------
Taxon:    Stenochironomus hilaris   (Walker)
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-ggs
jarvae
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Adults
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leutral
Mkaliphilous
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3olyhalobous
luhalobous
•lesohalobous
JliEohalobous
-uryhalinous
Eutrophic
lesotrophic
Dlieotrophic
^ystrophic
Saprophilic
Facultative
Saproxenous
Saorophobic
Euoxvphilous
fesoxvphilous
Dlieoxvphilous
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Dligothermal
Stenothermal
'•letathermal
Eurythermal
Eulichotophilous
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Dligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
EUieobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Epibenthic
Embenthic
Eoioe] ic
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Enilithic
Epixvlous
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Unattached
Winter

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Scavenger
Region I
Region II
Reelon III
Region IV
Region V
Region VI
Region VII
Rezion VIII
Region IX
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Concensus
&
Notes






















































Burrows in submerged wood
and leaf drift.

















                              221
-------
Taxon: Stenochironomus  macateei   (Malloch)

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leutral
Ukaliphilous
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Polyhalobous"
Euhalobous
fesohalobous
Dligohalobous
Suryhalinous
Eutrophic
fesotrophic
Dlieotrophic
Dystrophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Euoxvpli ilous
fesoxvphilous
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Taxon:
          Stietochironomus devinctus  (Say)

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Taxon:   Stlctochironomus  varius  Tomes
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Stenochermal
letathermal
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lesolichtophilous
Polylichtophilous
aligolichtophilous
Limnobiontic
Amnophilous
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Uieobiontic
larine
Estuary
Lake
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River
Scream
Sprine
Other
Epibenthic
Enbenthlc
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Scaveneer
Region I
Reeion II
Region III
Reeion IV
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Reeion VI
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Reeion VIII
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Notes













































































                               221  .
-------
Taxon:
           Svmpotthastia   Pagast

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Distribution
a
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Eggs
Larvae
Pupae
Adults
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Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
Oligohalobous
Euryhalinous
Eutrophic
tesotrophic
Oligotrophic
Dys trophic
Saprophilic
Facultative
Saproxenous
Saorophobic
Euoxvphilous
•lesoxvphilous
31ieoxvphilous
Anoxvphilous
^uthermal
•lesot-hBrmal
OligotherraaJ.
Stenothermal
Metathermal
Eurythermal
Eulichotophilous
Hesolichtophilous
Polylichtophiious
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Elheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Epibenthic
Embenthic
Enipelic
Episabulic
Epilithic
Epixvlous
Epizooic
Epiphytic
Attached .
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II.
Region III
Region IV
Region V
Region VI
Region VII •
Region VIII
Reeion IX
Reeion X
1

X







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X

X



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X
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X
X
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X

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Concensus
&
Notes















































                             225
-------
Taxon:    Tanypus  carinatus  Sublette
1
o
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3
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-------
Taxon-:
           Tanypus clavatus  Beck
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a cu
h n
Geographical
Distribution
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
lesohalobous
Olieohalobous
Suryhalinous
Eutrophic
'lesotrophic
Oliaotrophic
)ys trophic
Saprophilic
'acultative
Saproxenous
Saproohobic
Suoxvphllous
•lesoxvphilous
Uieoxvohilous
Anoxvohll ous
Huthermal
•^eso^hBrma^
Oligothermal
Stenothermal
letathermal
Eurj'thermal
Eulichotophilous
lesolichtophilous
Polylichtophilous
Oligolichtophilous
.imnophilous ''
ndifferent
heophilous
Rheobionbic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Epibenthic
Embenthlc
F.pipelic
Episabulic
Epilithic
Epixvlous
Eoizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Aueumn
Predator
Herbivore
Omnlvore
Scavenger
Region I
Region II
Region. Ill
Region IV
Region V
Region VI
Keelon.VII
Resion VIII
.Reeion IX
Region X
1

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X

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Concensus
&
Notes














































                             227
-------
Taxon:    Tanypus  grodhausl  Sublette
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General
Habitat
Specific
Habitat
to
u
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Geographical
Distribution
KES
nrvae
upae
dults
cidoblontic
Lcidophilous
eutral
dkallphilous
IkalibionCic
ndifferent
olvhalobous
*uhalobous
tesohnlobous
)lieohalobous
^uryhnlinous
:utrophic
lesotronhic
)lieotroDliic
)vstroi)hic
>aprophilic
"acultative
Saproxenous
aapronliobic
'qo}-------
Taxon;     Tanypus parastellatus  Sublette

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Taxon!   Tanvous neopunetipennls  Sublette
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Geographical
Distribution
-KRS
^arvae
upae
id u Its
icidobiontic
tcidophilous
eutral
akaliphilous
Ukalibiontic
[ndi£ferent
'olyhalobous
'uhalobous
iesohalobous
3llgohalobous
luryhalinous
"utrophic
•lesotroDhic
)li'eotroDhic
3vs trophic
Saprophilic
Facultative
Saproxenous
Snproplioblc
Euoxvphilous
esoxvphilous '
31iBOXVphilous

Rtt thermal
feso thermal
Dllgo thermal
Stenothermal
etathermal
iurytherraal
^ulichotophilous
^tesolichtopliilous
•olylichcoph llous
)Llgolichtophilous
jirMiobiontic
jimnoph llous
Endifferent
Uieophilous
Uieobiontic
larine
Estuarv
Lake
Pond
River
Stream
Sorine
Other
Eoibenthic
Embenthic
Knioelic
Eoisabulic
Epilithic
Eoixvlous
Epi^ooic
Eoinhvtic
Attached
Unattached
Winter
Pprt^p
Guqpier

Predator
Herbivore
Omnivore
Scavenger
Region I
Reeion II
Reeion III
Region IV
Reeion V
ReEion VI
Reeion VII
Reeion VTII
Re c ion IX
Region X 	
11

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Concensus
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                               230
-------
Taxon;    Tanypus punetipennis  Meigen

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IEggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
A3kalibiontic
Indifferent
Polyhaloboua
Euhalobous
Mesohalobous
Olinohalobous
Suryhalinous
Eutrophic
•lesotrophic
Olieotrophic
Dystrophic
facultative
Saproxenous
Saprophobic
luoxvphilous
lesoxvohilous
)lieoxvDhilous
Inoxvphllous
Cuthermal
•tesofhormal
Oligothermal
Stenochermal
letathermal
Eurythermal
Eulichotophilous
lesolichtophilous
Folylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Kheophilous

Estuary
Lake
Pond
River
Stream
Sprinz
Other
Epibenthic
Embenthic
Epipelic
Episabulic
Epilithic
Epixvlous
Epizooic
Epiphytic
Attached
Jnattached
•/inter
Soring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
ReRion I
Reeion II
Jeglon III
Reeion IV
teeion V
Reaion VI
Reeion VII
leeion VIII
^ei-ton IX
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                            231
-------
Taxon:
           Tanypus stellatus  (Coquillett)



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                               232
-------
laxon: Tanytarsus buckleyi  Sublette


c
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General
Habitat
Specific
Habitat
00
M
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ii
Bographical
Istribution
U 0



Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
AJkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
OliEohalobous
Suryhalinous
EutroDhic
•lesotroohic
Olieotrophic
Dys trophic
Sapronhilic
facultative
Saproxenous
Saproohobic
luoxvohilous
•tesoxvphilous
Olieoxvohilous
Anoxvphilons
!uthermal.
•fesothermal
Qligo thermal
Stenothermal
[etathermal
Eurythermal
Eulichotophilous
Mesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
ndif ferent
Rheophilous

Marine
Estuary
Lake
Pond
River
Stream
Spring
Dther
Epibenthic
Embenthic
Epipelic
Episabulic
Epilithic
Eoixvlous
Eoizooic
EoiDhvtic
Attached
Unattached
i-Hnter
Sorintf
Summer
Autumn
?redator
Herbivore
Omnivore
Scavenger
Region I
Region II
Segion III
legion IV
teeion V
JeKion VI
leEion VII
leeion VIII
leeion IX
Jeeion X
12

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X

X


X







X




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X


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Concensus
&
Notes















































                            233
-------
Taxon: Tanvtarsus confusus  Malloch
-------
Taxon: Tanytarsus dendyi  Sublette
I
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Habitat
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Habitat
to
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Geographical
Distribution
Eggs
Larvae
Pupae
Idults
Acidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
lesohalobous
Olieohalobous
Suryhalinous
iutrophic
lesotroohic
OliEOtrophic
)ys trophic
Saoroohilic
Facultative
Saproxenous
Saoronhobic
.uoxvphilous
fesoxvohilous
lli'eoxvphilous
inoxvphilmis
Suthermal
fesofhermal
Oligothermal
Stenothermal
lecathermal
Eurythermal
Cuiichotophilous
lesolichtophilous
Polylichtopltilous
Oligolichtophilous
Limnobiontic
Limnophilous
ndif ferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sorina
Other
Epibenthic
Embenthic
Eoioelic
Eoisabulic
Eoilithic
Eoixvlous
Epizooic
Epiohvtic
Attached
Unattached
Winter '
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Kezion V
Region VI
Region VII •
Region VIII
Reeion IX
Rezl,on X
12

X







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Concensus
&
Notes















































                            235
-------
Taxon: Tatiytarsus neoflavellus  Malloch
 J3
££
Geographical
Distribution
:RES
^arvae
}upae
.dults
Icidobiontic
kcidophilous
leutral
Ukaliphilous
Vlkallbiontic
[ndifferent
'olyhalobous
^uhalobous
•tesohalobous
)licohalobous
Zuryhalinous
^utrophic
•tesotrophic
31iEOtrophic
Jystrophic
Sap'rophilic
Facultative
5'aproxenous
Saprophobic
Euoxvphilo'us
^fesoxvphilous
31i2oxvphilous



Dligothermal
Stenothermal
Mecachermal
iurychermal
Eulicho tophilous
lesolichtophilous
Polylichtophilous
HlgolichCophilous
Limnobioncic
Limnophilous
Indifferent
Uieophllous
Uteobiontic
tarine
Estuary
Lake
Pond
River
Stream
Sorine
Other
Eoibenthic
Embenthic
Eoloelic
Episabulic
Epilithic
Epixvlous
Epizooic
Epiohvtic
ACT^flched
Unattached
Winter

puroflier"
Autumn
Predator
Herbivore
Otnnivore
Scavenger
Region I
Region II
Reeion III
Region IV
Reeion V
Reeion VI
Reeion VII-
Reeion VIII
Region IX
. £szi2n_X 	
2

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Concensus
&
Notes













































































                               236
-------
Taxon!  Tanytarsus quadratus  Sublette
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u
General
Habitat
Specific
Habitat
6C
a
1
23
•H >
•a cu
s-s
h n
Geographical
Distribution
Eggs
Larvae
Pupae
Adults
Aeidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Indifferent
Polyhalobous
Euhalobous
Mesohalobous
Olisohalobdus
Suryhalinous
Eutrophic
Hesotrophic
Olieotrophic
Jystrophic
Saprophilic
facultative
Sap roxenous
Saorophobic
Luoxvphilous
Mesoxvphilous
Oli^oxvohllous
Anoxvphilous
Euthermal
^fesot•hf!n^Al
Ollgothermal
Stenothermal
Hetathermal
Eurytherraal
Eulichotophilous
Hesolichtophilous
Polylichtophilous
Oligol ichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Eoibenthic
Embenthic
Eolpelic
Ep'isabulic
Eoilithic
Eoixvlous
Epizooic
Epiohvtic
Attached
Unattached
Winter
Sorine
Summer
Autumn
Predator
Herbivore
Omni vo re
Scavenger
Region I
Region II
Region III
Region IV
Region V
Reeion VI
Region VII
Region VIII
Region IX
Region X
12

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X
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X





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Concensus
&
Notes















































                            237
-------
Taxon: Tanytarsus recens   Sublette
01
jj
i

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Habitat
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Habitat
u
it
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Geographical
Distribution
:RRS
jarvae
'upae
Idults ,
•cidobiontic
Icidophilous
ieutral
Ukaliphilous
.Ikalibiontic
:ndifferent
3olvhalobous
"uhalobous
lesohalobous
Higohalobous
luryhalinous
'utrophic
•leso trophic
)ligotrophic
Jvstronhlc
Saprophilic
Facultative
Saproxenous
Saproohobic
Euoxvphilous
^lesoxvohilous
31i?oxvphllous

Rut-hprmnl
*feqo tTipnnal
Bligothermal
Stenochermal
letathermal
tarythermal
Bulichotophilous
Hesollctltophilous
Polylichtophilous
)ligolichtophilous
Limnobiontic
jimnophilous
Indifferent
Rheophilous
Uieobiontic
Marine
Escuarv
Lake
Pond
River

Sprine
Other
Epibenthic
Embenthic
ppipe3.ic
Episabulic
EoiHthic
Epixvlous
Epizoolc
Epiohvtic
Attached
Unattached
Winter



Predator
Herbivore
Omnivore
Scavenger
Region I
Ree-ion I£
Reeion III
Region IV
Reeion V
Reeion VI
Region VII
Region VIII
Region IX
Reeion_X 	
1

X







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Concensus
&
Notes












































































                               238
-------
Taxon:  Tanytarsus  xanthus   Sublette
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.Geographical
Distribution
Rgs
arvae
upae
adults
icidobiontic
Lcidophilous
leutral
akaliphilous
ilkalibiontic
ndifferent
>olyhalobous
^uhalobous
lesohalobous
)ligohalobous
]uryhalinous
'utrophic
lesotrophic
)ligotrophic
)ys trophic
Saprophilic
?acultative
Saproxenous
Saorophobic
:uoxvphilous
•fesoxyphilous
Dlieoxvohilous
\noxvph-f.1piis

^esnt-hprmal
31igothermal
Steno thermal
letathermal
iurythermal
Eulichotophilous
lesolichtophilous
Polylichtopliilous
pligolichtophilous
Limnobiontic
jimnophilous
[ndifferent
Rheopliilous
Uieobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sorine
Other
Eoibenthic
Embenthic
Epioelic
Eoisabulic
Enilithic
Epixvlous
Eoizooic
Epiphytic
Attached
Unattached
Winter
Spring


Predator
Herbivore
Omnivo re
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IX
Reeion X 	
12

X







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X
X
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X
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Concensus
&
Notes










































































                              239
-------
Taxon:	Telmatogeton japonicus  Tokunaga



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Larvae
Pupae
Adults
Acidoblontic
Acidophilous
Neutral
Alkallphllous
Alkallblontic
Indifferent
Polyhalobous
Euhalobous
Kesohalobous
Olieohalobous
Euryhalinous
Eutrophic
tfesotrophlc
OliEO trophic
Bys trophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Euoxvphilous
^esoxvohilous
Olieoxvphilous
Anoxvphilons
^ti thermal
Mesothormal
Oligo thermal
Steno thermal
Met a thermal
Eury thermal
Eulicho tophilous
^ esolichtophiious
Polylichtophilous
Oligolich tophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobioncic
Marine
Estuary
Lake
Pond
River
Stream
Sprine
Other
Enibenthic
Erabenthic
Epioelic
Eoisabulic
EoiliChic
Eoixvlous
Eoizooic
Eoiohvtic
Attached
Unattached .
H'fnter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Reeion III
Region IV
Reeion V
ReRion VI
Reeion VII
ReKion VIII
Reeion IX
Region X
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Concensus
&
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                             210
-------
Taxon;   Thalassomyia bureni  Wirth

13 (tf
aj.j:
££
Geographical
Distribution
Eggs
jarvae
'upae
\dults
kcidobiontic
\cidophilous
Neutral
\lkaliphilous
\lkalibiontic
Indifferent
Polyhalobous
Duhalobous
^lesohalobous
Dligohalobous
Euryhalinous
Eutrophic
^lesotrophic
Sligotroohic
Dystrophic
Saprophilic
facultative
Saproxenous
Sapronhobic
Euoxyphilous
^fesoxyphilous
Oligoxvphilous
Anoxvphilous
Euthermal
'•Tesothermal
Oligothermal
Stenothermal
4etathermal
Eurythermal
Euliciiotophilous
^lesolichtophilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
jimnophilous
Indifferent
Rheophilous
Elheobiontic
•larine
Estuary
Lake
Pond
River
Stream
Soring
Other
Eoibenthic
Embenthic
Eoioelic
Eoisabulic
Eoilithic
Epixvlous
Epizooic
Epiphytic
Attached
Unattached
1-Hnter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Reeion V
Region VI
Region VII
Region VIII
Reeion IX
Region X _ 	
31

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-------
laxon;    Thienemannimyia barberi  (Coquillett)
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Geographical
Distribution
?8SS
^arvae
'upae
\dults
\cidobiontic
Vcidophilous
{eutral
JLkaliphilous
Mkalibiontic
indifferent
'olvhalobous
Suhalobous
fesohnlobous
31it*ohalobous
Suryhnlinous
Sutrophic
(esotrophic
Dlieotrophic
Oys trophic
Saprophilic
Facultative
Saproxenous
Saprophobic ••
Euoxvph tlous
fesoxvph ilous
alicoxvphilous
^noxvph^^o^lR
Ruthermal
ff»e;nr!if»rmal
DligoChormal
Stenothermal
lletathermal
Eurythermal
Eullcho cophilous
Hesolichtophilous
Poiylichtophilous
Oligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Kheophilous
Uieobiontic
Marine
Estuary
Lake
Pond
River
Stream
Sprinz
Other
Eoibenthic
Embenthic
Eplpel.ic
Episabulie
Eoiliehlc
Epizooic
Eninhvtic
AttSchet?
Unattached
Winter
Spring
Summer

Predator
Herbivore
Omnivore
Scavenger
Region I
Region It
Recion III
Reaion IV
Region V
Reeion VI
Reeion VII •
Reeion VIII
Rot! Ion TX
Re(!ion_X 	
27

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Concensus
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Notes





































































                              242
-------
Taxon!   Thienemanninyia senata  (Walley)
1
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Habitat
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Habitat
at
n
00 O
ss
91 J3
m m
fo n
Geographical
Distribution
3RKS
jarvae
Pupae
Wults
\cidobiontic
\cidophilous
Neutral
Mkaliphilous
Ukalibiontic
Indifferent
Polyhalobous
Suhalobous
lesohalobous
Dligohalobous
Euryhalinous
Sutrophic
lesotrophic
Dlieotrophic
Dystrophic
Saprophilic.
Facultative
Saproxenous
SaDrophobic
Euoxvohilous
^esoxvDhilous
Oli^oxvohilous
Anoxvphilniis
Ruchermal
^esofhemiat
Oligothertnal
Stenothermal
Metathermal
Eury thermal
Eulichotophilous
Mesolichtoptiilous
Polylichtophilous
Oligolichtophilous
Limnobiontic
Litnnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
2s tiiarv
Lake
Pond
River
Stream
Spring
Other
Eoibenthic
Embenthic
Enioelic
Eoisabulic
Eoilithic
Eoixvlous
Eoizooic
Epinhvtic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region 'III
Region IV
Region V
Region VI
Region VII
Region VIII
Region IX
Region X
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Concensus
&
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                              243
-------
Taxon:  Thlenemaniella^xena	Roback
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Habitat
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Geographical
Distribution
:RRS
Larvae
?upae
Mules
Acidobiontic
/^cidophilous
Neutral
AlkaliEhilous
Wkalibiontic
Indifferent
^olyhalobous
Euhalobous
iesohalobous
OliEohalobous
Eurvhallnous
^utrophic
iesotronhic
Oliizotrophic
)ystrophic
Saprophilic
Facultative
Saproxenous
Saorophobic
Euoxvphtlous
Mesoxvphllous
Olieoxvohilous
Anoxvphl 1 oils
Euthermal
WesotTiprmal
)ligo thermal
Stenothermal
Ketathermal
Burythermal
Eullchocophilous
Mesolichtophilous
Polylichtophilous
Oligolichtophilous
t7imnobiontic
Linnophilous
Indifferent
Elheophilous
Rheobiontic
•larine
Estuary
Lake
Pond
River
Stream
Spring
Other
Eoibenthlc
Embenthic
Eoipelic
Eplsabulic
Enllfchic
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Auttimn
Predator
Herbivore
Omnlvore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Reeion VIII
Reclon IX
Reeion X
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                             211
-------
      Taxon:    Trlbelos sp. A  (Beck)*
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Habitat
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Habitat
60
n
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-------
Taxon: Tribelos fuscieornis  (Malloeh)


ex
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Taxon:     Tribelos jucundus  (Walker)
-------
Taxon: Tribelos quadrlpunctatus   (Malloch)
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Habitat
Specific
Habitat
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-
Geographical
Distribution
ERRS
Larvae
Pupae
Adults
Acldobiontic
Acldophllous
Neutral
Alkallphllous
Alkallbiontic
Indifferent
Polyhalobous
luhalobous
lesohalobous
Oligohalobous
Eurvhalinous
Sutrophic
•fesotrophic
OliEOtrophic
Dystrophlc
Saprophllic
Facultative
Saproxenous
Saprophobic
Euoxvphilous
Mesoxvphilous
Olieoxvphiloua
Anovvph-tloiis
Euthermal
MCSOthPTTHfll
Oligotherrcal
Stenothermal
Metathermal
Eurychermal
Eullcho tophllous
Kesolichtophllous
Polylichtophilous
Dllgollchtophllous
Llmnobiontlc
Llmnophllous
Indifferent
Rheophllous
Rheoblontlc
Marine
Estuary
Lake
Fond
River
Stream
Spring
Other
Epibenthlc
Embenthic
Epinellc
Eplssbullc
Epillthlc
Epixvlous
Eoizoolc
EpinhvCic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Reelon III
Region IV
Reeion V
Reeion VI
Reeion VII
Reeion VIII
Reelon IX
Reeion X
1

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Concensus
&
Notes











































Burrows In the petioles of
the yellow pond lily, Nuphar.



                             218
-------
Taxon:
           Trissocladius  Kieffer
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Habitat
Specific
Habitat
60
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Geographical
Distribution
~'KRS
jarvae
'upae
Vdults
Vcidobiontic
Vcidophilous
leutral
Ukaliphilous
U.kalibiontic .
Indifferent
'olyhalobous
^uhalobous
lesohalobous
)ligobalobous
Zuryhalinous
^utrophic
fesotrophic
HiKotroohic
Dystrophic
Saprophilic
Facultative
Saproxencus
Saprophobic
Suoxvphilous
fesoxvohilous
31ipoxvphilous
\noxvphilotis
Sutiierinal
"lesothermaL
Dligothermal
Stenothermal
letathermal
Eury thermal
Bulichotophilous
lesolichtophilous
Polylichtophilous
Dligolicheophilous
Uimnobiontic
Ijimnophilous
Indifferent
Rheophilous
Rheobioneic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Epibenthic
Embenthic
Epipelic
Episabulic
Eplliehic
Eoixvlous
Epizooic
Epiphytic
A.t tached
Una'tt'ached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omni vo re
Scavenger
Region I
R'eeion II
Region III
Region IV
Region V
Region VI
Region VII-
Region VIII
Reeion IX
Region X
1

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Concensus
&
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                              219
-------
Taxoni    Xenochironomus rogersi   (Beck S Beck)
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Geographical
Distribution
:KRO
.arvae
'upae
Idults
\cidobiontic
icidophilous
Neutral
Mkaliphilous
Mkalibiontic
Indifferent
Polyhalobous
Euhalobous
Icsohalobous
Dlleohalobous
Euryhalinous
Eutroohic
lesotroohic
OliBotroohic
)vstroohic
Saprophilic
Facultative
Saoroxenous
Saoroohobic
Euoxvohilous
^lesoxvohilous
Dlicoxvphilous
Anoxvphil oils
Euthcrmal
MpBof hprmol
Oligothermal
Steno thermal
ttetathermal
Eurythermal
£ulichotophilous
Hesolichtophilous
Polylichtopliilous
Qligolichtophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
larine
Estuary
Lake
?ond
River
Stream
Sorine
Other
Eoibenthic
Erabenthic
F.pipelic
Eoisabulic
Enilithic
Epizooic
Epinhvtic
Attached
Unattached
Winter
Spring
^^inim(>r
Autumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Reeion VII
Regio'n VIII
jjLeeion IX
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Concensus
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Notes








































































                              250
-------
Taxon;   Xenochironomus scopula  Townes
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Habitat
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Habitat
M
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ca
60 O
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fo m
Geographical
Distribution
Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Mkaliphilous
Mkalibiontic
Indifferent
'olyhalobous
iuhalobous
^lesohalobous
Dliebhalobous
Suryhalinous
Sutrophic
^tesotrophic
Oliaotrophic
)ystrophic
Saprophilic
Tacultative
Saproxenous
Saprophobic
Euoxvphilous
Mesoxvphilous
Olisoxvohilous
Anoxvphllous
Euthermal
^feso^^lprT^la^
Oligotherraal
Stenothermal
Hetatherraal
Eurythermal
Eulichotophilous
Mesolichtophilous
Folyiichtophiious
Oligoliciitophilous
Limnobiontic
Limnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
is tuary
Lake
Pond
River
Stream
Soring
Other
Epibenthic
Embenthic
Eoioelic
Epissbullc
Enilithic
Enixvlous
Epizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Aufcumn
Predator
Herbivore
Omnivore
Scavenger
Region I
Region II
Region III
Region IV
Region V
Region VI
Region VII
Region VIII
Recion IX
Region X 	
8

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Concensus
&
Notes





























































                             251
-------
Taxon:    Xenochironomus taenionotus  Say
to
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Habitat
ti
u
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£S
Geographical
Distribution
SBKS
jarvae
?upae
Adults
Icidobiontic
Acidophilous
Neutral
Alkaliphilous
Alkalibiontic
Cndifferent
?olvhalobous
!uhalobous
lesohalobous
Olieohalobous
Euryhalinous
Eutroohic
Hesotrophic
Olieotroohic
Dystrophic
Saprophilic
Facultative
Saproxenous
Saprophobic
Euoxvphilous
Kesoxvphilous
OHrB
-------
Taxon:
          Xenochironomus xenolabis  Kieffer


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Geographical
Distribution'
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§
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Eggs
Larvae
Pupae
Adults
Acidobiontic
Acidophilous
Neutral
Mkaliphil'ous
Alkalibiontic
Indifferent
'olyhalobous
Euhalobous
Mesohalobous
Olicohalobous
Suryhalinous
Eutrophic
Mesotrophic
Olieotirophic -
)ystrophic
Saprophilic
Tacultative
Saproxenous
Saprophobic
Suoxvphilous
Mesoxvphilous
Olisoxvphilous
Anoxvphilons
Rufchermal
^tesot-hprmal
Oligothermal
Stenothermal
Ketathermal
Eurythermal
Eullcho tophilous
^lesolichtophilous
Poly lich tophilous
Oligolichtophilous
Limnobiontic
[.imnophilous
Indifferent
Rheophilous
Rheobiontic
Marine
Estuary
Lake
Pond
River
Stream
Spring
Other
Epibenthic
Etnbenthlc
Epipelic
Episabulic
Epilithic
Epixvlous
Eoizooic
Epiphytic
Attached
Unattached
Winter
Spring
Summer
Autumn
Predator
Herbivore
Omni vo re
Scavenger
Region I
Region II
Region III
Regitfn IV
Region V
Region VI
Region VII-
Region VIII
Region IX
Reeion X
1

X







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                              253
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Taxon:
         Zavrelia  Kieffer
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Taxon:
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Autumn
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-------
                                    TECHNICAL REPORT DATA
                            (Please read Instructions on the reverse before completing)
i. REPORT NO.
  EPA-600/4-77-024
                              2.
                                                            3. RECIPIENT'S ACCESSION-NO.
4. TITLE AND SUBTITLE
  ENVIRONMENTAL REQUIREMENTS AND POLLUTION TOLERANCE
  OF COMMON FRESHWATER CHIRONOMIDAE
                                                            5. REPORT DATE
                                                              April  1977 issuing date
                                  6. PERFORMING ORGANIZATION CODE
7. AUTHOR(S)

  William M. Beck,  Jr.
                                                            8. PERFORMING ORGANIZATION REPORT NO.
9. PERFORMING ORGANIZATION NAME AND ADDRESS

  Florida A & M University
  Tallahassee,  Florida  32307
                                  10. PROGRAM ELEMENT NO.
                                     PE 1BD612
                                  11. CONTRACT/GRANT NO.

                                    R-802754
12, SPONSORING AGENCY NAME AND ADDRESS
 Environmental Monitoring and  Support Laboratory-Gin.,OH
 Office of Research and Development
"U.S. Environmental Protection Agency  ,
 Cincinnati, Ohio  45268
                                  13. TYPE OF REPORT AND PERIOD COVERED
                                   Final  4/8/74 to 12/31/75
                                  14. SPONSORING AGENCY CODE
                                      EPA/600/06
15. SUPPLEMENTARY NOTES
16. ABSTRACT
       Data  on the environmental requirements  and pollution tolerance of  230  taxa
  of freshwater chironomids were compiled from 33 references.   This compilation was
  prepared to assist biologists  in evaluating  data from macroinvertebrate samples
  collected  for the assessment  of water quality.   The following parameters were
  considered:   developmental  stage, pH, salinity, nutrients,  degradable dissolved
  organics,  oxygen, temperature, turbidity,  current, general  habitat, specific
  habitat, seasonal distribution.
17.
                                KEY WORDS AND DOCUMENT ANALYSIS
                  DESCRIPTORS
                                               b.lDENTIFIERS/OPEN ENDED TERMS
                                                c. COS AT I Field/Group
   Indicator  species
   Aquatic Biology
   Water Quality
   Environmental requirements
   Larvae
   Benthos
   Ecology
Fresh water
Chironomidae
Pollution  tolerance
                                                      6C
18. DISTRIBUTION STATEMENT
   RELEASE TO PUBLIC
                     19. SECURITY CLASS (ThisReport)'
                       UNCLASSIFIED
                          21. NO. OF PAGES
                              267
                     20. SECURITY CLASS (Thispage)
                       UNCLASSIFIED
                                                22. PRICE
EPA Form 2220-1 (9-73)
                                            261
                                                       U.S. GOVERNMENT PRINTING OFFICE: 1977-757-HIO/6581 Region No. 5-11
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