INVESTIGATIONS INTO BIOFOULING PHENOMENA
       IN FINE PORE AERATION DEVICES
                      by
           W. Jansen and J.W. Costerton
                 Microbios Ltd.
            Calgary, Alberta T2L 1G9

                     and

                 Henry Melcer
           Wastewater Technology Centre
           Burlington, Ontario L7R 4AD
        Cooperative Agreement No. CR812167
                 Project Officer

               Richard C. Brenner
Water and Hazardous Waste Treatment Research Division
       Risk Reduction Engineering Laboratory
              Cincinnati, Ohio 45268
 RISK REDUCTION ENGINEERING LABORATORY
   OFFICE OF RESEARCH AND DEVELOPMENT
  U.S. ENVIRONMENTAL PROTECTION AGENCY
           CINCINNATI, OHIO 45268

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                                  DISCLAIMER
    Development of the information in this report has been funded in part by the U.S.
Environmental Protection Agency under Cooperative Agreement No. CR812167 by the
American Society of Civil Engineers.  The report has been subjected to Agency peer and
administrative review and approved for publication as an EPA document Mention of trade
names or commercial products does not constitute endorsement or recommendation for use.
                                     ii

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                                    FOREWORD
    Today's rapidly developing and changing technologies and industrial products and
practices frequently carry with them the increased generation of materials that, if improperly
dealt with, can threaten both public health and the environment The U.S. Environmental
Protection Agency (EPA) is charged by Congress with protecting the Nation's land, air, and
water resources.  Under a mandate of national environmental laws, the Agency strives to
formulate and implement actions leading to a compatible balance between human activities
and the ability of natural systems to support and nurture life.  These laws direct EPA to
perform research to define our environmental problems, measure the impacts, and search for
solutions.                                                               ;            .

    The Risk Reduction Engineering Laboratory is responsible for planning, implementing,
and managing research, development, and demonstration programs to provide an authoritative,
defensible engineering basis in support of the policies, programs,  and regulations of EPA with
respect to drinking water, wastewater, pesticides, toxic substances, solid and hazardous
wastes, and Superfund-related activities. This publication is one of the products of that
research and provides a vital communication link between the researcher and the user
community.

    As part of these activities, an EPA cooperative agreement was awarded to the American
Society of Civil Engineers (ASCE) in  1985 to evaluate the existing data base on fine pore
diffused aeration systems in both clean and process waters, conduct field studies at a number
of municipal wastewater treatment facilities employing fine pore aeration, and prepare a
comprehensive design manual on the subject This manual, entitled "Design Manual - Fine
Pore Aeration Systems," was completed in September 1989 and is available through EPA's
Center for Environmental Research Information, Cincinnati, Ohio 45268 (EPA Report  No.
EPA/625-1-89/023). The field studies, carried out as contracts under the ASCE cooperative
agreement were designed to produce reliable information on the performance and operational
requirements of fine pore devices under process conditions. These studies resulted in  16
separate contractor reports and provided critical input  to the design manual.  This report
summarizes the results of one of the 16 field studies.
                  E. Timothy Oppelt Director
                  Risk Reduction Engineering Laboratory
                                         m

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                                      PREFACE
    In 1985, the U.S. Environmental Protection Agency funded Cooperative Research
Agreement CR812167 with the American Society of Civil Engineers to evaluate the existing
data base on fine pore diffused aeration systems in both clean and process waters, conduct
field studies at a number of municipal wastewater treatment facilities employing fine pore
diffused aeration, and prepare a comprehensive design manual on the subject  This manual,
entitled "Design Manual - Fine Pore Aeration Systems," was published in September 1989
(EPA Report No. EPA/725/1-89/023) and is available from the EPA Center for Environmental
Research Information, Cincinnati, OH  45268.

    As part of this project, contracts were awarded under the cooperative research agreement
to conduct 16 field studies to provide technical input to the Design Manual. Each of these
field studies resulted in a contractor report.  In addition to quality assurance/quality control
(QA/QC) data that may be included in  these reports, comprehensive QA/QC information is
contained in the Design Manual.  A listing of these reports is presented below.; All of the
reports are available from the National  Technical Information Service,  5285 Port Royal Road,
Springfield, VA 22161 (Telephone: 703-487-4650).

1.     "Fine Pore Diffuser System Evaluation for the Green Bay Metropolitan Sewerage
       District" (EPA/600/R-94/093) by JJ. Marx

2.     "Oxygen Transfer Efficiency Surveys at the Jones Island Treatment  Plants, 1985-1988"
       (EPA/600/R-94/094) by R. Warriner

3.     "Fine Pore Diffuser Fouling:  The Los Angeles Studies" (EPA/600/R-94/095) by M.K.
       Stenstrom and G. Masutani

4.     "Oxygen Transfer Studies at the Madison  Metropolitan Sewerage District Facilities"
       (EPA/600/R-94/096) by W.C. Boyle, A. Craven, W. Danley, and M. Rieth

5.     "Long Term Performance Characteristics of Fine Pore Ceramic  Diffusers at Monroe,
       Wisconsin" (EPA/600/R-94/097) by D.T. Redmon, L. Ewing, H. Melcer, and G.V.
       Ellefson

6.     "Case History of Fine Pore Diffuser  Retrofit at Ridgewood,  New Jersey"
       (EPA/600/R-94/098) by J.A. Mueller and P.O. Saurer                 i

7.     "Oxygen Transfer Efficiency Surveys at the South Shore Wastewater Treatment Plant,
       1985-1987" (EPA/600/R-94/099) by  R. Warriner

                                       iv                              i

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8.     "Fine Pore Diffuser Case History for Frankenmuth, Michigan" (EPA/600/R-94/100) by
       T.A. AUbaugh and SJ. Kang

9.     "Off-gas Analysis Results and Fine Pore Retrofit Information for Glastonbury,
       Connecticut" (EPA/600/R-94/101) by R.G. Gilbert and R.C. Sullivan

10.    "Off-Gas Analysis Results and Fine Pore Retrofit Case History for Hartford,
       Connecticut" (EPA/600/R-94/105) by R.G. Gilbert and R.C. Sullivan   ,

11.    "The Measurement and Control of Fouling in Fine Pore Diffuser Systems"
       (EPA/600/R-94/102) by E.L. Barnhart and M. Collins

12.    "Fouling of Fine Pore Diffused Aerators: An Interplant Comparison"
       (EPA/600/R-94/103) by C.R. Baillod and K. Hopkins

13.    "Case History Report on Milwaukee Ceramic Plate Aeration Facilities"
       (EPA/600/R-94/106) by L.A. Ernest

14.    "Survey and Evaluation of Porous Polyethylene Media Fine Bubble Tube and Disk
       Aerators" (EPA/600/R-94/104) by D.H. Houck                      ;

15.    "Investigations into Biofouling Phenomena in Fine Pore Aeration Devices"
       (EPA/600/R-94/107) by W. Jansen, J.W. Costerton, and H. Melcer

16.    "Characterization of Clean and Fouled Perforated Membrane Diffusers"
       (EPA/600/R-94/108) by Ewing Engineering Co.

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                                         ABSTRACT
    Microbiologically-based procedures were used to describe biofouling phenomena on fine pore
 aeration devices and to determine whether biofilm characteristics could be related to diffuser process
 performance parameters.  Fine pore diffusers were obtained from five municipal wastewater treatment
 plants in Wisconsin, one in Michigan, and one in California. Biofilm thickness and carbohydrate
 content were measured as were the film's ability to retain air bubbles.  Scanning electron microscopy
 (SEM) procedures were used to examine biofilm structure.  Diffusers were also tested for standard
 oxygen transfer efficiency (SOTE), bubble release vacuum (BRV), and dynamic wet pressure (DWP).
 Subsequently, one of the plants in Wisconsin was revisited where the three most useful of the biofilm
 characterization techniques were deployed to determine any relationship between biofilm thickness and
 content and operational parameters such as SOTE.  The effect of four different cleaning procedures on
 diffuser characteristics was also measured. A parallel laboratory-scale  investigation simulated
 microbially induced fouling of small-scale difrusers and the effect of bleach and acid cleaning.

    Microbial fouling of the difrusers occurred irregularly at all seven sites. Biofilm thickness and its
 distribution over a diffuser appeared to vary randomly. Highly structured biofilms were observed by
 SEM and are thought to interfere with the passage of an air bubble, such that the size of the bubble
 released from a fouled diffuser surface is probably not the same as that from a cleaned iand unfouled
 diffuser surface. Two types of biofoulant were observed. One appeared to reduce SOTE with
 minimal impact on diffuser headless as measured by DWP. The second appeared to increase DWP
 with minimal impact on SOTE.                                                  ;

    In general, measurements of biofilm thickness and carbohydrate content do not appear to be
 directly related to measurements of SOTE, DWP, and BRV. However, the microbiological and
 process characteristics are complementary and support the trends and observations made, particularly
 when supported by foulant properties such as mass per unit area, volatility, and acid soluble content
 Taken collectively, the data contribute to an improved understanding of the plant diffuser operation.

    After cleaning in the laboratory, the diffuser SOTEs were restored  to values ranging from 16.3 to
 20.9% at 1 cfm/diffuser. Inadequate data were collected to  demonstrate statistically significant
 differences, but the data show little difference in SOTE improvement between the cleaning processes
 used. Values of BRV, coefficient of variability, DWP, and  BRV/DWP were restored to clean water
 equivalent values.

    The laboratory investigations showed that the progressive development of a bacterial biofilm  on
the surface of a fine bubble diffuser can increase the size of bubbles released from that fouled surface.
5% bleach was found to be more effective in removing the biofilm than 14% HC1 on both artificially
induced and naturally occurring biofilms, and a combination of bleach  followed by acid  returned  the
stone to its original condition. Bleach cleaning from the air side was effective hi removing the
diffuser biofilm in the laboratory-scale model system.

    This report was submitted in partial fulfillment of Cooperative Agreement No.  CR812167 by  the
American Society of Civil Engineers under subcontract to Microbios Ltd. under the partial sponsorship
of the U.S. Environmental Protection Agency. The work reported herein was conducted over the
period of 1987-1988.
                                             VI

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                                  TABLE OF CONTENTS
Foreword	   ill
Preface	   iv
Abstract	   vi
Figures	  viii
Tables	    x
Acknowledgements	   xi

Introduction	    1
Field Investigations - Initial Characterization	,	    1
     Biofilm Characterization	...:	    1
          Procedures	    1
          Results	    5
          Discussion	    5
     Evaluation of Diffuser Process Characteristics	   19
          Procedures	   19
          Results	   23
          Discussion	   23
     Conclusions - Preliminary Field Studies	   25
Field Study - Verification	   28
          Procedures	   28
          Results	   28
          Discussion	   28
     Conclusions - Verification Study	   39
Laboratory Scale Simulation of Microbial Fouling and Cleaning Procedures	j	   39
          Procedures	   39
          Results	,	   41
          Discussion	   52
          Conclusions	   52
References	,	   54
Appendix A	   55
                                           vu

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                                          FIGURES
Number

Figure 1.  Three diffusers from a four-lunger after 15 months of service at
          Whittier Narrows	,	    2

Figure 2.  SEM of the surface of a clean Sanitaire ceramic disc diffuser.	    6

Figure 3.  SEM of the surface of a clean Norton ceramic dome diffuser.	    7

Figure 4.  SEM of the surface of the biofilm on a Sanitaire ceramic disc
          diffuser with detail of a large, highly structured aperture that
          appears to communicate with an air passage of similar dimensions
          within the adherent biofilm	    8

Figure 5.  SEM of the throat of an air passage within the same biofilm shown
          in Figure 4 showing the connection between the air passage and its
          highly structured surface aperture	    9

Figure 6.  SEM of elliptical aperture in the biofilm of the Norton dome diffuser.
          Bar indicates 0.5 mm	   10

Figure?.  Low power light micrograph of a section of a frozen preparation of
          biofilm from a diffuser after 13 months' service at the Green Bay
          plant. The section was taken at right angles to the fouled diffuser
          surface. Shown are large numbers of small (
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Figure 10. SEM of the surface of a biofilm on a Norton dome diffuser after
          16 months' service. The large air passage is partially occluded by
          a highly-ordered diaton skeleton and a loop of fungal mycelium.
          Bar indicates 0.5 mm.	•	  14

Figure 11. SEM of the surface of a biofilm on a Norton dome diffuser after
          16 months' service. Examination at this shallow angle shows an
          amorphous biofilm separated from the ceramic diffuser surface.	  15

Figure 12. Schematic representation of micrograph in Figure 11	  17

Figure 13. Proposed model of biofilm structure (not to scale)	.	  18

Figure 14. Sampling locations at the Whittier Narrows plant	  24

Figure 15. Diffusers taken from a four-lunger after 15 months' service at the
          Houghton plant showing relatively homogenous accretion of biofilm material	  26

Figure 16. Diffusers taken from an adjacent position in the same four-lunger
          at Houghton (Figure  15) showing the irregular accretion of a
          microbial biofilm.	  27
                                                                          i
Figure 17. Drained aeration tank at the Monroe plant	  29

Figure 18. Sampling locations for diffusers at the Monroe plant	  30

Figure 19. Detail of the fouled diffusers hi drained aeration tank at the Monroe plant	  31

Figure 20. Distribution of biofilms on diffusers taken from the Monroe plant	  32

Figure 21. SEM of the surface of the biofilm on a Sanitaire ceramic disc
          diffuser with detail of a large aperture. Bar indicates 0.5 mm	...i	  33

Figure 22. SEM of the surface of the biofihn on a Sanitaire ceramic disc diffuser
          with detail of a large aperture. Bar indicates 0.5 mm	  34

Figure 23. How schematic of laboratory diffuser system	  40

Figure 24. Bubble size determination. Each division in the grid equals 1 mm	;.	  42

Figure 25. Biofilm development after 3 weeks (30 October)	  45
                                            IX

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Figure 26. Biofikn development after 6 weeks (22 November)	  46

Figure 27. Average bubble size at 0.03 cfm	,	.	  47

Figure 28. Biofilm development after 6 weeks (20 April), showing
          air channels in biofilms	  49

Figure 29. SEM of the surface of a biofilm on the laboratory diffuser showing
          detail of a large aperture. Bar indicates 0.5 mm.	,	  53
                                          TABLES

Table 1.   Biofilm Characteristics for Diffusers - Preliminary Study	',	    4

Table 2.   Comparison of Plants with the Thickest Biofilms	   20

Table 3.   Summary of Diffuser Characteristics	   21

Table 4.   Comparison of Biofilm Properties with Diffuser Characteristics	   35

Table 5.   Summary of Monroe Diffuser Characteristics	;.	   36

Table 6.   Summary of Bubble Data for Bleach Cleaning	j	   43

Table 7.   Summary of Bubble Data for Acid and Bleach Cleaning	.>	   48

Table 8.   Cleaning of Full Size Diffuser	   51

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                                     INTRODUCTION

               Microbial films can establish themselves on surfaces in all aquatic systems that
support bacterial growth.  They can occur in natural aquatic systems whether they be aerobic or
anaerobic,  in industrial situations such as  cooling water  systems, heat exchanger surfaces and
oil recovery injection wells,  and in high organic level systems  such  as  wastewater treatment
plants.   Approximately half of the wastewater treatment plants  in  North  America that use
activated   sludge  systems  have  installed  diffused  aeration  facilities.   Very  early  systems
incorporated fine bubble ceramic plates as air diffusers and, in a few cases, the original systems
are still in use today.  Low cost,  coarse bubble aerators  were introduced later.  These systems
were  not as subject to  clogging  and resulted in few process  interruptions for  maintenance.
However,  because of the increased bubble  size, the oxygen transfer  efficiency (OTE)  of these
systems is much reduced and with the  oil embargo and energy crisis in the 1970s,  fine  pore
aeration  was once again revisited.  New generation fine pore systems  were developed  and have
been  successfully implemented  in  many plants.   Unfortunately, the application of  fine  pore
diffusers has been marred by the incidence of fouling in  the form of microbial films which can
have  a deleterious effect on  OTE.  The  effect of the films  on  OTE  has been irregular and the
frequency of occurrence has been random.  The reason(s)  for the presence of the  microbial films
in this  situation is(are)  not completely  understood and  therefore  it  has  not been possible  to
predict  any potential incidence  of microbial  fouling.  This investigation was  undertaken  to
provide  potential users of fine pore  diffusers with information that  could assist them in an
improved understanding of microbial fouling phenomena.

               This  report is presented in  two parts.  The first addresses  the  field work,  in
which diffusers  from seven municipal wastewater treatment plants were  examined by several
different biofilm characterization techniques.   Subsequently,  one of the seven  plants  was
revisited where the three  most useful of these  techniques  were used  to measure the thickness of
the fouling layers, to relate biofilm thickness to operational parameters  such as OTE and  to
examine  the effect  of four  different  cleaning procedures  on  diffuser characteristics.   In the
second part, laboratory simulation  of microbially induced fouling and the effect of bleach and
acid cleaning are reported.


                 FIELD INVESTIGATIONS -  INITIAL CHARACTERIZATION

BIOFILM CHARACTERIZATION

               Existing procedures such as light microscopy and scanning electron microscopy
(SEM) were used to define biofilm  shape and structure.  Carbohydrate content and the number
of living cells in  the biofilm were estimated to define the biofilm content.  New characterization
methods, that is, air bubble release volume (ABR) and the trapped air pocket size (TAPS), were
developed  to measure the  biofilm's capacity  to  retain or hinder the release of air bubbles.
Biofilm thickness was also estimated.
      ^
Procedures

               Fine pore  diffusers  were  obtained  from  five  plants  in Wisconsin,  one  in
Michigan and  one in California. Except for the Californian plant, the diffusers were drawn  from
four-lungers (Figure  1),  groups of four  diffusers that were temporarily installed in  the main
process aeration basin but separated from other diffusers on an individual manifold with its own
air supply.  The  location of diffuser and  months in service of these diffusers are shown in Table

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Figure 1.  Three diffusers from a four-lunger after 15 months of service at Whittier Narrows.

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1.   For the Whittier Narrows, California plant, difrusers were taken directly from aeration grids
in tanks that were  partially drained for  cleaning.   In all  cases, the diffusers  were examined
immediately by low power light microscopy to determine the size of any trapped air pockets and
a penetrometer  was  used to determine biofilm  thickness.  When the biofilm  was  sufficiently
thick (>3 mm), air was injected under the fouling layer at tankside to determine the air bubble
release volume.    Specific  areas of the  fouled difrusers were scraped and the  scrapings were
frozen for  later  carbohydrate  analysis, or  dispersed  in  phosphate  buffer at  tankside before
dilution and plating (in the nearest enclosed facility) to determine the number of colony-forming
units (CFU) per unit area.   Fragments of a fouled difniser were treated  with chemical fixatives
at  tankside  to  prepare  them for  SEM  and  whole,  unfixed difrusers  were  packaged  and
transported  to Ewing Engineering  for determinations of OTE, bubble release  vacuum  (BRV),
dynamic wet pressure (DWP), and volatile  and acid-soluble foulant fractions.  ;


       Light Microscopy                                                    ;
       Light microscopy is  particularly valuable in studies  of bacterial  biofilms because these
       highly  hydrated structures  can  be  examined  without prior dehydration  required  for
       scanning electron microscopy work.   Consequently, air bubbles and aqueous  channels can
       be  seen without dehydration  distortion.   Biofilms  or opaque surfaces, such as those  of
       these difrusers, can only be seen if illuminated from  above  (epi-illumination) because the
       surfaces themselves  interfere  with  the resolution of  biofilm structures in the innermost
       millimeter of the biofilm.  This technique  is used extensively to examine  the structure  of
       bacterial  biofilms  on opaque surfaces in natural  environments  (Geesey  ej;  al., 1977).
       Details of the procedures are given in Appendix A.

       Scanning Electron Microscopy
       Scanning electron microscopy yields a very useful topographic surface view  of the fouled
       surface to  be examined, but  the image is distorted by extreme dehydration (99%) of the
       exopolysaccharide matrix of the biofilm, while the bacterial cells are shrunk only 10 to  15
       percent. Once the limitations of this direct visualization technique are fully appreciated,
       it can yield very valuable  data on  biofilm morphology (Costerton,  1980).  Details of the
       procedures  are given in Appendix A.

       Biofilm Thickness                                                   ;
       The tendency of a  biofilm to deform upon touching with a ruler and a lack of resolution
       precluded the  use  of the ruler for measuring biofilm thickness.  Instead, thickness  was
       measured with a penetrometer as outlined in Appendix A.             j

       Number of Living Cells
       This  is a standard microbiological  procedure which  is widely  applied  in  biofilm
       characterization work. It is  described in  detail in  Appendix A.  The  test yields  a value
       expressed in colony-forming units per cm2.

       Trapped Air Pocket Procedure
       A technique was developed to measure the size of air bubbles that were  trapped within
       the biofilm.  Details are given in Appendix A (Light Microscopy).

       Air Bubble  Release Volume Procedure
       This procedure was  developed to measure the resistance of a  biofilm to passage of air
       bubbles through its mass.  A 0.5 ml syringe was used to inject air beneath the surface of
       a biofilm on a submerged diffuser.  The more resistant  a  biofilm, the larger the volume
       of air that could be  injected below the biofilm surface. The test gives  a  value of the
       maximum  volume of air  that was held by the biofilm before being released to the water
       above  the  biofilm.   As a. further measure of biofilm  resistance,  distinction  was made
       between immediate and delayed release of bubbles.

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       TABLE 1.   BIOFILM CHARACTERISTICS FOR DIFFUSERS - PRELIMINARY STUDY
PLANT
Monroe, WI

Houghton,
Houghton,
Green Bay,

Date
Sampled
- centre1
- periphery1
MI
MI
WI'

Madison, WI
Madison, HI .
Milwaukee,
WI
- South Shore


Jones Island
Whittier,
Tank 3
- Grid
- Grid
- Grid

- Grid
- Grid
Tank 1

Tank 2
- Grid
- Grid

- Grid

- Grid

- Grid
CA
(Control)
1
2
3a

3b
3c


(Acid)1
1
2a

2b

2c

3
7
7
3
3
3

4
4

5

5


9
9
9

9
9
9


29
29

29

29

29
July
July
Aug
Aug
Aug

Aug
Aug

Aug

Aug


Sept
Sept
Sept

Sept
Sept
Sept


Sept
Sept

Sept

Sept

Sept
87
87
87
87
87

87
87

87

87


87
87
87

87
87
87


87
87

87

87

87
Months
BIOFILM CHARACTERISTICS :
Diffuser in
Service ABR1
12
12
4
15
13

4
12

13

13


15
15
15

IS
15
12


15
15

15

15

15
1
d
O.Oli
O.Oli
O.OSd

O.Oli
0.02i

0.02i.

0.12d


d
d
d

d
d
d


d
d

d

d

d
TAPS*
run*
nm
nm
nm
Av 1

nm
Av 1

nm

Av 1.7
(Max 14x10)

Av 0.5-1.0
Av 1.0
Av 0.5-1.0
(Max 2.0)
Av 0.5-1.0
nm
Av 1.0
(Max 1.5)

Av 1.0
Av 0.5

Av 1.0

0.5x1.0
(few)
Av 0.5
Thickness
(mm)
nm
3-4
Max 1.0
Max 1.5
Av 1.5
(Max 3.7)
Av 1.5
Av 2.0

Av 1.5
(Max 2.6)
4.8-7.6


3
3
1

2.0
nm
1-3


3
Av 1
(Max 3)
Av 1.5
(Max 4.5)
Av 2
(Max 3)
2
CFU1 ,
0.7 x 10'
1.0 x 10'
5.4 x 10'
8.3 x 10'
4.5 x 10' i

3.1 x 10'
1.6 x 10';

1.6 x 10'

3.6 x 10 '-


1.2 x 10?
2.7 x 10*
1.9 x 10'

6.5 x 10'
4.6 x 10*
2.6 x 10'
i

7.5 x 10'
3.5 x 10?

4.5 x 10'

>7.5 x 10?

4.2 x 10'
Carbohydrate
Content (CH)'
320
3,500
340
380
400

480
540

1,100

1,700 •


800
720 • .
1,000

940
190
680


1,900
980

1,120

1,800

680
1 Discs unevenly fouled - heavy at periphery (81 cm )
  and  sparse at centre (298 cm )
* Tank dry for 1 week, 3 weeks, prior to sampling
1 Control tan): cleaned with high pressure hoses and
  acid tanks cleaned with high pressure - acid -
  high pressure
' Air bubble release volume in ml - i - immediate,
  d -  delayed
* Trapped air pocket  size in mm - measured microscopically
' Colony Forming Units per cm  as determined by serial  dilution
  and culture methods
 Carbohydrate in |lg  glucose equivalent/cm*
  Not measurable

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       Carbohydrate Content Assay                                        '•
       The phenol-sulphuric acid test may be used in microbiological work to measure the level
       of carbohydrate present in a biomass sample.  Details are given  in Appendix A,  The test
       yields  a value in  mass  of  glucose  equivalents  per  cm2.   All  adherent  biofilms are
       cemented together by bacterial or fungal exopolysaccharides which  form a large portion of
       the biofilm  mass.  Therefore, the  test will discriminate between  biofilms containing
       significant quantities of inorganic matter,  such as sand,  from  those biofilms which are
       made of cell matter and carbohydrate connecting tissues only.


Results

               The results of SEM analysis are presented as a series of micrographs in Figures
2 through 11.  Data for the other film characteristics are presented in Table 1.


Discussion

               When unfouled Sanitaire (Figure 2)  and Norton Dome (Figure 3) fine pore stone
diffusers were examined by SEM, rough, largely crystalline, surfaces with  irregular pores  were
observed.   The maximum dimensions of  these  irregular pores in the rigid  surface  of the
Sanitaire diffuser was 0.2 x 0.3  mm while that of the Norton Dome diffusers was 0.2 x 0.6 mm
and the  pores in both  diffusers  tended  to be  approximately rectangular.    Biofilms  with
thicknesses less than 1 mm (as determined by direct measurement of the  hydrated biofilm)  were
excluded from this  analysis because their structure  could not be confirmed by light microscopy.
Further, when these thin films were dehydrated for  SEM, a confusing set of cracks and fissures
were  produced.  All the biofilms  with thicknesses greater  than 1 mm  showed  large round
apertures (Figure  4) of highly  structured  air passages  (Figure 5) at their   surfaces.   The
frequency of occurrence of these apertures  was  random  and difficult to assess.  They occurred
most  frequently in the  thicker  biofilms of Milwaukee Jones  Island  and  Green  Bay.   SEM
examination  at varying tilt angles appeared to show that these air passages extended into the
biofilm  matrix.  Air passages were seen to penetrate the biofilms on both  Sanitaire (Figures 4
and 5) and  Norton Dome (Figure 6) diffusers.  Examination of frozen sections from these same
biofilms  by  light  microscopy  showed branching  air passages  throughout the thick biofilms
(Figure 7).  As a result of interference from the diffuser surface and from entrained particles,
the branching of these air passages  could not be effectively photographed or measured by  light
microscopy but trapped air bubbles as large as  5  mm in diameter were  frequently seen within
thick  biofilms.  One very large (10 x 14 mm) air bubble was  seen within the thick biofilm of a
heavily fouled diffuser from the Jones Island plant.

               As  a result  of the  shrinkage  of biofilms during  the  dehydration  stages  of
preparation  for SEM  all  dimensions must be treated  as minimal values.    The maximum
dimensions of the apertures of the air passages seen in biofilms on Sanitaire diffusers (Figures
4 and 5) were 1.5 x 1.5 mm and those seen in the biofilms on Norton Dome diffusers (Figure 6)
were  0.5 x  0.9 mm.  The apertures were round,  they were seen to  communicate  with air
passages within the biofilm, and their side  walls were composed largely of linear bacterial  cells
(Figure 8) embedded in  their  own  amorphous exopolysaccharide  matrices.   These  cells can be
identified as bacteria by their  rod-like shape and their  dimensions (0.5 to 1.0 um in width).
These  sticky biofilms trapped  regularly-structured diatom  skeletons  (Figure 9) and  other
inorganic particles (Figure  10),  and occasionally contained fungal mycelia (arrow in Figure 10),
but they were primarily composed of biofilm bacteria, which are highlighted by  Figure 8, in the
"throats" of the air  passage apertures.                                      >

               When the biofilm-diffuser interface was seen at a tangential: angle (Figure 11),
at the broken edge  of a diffuser fragment, it was observed that the biofilm was not intimately

-------
Figure 2.  SEM of the surface of a clean Sanitaire ceramic disc diffuser.

-------
Figure 3.  SEM of the surface of a clean Norton ceramic dome diffuser.

-------
Figure 4.  SEM of the surface of the biofilm on a Sanitaire ceramic disc diffuser with detail of a large,
           highly structured aperture that appears to communicate with an air passage; of similar
           dimensions within the adherent biofilm.

-------
Figure 5.  SEM of the throat of an air passage within the same biofilm shown in Figure 4 showing the
          connection between the air passage and its highly structured surface aperture.

-------
Figure 6.  SEM of elliptical aperture in the biofilm of the Norton dome diffuser. Bar indicates 0.5 mm.
                                            10

-------
Figure 7.  Low power light micrograph of a section of a frozen preparation of biofilm from a diffuser
          after 13 months' service at the Green Bay plant. The section was taken at right angles to the
          fouled diffuser surface. Shown are large numbers of small (
-------
Figure 8.  High power SEM of the surface of a biofilm on a Norton dome diffuser after 16 months'
          service. Linear bacterial cells are partially buried in the dehydration-condensed residue of
          their exopolysaccharide matrices. Bar indicates 0.005 mm.              ;
                                              12

-------
Figure 9.   SEM of the surface of a Sanitaire ceramic disc diffuser after 14 months' service showing a
           highly ordered diatom skeleton trapped within the biofflm matrix. Bar indicates 0.05 mm.
                                               13

-------
Figure 10. SEM of the surface of a biofilm on a Norton dome diffuser after 16 months' service. The large
          air passage is partially occluded by a highly-ordered diaton skeleton and a loop of fungal
          mycelium. Bar indicates 0.5 mm.
                                              14

-------
Figure 11. SEM of the surface of a biofilm on a Norton dome diffuser after 16 months' service. Exami-
          nation at this shallow angle shows an amorphous biofilm separated from the ceramic diffuser
          surface.
                                             15

-------
connected  to  the  diffuser  surface  at  all  points  and that  large  spaces  existed  at  the
diffuser-biofilm  interface.-   Figure 12  is  a schematic  based upon the  interpretation  of  the
micrograph shown in Figure 11.

               There  was no evidence  of  biofilm  caramelization.    The; intensity  of  acid
treatment  during  cleaning was not as  severe as  that  experienced during bore hole cleanout
when a sticky residue of biofilms was observed (Watkins  and Costerton, 1984).

               Presently used methods  for  light microscopy and SEM did  not  resolve  the
structure of bacterial biofilms with thicknesses of less than 1 mm.  The studies of biofilms with
thicknesses greater than 1 mm show that, at all seven  treatment plants  sampled, the biofilms
were  composed primarily  of  bacterial  cells enmeshed  in a matrix  of  their  own amorphous
exopolysaccharides. This composition has  been observed in  many biofilms in both natural and
industrial  aquatic  systems (Costerton  ej; al, 1987).  These  leathery bacterial biofilms  did  not
appear to be intimately connected at all locations on the diffuser surface but sometimes formed
large  spaces  by local disconnection (Figures 11 and 12).  The unique  feature of these bacterial
biofilms is that they were traversed by large (greater  than 0.5 mm  diameter) structured  air
passages that appeared to originate at the diffuser surface and were seen  to extend and branch
towards  the  biofilm surface  where they  terminated in large (approximately  1.5 mm) round
apertures.  When  a coherent  bacterial biofilm of this structure has developed on a fine bubble
diffuser it might  be  supposed that the  bubble  release surface has  changed  from  the small
irregular pores of the rigid diffuser surface to the larger, more rounded  apertures of the leathery
biofilm surface which might be expected to cause a  change in the  size of bubbles released.

               A  model is proposed in Figure 13 of  the structure of thick biofilms, based upon
these  SEM and light microscopy observations.  The  figure suggests how  the biofilm structure
may develop  to create a deteriorated surface for the release of evenly-sized small bubbles.  It is.
proposed that the  partial attachment of the biofilm allows large  spaces to develop between  the
diffuser surface and the biofilm. Air bubbles could coalesce  in these  spaces.  The existence of
large  diameter branched air passages in the biofilm would  subsequently convey coalesced  air
bubbles to  the biofilm surface, at which point they would leave via larger external apertures.

               Values  for the film characteristics  shown in Table  1 occur  in an  apparent
random manner.  A major difficulty experienced in  carrying out these measurements was  the
inconsistency of film occurrence.  On occasions the  film was thick, mostly  it was very  thin (less
than 3 mm);  sometimes it covered the whole diffuser,  often it did  not; sometimes the film was of
a leathery  consistency and able to  retain air bubbles, sometimes it was soft and fluffy-like and
it was difficult to implement the measurement of ABR and TAPS.

               It  was  possible to  measure TAPS in thick  biofilms (greater than 3  mm) but
resolution was poor in thinner biofilms.  ABR measurement was also  simpler  in biofilms of 3
mm thickness or greater.  In  the  case of the Monroe diffuser,  there  was considerable biofilm
growth  on the  periphery  whereas the centre  portion  was almost devoid  of biofilm.   Two
measurements  were reported for most  parameters  to  illustrate differences in characteristics
across the diffuser.  This irregular deposition of biofilm would suggest a^ random  sampling
technique such as the four-fold box  technique (Southwood, 1978) be implemented.

               The utility of CPU data  is  also  compromised  by the nature of the  biofilm
surface.  The determination of the  number of living cells in a leathery biofilm  is more difficult
than in a soft biofilm because the  organisms are not as easily dispersed in the former  case as
they are in the latter.  Consequently, one CPU may equal one cell in a soft biofilm, or one CPU
may equal an  aggregate of thousands of cells in a  leathery biofilm.   If  leathery biofilms  are
subjected to  extra  sonication  to increase cell dispersion,  this procedure begins to kill biofilm
bacteria and may  actually decrease the CPU values.   In soft  biofilms,  this technique yields
values one order of magnitude  lower than direct microscopic counts and, in leathery biofilms,
                                         16

-------
Figure 12. Schematic representation of micrograph in Figure 11.
                           17

-------
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the CPU values are often  three orders of magnitude lower than the direct counts.  Thus, these
determinations of live bacterial cells may have a comparative value but appear to have little
absolute value in this study.   On the other hand, the leathery consistency of some  biofilms
allowed the accurate determination of thickness through the use of a penetrometer.

               In  view of the limitations  of  some of the methods used  to  measure  biofilm
properties, Table 2 was created by editing the data in Table 1 to retain only those plants  with
biofilms greater than  3 mm.   All  CPU data for these plants  were also omitted.  The  thickest
biofilm (Milwaukee, Jones  Island) corresponded to the 4th highest carbohydrate content reading
and it was seen to contain large  air  pockets (10 x 14 mm)  and to hold 0.12 ml of air  in  a
delayed response to the ABR test  The five  thickest biofilms (shown in Table 2) correspond to  3
of the 4 highest carbohydrate contents [Milwaukee - Jones Island, Monroe (periphery), Whittier
Acid! grid  1 and Whittier Acid grid 2c].  In contrast,  the two lowest biofilm thicknesses [Monroe
(centre), and Whittier Control grid 3c] correspond to the two lowest carbohydrate values (same
sites) (Table 1).

               While carbohydrate  analysis emphasizes  the organic  content of  the  biofilm,
foulant  data from  Table 3  show  that trapped non volatile material  makes up a significant
fraction of the  biofilm.                 .
EVALUATION OF DIFFUSER PROCESS CHARACTERISTICS                '

Procedures

               Duplicate diffusers from each of the seven sites were tested for standard oxygen
transfer  efficiency (SOTE), dynamic wet pressure (DWP) and bubble release volume (BRV).
Both  DWP  and BRV provide a measure of the bubble release  pressure.  DWP is the pressure
differential  across the entire diffuser when operating in a submerged condition.; It measures the
overall pressure  that  is available to form bubbles at a  specific flux over the  entire diffuser
surface.  Since only a small proportion of pores are emitting air at any given time,  DWP is a
measure of the resistance  to passage of air.  In cases where the diffuser is partially fouled, air
may  short  circuit the fouled area, that is, DWP may not indicate  the  true condition of the
media.   BRV, on  the  other hand,  measures the  average pressure  that is  required  to  form
bubbles at a specific air flux over a limited region of the difruser.  In all probability, the  BRV
test will force air through  some pores that  were inactive in a DWP test and gives an indication
of the uniformity achieved during diffuser manufacture  and of the non-uniformity developed
during difruser operation.  It is, therefore,  a  more sensitive test than DWP.   The ratio of BRV
to DWP  is also an effective parameter to characterize the condition of a diffuser. The ratio  is
closely related to the  fraction of the diffuser area that is actually emitting  bubbles.  As this
ratio  decreases,  less  effective  area is  available for  the same  air flow.   
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               The same tests were carried out for the diffusers from the seventh site, Whittier
 Narrows, excepting that diffusers were taken from two tanks.  The diffusers had passed through
 different  cleaning regimes.   The locations from which  diffusers  were sampled are shown in
 Figure  14.  Tank 3 had previously been cleaned by  high pressure hosing only.  The diffusers
 had been in service for 15  months prior  to the time of sampling.  Tank 2 had earlier been
 cleaned by the Milwaukee method; high pressure hosing, acid wash and  high pressure hosing.
 It,  too, had been in  service for 15  months  prior to the  time of sampling.  Diffusers  were
 collected from each of the three  grids in each tank to observe  any change in SOTE along the
 length of each tank and any corresponding change in biofilm properties.       i


 Results

               Data for SOTE, BRV,  DWP, BRV/DWP, EFR  and  foulant characteristics for all
 test sites are reported in Table 3.  The diffusers from Houghton were broken in transit limiting
 analysis to  BRV only.  The diffusers  from Monroe were  not  cleaned although  data for a new
 diffuser are included for comparative purposes.  Data  for plants  with biofilm thicknesses greater
 than 3 mm, Milwaukee, Jones  Island, Monroe and Whittier are  also  recorded in Table  2 to
 provide a direct comparison  with  data on biofilm properties.


 Discussion

               The  diffusers from Monroe, Green Bay, Madison, Milwaukee,; Jones Island  and
 Milwaukee,  South Shore, were all  Sanitaire  ceramic  discs.  Except for the ' Milwaukee, Jones
 Island diffusers, they demonstrated an SOTE of 18 to 19% at  1  cfm/diffuser in the fouled  state
 reducing to a value of 15  to 16%  at 3 cfm/diffuser (Table  3).  The Jones Island diffuser was
 heavily fouled registering an SOTE of 10.5 and 8.3% respectively.  The  Jones Island  diffuser
 also registered a very high BRV,  COV and EFR and a low value of DWP/BRV.  However, DWP
 at  1 cfm  was at a normal  value indicating that the biofilm had  not significantly affected the
 diffuser head loss  despite  significantly  affecting  uniformity  of bubble formation and SOTE.
 Upon cleaning, all the diffusers were restored  to almost new  conditions with SOTE, DWP and
 BRV values close to that in  clean water.

               The  diffusers from Whittier  Narrows were all  Norton  ceramic domes.   They
 demonstrated an  SOTE of 14 to 16% at 1 cfm/diffuser in the fouled state, reducing to a value of
 10  to 13% at 2.7 cfm/diffuser (Table 3). All the diffusers had  a  thick growth ;of biofilm (greater
 than 3 mm).  The tank 3 diffusers, which had historically been cleaned by hosing with water,
 demonstrated a  head loss effect  that was  greater than in tank 2 perhaps  indicating  that the
 acid cleaning procedures followed in tank  2 had delayed a head loss increase as compared to
 tank 3.                                                                   ,

               Upon cleaning the Whittier Narrows  fouled diffusers in the  laboratory, SOTEs
 were not  restored to clean  water values.  They  remained the  same or were slightly  reduced.
 BE,V and DWP  were reduced to  almost clean  water  values.  This difference in behaviour (as
 compared to diffusers  from the other sites)  may be attributed to the fact  that  the  Whittier
 Narrows diffusers were cleaned in the  laboratory by hosing only, unlike  the other diffusers that
 were cleaned in the laboratory by the Milwaukee method.

               When comparing  biofilm properties with diffuser process  characteristics (Table
2),  the lowest SOTE and the highest BRV measured were those at Milwaukee,  Jones Island
 which also reported  the thickest biofilm.   On  the other hand,  the DWP at Jones Island was
relatively  low indicating that the biofilm was  interfering sufficiently with bubble formation to
 depress SOTE  but did  not  affect the head loss across  the diffuser as measured by DWP.
Whittier Narrows also reported thick biofilms but relatively high  SOTEs were measured for both
                                         23

-------


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3b
3a

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<
    Inlet

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    (Tank 3)









3

2c
2b
2a

1

nlet








Acid-Cleaned   Tank
          (Tank 2 )
   Figure 14. Sampling locations at the Whittier Narrows plant.
                       24

-------
tanks 2 and 3.  However, BRVs and DWPs for tank 3 were higher than  observed at the other
plants.  The biofilm at Whittier Narrows  appeared to have only minimal impact on SOTE  but
some  effect on DWP and BRV was observed.  Although the biofilm thickness did not change
along the tank length, its carbohydrate content declined  slightly.  In contrast, SOTE increased
slightly but  measures of head loss and uniformity, BRV and DWP  deteriorated  slightly.  The
different cleaning histories of tanks 2 and  3 appeared to have had no  effect on biofilm thickness
and SOTE.  Higher carbohydrate values were observed for  the  acid-cleaned tank  (2) versus  the
tank hosed with water (3).  Lower values  of BRV and DWP were measured for  diffusers from
the acid-cleaned tank (2) as compared to the tank hosed  with water (3).  There appeared to be
no relationship between ABR and the diffuser process characteristics nor between volatility and
biofilm thickness or carbohydrate value.

               In the case of Monroe,  biofilm properties  were measured on the thickest portion
of biofilm that occurred on the diffuser periphery.  However, measurements  of diffuser process
characteristics  were measured  in  the  centre of  the  diffuser as required and  show  that  the
difiEusers were operating well at high SOTE and low DWP and BRV.   Unfortunately, the biofilm
in the centre of the diffuser was not as thick as that at the periphery.  Rather, it was very thin
and as a result, did not impair SOTE nor affect DWP and  BRV.  A comparison of biofilm
properties and  diffuser process  characteristics  for  the  Monroe diffusers  is  probably  not
appropriate in  this case  in  view of the differences in biofilm thickness at the centre and  the
periphery of the diffusers.

               Observations of biofilms on  operational  fine pore diffusers| have  shown  that
biofilm formation is sometimes relatively uniform  (Figure 15) but sometimes markedly  irregular
(Figure 16),  even at adjacent  sites on  the same four-lunger.   It  is possible  that  localized
differences in  shear forces  and  periodic sloughing of parts  of the  biofilm produce variations in
biofilm thickness and distribution.  This behaviour will require careful  sampling  in  subsequent
investigations.  Comparisbn of diffusers that appeared to be equivalent showed that only limited
trends appeared to exist between biofilm  characteristics and diffuser process effects.   Biofilm
thickness and carbohydrate content are two quantitative estimates of biofilm. accretion  and they
can be easily measured.  There  appeared  to  be two effects that the biofilm 'exerted on diffuser
process characteristics.   The biofilm  can  reduce SOTE  but  have  minimal  effect  on  DWP
(Milwaukee, Jones Island)  and it can have minimal   impact on  SOTE but  increase  DWP
(Whittier Narrows).


CONCLUSIONS - PRELIMINARY FIELD STUDIES
p.»n»»m.i.. in .»• nM- !-•• i Mi n -• n i  PIWi . . - •TiTiTghTB im • i* n .^•^iCTaaTT^r i» -T.n»i.Tf - i  f FM,                        j


       Microbial fouling  of fine pore  diffusers occurred irregularly  at the  seven  wastewater
       treatment plants  examined.  The  thickness of  a  biofilm and  its distribution over  a
       diffuser  appeared to vary randomly.

       Biofilms  appeared to reduce oxygen transfer  efficiency  but have  minimal effect on
       dynamic  wet pressure for  diffusers at some  locations.   However  at other  locations,
       biofilms had  minimal impact  on oxygen  transfer  efficiency but  appeared to increase
       dynamic wet pressure.

       Scanning electron microscopic examination of biofilms with thicknesses greater than  1
       mm showed that the biofilms were  largely  composed of linear bacterial cells enmeshed in
       their own exopolysaccharide matrices.  They did not  appear to be intimately connected to
       all areas of a diffuser surface and occasionally  large spaces occurred between the biofilm
       and diffuser surface allowing the accumulation of  air bubbles in  pockets.   These biofilms
       were  traversed by structured air passages that terminated in large  (approximately 1.5
       mm) round apertures at the biofilm  surface.
                                                                        i

                                                                                        10

                                         25

-------
Figure 15. Diffusers taken from a four-lunger after 15 months' service at the Houghtonplant showing
          relatively homogenous accretion of biofilm material.
                                              26

-------
Figure 16. Diffusers taken from an adjacent position in the same four-lunger at Houghtpn (Figure 15)
          showing the irregular accretion of a microbial biofilm.                  \
                                             27

-------
       The highly structured biofilms are thought to interfere with the passage of an air bubble
       such that the  size of the bubble released  from a fouled diffuser  surface is probably not
       the same as that from a  clean and unfouled diffuser surface.            i

       The  phenol-sulphuric acid  procedure for  measuring carbohydrate  content provided a
       useful measure of biofilm mass.

       The trapped air pocket and air bubble release procedures  provided a measure of the air
       holding capacity of biofilms with thicknesses greater than 3 mm.

       The  enumeration of  living bacteria  in  the biofilms was not  possible by conventional
       methods due to the leathery nature of the biofilms.
                              FIELD STUDY - VERIFICATION

Pitxzedures

               On June 13, 1988, tank  #1 at the Monroe, Wisconsin  wastewater treatment
plant was drained for cleaning (Figure 17).  Tank 1 contained Sanitaire  ceramic disc diffusers.
Before cleaning, four representative diffusers were taken from each of four  grids at locations
shown in Figure  18.  They were identical except that the diffusers at location 1 had a  specific
permeability of 50 units while the remainder had a specific permeability of  26 units.   Two of
these diffusers were retained  for  microbiological  analysis.   The diffusers were  photographed
(Figure 19)  to record visual observations of the evenness of their biofilm  accretions (Figure 20).
The biofilm  thickness of one of each of the diffusers was measured at 7 randomly  chosen points
on the diffuser surface using a  simple penetrometer.  The biofilm of each diffuser was  scraped
from two measured 4 cm"  sections and transported to Calgary, one  for analysis by  the  phenol-
H2SO< method and  the second  sample was prepared  for  SEM by the  methods  outlined  in
Appendix A.  Four other diffusers were transported to Milwaukee where  DWP, BRV, EFR, and
SOTE were  determined "as received" and after cleaning by 4 methods:
                     (i)      Hosing at 65 psi.
                     (ii)     Hosing, bleach treatment and hosing.
                     (iii)    Hosing, acid treatment and hosing.
                     (iv)    Hosing, acid treatment, brushing, and hosing.


Results
                                            x

               SEM photographs are presented in Figures 21 and  22.   Biofilm thickness and
carbohydrate content values are given in Table 4. Data for diffuser process characteristics  are
recorded in Table 5.


Discussion                 .                                               :
                                                                           i
               The  biofilm thickness  and the biofilm carbohydrate content were more variable
on the diffusers at location 1 (Table 4) and visual observations indicated  heterogeneity between
individual diffusers  at  this location.    Bacterial biofilms were consistently thick  at location 2
(Figure 15)  and these values decreased slightly and  stabilized as the  process fluid moved past
locations 3 and 4, towards  the outlet.  Apart from the heterogeneity seen in location 1, near  the
inlet, the biofilm thickness and biofilm carbohydrate  values were similar on pairs  of diffusers
from each location.  On  all diffusers sampled, there was good agreement between the measured
biofilm thickness  and the carbohydrate content per cm" of surface  area (Table 4).
                                           28

-------
Figure 17. Drained aeration tank at the Monroe plant.
                      29

-------
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      Figure 18. Sampling locations for diffusers at the Monroe plant.
                                30

-------
Figure 19.  Detail of the fouled diffusers in drained aeration tank at the Monroe plant
                                        31

-------
   PFV-.  :"y,--~'' -•?»•' "• TB*V '
Figure 20. Distribution of biofilms on diffusers taken from the Monroe plant
                                  32

-------
Figure 21. SEM of the surface of the biofilm on a Sanitaire ceramic disc diffuser with detail of a large
          aperture. Bar indicates 0.5 mm.
                                               33

-------
Figure 22.  SEM of the surface of the biofilm on a Sanitaire ceramic disc diffuser with detail of a large
           aperture. Bar indicates 0.5 mm.
                                               34

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                Visual  observation  of all  of  the  diffusers  in  a  particular  grid showed
heterogeneity at location 1  and increasing homogeneity at locations 2  to 4  (Figure  19).  The
bicifilms  at all locations were soft  but  they were sufficiently coherent that 4  cm2 sections could
be cut and lifted out without breakage.  Few trapped air pockets could  be seen and none were
seen  that were larger  than 1.5 mm in diameter.  Visual  examination  of the biofilm surfaces
showed a large number of  crater-like  holes ranging  in  size from  1  to  10  mm.   SEM showed
these crater-like holes  very  clearly (Figures 21  and 22)  and measurements indicated that they
were  as  large  as  1.5 mm.   Because of the smaller areas that can be observed by  SEM, other
larger holes that were observed visually were not seen by SEM.

                The microbiological indices of biofilm formation, biofilm thickness  and biofilm
carbohydrate per cm2, appear to be well chosen  because  results  are consistent between methods
and because  they confirm visual observations. The indices  were successfully applied for biofilms
greater  than  2.0  mm.   These  data indicate  that  biofilms formed   unevenly  on  the  high
permeability  diffusers at location 1 near the inlet  of Tank  1 of the Monroe plant, but that they
formed evenly  on the less permeable diffusers at location 2 and decreased in  thickness towards
the outlet (locations 3  and  4).   The observation of biofilm heterogeneity at location 1 may  be
linked to the higher organic loading and non-uniform mixing, effects typically  found  at the inlet
end of plug flow aeration tanks rather than to differences in diffuser permeability.

                The  morphological data  confirmed  previous  observations  that the size  of
apparent bubble releasing structures on  the surface  of biofilms on the  fouled diffusers ranged
from  0.5 to 10 mm, in wet  and SEM  samples.   Further evidence of air passages within these
fouling  layers  (e.g. trapped air  bubbles)  indicated  that  the biofilms  on these air-releasing
surfaces  are  traversed by a  network of air  passages and  apertures that increase the bubble size
produced by fouled diffusers.                                              :

                The diffusers from Monroe were  Sanitaire  ceramic diffusers  and had been  in
sei-vice for two years.   Interim measurements of SOTE, DWP and BRV had been made of these
diffusers during this  period  and are reported by Redmon (1989).  After two years, the diffusers
demonstrated an SOTE of 15 to 19% at 1  cfm/diffuser depending upon  location.  There did not
appear to be a change in SOTE along the  length of  tank  1.  Rather, the differences in SOTE
appeared to be random nor did they appear to be related to  differences in diffuser permeability.

                Moderate organic fouling was observed for all diffusers as shown by  the mass of
foulant that ranged from  10.8 to 21.9 mg/cm2 with percent volatilities ranging  from 54.6 to 68.7.
Although the variability appeared to occur at random, when expressed as mean values as shown
in Table  4, the foulant  mass declined along the tank length  similar to biofilm thickness and film
carbohydrate values.   However,  mean volatility  increased along the  tank  length indicating
pei-haps  that measurement  of volatility includes  components in  the  biofilm other  than  the
organic matter. The biofilm did affect  the  uniformity of bubble formation with moderate values
of BRV  and  DWP/BRV being recorded (9.6  to  18.1 in.  and 0.46 to  0.60 respectively).   When
expressed as mean values as  shown in Table 4,  there was a steady trend of increasing BRV
values along  the  tank length  despite the declining biofilm  thickness  and film carbohydrate
values.   The  average values  of percent acid soluble content  also shows a decline along the tank
length.  When  coupled  with  a corresponding increase  in percent volatility along the  tank length
the trends perhaps indicate that the nature of the biofilm is changing along the tank length.  It
would suggest  that the biofilm contains more inorganics at the head of the tank, an area  which
is heavily influenced by the  properties of the influent.  The  inorganics may contribute to a more
open  structure thereby minimizing the increase in BRV at the  head of the  tank.   Along the
tank,  the biofilm changes in nature to  that of reduced biomass content (as expressed by biofilm
thickness and foulant mass), but more  importantly, reduced inorganic content which may affect
the openness of the structure and deteriorate the uniformity of bubble release as expressed  by
the increasing  value of BRV.  Moderate values  of COV  and EFR were  also: observed, although
these  were reduced to clean  water  equivalent values at the  end of the second pass.
                                          38

-------
               On  the other hand, values of DWP at 0.75 cfm were relatively low with values
close to clean water equivalents, indicating that the biofilm had not significantly  affected the
diffuser head loss despite  affecting uniformity of bubble formation as  measured by  BRV and
BRV/DWP.

               Upon cleaning  in  the  laboratory,  the diffuser SOTEs  were restored  to values
ranging from 16.3 to 20.9% at Icfm/diffuser, an increase of approximately 10%.   Inadequate data
were  collected  to demonstrate  statistically  significant  differences but the  data  show  little
difference in SOTE improvement between  the  cleaning processes used.   Values of BRV, COV,
DWP  and BRV/DWP were restored to clean water equivalent values.  It appears that after two
years' service which included a  cleaning in situ by the Milwaukee method  after one year, the
biofilm had  minimal impact on  SOTE but a moderate impact on  diffuser head  loss as  expressed
by DWP.                                                                 ;

CONCLUSIONS - VERIFICATION STUDY                                 '
                                                                    . •    i
      The morphological characteristics of biofilms  formed on the diffusers from  the Monroe
      activated sludge plant agree well  with those seen in other plants examined  in the
      preliminary study.   They  lend  support to a physical  mechanism  by  which bacterial
      biofilms  can alter the size of bubbles released by diffusers  as described in Figure 13.

      The  utility  of  measuring  biofilm  thickness  with a penetrometer  and  the  chemical
      measurement of the bacterial component of biofilms by the phenol -H2SO4 method as used
      in the preliminary stuHy has been confirmed in this phase of work.

      Measurements of biofilm thickness and  carbohydrate content do not appear to be directly
      related to measurements of SOTE,  DWP  and BRV.  However,  the  microbiological  and
      process characteristics are complementary and support the trends and observations made,
      particularly when supported by  foulant properties such as mass  per unit area, volatility
      and  acid  soluble content.   Taken  collectively,  the  data contribute  to  an  improved
      understanding of the plant diffuser operation.                         j

      For the  Monroe  plant,  biofilms were  of moderate  thickness  and were  found  in  a
      homogeneous  manner along the tank length except for some variability at the  head end
      of the tank.  The biofilms appeared to have only a slight impact on  SOTE whereas they
      had a progressively  deteriorating influence on BRV along the tank length. Diffuser head
      loss (DWP) was  not affected by the biofilm.                    .  ..     ;


                          LABORATORY SCALE SIMULATION OF
                  MICROBIAL FOULING AND CLEANING PROCEDURES


               A  laboratory  scale  system in  which  small ceramic   discs were  exposed to
wastewater  and progressively fouled was  developed to determine whether  the formation of a
bacterial biofilm on an air  releasing surface had the capacity to increase the size  of air bubbles
released from that microbially fouled  surface.

Procedures                                                             ;

               Five small  (1.25 cm diameter x 1.3 cm) fine bubble diffusers were mounted on a
manifold and placed in a tank  72 cm x 30 cm x 40 cm (Figure 23).  The  diffusers were made
from a porous stone medium used in the manufacture of Sanitaire ceramic disc diffusers.  The
tank was filled with mixed liquor obtained from the Bonneybrook Wastewater Treatment plant
in Calgary,  Alberta.  Sludge was settled in the clarifier section and  recycled to the aeration
                                         39

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tank.   500 g  of powdered skim milk was added  daily  in  50 L of feed giving an hydraulic
retention time of 0.58 days.   Utility air at a pressure of 10 psi was  regulated to provide a flow
of 0.03  cfm at  each diffuser.   This  value was chosen  because it provided  the most  even
distribution of bubbles across  the diffuser surface.                           :

                Three experiments were completed and  twelve  diffusers  examined.  In the first,
the air flow rate was not controlled and progressively decreased as a result of the back pressure
induced by the  biofilm.  At  each sampling interval, the air  flow rate and bubble size were
measured  at  this decreased  air flow and again after  the air flow rate was  returned to  the
original setpoint of 0.03 cfm.  In the  second and third experiments, new diffusers were installed
and  the air flow rate was calibrated  weekly to maintain a constant air flow of 0.03 cfm for the
duration of each experiment.

                Biofilm development was monitored visually.  Periodically, the diffiiser manifold
was  transferred  to a clear water tank and air bubble  diameters were measured using a clear
plastic  cylinder (30 cm x 12 cm  x 3 cm).  The cylinder was inverted, submerged and held over
each diffuser allowing bubbles to enter the open end.  The bubbles were photographed  and the
bubble  size was  measured against a  grid  on the back wall of the cylinder (Figure  24).  In  the
first experiment,  bubble  size  was  measured initially with no  biofilm, at intervals during  the
development of the biofilm, and  again when the biofilm completely covered the  diffuser surface
(after 6  to 8 weeks) at which  time the biofilms were then characterized  by the  phenol-sulphuric
acid test for  carbohydrates. The diffusers were then cleaned by soaking in 5%  bleach for 24 hr
and bubble size was measured again for comparison with initial bubble sizes.  '

                In the second experiment, the effects on bubble size after cleaning with bleach
and  acid were examined.  Following the  growth of a mature  biofilm, one diffuser  was left to
soak in  14% HC1 for 24 hr while  a  second was soaked in 5% bleach for 24 hr.  A third was
placed in a tank  of clean water, 100 ml of 5% bleach was poured into  the air  line which was
reconnected to  the diffuser and the bleach was allowed to  remain  in contact with the air side of
the diffuser for 24 hr.  The carbohydrate content of all three  cleaned diffusers was measured as
well  as that of a control diffuser which had not been cleaned.  A fifth diffuser was retained for
measurement of colony  forming  units  and  SEM analysis.   After cleaning,  bubble sizes were
measured  as  in the first experiment,  along with  carbohydrate  content, for comparison with
initial values for clean diffusers.
                                                                          !
               A mature biofilm was grown in the third experiment  and then exposed to either
14%  HC1 or 5% bleach  that was  delivered in liquid  form to the air side  of the diffusers via  the
air line. Bubble size and carbohydrate content were not determined in this third experiment.

               To compare the  effects of 14% HC1  and 5%  bleach cleaning on the artificially
generated biofilm with  those  on  a naturally occurring biofilm, a fouled  Sanitaire ceramic disc
from the Green Bay, WI. sewage treatment plant was obtained.  The diffuser was broken into
approximately  4  cm2 pieces.   Different pieces were placed in 5% bleach,  in 14%  acid or both in
sequence to  see how each would affect  the biofilm.   The biofilm that  remained was then
characterized using the carbohydrate  assay and by measuring the biofilm solids content from a
4 cm2 area. Atomized bleach  or acid could not be placed in the air line to the'. diffuser to attack
the large diffuser from the air side, as was done on the small diffusers.


Results

Bleiach Cleaning
               The average bubble size produced by the  unfouled fine bubble diffusers, at  the
beginning  of the  experiment  (9  October),  ranged between 2.54  and 3.40 mm with standard
deviations ranging from 0.68 to 0.96 mm (Table 6).  Direct observations through the submersible
                                          41

-------
Figure 24. Bubble size determination. Each division in the grid equals 1 mm.
                                 42

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TABLE 6. SUMMARY
Date
9 Oct 87



30 Oct 87







22 Nov 87

•











6 Dec 87














After bleach
cleaning

Carbohydrate

av. dia.1
st. dev.
no. of dets.
cfm
av.dla.
st. dev.
no. of dets.
cfm
av. dia.
st. dev.
no. of dets.
cfm
av. dia.
st. dev.
no. of dets.
cfm
av. dia.
st. dev.
no. of dets.
av. dia.
st. dev.
no. of dets.
cfm
av. dia.
st. dev.
no. of dets.
av. dia.
st. dev.
no. of dets.
cfm
av. dia.
st. dev.
no. of dets.

av. dia.
st. dev.
no. of dets.
cfm
av. dia.
st. dev.
no. of dets.
av. dia.
st. dev.
no. of dets.
(Jig/era1)
OF BUBBLE DATA FOR BLEACH CLE
8
2.54
0.68
82
0.03
2.89
0.66
46
0.01
2.81
0.83
37
0.03
2.6 S
0.56
25
0.01
6.5 L1
1.29
4
2.48 S
0.81
32
0.03
6.1 L
1.47
5
2.87 S
0.79
23
0.002
5.33 L
0.58
3

2.64 S
0.92
41
0.03
5.0 L
1.17
6
2.64
0.50
60
NA
C
3.03
0.95
43
0.03
3.47
0.81
42
0.005
3.25
0.77
50
0.03
4.8
0.42
10
0.002



4.12
0.74
47
0.03



4.54
0.50
12
0.002




4.38
1.53
36
0.03



2.95
0.68
69
31900
0
3.05
0.86
49
0.03
3.21
0.69
58
0.02
3.27
1.01
41
0.03
2.58
0.45
19
0.01
5.0 L
0.89
6
2.85
0.35

0.03
4.65
1.12
6
4.31
,0.72
13
0.003




4.05
1.26
48
0.03



2.91
0.60
87
38200
E
3.40
0.83
46
0.03
3.90
0.49
35
0.01
3.77
0.71
46
0.03
S 5.18
0.96
11
0.005



S 4.76
0.69
32
0.03
L


NA'






(no biofilm)
3.33
1.08
55
0.03






NA
F
3.19
0.96
39
0.03
3.41
0.54
56
0.02
3.35
0.94
42
0.03
4.69
0.94
21
0.01



4.07
0.71
55
0.03



5.75 ,
0.65
8
0.003




4.21
1.63
33
0.03



3.51
0.71
68
21200
Average diameter in ran
S refers to small diameter bubble series
L refers to large diameter bubble series
Not available
                                      43

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plasitic tube showed no obvious heterogeneity of bubble sizes from different areas of these small
diffusers.

               After 3 weeks of fouling (30  October), fairly uniform biofilms had developed on
all five diffusers (Figure 25) and the  average bubble  size had increased in each case, at the
reduced  air flow  rate produced by progressive bacterial fouling and at an air flow rate setpoint
of 0.03 cfm.  Large differences in. air flow rate (see diffuser C on 30 October, Table 6) did hot
radically affect measured bubble size.

               After 6  weeks (22  November),  all five  of the  diffusers had developed thick
biofilms  but  those  of diffusers B and D were very irregular in thickness (Figure 26).  Bubble
size had increased  in the  more  evenly fouled diffusers (C, E  and F) at both the decreased  air
flow rate produced by fouling  and the  air flow rate setpoint.  Close examination of diffusers B
and D through the submersible plastic cylinder  showed  that  some areas were fouled by thick
biofilms  while  other areas of the diffuser surface were almost clean.  The heavily fouled areas
produced smaller numbers of very large bubbles while the cleaner areas produced large numbers
of much smaller  bubbles.  For this reason,  averages of the large and  small bubble  sizes  were
not calculated but were reported separately (Table 6).                         '

               After 8 weeks (6 December), thick biofilms had  developed on  all five diffusers
but the biofilm on diffuser E was accidentally lost during transfer to the clear water tank.  The
thick biofilms  on diffusers  C,  D and F had produced  increases in  bubble size  at both  the
reduced  air flow  rate  and the  air flow rate  setpoint.  The biofilm  on diffuser B  was seen to
release large bubbles from its  fouled area and small bubbles from its almost clean areas, when
observed directly using  the  submersible plastic  cylinder.  Therefore,  bubble sizes  were again
reported in large  and small size categories (Table 6). At  the end of the experiment, the mature
biofilms from diffusers B, C,  D and F only were removed, assessed for carbohydrate content and
bleaich cleaned.

               In  every instance in  which  bubble  size was measured at the reference air flow
rate of  0.03 cfm,  there  was  a progressive increase  in bubble  size  with  increasing  biofilm
accretion, and  a  return to near initial values when these fine  bubble diffusers  were cleaned
(Figure 27).   The  data from  diffuser  E indicate  that the spontaneous removal  of  a  fouling
biofilm returns bubble size to very  near the initial unfouled  value.  The direct observation of
unevenly fouled diffusers (B  and D)  indicate that heavily  fouled areas of these; diffusers produce
larger  bubbles  than almost clean.areas of the same diffusers observed at the same time and at
the same air flow rate values.
                                                                  t
Bleach and Acid Cleaning                                                   \

               The initial,  diameter  of the  bubbles from the unfouled diffusers  (7 March),
ranged between 2.82 and 3.09 mm  with standard deviations ranging from 0.49  to  0.69 mm
(Table 7).  After  3  weeks (29 March), a biofilm had started to form  on all five diffusers but no
air channels were visible except for one that had  formed on diffuser 3.

               After 6 weeks (20 April), all five diffusers had thick biofilms (approx. 13  mm)
and all had visible  air channels (Figure 28).   The average bubble size with biofilms present had
increased and ranged between  3.77 and 4.62 mm with standard deviations ranging from 1.05 to
2.15 mm.  The smallest average bubble size from diffuser #2 (3.77  mm) was  also observed as
having two  small air bubble  streams  coming from one area of the  diffuser, with no  biofilm
present which could account  for a relatively smaller increase in average bubble size.

               On 25 April, the diffusers were  cleaned.   Diffuser #5 was left to soak in 14%
HC1 for 24 hours, diffuser  #4 was soaked in 5% bleach for  24  hours  and the  air side of diffuser
#3 was soaked  with 5% bleach for 24 hours.
                                          44

-------
Figure 25. Biofilm development after 3 weeks (30 October).
                           45

-------
Figure 26. Biofilm development after 6 weeks (22 November).
                           46

-------
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                              47

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    TABLE 7.  SUMMARY OF BUBBLE DATA FOR ACID AND BLEACH CLEANING
Date
07 March


29 March


20 April


25 April


Cl waning


88 av.
St.
no.
88 av.
St.
no .
88 av.
St.
no.
88 av.
St.
no.

dia.1
dev.
of dets.
dia.1 '
dev.
of dets.
dia.1
dev.
of dets.
dia.1
dev.
of dets.
1
2.92
0.52
51
3.06
0.73
37
4.62
1.27
37
3.07
0.78
35
Procedure



2
2.82
0.69
42
2.97
0.60
29
3.77
1.05
35
2.94
0.64
31
control

3
3.09
0.60
46
3.30
1.18
42
4.59
1.21
24
3.14
0.87
32
100 ml 5%
bleach in line
4
2.90
0.57
2L
'3.01
0.69
45;
4.47
1.56
38;
2.97
0.5Q
23
5% bleach
soak
5
2.98
0.49
30
3.12
0-74 . ,
31 :
4 .27
2.15
22 ••
3.59
1.05
40 I
14% KCi
soak
Carbohydrate (pg/cm*)
                                 37,200
                      1,920
680 i
                                                                   15,200
cfu'a/diffuoar
3.70x10*
  Average diameter in mm
                                 48

-------
Figure 28. Biofilm development after 6 weeks (20 April), showing air channels in biofilms.
                                      49

-------
               The upper two-thirds of the biofilm on diffuser #5 had been detached and was
found floating in the tank.  One small  section of the cleaned diffuser had no 'biofilm but in the
remaining areas  the  air passages seemed  to be  partly occluded by a condensed biofilm residue
which  may have accounted for  the  failure of the bubble  size to return to  initial clean values
(Table 7).  The carbohydrate content of  diffuser #5, after cleaning, was 15200 ng/cm2.

               Diffuser #4 was soaked  in  5%  bleach  for 24  hours.    After 30  minutes,
approximately one  quarter  of the biofilm  had been removed and some large pieces  of biofilm
were found floating at  the  top of the tank.   After 4 hours, two-thirds of the biofilm had  been
removed.   After 24 hours, approximately 95%  of the biofilm was  gone, the  average bubble size
had decreased to values almost the same as  initially measured, and the carbohydrate  content of
the cleaned surface was 680 ug/cm2 (Table  7).

               Diffuser #3  received 100 mis of 5% bleach from the air side.. ; Almost all of the
bleach went through immediately with a  small  amount remaining in-line.  After 4 hours, the
biofilm thickness had declined and within  24 hours approximately 75% of the biofilm had  been
removed and the average bubble size was  approximately the same as measured with the clean
diffuser.   The remaining 25% of the biofilm  was still attached at the outer  edge  of the diffuser
that, had not been reached by the bleach.  This made the carbohydrate content reading of  1920
ug/cm2 higher than  it should  have  been in  terms of biofilm fouling (Table 7),  The cfu  data
showed that there were many (3.70 x 109) living bacterial cells per cm2 of the fouled surface and
the SEM showed a  thick biofilm with visible  air passages (Figure 29).

Air Side Acid and Bleach Cleaning                                          ;

               In the third experiment only two diffusers were used, because the other three
had sloughed or lost  their biofilm, one  when removing the apparatus from the tank.  With the
two remaining biofilms, diffuser #3 was cleaned with -100 ml of 5% bleach and diffuser  #4  with
100 ml of 14% HC1.  Approximately 90 ml of both cleaning agents passed through the  diffuser
immediately with the remaining 10 ml of bleach or acid being carried through the diffusers over
a period of 2 to 3 hours.  After 4 hours, the biofilm on  diffuser #3 was visibly Deduced and  after
7 hours, it had almost completely disappeared, with a carbohydrate reading of only 250 ug/cm2.
After 24   hours, diffuser  #4  was  still  75%  covered with  biofilm  and  yielded  a  surface
carbohydrate value of 8500 ug/cm2.                                          ,

Cleaning of Full Size Diffuser

               Cleaning with bleach and  acid was carried out on pieces  of a  naturally fouled
ceramic disc. The control piece  which had not been cleaned, registered a carbohydrate value of
3200 ug/cm2 (Table  8), and had  a biofilm dry weight of 64 mg/cm2.  One section of the  diffuser
was placed in 5% bleach for 4 hours and another for 24 hours. After soaking in 5% bleach for
4 hours, a piece of the diffuser registered a carbohydrate value of 680 ug/cm2 and a biofilm dry
weight of 20 mg/cm2, but after 24 hours the carbohydrate and biofilm dry  weight values  were
both zero.  Some  non-measurable but visible residue remained on the  bleach-cleaned diffuser but
after being placed  into 14% HC1 for one hour it  was  visibly clean.  Soaking pieces  of the
diffuser in 14% HC1 for 24 hours resulted  in a carbohydrate value of 1920 ug/cm2 and a biofilm
dry weight  of 43 mg/cm2.  The  remaining biofilm although very soft was still attached to the
diffuser.  The diffuser was then placed in 5% bleach for one hour and the remaining biofilm was
removed with carbohydrate and biofilm dry weight values reduced to zero.
                                          50

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                TABLE 8.  CLEANING OF FULL SIZE DIFFUSER
                                            Cleaning Procedure
             Control      5% Bleach    5%  Bleach     Cone. HC1
                           4 hr        24  hr        24 hr
                                    Cone.  HC1-24 hr
                                    + 5% bieach-1 hr
Carbohydrate   3200
 tUg/crn2)
680
             0.0
                         1920
                                       ;0.0
 Biofilm
Dry Weight
 (mg/cm2)
 20
             .0.0
                          43
                                       :0.0
                                      51

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Discussion
                It  is  recognized  that  this  laboratory  scale  test system  cannot  duplicate
conditions in a full scale activated sludge plant equipped with fine bubble diffusers.  The depth
of the laboratory tank is too shallow to simulate large  tank conditions and the nutrient factors
affecting  biofilm  formation  are  probably  different from those obtained  in  most  regions of
activated sludge tanks.

                The  objective of these laboratory experiments  was to determine whether the
development of a fouling biofilm on an air-releasing surface could change  bubble size. This  was
considered to be very important in any analysis of the effects of biofilm formation  on  bubble  size
in activated sludge plants and it can be addressed only  in a simple system in which the bubbles
produced by individual diffusers can be measured separately and related to the degree of fouling
of the diffuser in question.  In  these experiments, the diffusers could be returned  to their initial
clean states  by a variety of cleaning methods so that each  diffuser was,  in  a  sense,  its own
control.

               The  gradual development  of bacterial biofilms  on the small  diffusers,  in  this
scaled-down tank model, significantly changed the size  of the  air bubbles released  from their
surfaces.   The  size  of the  released  bubbles increased progressively  as  the  bacterial  biofilm
developed, and  returned to initial values  when  the  biofilms were removed from the  diffuser
surfaces.   In unevenly fouled diffusers, large bubbles were  released from surface  areas covered
by  thick biofilms  while much smaller bubbles were released from clean  surface  areas.  These
data show that the development of bacterial biofilms on an air-releasing surface, in a laboratory
scale  model  system,  consistently  causes significant increases in  the  size  of the air  bubbles
released from these surfaces.  Thus, in a reproducible laboratory-scale model, the  principle that
bacterial  biofilm development  can increase the  size of  bubbles  released  from   a fine  bubble
diffuser has been established.                                               ;

               The  biofilms  developed  in  this  laboratory-scale  model system (Figure  29)
resemble those found  on fine bubble diffusers (Figure 21) in activated sludge tanks, in that they
are thick, largely bacterial accretions, of  cells  and exopolysaccharides that are traversed by
branching air channels.  These laboratory data suggest  that the decrease in SOTE that is  seen
in actual operating activated sludge plants  may be related to increases in bubble  size resulting
from the development of the thick bacterial biofilms, honeycombed  with air passages, that have
been observed on operating fine bubble diffusers in this investigation at several locations.

               These  data  on diffuser  cleaning  support the developing understanding of the
nature  of  the biofilms that  can  be established  on air-releasing surfaces.   Air pressures  that
operate on these biofilms are directed  towards the surrounding fluid, tending to remove accreted
bacteria from the fouled  surface.   Consequently, the biofilm  can be retained !only if  it develops
carbohydrate "tethers" that hold it on to the fouled surface.  Bleach and acid were both effective
in cleaning fouled diffusers,  if they are applied to the process water side in a soaking or hosing
mode,  but bleach  was especially  effective if introduced from the air  side and  allowed to
hydrolyze the biofilm-retaining  carbohydrate tethers. However, a practical method of delivering
bleach to the air side of fine bubble diffusers in operating activated sludge plants is needed to
implement in process cleaning of these systems.                             !
Conclusions

                1. The progressive  development  of a bacterial biofilm on the surface of a fine
bubble diffuser can increase the size of bubbles released from that fouled surface.
                                           52

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Figure 29. SEM of the surface of a biofilm on the laboratory diffuser showing detail of a large aper-
          ture. Bar indicates 0.5 mm.
                                               53

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                2. 5% bleach is  more effective in removing the biofilm than  14% HC1 on both
artificially induced and. naturally  occurring biofilms and  a  combination  of bleach followed by
acid will return the stone to its original condition.

                3. Bleach cleaning from the air side is effective in removing the diffuser biofilm
in a laboratory-scale model  system.
                                       REFERENCES

Cossterton, J.W. (1980).  Techniques  for the study of adsorption of microorganisms to surfaces.
Absorption of Microorganisms to Surfaces.  K.C. Marshall and G. Bitten, Eds., Wiley Interscience
50-70.

Coisterton, J.W. ei al  (1987).  Bacterial films in nature and disease.  Ann. Rev. Microbiol. 41
435-464.                                                                     	fcjuu^ -i,

Gessey,  G.G.,  W.T.  Richardson,  H.G.   Yoemans,  R.T.  Irvin and  J.W.  Costerton  (1977).
Microscopic examination of natural sessile bacterial populations  from an alpine stream.  Can. J.
Microbiol.. 23,  1733-1736.                                                  !

Redmon,  D.T.  and  L.  Ewing (1989).  Report  on  the effect of pore size  on oxygen  transfer
capabilities, fouling  tendencies, and  cleaning amenability  of ceramic  diffusers at Monroe, WI.
Prepared  for American Society of Civil Engineers.  Ewing Engineering, Milwaukee, WI. •

Southwood, T.R.E.  (1978).   Ecological Methods (Second Ed.),  Chapman-Hall  Publishers, New
York.

Watkins,  L. and J.W.  Costerton  (1984).  Growth and biocide  resistance of bacterial biofilms in
industrial systems.   Chemical Times and Trends, October, 1984,  35-40.
                                          54

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                                       APPENDIX A
                    DETAILS OF CHARACTERIZATION PROCEDURES

1.      Light Microscopy
       To observe  aqueous channels, the fouled diffuser was  broken with a cold chisel and  a
       fragment was placed,  biofilm uppermost, on  the stage of a Zeiss dissection microscope
       and examined between  7x - 30x magnification with ambient light  as  epi-illumination.
       The size  of trapped air pockets was observed by cutting 3 to 8 segments  through  the
       biofilm, at right angles to the fouled surface, and mounting these segments on their sides
       between a glass slide and a  glass coverslip.  A variable number of; air pockets seen
       within these biofilm segments and air refractive pockets within them were measured by
       reference to an  optical  grating.  If air  pockets were  numerous,  a  smaller number of
       segments were examined.                                           !

2.      Scanning Electron Microscopy                                       ;
       The diffusers  were washed with distilled  water to remove components of the bulk fluids
       and were then  fractured with  a cold chisel  to obtain  fragments with  surface areas of
       approximately 2 cm2.  The biofilms were then  preserved by fixation for 2 hours in  5%
       glutaraldehyde (a tanning agent used, on animal skins) in 100 nM cacodylate buffer at
       20°C.   Following two washed in cacodylate  buffer  to  remove the fixative  but  not  the
       biofilm, the specimens were shipped to Calgary where they were  prepared  for SEM by
       dehydration  in  ethanol  and coating  with platinum in  a  Balzers  300  freeze  etching
       apparatus.   These biofilm specimens were then examined in a  JEOL 450S scanning
       electron microscope with a  45 degree tilt stage  so that  they could be rotated for  viewing
       from a series of favorable angles.

3.      Biofilm Thickness                                                  ,
       The pointed probe of a universal penetrometer was placed on the top of the biofilm  and
       the gauge was zeroed. The rod was then released and pushed down, if necessary, until  it
       reached the surface of the diffuser. The biofilm thickness was then recorded  in mm.

4.      Enumeration of Live Bacterial Cells
       A 4 cm2 area of the diffuser was aseptically scraped with a sterile scalpel  blade into  a
       phosphate buffer solution (pbs) and serial dilutions are made to 10s.  A 1 ml sample  is
       taken  from each dilution and plated in duplicate on Brain Heart  Infusion (BHI) plates.
       Plates with significant numbers (between  30 and 300 colonies) are counted.
                                            55

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      To calculate the number of bacteria  per cm",  the  bacterial  count is, multiplied by the
      reciprocal of the dilutions and divided  by the initial area used.        i

5.     Carbohydrate Assay
      Measured against glucose standard curve of 200 ug/ml  according to Dubois, M. ej al (1).
      A 4 cm2 area' on  the  diffiiser surface is aseptically scraped with  a sterile scalpel  blade
      into 4 ml of 2 N-HaSO4 and kept frozen for analysis in the laboratory. >

   Reagents                                                            •
   (1) Concentrated sulphuric acid
   (2) Phenol (5%) in distilled water                                       ;
                                                                         i
   Procedure                                                            ;
      (1)      Frozen vials are hydrolized for 20 to  30 minutes in a' beaker of boiling water.
      (2)      Samples are serially diluted  1/10, U100
      (3)      The "standard glucose series" are prepared as follows:
             concentration
             of glucose

             distilled
             water

   Assay
       0.5 ml of phenol reagent is added to 1.0 ml of sample followed by  2.5 ml of concentrated
       sulphuric acid and vortexed immediately. The  addition of sulphuric acid must be rapid.
       Slow  or  sloppy addition leads to poor standard curves and results.   Samples  are
       incubated  in  darkness  for  60  minutes  and  then  read  at  490  nm  on  a  UV
       spectrophotometer.
0
0
1.0
20
0.1
0.9
40
0.2
0.8
60
0.3
0.7
80
0.4
0.6
200, ug/ml
1.0 ml
0 ml
       (1)     Colorimetric method for determination of sugars and related substances.   Anal
              Chem. 22 (3): 350-356, 1956.
                                         56

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