United States
Environmental Protection
Agency
Technology Transfer
EPA/625/8-89/015
Biomonitoring for
Control cjf Toxicity in
Effluent Discharges to
the Marine Environment
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                                                          EPA/G2S/8-89/015
  Biomonitoring for
  Control of Toxicity in
  Effluent Discharges to
  the Marine
  Environment
September 1989
Center for Environmental Research Information
Office of Research and Development
U.S. Environmental Protection Agency
Cincinnati, OH 45268

Environmental Research Laboratory
Office of Research and Development
U.S. Environmental Protection Agency
Narragansett, Rl 02882

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This document was written by Pamela DiBona, William
Heyman, and Heidi Schultz of Eastern Research Group, Inc.,
Arlington, Massachusetts. Production assistance was provided
by Susan Edwards. Technical assistance was provided by
Steven Schimmel, William Nelson, and Donald Phelps of the
EPA Environmental Research Laboratory (ERL) in
Narragansett, Rhode Island; Orville Macomber of the EPA
Center for Environmental Research Information; William Peltier,
EPA Region IV, Athens, Georgia; and Rod Parrish, ERL-Gulf
Breeze. Florida.

The principal investigator at ERL-Narragansett was Steven
Schimmel. Assistance was provided in the development of the
Complex Effluent Toxicity Testing and Biomonitoring  Programs
at ERL-Narragansett by Donald Phelps,  William Nelson,
George Morrison, Suzanne Lussier, Richard Steele, Margarete
Heber (now at EPA Headquarters, Washington, DC), Diane
Nacci, Glen Thursby, Elise Torello, Ruth Gobell-Gutjahr,  David
Bengtson, and Walter Berry. Contributions were provided by
Rod Parrish, ERL-Gulf Breeze, relating to toxicity testing of
drilling muds and produced waters. William Peltier and staff at
EPA Region IV (Athens, Georgia) conducted case studies and
provided data and assistance at two Florida locations.
Photographs were provided by  ERL-Narragansett. This
document is ERL-Narragansett Contribution Number  1036.

Technical review of the document was provided by Dr. Gary
Chapman, ERL- Newport, Oregon; Margarete Heber,  EPA
Office of Water Enforcement and Permits; Peter Nolan, EPA
New England Regional Laboratory, Lexington, Massachusetts;
Rod Parrish, EPA ERL-Gulf Breeze, Florida; William Peltier,
EPA Region IV, Athens, Georgia; and Landon Ross, Florida
Department of Environmental Regulation, Tallahassee, Florida.

This report has been reviewed by the U.S. Environmental
Protection Agency and approved for publication. The process
alternatives, trade names, or commercial products are only
examples and are not  endorsed or recommended by the U.S.
Environmental Protection Agency. Other alternatives  may exist
or may be developed. In addition, the information in this
document does not necessarily reflect the policy of the
Agency, and no official endorsement should be inferred.

This guidance was published by

 U.S. Environmental Protection Agency
 Center for Environmental  Research  Information
 Office of Research and Development
 Cincinnati, OH 45268

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  Foreword
This document describes the use of biological monitoring
as an effective water quality -based approach to
controlling the toxicity in discharges to estuarine and
marine waters. The development of these methods was
conducted primarily at EPA's Environmental Research
Laboratory in Narragansett, Rhode Island (ERL-N).  This
document is intended for Federal and State National
Pollutant Discharge Elimination System (NPDES) permit
writers, water quality specialists, managers of pollution
control systems, regulatory decisionmakers, research
scientists, and  informed laypersons.

The document is organized into seven sections. Section
1,  Introduction, is an overview of the regulations that
mandate a change to water quality-based permit limits
and the testing programs that support that change.
Section 2, Marine/Estuarine Complex Effluent Toxicity
testing  Program (CETTP), describes EPA's efforts to
establish the validity of whole-effluent toxicity testing for
marine/estuarine environments. The section describes the
test methods developed to determine the effects of whole
effluents on survival, growth,  and reproduction of several
test species. Section 3, In Situ  Biomonitoring, presents a
program to determine the biological responses of marine
and estuarine test species to pollutants. Section 4,
Integration of Effluent and In Situ Biomonitoring Programs,
describes how  the complex effluent and in situ
biomonitoring programs may be used in tandem to control
toxic discharges to marine waters.  Section 5, Application
of  the Biomonitoring Strategy: New Bedford Harbor Pilot
Dredging Program, describes how  ERL-N employed the
tandem approach described in the  previous section to
monitor cleanup options for one of the most polluted
industrial harbors in the country. Section 6, Conclusions
and Future Directions, discusses possible refinements to
existing marine biomonitoring programs and their use in
NPDES permitting. Appendix A summarizes the results of
five case studies in which the CETTP and in situ
biomonitoring programs were evaluated, and Appendix B
is a glossary of terms useful in  understanding the
monitoring programs.
                                                                                                           in

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   Contents
 1.
2.
3.
Introduction      1
Regulatory Background        1
Special Considerations for Marine and Estuarine
       Waters
                  1
Complex Effluent Toxicity Testing
   Program        2
In Situ Marine and Estuarine Biomonitoring
   Program        2

Marine/Estuarine Complex Effluent Toxicity
Testing Program        5
General Introduction to the Marine/Estuarine
   Complex Effluent Toxicity Testing
   Program        5
Red Algal Sexual Reproduction Test
   Method         5
Kelp Sexual Reproduction Test
   Method         6
Sea Urchin Fertilization Test Method        8
Mysid Survival, Growth, and Fecundity Test
   Method         9
Sheepshead Minnow Embryo/Larval Survival and
   Teratogenicity Test         9
Sheepshead Minnow Larval Survival and Growth
   Test       9
Inland Silverside Larval Survival and Growth
   Test       9
A Special Case: The Toxicity of Drilling Fluids -
   Mysid Static, Acute Toxicity Test
Analysis of Test Results        11
Field Verification of Test Methods
Test Precision     12
Relative Sensitivity of the CETTP Tests
Incorporating Test Results into NPDES
   Permitting        13
In Situ Biomonitoring     15
Historical Perspective      15
Biological Methods        16
Chemical Methods        18
Validation of Test Methods with Case
      Studies     18
                                             11
                                             12
                                             13

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  Contents (continued)
     Comparisons among Various Levels of Biological
       Organization           20
     The Integration of Biomonitoring into Marine
       Discharge Regulations      20
     Current Research          22
     Summary
22
4.   Integration of Effluent and In Situ Biomonitoring
       Programs      23
     Complex Effluent Toxicity      23
     Receiving Water Toxicity Estimates        23
     Integrated Approach for Toxic Discharge
       Control         24
5.   Application of the Biomonitoring Strategy: The
       New Bedford Harbor Pilot Dredging
       Project        25
     Introduction       25
     Dredging Project Description         25
     Monitoring Strategy       25
     Summary         28
6.   Conclusions and Future Directions      33
     Complex Effluent Toxicity Testing          33
     Future Research and Refinements to Complex
       Effluent Toxicity Testing Program       33
     In Situ Biomonitoring         33
     Future Research and Refinements for In Situ
       Biomonitoring   33
     Integrated Approach for Toxic Discharge
       Control - Integration with Large-Scale Coastal
       Monitoring      34
     The Integrated Approach - Field Tested     34
7.   References
35
 Appendix A. Case Studies        39
     CETTP Case Study I: Fernandia Beach,
        Florida         39
     CETTP Case Study II: East Greenwich, Rhode
         Island         40
VI

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  Contents (continued)
    CETTP Case Study Ml: Panama City,
      Florida        43
    In Situ Biomonitoring Case Study: East Greenwich,
       Rhode Island *       44
Appendix B.  Glossary
57
                                                                                      vii

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   1.     Introduction
Regulatory Background
In 1972, the National Pollutant Discharge Elimination
System (NPDES) was established under the Clean Water
Act to control point source pollution to the Nation's
surface waters. Point sources of pollution can be directly
attributed to a specific source, such as a factory or
publicly owned treatment works. Every point source must
apply for and obtain an NPDES permit that limits the
concentrations of pollutants that may be discharged in its
effluent. EPA has delegated permit-setting activities to
States  whose permit programs meet applicable EPA
requirements; these States operate their own NPDES
programs with EPA overview.

In 1984, EPA issued a policy statement recommending
effluent biomonitoring as one part of a "water quality-
based approach to controlling toxic pollutants" (1). The
policy directed States to use data from biological testing
of complex effluents discharged from point sources  for
setting NPDES permit limits. EPA also prepared a
technical support  document (2) that described ways to
implement the policy. This policy made biological testing
an integral part of the NPDES program under the Clean
Water Act. and effectively changed the permit-setting
emphasis from a technology-based approach to a water
quality-based approach. The 1987 amendments to the
Clean Water Act (now called the Water Quality Act)
reinforced this policy by stating that water quality
standards should  be chemical-specific numbers when
such numbers have been established, however biological
monitoring  should be used in areas where no criteria for
aquatic life exist and to address  the effect that the
interaction of chemicals has on toxicity (3). EPA made a
more definite step toward water quality-based control in a
proposed rule published in the Federal Register in
January, 1989 (4), which  states that "controls for whole
effluent toxicity are an essential component of EPA's
integrated approach to toxics control."

To control the discharge of toxics under the toxicity-based
approach, EPA recommends that complex effluent limits
be used in conjunction with limits on specific pollutants. In
many cases aquatic toxicity data for specific chemical
compounds are often not available for use in setting
standards. Also, wastewaters usually contain mixtures of
chemicals that are difficult for industries or municipalities
to characterize. Compounding these problems is the fact
that even if pollutants are discharged at toxic
concentrations below analytical detection limits, the
effluent may still be toxic. This toxicity may be due to a
variety  of factors:  an excess of nonconventional pollutants
(such as total suspended  solids); conventional pollutants;
unanticipated interactions between the chemical mixtures
in the effluent; and/or unanticipated interactions between
the effluent and the receiving water. Complex effluent
toxicity testing takes these factors into account and
complements the chemical-by-chemical approach.

Special Considerations for Marine and Estuarine
Waters
Since the 1984 shift to a water quality-based approach,
biomonitoring methods in freshwaters such as lakes and
rivers have been well established. Translating these
methods from freshwater to marine environments,
however, is not a simple task. Differences between the
two systems require alternative testing methods and test
species. In addition, the physical differences  between
streams and estuaries, rivers and bays, as well as the
chemical differences between saltwater and freshwater,
require special consideration.

Physical Differences
Many physical conditions affect the mixing of effluents in
marine/estuarine and freshwater systems. Marine and
estuarine systems, however, are much more complex -
water depth and current direction vary with the tides, and
tidal volume varies on a monthly and seasonal basis.
Models that predict effluent mixing must take these and
other factors into account. For example, an outgoing tide
may quickly carry the effluent plume into deeper waters,
but if little vertical or lateral mixing occurs along the
plume, then a concentrated effluent could be delivered to
a critical resource area (oyster beds, spawning grounds,
etc.). In addition, estuaries tend to act as chemical
"sinks." Stratification can trap effluents below horizontal
temperature gradients in the water, and the tide's
oscillation can restrict the plume from significant
horizontal movement (2). The contaminants sit and collect
in what becomes a basin for pollution. Finally, effluent
plumes may mix or disperse less easily in saltwater than
in freshwater because of differences in salinity and
specific gravity between the effluent and the surrounding
waters.

Chemical Differences
The buffering capacity of saltwater is much greater than
that of freshwater. Certain chemicals either break down
more easily or remain in their original toxic form longer in
saltwater than they would in freshwater. Also, the
speciation or form of the individual chemical mixture
components (e.g., solid, liquid, ion, or complexed) varies
depending on whether the effluent is in saltwater or

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 freshwater.  This is an important factor because some
 chemicals are accumulated or degraded by organisms in
 one form, but not in another.

 Biological Differences
'Marine and estuarine waters,  in general, support a greater
 diversity of species than do freshwater rivers and lakes.
 In addition, unlike rivers and lakes, in which fish may
 spawn and live the rest of their lives in one area,
 estuaries and bays act as nursery grounds for many
 species that later move out into open waters. Exposure to
 toxic effluents is more likely to affect these immature,
 more sensitive life stages.

 Resulting Differences in Approach
 These physical, chemical, and biological differences
 between marine and freshwater environments require
 different effluent and organism sampling methods as well.
 Because of a marine or estuarine water body's depth and
 expanse, and the possibility of disturbances from violent
 storms, marine biomonitoring  logistics are more difficult to
 plan and carry out. In freshwater biomonitoring, benthic
 surveys  are frequently used. These surveys involve
 cataloging all of the species inhabiting the  benthos, and
 characterizing the "health" of the water body on that
 basis. However, the benthic zone in marine environments
 often may not be directly affected by toxic  discharges.
 The discharge, if released in a freshwater plume, flows up
 through the water column to the surface of the water  and
 usually does not flow  along the bottom. Suspended
 solids, however, may  settle to the bottom.  Furthermore,
 findings from biological surveys of benthic  species may
 indicate  residual effects from  past pollution. Estuaries
 tend to serve as depositional  areas, therefore many
 discharges, particularly municipal discharges, superficially
 enrich the benthic sediments with organic carbon. This
 enrichment by itself, in the absence of toxic chemical
 contamination, can substantially alter the biological
 community.

 Populations of species within the marine community also
 change naturally as they move, die out, or populate the
 benthos in a patchy or irregular fashion, making the
 detection of adverse effects from pollutants more difficult.
 The ambient receiving water toxicity tests used in marine
 and estuarine environments measure the effects of the
 suspended plume on  indigenous species. In situ
 biomonitoring methods provide another alternative for
 measuring the effects of effluents on marine species. This
 permits the effects of any toxicity present in the waters
 surrounding the discharge to  be measured directly,
 without relying on laboratory exposure systems.

 The great diversity in species present in the marine
 environment affects species selection for toxicity testing
 as well.  Animal and plant kingdoms both must be
 represented in testing. All of these factors  were
considered when adapting freshwater methodologies to
marine conditions and species.

Complex Effluent Toxicity Testing Program

The Complex Effluent Toxicity Testing Program (CETTP)
was initiated as a means of testing whole effluents using
aquatic organisms as indicators of effect. The overall goal
of the CETTP was to enable regulators to use the
parameter toxicity data to set NPDES permit limits based
on water quality protection. Researchers first applied the
technology to freshwater systems. From the program's
outset in 1982, EPA has studied the toxic effects of
effluents in several rivers in the United States, with five
goals in mind:

  •  To verify that complex effluent toxicity tests predict
     and quantify adverse impacts to receiving waters.

  •  To identify complex effluent toxicity test procedures
     that support the NPDES regulatory process.

  •  To demonstrate, through case  studies, the
     effectiveness of tests in a variety of discharge
     scenarios.

  •  To field test short-term chronic toxicity tests.

  •  To compare the receiving water toxicity with the
     effluent toxicity observed at known concentrations
     (5).

The freshwater CETTP resulted in several  biological tests
that established the validity of whole effluent toxicity
testing.

EPA's marine CETTP set out to achieve the same five
goals described above for the freshwater portion of the
CETTP. To adequately address the differences between
freshwater and marine/estuarine water systems, the
number of test species was expanded, and ambient
receiving water toxicity tests were used rather than
biosurveys. The biomonitoring tests  described here were
designed for the measurement of toxicity to the
environment where relatively small ratios of effluent
volume to receiving water volume exist. Freshwater
receiving streams may contain more than 6>0 percent
effluent after discharge, but most estuaries and bays, by
virtue of their potentially greater volume, generally provide
higher effluent dilution.

In Situ Marine and Estuarine Biomonitoring
Program
The in situ marine biomonitoring  program focuses  on
determining not only contaminant concentrations, but also
contaminant effects. This program extends the CETTP
policy of emphasizing specific effects rather than specific
chemicals. The five objectives of the program are:

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                                                      I
•  To identify general problem areas or conditions of
   the Nation's bays and estuaries and other near-
   coastal waters.

•  To track dischargers' response to regulations by
   monitoring point source discharges and nonpoint
   sources of pollution.

•  To identify links between chemical pollutants and
   biological responses.

«  To identify links between impacts at different levels
   of biological organization.

•  To predict population- and community-level effects
   from the specific effects of chemicals on individual
   organisms.
In situ biomonitoring measures biological response in
individual organisms. Eventually, the tests will monitor
biological parameters at four levels of biological
organization: cellular, tissue/organ, individual, and
population.

The biomonitoring strategy utilizes a stepwise progression
of sensitive tests, from simple to more complex and time-
consuming; the first tests are performed on sensitive
species, and provide information that enables the
investigator to determine if other, more complex and
expensive tests are needed. The strategy brings together
regulators and scientists  in a single decisionmaking
process. Together, they decide whether the problem
justifies additional testing, and if so, which tests should be
used  (6).

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   2.     Marine/Estuarine Complex
          Program
General Introduction to the Marine/Estuarine
Complex Effluent Toxicity Testing Program

The 1984 amendments to the Clean Water Act
recommend  the use of biological test methods to help
.prevent the "discharge of toxic materials in toxic
amounts." The 1987 Water Quality Act further supports
EPA's increased use of biological test methods in setting
National Pollutant Discharge Elimination System (NPDES)
permit limits to protect our Nation's surface waters for
their designated uses.  In response to the  1984
amendments, researchers at EPA's Environmental
Research Laboratory in Narragansett,  Rhode Island (ERL-
,N) developed the marine component of the Complex
'Effluent Toxicity Tesjing Program (CETTP). The program
includes tests that estimate "safe" or "no effect"
concentrations for toxic effluents and thus predict
allowable effluent concentrations that should provide for
the normal propagation of fish and other aquatic life in
marine receiving waters.
U.S. EPA's Environmental Research Laboratory in Narragansett,
Rl.
Historically, effluent toxicity tests measured only acute or
short-term (generally less than 96 hours) effects with
lethality as the test endpoint. Such methods and their
application are provided in "Methods for Measuring the
Acute Toxicity of Effluent to Freshwater and Marine
Organisms" (7). Since actual field exposures often occur
at sublethal concentrations, EPA realized the need to
directly test the effects of chronic (longer-term, lower
concentration) exposures. The 1985 acute methods
manual provides a means to protect for chronic effects by
Effluent Toxicity Testing
 extrapolation from the acute toxicity determinations.
 Previously definitive chronic toxicity testing required a
 minimum of 30 days up to a full year to measure the
 actual effects of low toxicant concentrations on the entire
 life cycle of an organism. These full life cycle tests were
 developed with several fish species and with mysids but
 virtually no other species. Since the observed  toxicity in
 these tests was most strongly manifested during the
 organisms early and most vulnerable life stages,  the tests
 were abbreviated to include only these life stages. In an
 effort to produce comparable, reliable results within  much
 shorter time periods and at reasonable costs, tests were
 developed that focused on  several  other organism's
 vulnerable life stages. Test endpoints were targeted to
 survival, growth, fecundity,  and reproduction. Eventually,
 abbreviated marine chronic tests were developed that
 estimate long-term toxicity in less than 9 days. The
 CETTP tests evaluate the biological (toxicity) effects of
 complex effluents or receiving waters. The tests,
 therefore, include the additive effects of complex effluents
 on several important life functions of exposed organisms.

 Researchers developed tests for six species in four
 phylogenetic groups under the CETTP (Table 2-1). The
 kelp toxicity test has just recently been developed for the
 CETTP. All of these tests may be used to develop permit
 limits and assess compliance for toxic discharges into
 marine/estuarine environments. The techniques in these
 tests are designed to minimize equipment and  sample
 volume so that they can be conducted either on site, in a
 mobile laboratory, or at an offsite lab, by shipping minimal
 volume samples of effluents or receiving waters.

 The following sections briefly describe these tests.
 Detailed descriptions of the test methods are provided in
 "Short-term Methods for Estimating Chronic Toxicity of
 Effluents and Receiving Waters to Marine and  Estuarine
 Organisms" (8) and "User's Guide  to the Conduct and
 Interpretation of Complex Effluent Toxicity Tests at
 Estuarine/Marine  Sites"  (9). General guidelines for
 conducting short-term chronic toxicity tests are presented
 in Table 2-2. EPA also is currently developing tests with
 additional marine plant and animal species from other
 phylogenetic groups and other geographic locations.

 Red Algal (Champia parvula) Sexual
 Reproduction Test Method
 This test method  estimates  the chronic toxicity of
 effluents and receiving waters on the sexual reproduction

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Table 2-1. Test Species, Response Parameters, and Test Duration of the
Marine/Estuarine Short-term Complex Effluent Toxicity Tests (8)
Species Test Duration Test Endpoint(s)
Champia parvula
(Red macroalgae)
7 to 9 days total;
2-day exposure, 5- to
7-day incubation
Cystocarp
production
                   Laminaria saccharina
                   (Kelp)
                   Arbacia punctulata
                   (Sea urchin)
5 to 9 days total;
2-day exposure, 3- to
7-day incubation

1.33 hours total;
1 -hour sperm exposure,
20-minutes fertilization
Sporophyte
production
Egg
fertilization
Mysidopsis bahia
(Mysid)
Cyprinodon variegatus
(Sheepshead minnow)
Cyprinodon variegatus
(Sheepshead minnow)
Menidia beryllina
(Inland silversides)
7 days .total;
7-day exposure
9 days total;
9-day exposure
7 days total;
7-day exposure
7 days total;
7-day exposure,
Survival, growth, and
egg production
Survival without
terata
Survival and
growth
Survival and
growth
 ERL-N's mobile laboratories.
of the marine macroalga, Champia parvula. Seaweeds
had previously been considered far less sensitive to
toxicity than aquatic animals, and thus not useful for
toxicity testing. However, this conclusion was based on
algal growth as the test endpoint. Algal reproduction tests,
including those with Champia, can sometimes be more
sensitive than tests with aquatic animals.

Unialgal stock cultures of Champia parvula should be
maintained, so that mature plants are available year round
for toxicity  testing. There are three macroscopic stages in
the life cycle of Champia, and all three look the same
superficially (Figure 2-1); however, only the male and
female plants are used in toxicity testing. The female
              production of cystocarps, which are the products of red
              algal sexual reproduction, is used as the endpoint.

              The toxicity test with Champia consists of first combining
              male and female plants and exposing them for 48 hours,
              during which time fertilization should occur, and  then
              allowing cystocarps to develop over a 5- to 7-day
              recovery period.  Five female branches and one male
              branch are placed together in each test chamber, with
              three or four replicate chambers for each treatment. After
              exposure to the effluent, females are removed and placed
              into clean seawater. At the end of the recovery period,
              the number of cystocarps per plant are counted. The
              results are compared to a control in a series of statistical
              tests to calculate the "no effect" concentrations of the
              effluent.

              Kelp (Laminaria  saccharina) Sexual
              Reproduction  Test Method
              ERL-N recently developed this test method for use in the
              CETTP (10); therefore, the method is not included in the
              original test manual (8). The method of estimating the
              chronic toxicity of whole effluents and receiving waters on
              the sexual reproduction of the brown macroalga
              Laminaria saccharina is similar to that of the Champia
              test.  This test is  an excellent complement to the Champia
              test because, while Champia is found in warm waters,
              kelps are primarily found in cold waters throughout the
              west coast of North America and in New England on the
              east coast.

              ERL-N maintains unialgal stock cultures of male and
              female Laminaria that are available for toxicity testing. Six

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Table 2-2.    General Guidelines for Short-term Marine Complex Effluent Toxicity Test Methods3 (8,9)
          Guideline
                                                              Description of Guidelines
 Health and safety

 Quality assurance
 Facilities and equipment

 Effluent samples
                                 Appropriate measures should be
                                 effluents or receiving waters.
aken to protect against possible health hazards derived from
 Dilution water
                                 All tests should be performed by aquatic toxicologists or under their supervision. Tests require high-
                                 quality test conditions to ensure confidence in test results. Quality considerations include: animal
                                 health, sample collection and han lling methods, instrument condition and calibration, adequate
                                 controls and replication, accurate recordkeeping, use of reference toxicants, and test acceptability
                                 criteria.

                                 Tests can be conducted in an appropriately equipped stationary or mobile laboratory.

                                 Water samples should be collected just prior to the start of tests. Water samples are generally
                                 collected as  24-hour composite samples,b which are appropriate when the components of an
                                 effluent remain relatively constant in volume and abundance.  (Note, however, that this sampling
                                 method may not adequately represent short-term peaks in toxicity. This type of toxicity must be
                                 addressed using acute tests on grab samples.)  Samples of whole effluents are collected at the
                                 discharge site,  while receiving wa ers are collected at a series of predetermined sampling stations,
                                 representative of varying degrees of impact.

                                 A range of test solutions is create i with varying effluent concentrations to determine the
                                 concentration of effluent that is toxic to the organism.  Five different effluent concentrations and a
                                 control containing no effluent are selected for each test. Careful steps are taken to ensure that
                                 salinities in all samples are identical, and remain constant throughout the experiment. Salinity is
                                 adjusted with the aid of either artificial  seawater (for some but not all species) or with hypersaline
                             brine, made from evaporating higr
                                                              quality, filtered seawater.  Seawater of desired salinity is created
                             by mixing a proportion of brine wit n deionized water.

 Endpoints for chronic toxicity    Endpoints in these tests include survival, growth, fecundity, and reproduction.
 tests                                                     I

 Statistical analysis of test       The statistical analysis of these tests should be conducted by or with the aid of a qualified
 results                       statistician. Through a series of statistical tests, two different estimates of the toxicity of an effluent
                             are made:  the NOEC (No-Obserjed-Effect Concentration),  an estimate of the highest concentration
                             that will not have a significant effect on the organism; and the EC (Effective Concentration), an
                             estimate of the proportion of orgarjisms that will show effects at any given concentration.

aThese guidelines refer to the conduct of all tests mentioned in this section.
bComposite sampling is generally conducted automatically with a co lection apparatus placed adjacent to a discharge. A 5-liter water
 sample is often collected over a 24-hour  period by collecting small volumes of water at regular intervals.


                                                            days prior to testing, males and females are placed
                                                            separately in a standard  blender and are blended to
                                                            provide smaller cell clusters,  filtered to remove remaining
                                                            large clusters, and  diluted. Males are  placed in petri
                                                            dishes and females on small  glass slides,  100 to  150
                                                            females per slide.  The cultures are maintained for 5 days
                                                            in  a high iron  medium to allow gametogenesis to occur
                                                            (Figure 2-2). If significant pregametic cells are identified
                                                            after 6 days, the water in male  dishes is replaced either
                                                            with treatment (various effluent dilutions) or with control
                                                            seawater. One slide of female parts is then  placed in
                                                            each of the male dishes.  Exposure is  for 48 hours, after
                                                            which the females are removed and placed  into clean
                                                            seawater. After an additional 4 to 7 days, the slides are
                                                            examined for sporophytes, the products  of Laminaria
                                                            sexual reproduction. The results  are expressed as the
                                                            number of sporophytes in test conditions compared to the
                                                            number of sporophytes that develop in control seawater.
                                                            The results are statistically analyzed and the "no effect"
                                                            concentrations of the effluents are  derived.
Robust sample of the marine macroalga Champia parvula.

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Tetrasporangia^
                                           Spermatia
                                           Fertilization
      Tetrasporophyte
                                           -Cystocarp
 Figure 2.1. Sexual reproduction in the Red Marine Macroalga
          Champ/a parvula (8).
  Cystooarps in a mature female Champ/a parvula.
 Sea Urchin (Arbacia punctulata) Fertilization Test
 Method
 This test method, adapted from a method by Dinnel et al.
 (11), estimates the chronic toxicity of effluents or
 receiving waters to the gametes of the sea urchin
 (Arbacia punctulata). Arbacia adults are hardy and can be
 held in aquaria for long periods. This species can be
 induced to spawn year round, making it especially
 convenient to work with.  Because this test can be
 conducted rapidly {1 day), it can be used to quickly test a
 multitude of effluent types and locations.

 Adult A. punctulata  can be collected easily along the
 Atlantic coast or purchased from commercial suppliers.
 Once obtained, they are  separated into male and female
 cultures. Male gametes are released by stimulating the
                                                                           Laminaria saccharina
   Microscopic

  Antheridia
                                                              Male

                                                              eggs
                                                           Figure 2-2.   Sexual  reproduction  in  the Kelp Laminaria
                                                                      saccharina (10).
                                                            Adult sea urchins Arbacia punctulata.
organism with a 12-volt electrode. The sperm are
collected and iced for not more than 1  hour. Female
gametes are collected  similarly and may be maintained at
room temperature for several hours before use.

Urchin sperm are exposed to effluents or receiving waters
for 1 hour. Then eggs are added to the sperm and
allowed to incubate. After 20 minutes of incubation, the
test is terminated by the  addition of a preservative. The
test is conducted with three or four replicates of each
treatment, each containing 2,000 eggs and about 5 million
sperm.

Percentage fertilization is calculated after examining a
subsample from each test vial under a dissecting
microscope. The test results are recorded as the
percentage of eggs fertilized in test solutions compared to
the percent fertilized under control conditions. The results

-------
 are used in a series of statistical tests to calculate the
; "no effect" concentrations of the effluent.

 Mysid (Mysidopsis bahia) Survival, Growth, and
 Fecundity Test Method

 This test method estimates the chronic toxicity of
 complex effluents and receiving waters to the vulnerable
, period of egg development of the mysid (Mysidopsis
 bahia). Maturing juveniles are exposed to various
 concentrations of effluents or receiving waters in a 7-day
 test. Each  day, new sample water is collected and
 prepared with appropriate effluent concentrations,
 salinities, and food concentrations, and maintained under
 appropriate environmental controls.  Eight replicates of
 each concentration are maintained with five mysids in
 each treatment to ensure the statistical significance of
 test results. The test is completed after the mysids are
 exposed to the test solutions for 7 days.

 At the termination of the  test all mysids are transferred to
 clean seawater for observation. The sexes are determined
 and the number of immature mysids, mature males,
 mature females with eggs,  and  mature females without
 eggs are recorded (Figure 2-3). Dry weights are
; determined to calculate growth. The test results compare
 mysid survival, growth (weight), and fecundity (the
 percentage of females carrying  eggs) in test solutions to
i mysid survival, growth, and fecundity in clean seawater
 controls. Statistical tests  are used to calculate the "no
 effect" concentrations of the effluent.

 Sheepshead Minnow (Cyprinodon variegatus)
 Embryo/Larval Survival and  Teratogenicity Test
 This test method estimates the  chronic toxicity of whole
 effluents and receiving waters to the early developmental
 stage of the sheepshead minnow (Cyprinodon
 variegatus). (This method was developed at EPA Region
,VI Laboratory, Houston, TX.) Newly  fertilized (less than 24
 hours old)  C. variegatus embryos are exposed to a range
 of effluent concentrations or receiving waters from shortly
 after fertilization, through hatching, and for 4 days into the
 larval period. Each day, new sample water is collected
 and prepared with appropriate salinity, effluent
 concentrations, and other environmental controls. No food
 is required  in this test because the extremely young
; larvae can  survive on the nutrients provided in their yolk
 sacs. Three or four replicates of each concentration are
 maintained with 10 to 15  embryos in each treatment to
 increase the significance  of test results. The test is
 completed,  after 9 days of total exposure or 4 days of
 post-hatch  exposure, whichever comes first.

 At test termination, all live fish larvae without obvious
 terata are counted to determine survival in each test
 chamber. Survival is counted as those healthy larvae
 without deformity (terata) at the  completion of the
 exposure period. There is no measurement of growth in
 this test. The test results  are acceptable if the larvae in
 control cultures show survival rates greater than 80
 percent. A series of statistical tests again are used to
 calculate "no effect" concentrations of the effluent from
 the results of survival and terata determinations.

 Sheepshead Minnow (Cyprinodon variegatus)
 Larval Survival and Growth Test

 This test method estimates the chronic toxicity of whole
 effluents and receiving waters to the sheepshead minnow
 (Cyprinodon variegatus). By focusing on the vulnerable
 early larval stage, the test estimates the impact on the
 entire life cycle of the fish.

 In the test, newly hatched (less than 24 hours old) fish
 larvae are exposed to various concentrations of effluents
 or receiving waters for 7 days (Figure 2-4). The salinity
 range suitable for conducting the test is 20 to 32 parts
 per thousand (ppt); an appropriate salinity is selected and
 is held constant throughout the experiment. Each day,
 new sample water is collected and prepared with
 appropriate dilutions, salinity, dissolved oxygen (DO), and
 temperature. Three or four replicates of each
 concentration are maintained with 10 to 15 larvae in each
 treatment. After the new water is added each day, the
 larvae are fed to excess with newly hatched  larval brine
 shrimp, Artemia sp. The test is completed after the
 minnows are exposed to the test solutions for 7 days.

 At test termination,  all healthy larvae are first counted to
 determine survival, then sacrificed, dried, and weighed to
 determine growth. The test results compare the larval
 growth or survival in test solutions to the larval growth
 and survival in clean seawater controls. The  test results
 are acceptable if the larvae in the control cultures (no
 effluent) show survival rates greater than 80  percent and
 acceptable dry weights (8). A series of statistical tests are
 used to calculate the "no effect" concentrations of the
 effluent.

 Inland Silverside (Menidia beryllina) Larval
 Survival and  Growth Test

 This test method estimates the chronic toxicity of whole
 effluents and receiving waters on the vulnerable larval
 stage of the inland silverside (Menidia beryllina)  (Figure 2-
 5). This fish species is an important dietary component of
 many commercially harvested species including  mackerel,
 bluefish, and striped bass. These minnows are tolerant of
 a large salinity range (<5 to 32 ppt) and can therefore be
 useful  for effluent testing in estuaries where salinities may
 be lower than in open ocean environments.

 Test methods for this species are nearly identical to those
for the sheepshead  minnow survival and growth  tests
described previously. Seven- to 11 -day-old larvae are
exposed to various concentrations of effluents or
receiving waters for 7 days. Results of this test are
recorded as the comparison  between larval growth

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                                       Eyestalk
             Antennule
                                                     Carapace
                                                             Developing Brood Sac

                                                         Oviducts with Developing Ova
Figure 2-3.   Mature female mysld Mysidopsis bahia with eggs in the brood sac (8).
                                                                                 Cyprinodon variegatus
                                                                                  Sheepshead Minnow
Conducting the sheepshead minnow toxicity test in  ERL-N's
mobile laboratory.
(weight) or survival in test solutions and larval growth and
survival in clean seawater controls. Test results are
acceptable if larvae in control conditions exhibit survival
rates greater than 80 percent and acceptable dry weights
        7-Days Old


Figure 2-4.   Early  development of the  juvenile sheepshead
            minnow Cyprinodon variegatus (8).
(8). The results of survival and growth determinations are
put to a series of statistical tests and the "no effect"
concentrations of the effluent are derived.
 10

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                    Menidia beryllina
                    Inland Silverside
1 Figure 2-5.   Inland silverside minnow Menidia beryllina (12).


 A Special Case: The Toxicity of Drilling Fluids -
 Mysid (Mysidopsis bahia) Static, Acute Toxicity
 Test

 Oil and gas drilling operations produce effluents that are
: frequently discharged to marine/estuarine environments
 (Figure 2-6). Drilling fluids, an added constituent used to
 lubricate the drill bit, contain primarily clays and minerals
 (for example barium sulfate, bentonite, lignite, and
1 lignosulfonate), but also contain materials used as
 dispersants and thinners, fluid loss reducers, lubricants
 and emulsifiers, defoamers, biocides, corrosion  inhibitors,
 and pH control/stability additives (13). Most commonly,
 the toxicity of drilling  fluids is assessed by determining
, the acute toxicity of the fluids to mysids, Mysidopsis
 bahia, according to the method  developed at EPA's
 Environmental Research Laboratory in Gulf Breeze,
' Florida (ERL-GB).

 In the 96-hour test, 3- to 6-day-old mysids are exposed to
 various concentrations  of drilling fluids. Samples are
 prepared by mixing the drilling fluid with seawater (1 part
 fluid: 9 parts seawater, v:v), allowing the sample to settle
 for 1 hour, and then decanting only the suspended
 paniculate phase (SPP) for toxicity testing. If necessary,
 the pH is adjusted to the seawater's original alkalinity and
 the dissolved oxygen concentration is increased by
; aeration to at least 60 percent of saturation. A range-
 finding test should be conducted with samples of
 unknown toxicity to determine the range of concentrations
 for the definitive test. The definitive test uses mysids from
 a single hatch that are randomly distributed into five
 concentrations of the SPP plus a control.  Three replicates
 of each concentration are maintained with 20 mysids in
 each. All treatments are gently aerated for the duration  of
 the test. Mysids are fed Artemia at a rate of 50 Artemia
 per mysid per day. Temperature, dissolved oxygen, and
 pH are measured and recorded  after 0, 24, 48, 72, and
 96 hours.
After 96 hours of exposure, the test is terminated and the
number of live mysids in each treatment is recorded. The
test results are acceptable if at least 90 percent of the
mysids in control conditions survived. A series of
statistical tests are then used to estimate the median
toxic concentration of the drilling fluid.

The toxicity of drilling fluids is not routinely assessed with
the chronic toxicity tests thus far described because of
the physical characteristics of the fluids which may
contain materials of varying densities  that separate upon
settling. A special test apparatus that  maintains particles
in suspension could be adapted so that the CETTP tests
could be used to assess potential impacts of drilling
fluids.

Analysis of Test Results

The results of the CETTP tests are used to estimate the
interval between the highest "no effect" concentrations  of
the effluents and the lowest effect concentrations of the
effluents tested. The statistical methods employed are
rather complex and should be handled by, or with the aid
of, a statistician. U.S. EPA has prepared  a complete
explanation of these methods (8). The environmentally
"safe" concentration of an effluent is the highest
concentration of that effluent that will not have any
adverse effects on the organisms in a receiving water.
This "safe" or "no effect" concentration of an effluent is
estimated with toxicity testing. By using statistical
methods on the same set of data from a  single  test, two
different expressions of toxicity are calculated.

The first method of statistical evaluation uses a threshold
model of toxicity. The method  can be  used to derive the
No-Observed-Effect Concentration (NOEC) and the
Lowest-Observed-Effect Concentration (LOEG) of the
effluent. The use of NOECs and LOECs assumes both of
the following  premises. First, the model assumes that a
true threshold concentration exists, below which there is
no adverse effect to the organism and above which  there
is an adverse effect. In addition, the model assumes that
adverse effects that are not statistically observable are
not significant from a biological standpoint. If a threshold
model of toxicity is accepted, NOECs  will express the
highest concentration of the effluent that will not
adversely affect the organism tested.

The second method of statistical analysis is based on a
continuous or concentration-dependent model of toxicity.
Alternatively to (but not inconsistent with) the threshold
model, the concentration-dependent model is used to
derive Effective Concentrations (ECs). (If the endpoint
being tested is mortality,  EC is called LC  or Lethal
Concentration). By definition, any EC or LC value is an
expression of some amount of adverse effect. For
example, EC50 is the effluent concentration that would
affect 50 percent of the organisms tested. Interpretation
of EC values, therefore, requires the judgment of a
biologist. By calculating both NOEC/LOEC and EC values
                                                                                                               11

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''.' • ' ••*," '•*'..'•.' ' '- ' •
— Convective Descent' 	 ^
•'• :.'•'. '.••.;•'! :.-.• Enco
•• . V:!" .""..', ' Neutr


unter
al
ancy

Diffusive Spreading
Greater Than
Dynamic Spreading
-•• ' 	 •-•...'•";• ». , ??.- 	 "**•• •-*" — •-••- — -'^*' 	 ° * * — """"' '"•• "*"
Figure 2-6.  Effluent discharge plume from an oil and gas drilling operation in a marine/estuarinc environment (14).
from the same results of each test, biologists can provide
regulators with sound, reproducible data and a solid basis
for informed decisionmaking.

Biologists should use caution in interpreting test results in
cases where the'site-specific effects of pH, salinity,
temperature, and other natural conditions may affect the
bioavailability of the toxicants in question. For instance
the bioavailability of free ammonia is largely dependent on
the pH of the ambient water, and thus pH can affect the
apparently "safe" or "no effect" concentration of the
effluent,  and skew test results. An example of interactive
effects between an effluent and a receiving water was
encountered during the evaluation of the toxicity of the
effluent discharged from a pulp and paper mill in
Fernandina Beach, Florida (see Appendix A).

Field Verification of Test Methods

Validation of short-term toxicity tests is a necessary first
step before they are incorporated into the environmental
regulatory process. ERL-N validated these new methods
first with extensive field testing. This was accomplished
by comparing effluent toxicity to receiving water toxicity at
comparable effluent concentrations. Scientists conducted
11 such field tests at a variety of sites with  an array of
discharges from chemical plants, sewage treatment
plants, shipyards, and an aircraft refurbishing facility. The
results of these tests indicated a good correlation
between the toxicity of receiving  waters and comparable
effluent concentrations in the laboratory. Tests that were
most sensitive to effluent concentrations in  the laboratory
were correspondingly most sensitive to the appropriate
receiving waters in almost all cases. Case studies
illustrating this correlation are provided in Appendix A.

Test Precision

Precision tests were performed within a single laboratory
(intralab, ERL-N) to analyze the variability introduced by
the methods and by individual species. Precision
(closeness of repeated measures) is a measure of test
validity. Accuracy (closeness to a standard) for toxicity
tests, however, cannot be determined. The results of
precision tests were compared to the precision results of
similar toxicity tests for freshwater organisms; the
precision of the marine tests were found to fall within the
commonly accepted range (15,16).

Researchers calculated EC50 values from each toxicity
test that regulators would use for setting NPDES permits.
EC50 values are more useful for precision comparisons
than NOECs because they are not expressed only with
reference to the effluent concentrations in the test; rather,
they are derived from a continuous, concentration-
dependent response curve. ECSOs, therefore, are
discrete estimates of toxicity and can be expressed as a
single number for each  replicate of a given toxicity test.

Morrison et al. (17) examined  EC values for five replicates
of each short-term chronic test and statistically compared
them for variation. Precision test results indicated that
while some tests are more variable than others, the
overall precision of the tests is within the commonly
 12

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                                                        I
 accepted range for similar toxicity tests with freshwater
 species. The red algal (Champia parvula) reproduction
 test and the sea urchin (Arbacia punctulata) sperm cell
 test gave the most variable (yet acceptable) results, while
 the sheepshead minnow (Cyprinodon variegatus) results
 were most reproducible. For reliable results, EPA
 recommends that toxicologists use a representative of the
 fish, invertebrate, and plant tests when testing for toxicity.
i The exact testing requirements for a discharger, however,
 are specified on a case-by- case basis in NPDES permits.

 An intralaboratory precision test of the mysid, drilling fluid
 acute toxicity test was conducted at the ERL-GB (18).
The researchers found that the sources of variability
 included the condition of the sample itself, the manner in
 which the SPP is prepared, the condition of the test
 animals, and the skill and experience of the people
 conducting the test. They also found that the
 interlaboratory variability of the test conducted at 10
 commercial laboratories was comparable to the variability
. of interlaboratory acute tests conducted with single
 chemicals.

 Both single-lab and multilab precision tests were
 completed during the validation of the marine and
;freshwater acute toxicity tests (2). Similarly, an interlab
 precision test of the sheepshead minnow short-term
 chronic toxicity test has been completed from a total of
 seven tests by four participating laboratories. The labs
 reported very similar results indicating good interlab
 precision. Similar "round robin" precision tests of the
 remaining marine CETTP tests  will be completed by
 spring,  1990.

 Relative Sensitivity of the CETTP Tests

 In adapting the CETTP from the freshwater to  the
, marine/estuarine environment, scientists considered the
 greater diversity of plant and animal life in
 marine/estuarine waters. In an attempt to account for the
 extreme natural variation in biological communities, as
 well as  constantly changing physical and chemical
 environments, marine/estuarine toxicity tests should
 encompass a wide variety of species to be representative
 of the environment. The  complex effluents discharged
 can react and change and, therefore, are almost
. impossible to quantify or regulate through analytical
 chemistry alone. To take into account all of these factors,
,the marine CETTP uses biological toxicity tests with a
 variety of species types and test endpoints.

 The relative sensitivity of the tests is difficult to assess
 because different combinations of conditions may affect
 each species differently. All test results, however, can be
 used to calculate NOEC and EC values that are  used
 directly to set NPDES effluent permit limits. While species
 and endpoints differ, NOEC and EC values are directly
Comparable, indicating the relative sensitivity of test
species. No individual test has proven consistently the
most sensitive. NPDES permit limits, however, should be
dictated by the results of the most sensitive species
tested,  i.e., the lowest NOEC from all the tests
conducted. Therefore, the more species tested, the more
protective the test results will be for the receiving waters.

The sea urchin sperm cell test is often  the most sensitive
and can be conducted most rapidly. Therefore, regulators
are encouraged to use this test for rapid screening in
conjunction with two other tests.

Incorporating Test Results into NPDES
Permitting

Delegated States have the responsibility for issuing
NPDES effluent discharge permits. The States are
provided considerable leeway in their use of Federally
approved test methods for this purpose. The "Permit
Writer's Guide for Water Quality-based  Permitting for
Toxic Pollutants"  (19) supports  EPA's shift in  emphasis
from the solely chemical-specific approach to  one
integrating both the chemical-specffic and whole  effluent
techniques as a basis for developing NPDES permits.
Presently, regulators developing NPDES permits can  refer
to the permit writers guide,  "Methods for Measuring the
Acute Toxicity of  Effluents to Freshwater and Marine
Organisms" (7), and "Short-term Methods for Estimating
Chronic Toxicity of Effluents and Receiving Waters to
Marine and Estuarine Organisms" (8). EPA strongly
recommends that States include both acute and chronic
toxicity tests as bases for NPDES permits.

Inherent in the design of many water quality-based
discharge permits is the recognition of the mixing that
occurs after the effluent is released. The "design flow" of
effluents into freshwater is very well understood and can
be described with a variety of computer models (2). While
the mixing of effluents in marine/estuarine waters is not
nearly as well understood, some models are discussed in
"Initial Mixing Characteristics of Municipal Ocean
Discharges" (20). EPA recommends using "mixing
zones"  for water quality-based NPDES  permits, if
appropriate. Acute toxicity criteria are applied within the
estimated mixing zone while chronic toxicity criteria are
applied  at the  edge of this zone with consideration of
critical mixing  conditions. Treatment systems should be
designed to meet the more stringent requirements (acute
or chronic)  in each site-specific case. While toxicity tests
are water-quality based, the permits are based on
allowable allocations of priority pollutants and effluent
volumes, both instantaneous and averaged over various
time periods. A complete description of these  permitting
techniques is provided in the "Permit Writer's  Guide to
Water Quality-based Permitting for Toxic Pollutants" (19),
and case studies  of their application in marine/estuarine
systems are provided in Appendix A.
                                                                                                             13

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   3.    In  Situ Biomonitoring
 The in situ biomonitoring program at ERL-N includes
 toxicity tests with aquatic plants and animals as well as
 biological and chemical tests of bioaccumulation. During
 the design of the program, it was recognized that
 measurements of water chemistry alone (without a direct
 biological comparison) may only allow inferences from
' chemical concentrations observed in the field to actual
. biological effects. ERL-N is,  therefore, developing
 techniques to  assess biological effects and chemical
 uptake simultaneously. The goal is to provide regulators
 with a suite of biological and chemical tests that can be
 integrated for an effective assessment of the receiving
 water effects of complex effluents discharged into
 marine/estuarine environments.

 Many biological methods for toxicity testing are
 comparable to chemical methods in terms  of time and
 resources.  A  recommended strategy for toxicity
 evaluations, therefore, is to conduct simple biological
 assessments first, and then, if  necessary, conduct more
: complex biological and chemical tests  for a complete
 assessment. Finally, the monitoring  data must be
; effectively incorporated into the regulatory  process.
 Chemical-specific assessment  should additionally be used
 when pollutant criteria are available. Most traditional
 biomonitoring efforts have been implemented as follow-
 ups to environmental enforcement measures, and thus
 the data  collected are frequently not used to their
 potential. A key goal of the in situ biomonitoring program
 is to link scientists and regulators during the regulatory
 effort. This will ensure that data are  gathered in concert
 with the needs of the regulators and are available soon
 enough for inclusion in the decisionmaking process.

 Historical Perspective

 Narragansett Bay, Rhode Island, has suffered from the
 coastal dumping of industrial pollutants longer than any
 other area in the United States. Contamination first began
 in 1793 when the Slater Saw Mill was built  upstream of
jthe upper bay. Over the past two centuries, as the
 Narragansett Bay area became more and more
 industrialized, textile mills, machine tool activities, jewelry
 and plating operations, and human waste disposal
 systems  all contributed effluents to the bay.

 This long history of effluent discharge,  combined with  the
 continual tidal flushing and riverine currents of the area,
 have created a contamination gradient  throughout the
bay. The upper reaches of Narragansett Bay (especially
 sediments) have been heavily impacted by  pollutants
while the lower portions of the bay are  relatively clean.
 This situation has established a relatively stable
 contaminant gradient which has provided an ideal location
 for field testing marine/estuarine biomonitoring
 measurements.

 Initial bioaccumulation studies first showed the link
 between contaminant concentrations in mussel tissues
 and proximity to industrial discharges. For regulatory
 purposes, however, stronger inferences had to be made
 directly linking the observed mussel tissue contaminant
 residues with actual biological toxicity. In the early 1970s,
 the Coastal Environmental Assessment Station (CEAS)
 program was developed at ERL-N as a multipurpose
 program to assess the relative environmental health of
 marine coastal waters. The CEAS program used the blue
 mussel Mytilus edulis as a biological indicator species.
The blue mussel Mytilis edulis has been selected for ERL-N's in
situ biomonitoring program (21).
The mussels M. edulis and M. californicus are indigenous
to and abundant in bays and estuaries around much of
the coastline of the United States. Mussels feed by
filtering particulates out of surrounding seawater, so they
are excellent indicators of ambient water quality. In
addition,  mussels feed continuously throughout the  year
(without a dormant period),  making them a particularly
convenient species for physiological study. Finally,
mussels  are excellent for transplanting because
large.healthy populations of animals can be found and
distributed easily into field cages at various sampling
sites.  Another species, the hard clam Mercenaria sp.,
was explored as a test species but found not to be
amenable to year-round  in situ testing.
                                                                                                            15

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While the mussel has many advantages, its primary
disadvantage for short-term toxicity testing is its
reproductive habits. Species that reproduce frequently
provide results that are useful for extrapolation to
population level effects. Mussels spawn sporadically and
infrequently and, therefore, are not useful for assessing
effects on reproduction. Other species are presently
under consideration as future test organisms to directly
predict the population effects of toxic marine discharges
through reproductive tests.

The CEAS program is an intensive effort to relate
biological effects in mussels to the concentrations of
contaminants observed in their tissues. An initial step in
the program involved transplanting mussels on a transect
along a pollution gradient in Narragansett Bay (Figure 3-
1). Physiological measurements on transplanted mussels
indicated a correlation between biological effect and
contaminant concentration along the Narragansett Bay
Transect. Subsequent testing with a variety of biological^
tests supports these initial findings,  thus providing a
strong basis for use  of Narragansett Bay as a "field
laboratory" for testing various biomonitoring
methodologies.
                              Narragansett Bay
                              Rhode Island
     Providence
     River
     Greenwich
     Bay
 Figure 3-1.  Narragansett Bay, Rhode Island. Transect stations
            1-4 (22).
Currently, the in situ biomonitoring program at ERL-N is
evaluating biological effects at various levels of biological
organization. Recent results obtained from extensive field
tests indicate that the biomonitoring program may provide
valuable techniques to use as regulatory tools in  the near
future by predicting contaminant-induced perturbations  in
marine environments at the population level.

Biological Methods
A wide variety of biological methods have been tested at
ERL-N for potential use in the biomonitoring program. The
methods vary in complexity, cost, stage of development,
and the level of biological organization tested (Table 3-1).
Of all the biological methods explored, the scope for
growth (SFG) index in  M, edulis was found to be one of
the most effective methods evaluated and now forms the
primary component of  ERL-N's in situ biomonitoring
program. SFG  (a measure of the energy budget  of an
animal) has been used to measure the physiological
condition of mussels transplanted to polluted areas
compared to those transplanted to "cleaner" conditions
(23). Similarly,  SFG measurements on mussels exposed
to contaminated sediments in a lab exposure system have
been compared favorably to those of mussels exposed to
control conditions (see Section on Validation of Test
Methods with Case Studies). Additionally, shell growth
measurements on mussels can be used to estimate
biological effects of contamination. After either field or lab
exposures, chemical analyses of a subset of mussels are
examined for correlation with biological effects.

Scope for Growth Procedures
Calculation of the SFG index for M. edulis requires the
measurement of three parameters: clearance rate,
assimilation efficiency, and respiration rate.
Measurements are completed under standardized
conditions that match exposure conditions as closely as
possible, with respect to temperature and salinity. An
algal food concentration of 0.5 mg/L is used because it
provides sufficient energy for the organism during the
tests. All SFG  measurements are  completed within 24
hours of collection from field or lab exposures to ensure
that results are indicative of experimental conditions.A
detailed description of these  methods is provided by
Nelson et al. (24); however, a brief summary of  each
procedure is provided here.

 Clearance rate (CR). Mussels are placed into individual
chambers through which filtered seawater, with a set
concentration of algae, flows at a  constant rate.  After a 1 -
 hour acclimation period, CR  is determined at three hourly
 intervals for each mussel by  measuring incoming and
 outgoing algal  concentrations with an electronic  particle
 counter.

 Respiration rate. Respiration rates are determined by
 isolating each  mussel in a glass respirometer vessel fitted
 with an oxygen electrode. The electrode is connected  to
 16

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      Table 3-1.    Methods in Use and Under Development for In
                    Situ E

                    vel of
Biomonitoring (8)
                  Method
 Level of
Complexity
  Lev
 Bioogical
Organization
  Tested
                     Description of Method
       Methods Discarded

       Adenylate Energy Exchange (AEC)         High
       Blood chemistry                       Medium
       Methods Requiring Refinement

       Biomarkers including metallothionein       High
       induction, sister chromatid
       exchange, immunosuppression

       Histopathology                          Low
       Tissue microbiological contamination      Medium


       Demographic analysis                    High;



       Quantitative benthic sampling              High



       Methods that Can Be Used
       Immediately

       Growth, survival, and fecundity tests        Low

       Scope for growth (SFG)                Medium




       Gill respirometry                      Medium
                                                            O
an oxygen meter which, in turn, is connected to a strip
chart recorder. The decline in dissolved oxygen is
monitored on the strip chart recorder for approximately 30
minutes. Seawater containing algae (0.5 mg/L) is pumped
into the vessel during an acclimation period at a rate of .
80 mL/min to ensure that food is present in the chamber
and that routine metabolic rate is  measured.

Assimilation efficiency. Mussels are allowed to feed
overnight in  the individual chambers used to measure
clearance rate. The fecal  material is removed from each
                  Cellular      The amount of energy available to an organism from the
                              pool of adenine nucleotides (ATP, ADP, and AMP) can
                              sometimes be affected by stress. AEC is not
                              significantly affected by contaminant concentration and
                              has been discarded.

                  Cellular      Changes in the relative abundance of several normal
                              blood components might have indicated pollution effects.
                  Ce lular      There exist several potential biomarkers or sublethal
                              biochemical indices which might be used as early
                              biological sentinels of pollution effects.

                   Organ      Visual examination of sensitive tissues such as gills, liver,
                              and reproductive organs may indicate structural tissue
                              damage in response to contaminant exposure.

               Orgarj, system   Various bacteria might be used as tags for contaminated
                              sediments.

                 Population     Population dynamics can be inferred from the computer
                              compilation of individual responses to contamination
                              along a gradient.

                 Population     Community structure and contaminant  effects can be
                              determined by estimating benthic species diversity and
                              relative abundance.
                 Ind vidual     See Section 2 of this document.

                 Ind vidual     The energy left for growth and reproduction after routine
                              metabolic costs.  SFG decreases with increased
                              contaminant exposure.  (See full explanation in this
                              section.)
                    •gan       Health of animals determined by the respiration rate of
                              excised gill tissues. (Produces results which are
                              redundant to SFG so the test is not used.)
                       chamber the following morning.  Fecal material is dried,
                       weighed, ashed, and  reweighed to determine the ash-free
                       dry weight to dry weight ratio. A similar procedure is
                       completed with the cultured algae to obtain the ash-free
                       dry weight to dry weight ratio of the food. Assimilation
                       efficiencies are calculated for each treatment (25).

                       Scope for Growth Calculations
                       After completion of the physiological measurements, the
                       length and volume of each mussel is measured and the
                       tissue excised,  dried, and weighed.  The clearance rates
                                                                                                                          17

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and respiration rates are standardized to the mean weight
of all the mussels used in the treatment.

The weight-standardized values for each mussel are then
used to calculate the SFG of each individual by
substitution into the following equation:

   Scope for Growth  = (C x A) - (R)

where

   C =   energy consumed (clearance rate x
           surrounding food concentration x energy of
           food).

   A =    assimilation efficiency.

   R =   energy lost through respiration.

Shell Growth
To determine if shell growth is correlated with exposure to
toxic contaminants, measurements of mussel growth are
taken concurrent with field and lab exposures. The shell
length of 15 individually marked mussels per cage is
recorded before and after field or lab exposure. Growth is
calculated as the change in shell length per unit time.

Field Exposure
Prior to a field deployment, mussels are collected  with a
scallop dredge 1 to 2 days before introduction into test
waters. The collection  site should be similar to the site of
deployment with respect to temperature (<4°C) and
salinity  (<  5 parts per 1000) and exhibit good water
quality. The mussels are sorted to obtain a narrow size
range and are distributed into polyethylene field cages
(Figure 3-2). The cages are then transplanted to field
sites along  a gradient of contaminant concentrations,  as
shown for the Narragansett Bay sampling stations 1
through 4 (Figure 3-1). This technique of transplanting
along a transect is the basis of in situ biomonitoring at
ERL-N. After a specified deployment period,  usually 7 to
30 days, the cages are retrieved and the mussels
returned to  the lab. A subset of mussels from each cage
is used for biological testing,  while the remainder of the
mussels are frozen for chemical analysis.

Lab Exposure System
In addition to in situ biomonitoring, a laboratory exposure
system was developed for the mussel which will provide
further information for evaluating NPDES permits.  Initial
exposures with the system indicate that the sensitivity of
the mussel  to copper,  a standard  contaminant, is
comparable to other species  currently used in the
CETTP.

Chemical  Methods
The two methods used most commonly for determining
the concentrations of specific pollutants in animal tissues
       Surface
         Plastic Float
       Polypropylene
       Line
            Mussel
            Baskets
            Anchor
                                    (Sub-Surface)
                                         Approx.

                                          1 m
Figure 3-2.
             CEAS Station

Mussel  field cages used to transplant mussels
along transects (22).
 are atomic absorption spectrophotometry (AA) and gas
 chromatography/mass spectrophotometry (GC/MS). AA is
 used to measure trace metal concentrations, while
 GC/MS, together with electron capture gas
 chromatography, is used to determine the concentrations
 of volatile and semi-volatile organic chemicals like
 polychlorinated biphenyls (PCBs) and^olycyclic aromatic
 hydrocarbons (PAHs) (26).

 Validation of Test Methods with Case Studies
 The sensitivity testing and validation of response
 parameters used within the in situ biomonitoring program
 have been completed through a series of case studies.
 One such study to field verify these methods was
 conducted in conjunction with the U.S. Army Corps of
 Engineers' Field Verification Program at Black Rock
 Harbor (BRH), Bridgeport, Connecticut. The approach
 taken was to evaluate the relationship between
 contaminant exposure, mussel tissue residue
 concentration, and subsequent  biological effect. Mussels
 were exposed to various concentrations of contaminated
 sediments in the laboratory, followed by measurements of
 the mussels' contaminant tissue residues, SFG, and shell
 growth. Growth measurements  as well as SFG were
 highly correlated with the concentration of contaminated
 BRH sediments to which test organisms were subjected
 (Figure 3-3).

 A second study was initiated to determine the exposure
 duration period required to assess toxicity  in the field. The
 SFG results of 1-week exposures were compared to SFG
 results of 1-month exposures along a contamination
 gradient in East Greenwich Cove (see Appendix A).
 Figure 3-4 shows that SFG measurements decreased
18

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5-

§ 4-

-------
with increasing proximity to the outfall. While 30-day
exposures induced more pronounced SFG declines than
1-week exposures, the short exposure period produced
significant and usable results, illustrating the utility of 7-
day exposures. This study also showed that both
clearance rate (CR) and respiration rate (RR) were
significantly affected by exposure to toxics.

One might assume from Figure 3-3 that SFG results are
redundant with simple growth measurements; however,
the two endpoints can reflect different effects. SFG is a
measure of all the energy available for growth, both
somatic (including tissue and shell) and reproduction.
Therefore, increases in shell growth reflect only  a portion
of the energy available to the organism. During certain
periods of the year, most of the organism's energy  may
be allocated to reproduction  and very little to shell growth.
However, even during periods of slow shell growth, the
organism must still maintain the physiological functions
necessary for survival. Because of this, the SFG index is
more reflective of the overall condition of an organism at
any time of the year. In addition, the findings in East
Greenwich indicate that both mussel CR and RR can be
significantly affected by toxic exposure. Therefore, the
SFG measurements illustrate not only the extent of the
effect, but may also provide  an indication of the biological
mechanism behind the observed effect.

Comparisons  among Various Levels of
Biological  Organization
Currently, research is being focused on developing tests
for assessing impacts at many levels of biological
organization to ensure that indicated toxicity is of real
ecological significance. As a goal, tests at any level of
biological organization (i.e., cellular, tissue, individual, or
population) should predict the community/population
effects of a discharge. The SFG index on individual
mussels is indirectly indicative of population effects of
toxic pollutants.  Contaminant exposure has been shown
to decrease SFG, thus lowering energy available for
somatic  and reproductive growth. This lowered energy
may impact reproductive output and cause an effect at
the population level.

The more scientists know about the mechanisms behind
the observed effects, the better they will  be able to
interpret individual, tissue, and cellular responses and
predict population  responses. For example, showing that
a toxicant can inhibit an organism's ability to reproduce is
a way to predict the population effects of that toxicant.

Future research will concentrate on developing test
methods focusing  on all levels of biological organization to
gain a comprehensive understanding of marine
ecosystems and thus better understand contaminant-
induced perturbations within these environments. Tests at
the subcellular level may produce results, but those
 results may be difficult to interpret and extrapolate  to
community level effects. Changes at the community level
are difficult to detect because of the often extreme natural
variations that can occur in marine ecosystems. Even if
detected, community level changes are extremely difficult
to interpret, especially for regulatory purposes. For
instance, if a change is observed in community structure
or species diversity, it may only reflect natural variations
within a complex and changing ecosystem,  and not
necessarily indicate contaminant effects. A
comprehensive understanding of the biology of marine
ecosystems will allow better predictions of contaminant-
induced perturbations within these  environments, thus
providing regulators with a more solid  base  from which to
make informed  decisions concerning marine discharges.


The Integration of Biomonitoring into Marine
Discharge Regulations

The implementation of marine toxicity  evaluations  requires
the combined expertise  of both  marine scientists and
regulators.  Past monitoring efforts  have been conducted
without a formal means  for regulators  to use the
monitoring data generated, so the  data have rarely been
effectively incorporated  into decisionmaking. Because of
the lack of a framework within which to interact with.
regulators during the implementation of toxicity
evaluations, a conceptual framework for regulator/scientist
interaction was developed. The framework follows  EPA's
principle of selecting the simplest reliable indicator of
biological impact. The framework provides two
opportunities for regulator/scientist interaction.  In the first
phase, regulators pose questions that scientists and
regulators together design experiments to answer. They
can select tests from the array of methods  in Table 3-2.
The sensitivity  of the test chosen should be in
accordance with the hierarchy of relative sensitivity, cost,
complexity, and environmental contamination. Rapid and
inexpensive biological tests with a  low rate  of false
negative results should  be used for initial site
assessment, and more sophisticated biological, chemical,
and physical tests should be used when indicated  by
toxicity.


When test methods have been  selected and monitoring is
in progress, the second phase of regulator/scientist
interaction is invoked. A set of criteria are developed by
the same team of regulators and scientists  who designed
the initial monitoring strategy. The decision criteria
provide the team with the means to interpret the data
being gathered, reevaluate the techniques and strategies
employed in monitoring, and formulate decisions
regarding changes and  additions to the monitoring effort.
The flowchart in Figure  3-5 depicts steps that might be
followed during the risk assessment of a marine/estuarine
discharge. The decisions made at each juncture are
guided by the decision criteria developed by the
regulator/scientist team at the studies initiation. Using
decision criteria to ensure that proper data are gathered
within appropriate time frames, scientists and regulators
can beneficially interact, streamlining  risk assessment and
 20

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    Table 3-2.   Classification of Biological, Chemical, and Physi
                 the Proposed Monitoring Strategy (28)

                                        Level 1
                               :al Analyses into Four Categories that Are Subsequently Used in
                                                              Level
                                 I 2
                                                                                       Level 3
                                                                                                                Level 4
     Analysis
Survival
Growth
Fecundity
Chemistry - inorganic
    water analysis
     Technical sophistication        Low

     Time required to obtain results  Hours to days
Scope for growth
Gill respirometry
Chemistry j- organic water
    analysis
                     Intermedia e-high

                     Days to weeks
Population dynamics
Chemistry - tissue and
    inorganic analyses
                         High

                         Weeks/Months
Chemistry - Tissue and
     sediment analysis
Physical
     Bathymetry
     REMOTS interface
     Side scan sonar

High

Weeks/Months
                                                    Archive Tissue Samples
                                                    for Chemical |Analyses
                                                    at Low Priority
                                                     Area Receives Low
                                                     Priority as ar
                               Environ-
                                                     mental Problem
                                                     Area Receives Medium
                                                     Priority as an Environ-
                                                     mental Problem
                       Implement Pollution Abatement Measures
Figure 3-5.    Flow chart depicting the proposed use of decision criteria in risk assessment and coastal monitoring (28)
                                                                                                                                21

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drastically reducing the cost of marine/estuarine
biomonitoring. Decision criteria were effectively used
during the toxicity evaluation of New Bedford Harbor,
Massachusetts, described in Section 5.

Current Research
Ongoing research is currently  focused on evaluating and
developing several new techniques for the in situ
biomonitoring program. Of particular importance is the
field verification of lab results with mussels.  If field
verification is successful, scientists will be able to conduct
reliable, rapid, and inexpensive lab studies of mussel
exposure. The methods may then be  suitable for
incorporation into the appropriate NPDES permits. Future
research will focus on:
   • Tests with other species from other geographic
     locations - to increase the range and species
     representation of the tests.

   • Tests with other levels of biological organization - to
     further demonstrate the relationship between
     different levels and thus  provide insight into
     population and community level  effects.

Toward these two research goals, ERL-N is currently
evaluating another species, the slipper shell snail
Crepidula fornicata, for in situ transplanting. C. fornicata
are found on the east coast from Maine to the Caribbean
and on the west coast from California to  Baha. Like
mussels, these filter feeders are easily transplanted and
convenient for physiological studies. Unlike  mussels,  C.
fornicata reproduce frequently and prolifically, which may
be extremely useful for reproductive studies. The results
of tests with this species may be useful for  extrapolating
to population level effects of toxic marine discharges.
The  slipper  shell snail  Crepidula fornicata is  under
consideration as another species for in situ biomonitoring (21).
Summary
Upon completion of laboratory-to-field comparisons of
these tests, short-term laboratory tests will be
complemented by in situ assessment of receiving water
conditions. In contrast to extrapolation of laboratory
estimates of toxicity, in situ tests will measure the actual
physiological effects of field exposure to a variety of
point-source discharges with mussels, thus  providing an
important regulatory tool for marine toxicity evaluations.
 22

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   4.    Integration  of  Effluent and  In  Situ  Biomonitoring
          Programs
 Complex Effluent Toxicity

 To develop NPDES permits for the control of toxic
 effluent discharges to marine/estuarine environments,
 regulators require a quick, reliable, and inexpensive
 procedure to estimate effluent toxicity. To accomplish this
 requires an estimate of both the effluent's absolute and
 relative toxicity. Absolute toxicity is defined as the
 laboratory-determined toxic concentration of the complex
 effluent, while relative toxicity is defined as the toxicity of
 the effluent after mixing with the receiving water.

 Absolute toxicity is estimated with short-term CETTP
 tests. Relative toxicity can be estimated in two practical
 ways. First, relative toxicity can be estimated by sampling
 the receiving water at various sites and testing the
 samples with the  same CETTP tests used to assess
 absolute toxicity. Alternatively, relative toxicity can be
 assessed with in situ biomonitoring by determining the
 uptake of various toxicants and the corresponding
 biological effects on organisms that are transplanted into
 the receiving water.


 Receiving Water Toxicity Estimates

 Use of Dye Studies to Estimate  Effluent
 Concentrations in Receiving Waters

 While laboratory toxicity tests on various concentrations
 of effluent provide an estimate of "end-of-pipe" toxicity of
 a discharge, for regulatory purposes it is essential that the
 toxic concentrations of complex effluents determined in
 the laboratory correlate with the actual effluent
 concentrations observed in the field. The concentration of
 effluent in receiving waters at field sites, however, is
 difficult to predict  due to the complex mixing and
 dispersion of the effluent after it leaves the pipe.

 To estimate actual receiving water concentrations of
 effluents, dye studies are performed. Prior to  sampling
 any receiving water, the effluent is saturated with
 rhodamine WT dye by continually adding the dye to the
 effluent for 3 days or until an initial steady state is
 reached. Subsequently, when sampling occurs at various
 receiving water sites, the dye should be in equilibrium
with the effluent. The concentration of dye in the effluent
can be measured  with fluorometry (fluorescence), and
used along with the dye infusion rate and the  effluent
discharge rate to calculate the concentration of the
 effluent at each sampling site. In relatively stable
 environments, dye studies can be a powerful tool for
 comparing known, effluent concentrations in the lab with
 effluent concentrations at various field sites. In certain
 site-specific conditions, however, dye studies can be
 drastically altered by local environmental  conditions. For
 example, biologists should use caution when interpreting
 the results of a dye study conducted in shallow or wind-
 driven estuaries.
Researcher collecting a water sample during a dye study.
CETTP Tests of Receiving Waters (Relative Toxicity
Estimates)

Once dye studies have been conducted and the effluent
concentrations at field sites have been estimated, short-
term chronic toxicity tests are conducted with receiving
waters. Sampling sites vary in their proximity to the
discharge site. Toxicity tests on receiving waters are
conducted with the same set of test organisms used with
complex effluent samples alone, and the results are again
used to estimate the toxic concentration of the effluent.

ERL-N found that the toxic concentration of the effluent
measured in the lab is almost always the same as the
toxic concentration of effluent in parallel receiving waters.
The high correlation between lab and field results enables
regulators to incorporate these tests into NPDES permits
with a high degree of confidence. While the tests are
attractive because they can be conducted rapidly, in situ
biomonitoring might provide supplemental information on
                                                                                                           23

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longer-term effects. One of the longer-term in situ chronic
toxicity tests is described in the following paragraphs.

Use of In Situ Biomonitoring with Mussels to
Complement CETTP Tests
The efficacy of marine/estuarine complex effluent tests in
rapidly estimating an effluent's toxicity makes them an
important regulatory tool. However, there is concern  that
the very speed of the short-term tests may compromise
their long-term accuracy. This becomes increasingly
important during implementation because Regions and
States exhibit a wide latitude in their monitoring
requirements. For example, if all samples are collected
during 1 intensive week, but the effluent discharge
continues throughout the year, this "grab" sample
technique may not account for:

   • Temperature and seasonal effects, and
     noncontinuous discharge rates from effluent tanks or
     reservoirs.

   • Differences in effluent components over time.

To ensure that short-term toxicity evaluations are
consistent with long-term trends, it is recommended that
long-term in situ biomonitoring supplement the current
CETTP tests in certain site-specific discharge situations.
Through periodic deployments throughout the year,
mussels can be used to monitor long-term trends
(Section 3). In situ toxicity tests with  mussels may
eventually complement CETTP tests, either by monitoring
a specific discharge or as a general monitor of receiving
water health.

Integrated Approach for Toxic  Discharge
Control

The integrated water quality-based approach of using in
situ monitoring with existing CETTP methods for toxic
discharge control may be a powerful tool for regulators.
This approach will determine ambient water  quality by
measuring SFG on mussels and thereby assess the
toxicity of an effluent (Section 3). The first step in this
approach will be to compare SFG response  of mussels
 from the laboratory exposure system with field-exposed
 mussels, over 7 days. A second step will include a
 comparison of the sensitivity of 7-day lab exposure  with
 mussels to the sensitivity of short-term CETTP tests.

 Point-Source Evaluations and Large-Scale Coastal
 Monitoring
 The integrated water quality-based approach, combining
 toxicity testing of effluents and receiving waters with
 ambient toxicity testing, could be further incorporated into
 large-scale coastal monitoring  systems. The National
 Oceanic and Atmospheric Administration's (NOAA)
 Mussel Watch Program already uses in situ biomonitoring,
with mussels as regional indicators of marine ecosystem
health. This monitoring program periodically collects
mussels at about 150 coastal and estuarine sampling
sites,  and measures mussel tissue residues of tracer
contaminants including PCBs, PAHs, and several heavy
metals (29).  High tissue residues might indicate potential
toxicity problems within a region. In this way, Mussel
Watch can identify problem areas that require additional
testing to pinpoint discharge sites or upstream runoff
problems. EPA's new Environmental Monitoring and
Assessment Program (EMAP) will be especially important
in addressing ecological status and trends. EMAP will be
phased into implementation in 1992 and will integrate the
existing ecological  monitoring programs concerning the
Nation's forest, wetland, near-coastal,  inland surface
water, and agricultural areas. The near-coastal
component of EMAP might be an extremely useful tool for
integrating the results of localized marine toxicity
evaluations and large-scale marine monitoring efforts.

Combining the integrated approach with large-scale near-
coastal monitoring  programs may allow the tracking of
pollutants from "end-of-pipe" through receiving waters to
estuaries and  open bays. This integration has the
potential to vastly increase the ability to  assess the
results of existing pollution problems.  Baseline data of
this kind also will help to assess the field effectiveness of
remediation efforts.

Hierarchical Framework for Biomonitoring Tests

To further increase the efficiency of marine/estuarine
biomonitoring, it is proposed that all biomonitoring
techniques be placed into a hierarchical framework of test
complexity.  Each test will, therefore, be incorporated into
toxicity evaluation, according to the severity of the
problem. Section 3 describes the hierarchy of existing
tests, and the framework within which researchers and
managers can interact to best use their limited resources
to select from the  array of tests and control pollution of
our Nation's estuaries and oceans.

The most effective approach to controlling toxic
discharges is for scientists and regulators to work
together closely to design appropriate control strategies.
 Biological tests that both rapidly estimate toxicity and
 monitor the long-term trends of point-source discharges
 can be used in a framework within which managers and
 regulators can evaluate the data and make decisions
 rapidly. This will ensure that scientists provide the
 answers most needed by regulators, in  a timely fashion.
 In addition,  a tandem approach for localized or point-
 source toxic control  can be integrated into large-scale
 coastal monitoring programs. The integration of these
 monitoring programs could provide a dynamic picture of
 the local effects of toxic discharges and subsequent
 remediation measures, as well as the larger-scale effects
 in near-coastal waters.
  24

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   5.     Application of  the Biomonitoring
          Bedford Harbor Pilot Dredging
 Introduction

 New Bedford Harbor is located between the city of New
 Bedford and the town of Fairhaven in Buzzards Bay,
 Massachusetts (see Figure 5-1). Since the 1940s,
 electronics and manufacturing companies in the area
 have discharged PCB-laden effluent into the Acushnet
 River and the harbor.  High PCB concentrations in the
 river and upper harbor sediments were first documented
 in 1974 (30). Over the past 15 years, nearly 18,000 acres
 of PCB- and heavy metals-contaminated sediment have
 been described, with PCB concentrations as high as
 1,000 parts per million (ppm) in some areas of the harbor
 (31). In 1982, the site was added to EPA's National
 Priorities List of hazardous waste sites  slated for cleanup
 under the Superfund law.

 EPA's 1984 feasibility study proposed several alternatives
 for the harbor cleanup. One of the alternatives included
 dredging contaminated sediments and disposing of them
 in either a land-based, confined disposal facility (CDF), or
 an aquatic, confined aquatic disposal (CAD) area.
 Federal, State, and local officials, as well as the public,
 expressed concern over these alternatives. Many thought
 that the sediments stirred up during the dredging  would
 affect the plants and animals that inhabit the harbor's
 waters. Others cited potential pollution problems from
 contaminated water (leachate) leaking from the proposed
 disposal site (32). To answer these concerns,  EPA
 decided to pre-test the dredging and disposal options.
 Working with the U.S. Army Corps of Engineers (COE),
 EPA designed a pilot study to determine the best
 remediation option for the Superfund site.

 Dredging Project Description

 The COE examined and compared the effects of three
 hydraulic dredges and two disposal methods during this
 project. The dredges were selected  based on their ability
 to remove sediment with the least amount of
 resuspension and to operate in shallow  water.  The latter
was important because the dredging area was only 4 to 5
feet deep at high tide. The first dredge was a cutterhead
dredge; the second, a horizontal auger dredge called a
 "mudcat"; and the third, a specially constructed dredge
called a "matchbox" (34). The two disposal methods
investigated were:  1) a confined disposal facility (CDF),
which consisted of a containment dike partially in water
          Strategy: The New
       Project
 and partially on land; and 2) a confined aquatic disposal
 cell (CAD), which was an in situ underwater disposal
 method.

 The COE dredged two areas in the pilot study cove
 (Figure 5-2). Contaminated sediments from dredge area 1
 were disposed of into the CDF (Figure 5-3). The
 underlying "clean" sediment was also dredged to the
 CDF to "cap" the contaminated material. The resulting
 depression in area 1 became the CAD cell (Figure 5-4).
 The dredged material from  area 2 then was pumped into
 the CAD cell, and a  second "clean" layer of underlying
 sediment from area 2 was used to cap the underwater
 disposal area.

 Monitoring Strategy

 The pilot project was an opportunity to test the
 biomonitoring research and management strategy
 described in Section 3. Before the COE began any
 operations, baseline  physical, chemical, and biological
 measurements were completed. The biological
 measurements were used to assess the effects of
 existing water quality on plant and animal survival, growth,
 and reproduction. These  tests served as a benchmark
 against which increased contamination and/or toxicity
 associated with the operational phases of the study were
 compared.

 The determination of whether the operation caused an
 unacceptable effect was complicated by the fact that
 State and Federal water quality standards for PCBs and
 certain heavy metals were already exceeded under
 preoperational baseline conditions. In addition, the U.S.
 Food and Drug Administration (FDA) action level for PCBs
 in seafood was exceeded before dredging started.

 In order to assess the effects of the dredging project,
therefore, it was necessary  to develop a set of site-
specific numerical values, called Decision Criteria (Table
5-1). These criteria were  compiled in a Decision Criteria
Document (8), and were established for measured
physical, chemical, and biological parameters, based on
the  preoperational data. During the dredging and disposal
operations, data on the same physical, chemical, and
biological parameters were collected and compared to the
Decision Criteria values. A Decision Criteria Committee,
                                                                                                       25

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comprised of representatives from  ERL-N, EPA Region I,
the COE,  and the State of Massachusetts, evaluated the
data on a daily basis. If the Decision Criteria values were
exceeded, the committee could require engineering
corrections to the dredging operation before work was
resumed.  In this way, the marine environment and its
resources were protected during the pilot study activities.
Water Quality Monitoring
Four stations were selected for water quality monitoring in
NBH, in addition to a reference station located in
Buzzards Bay (Figure 5-5). Station NBH-1  was located
north of the dredge site; Station NBH-7 was adjacent to
the cove where dredging occurred; Station NBH-2 was at
the Coggeshall St. Bridge, the transition point between
                                            Wood St.
                                                                     Canada
                 Upper Harbor
                                                 Pilot Study
                                                 Area
                                              Coggeshall St.

                                               Rte. 195
               Lower Harbor
                            Atlantic Ocean



                           Cap Code

                          Buzzards Bay
                                                                           New Bedford Harbor
                                                                 / Long Island Sound	
                                                      • Butler Flats
                                                       Lighthouse
                                                  Upper Buzzards Bay


                   '3,000    ' 6,000 feet


  Figure 5-1.   New Bedford Harbor, in Buzzards Bay, Massachusetts (33).
 26

-------
 Aerial view of the New Bedford Harbor confined disposal facility
 (CDF). The silt curtain surrounding the study area extends from
 the far edge of the COF dike.
the more severely polluted upper harbor and the lower
harbor; and Station NBH-4 was at the hurricane barrier,
the transition point between NBH proper and Buzzards
Bay. The reference station for all water quality monitoring
was NBH-5, located at West Island in Buzzards Bay.
Mussels were deployed at stations NBH-2, NBH-3, NBH-
4, and NBH-5. Station NBH-6 was used as a mussel
reference station for the first deployment only;
subsequent deployments  utilized NBH-5 to facilitate
mussel collections.

Seawater was collected separately for the ebb and flood
tide at each NBH station.  This method allowed estimation
of net transport of PCBs and metals over time, especially
at NBH-2. Each water sample was analyzed for various
physical (total suspended  solids) and chemical
constituents {PCBs, copper, cadmium, and lead). In
addition, a suite of biological  tests were conducted to
assess acute effects (in fish, mysids, mussels, sea urchin
sperm cells, and a red alga) and chronic effects (fish
growth, mysid growth and reproduction, mussel scope for
growth, algal reproduction). A typical 7-day monitoring
cycle is presented in Figure 5-6.

Once the operational phases of the Pilot Project began,
water samples were collected in a manner similar to that
of the preoperational phase. However, during the
operational phase the ebb samples were returned to ERL-
N immediately and chemical analyses and acute biological
tests were completed overnight. These results were
transmitted to the Decision Criteria Committee prior to the
start of that day's dredging. The committee would then
assess any adverse environmental impacts and make any
necessary adjustments to  the operation. This "real-time"
monitoring allowed the project managers to make timely
decisions based on actual data, thus ensuring the best
degree of environmental protection possible.
 Summary of Results - Operational Phase
 Physical measurements. The temperature and salinity
 data were similar among the NBH stations during this
 study. Within the harbor, temperature decreased slightly
 with depth, and salinity increased minimally with depth,
 reflecting input from the Acushnet River at the head of
 the estuary. Total suspended sediment (TSS)
 measurements at the sampling stations indicated that the
 dredging operation  resulted  in  little or no increase in TSS
 concentrations over background.  On one occasion an
 increase in TSS concentration occurred at NBH-2. This
 was caused by the  opening  of the silt screen around the
 operation site which normally worked to contain
 suspended particles within the study area. This resulted in
 the discharge of a plume from the construction site. The
 problem was quickly rectified and never recurred.

 Water Chemistry. The PCB concentrations in NBH
 exceeded the Decision Criteria values on only four
 occasions (at NBH-2) during the operational phase of the
 project. Three of these instances could be directly
 attributed to a specific operational event. In all cases,
 modification of the operation resulted in lowered PCB
 concentrations in the water the following day. The only
 other exceedence was caused by a meteorological event:
 50 mph winds at low tide caused resuspension of
 contaminated sediment from the upper harbor.  At no time
 did the concentrations of copper, cadmium, and lead
 exceed the Decision Criteria values  as a result of any
 operational event.

 Mussel Chemistry. Mussels deployed in NBH showed a
 distinct spatial and temporal  pattern  with respect to PCB
 uptake. The PCB tissue residues from mussels deployed
 before dredging indicated that mussels located  in the
 upper harbor exhibited the highest PCB concentrations in
 their tissues;  the concentration decreased moving down
 the harbor (Figure 5-7). This pattern was consistent
 regardless of length of field exposure. The data indicated
 that PCB tissue residue concentrations also increased
 with  length of exposure. These preoperational data
 demonstrated the bioavailability of PCBs in NBH under
 background conditions. Analysis of mussel tissues
 exposed during operational phases of the project showed
 the same pattern.as the preoperational exposures.
 Therefore,  no additional increase in PCB bioavailability
 could be attributed to the dredging operation.

 Biological Tests. The sea urchin (Arbacia punctulata)
 sperm cell test and the growth  and survival tests with the
 sheepshead minnow (Cyprinodon variegatus) indicated no
 toxicity at any of the stations. The preoperational red
 algae (Champia parvula) reproductive tests proved
 inconclusive because of problems with stock cultures.
 Toxicity to this species was observed on several
occasions during operation. However, it was not
correlated with measured PCB  concentrations and may
have been caused by elevated  copper concentrations in
the water from a source other than the dredging

-------
operation, as the red macroalgae are particularly sensitive
to copper. Growth and reproduction in the mysid
(Mysidopsis bahia) indicated no effects as a result of the
dredging operation.

Mussel SFG and shell growth values from the first
preoperational deployment demonstrated a significant
inverse relationship with PCS tissue residue
concentrations in NBH (Figures 5-7 and 5-8). In addition,
SFG values on day 7 were reflective of results obtained
on day 28. This pattern was not evident during a second
deployment and was attributed to the fact that mussels
were at the peak of the gametogenic cycle at this time.
During gametogenesis, the mussels would expend extra
amounts of energy and the SFG measurement would be
affected. The SFG values of mussels collected during the
operational phase of the project never exceeded the
      Decision Criteria values; therefore, no adverse impact
      could be attributed to the operation;

      Summary

      The New Bedford Harbor Pilot Dredging Project was a
      unique opportunity to use biomonitoring techniques,
      ranging from the CETTP tests to in situ exposures, to
      evaluate a potentially environmentally damaging dredging
      operation on a "real-time" basis. A set of site-specific
      criteria was  developed and utilized by environmental
      managers to assess, on a day-to-day basis, the effects of
      this dredging operation on water quality in NBH. The
      results indicated that the dredging operation had a
      minimal effect on existing water quality. On those
      occasions when elevated PCB concentrations were
      detected, they could be directly attributed to a specific
      operational procedure and modified before environmental
                              CAD
                      Top of Bank

                         EL.+ 6'
 Dredge Area 2



CDF Discharge
                                                                  MLW
                   Scale: 1" - 400'
                   Datum: Mean Low Water (MLW)
  Figure 5-2.  Dredging areas and disposal facilities at New Bedford Harbor pilot study area (35).
 28

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Secondary Cell

     90' 	f
                 Scale:      1" = 70' Horizontal
                           1" = 14' Vertical
                 Datum:     Mean Low Water
                             CDF Discharge
                                                              Primary Cell

                                                             - 380' 	
                                                         	EL. + 10'	
                               Secondary
                               Settling
                               Cell
                                            Primary Settling
                   Cell
                                                                            /
                                                  Top of D
                                                                                           I
Figure 5-3.  Cross section and top view of confined disposal facility (CDF), showing the two settling cells (35,34).
damage was done. The result was that no acute or
chronic biological effects were observed as a result of
this operation. While the approach to real-time monitoring
can be labor intensive, its application is recommended
when an operation, such as dredging a Superfund  site,
                      can potentially have severe environmental impacts. The
                      Corps of Engineers is presently reviewing the data
                      gathered during the operation, and will decide on the best
                      dredging and disposal option for the more contaminated
                      portion of New Bedford Harbor
                                                                                                                 29

-------
                                                                                          -0.5'
                                                                                          -0.5'
                                         Datum: Mean Low Water g$ Clean Sediment
                                         Tide Range:  3.7'         :->: PCB-Contaminated Sediment
                                       Figure 5-4.    Cross section of confined aquatic disposal
                                                     (CAD) cell, unfilled and filled (35).
    Table 5-1.    Numerical Decision Criteria for New Bedford Harbor (36)
                         Endpoint
                                                     Time to
                                                      Obtain
                                                       First
                                                       Value
                          New Bedford Harbor Station Location
                    Coggeshall St. Bridge
                          (NBH-2)
                                            Hurricane Barrier
                                                (NBH-4)
      Water Chemistry
      PCB (total)
      Cd
      Pb
      Cu
      Mussel PCB Tissue
      Residues
        7-day exposure
      28-day exposure
      Biological Responses

      Acute  (% Survival)

      Fish
      Mysids
      Mussels
      Sea Urchins (% fort)
      Red Algae

      Chronic Effects

      Fish (dry wt, mg)
      Mystds (dry wt, mg)
           (reproduction)
      Mussels:
           scope for growth  (J/h)
           shell growth (mm)
      Sea Urchin (% fert)
      Red Algae  (# cystocarps)
  24 hr
  24 hr
  24 hr
  24 hr
 10 days
 30 days
 All tests
monitored
  daily
  8 days
  8 days
  8 days
 28 days
  1 day
  8 days
Ebb (itg/L)
1.4
9.3"
7.2
13.0
Net Transport (kg)
0.71/cycle
                  80
                 160
                 Ebb

 Mortality > 20% of control value for two
    species or > 50% for one species
Ebb
   0.44
    9.3*
   15.0
    6.0
                                19.0
                                23.0
                                                  Ebb"*

                                               20% (40%)
                                               20% (40%)
                                              50% (100%)

                                              7.5 J/h (15 J/h)
                                              50% (100%)
                                               25% (50%)
                                              50% (100%)
         "U.S. EPA water quality criterion; criteria-continuous concentration.
         This value represents a statistically and biologically significant reduction from control values. A significant reduction in two endpoints, or a
         twofold reduction in one endpoint (in parentheses), requires that the decision criteria committee evaluater the monitoring dataand other
         information related to the operation prior to the resumption of operations on the next day.
30

-------
Figure 5-5.   New Bedford Harbor sampling station locations for
            biological, physical, and chemical  tests  during
            preoperational and operational monitoring (37).
                                                                     Day 1   Day 2   Day 3   Day 4   Day 5    Day 6    Day 7
(x)
A
A
®
A
A
®
A
A
0
A
                                                                   Key:     x - Chemistry Samples
                                                                                Suspended Solids
                                                                                Sperm Cell Test

                                                                          (x) - Same as Above With 24 Hour Data Turnaround.


                                                                          fxl - Same as Above With Data Available on the First
                                                                          —   Day of the Next  Operation.


                                                                          A . Water Samples for Seven-Day Chronic Toxicity
                                                                          *-*   Tests.

                                                                          A - Water Samples for Seven-Day Chronic Toxicity
                                                                          —•   Tests 'Plus Champia.
                                                                  Figure 5-6.   Sampling scheme for a generic 7-day sampling
                                                                               period during the New Bedford  Harbor pilot
                                                                               Study (38)
Q.

m
                                                                         100 -
                                                                          80-
     60-
                                                                          40 -
                                                                          20-
                                                                                   NBH-2
                        NBH-3    NBH-4

                        Station Location
                                                                                                               NBH-6
                                                                   Figure 5-7.   PCB concentrations  in mussel tissues over  time
                                                                               versus total  PCB  concentration as  determined
                                                                               from receiving water  analysis (39).
                                                                                                                           31

-------
       15
       10
        0 •
      -10
             _*Day7
             *-* Day 28
                NBH-2    NBH-3    NBH-4    NBH-6

                          Station Location
      1.5
      0.5
               • Day 28
                NBH-2    NBH-3    NBH-4
                          Station Location
NBH-5
Figure 5-8.   Mussel scope for growth (SFG) in New Bedford
            Harbor at 7 and 28 days, mussel shell growth at 28
            days. Note that 7-day SFG tests are predictive of
            the 28-day trends in SFG and shell growth (37).
 32

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   6.     Conclusions and  Future  Directions
The Clean Water Act emphasizes the use of biological
monitoring as the basis for maintaining national water
quality standards. Inherent in the process of
environmental protection is the definition of the baseline
health of the ecosystem under scrutiny. Only when such
a baseline is established can trends be identified and
subsequently correlated with natural or anthropogenic
causes.

A Complex Effluent Toxicity Testing Program has been
well established for freshwater systems, but modifications
were necessary to adapt the methods to marine/estuarine
environments. EPA has designed state-of-the-art marine
toxicity test methods to serve as the marine counterpart
to the freshwater program, as presented in  previous
sections.

In developing the toxicity test methods, two distinct, yet
related, goals were defined: 1) to develop tests that
rapidly estimate the toxicity of effluent discharges so that
regulatory actions can proceed and 2) to enhance the
comprehensive understanding of ecological systems and
trends and thereby contribute  to long-term environmental
protection.

Complex Effluent Toxicity Testing
The Marine/Estuarine Complex Effluent Toxicity Testing
Program is currently used to help develop NPDES
permits. Test species include sheepshead minnows;
inland silversides; mysids; a sea urchin;  and two
macroalga! species, Champia  and Laminaria. These
species were selected for the  following reasons:

  • Laboratory cultures can be easily .cultivated.

  • Gametes or larvae can be obtained year round for
     shipping to onsite locations.

  • Gametes or larvae require small volumes of
     exposure water.

  • Tests are inexpensive to conduct and can be easily
     taught.

Future Research and Refinements to Complex
Effluent Toxicity Testing Program

To increase the range  and accuracy of the
marine/estuarine CETTP, new species from other
 phylogenetic groups and other geographic areas will be
 added to the program (specifically, from the west coast).
 In addition, future research will focus on special
 environments, including brackish (low salinity) and tropical
 waters. Biomonitoring techniques for brackish water
 would be useful for discharges at estuary headwaters
 including many areas within the  Chesapeake Bay and the
 Gulf of Mexico. Tropical species are needed to test
 effluent discharges in the Hawaiian Islands, Virgin  Islands,
 Puerto Rico, Guam, and other U.S. protectorates.  This is
 particularly important because most States do not  allow
 the introduction of nonindigenous species.

 In Situ Biomonitoring

 ERL-N has developed in situ biomonitoring methods to
 enhance and augment the short-term test methods and
 monitor long-term trends of marine discharges. The in situ
 program relies on mussels transplanted along dilution
 gradients that start near the discharge site and end in
 clean water. By correlating the contaminant levels  in the
 mussel tissues and subsequent biological effects, over
 time,  scientists can determine environmental trends near
 to and at various distances from  a discharge site.
 Additionally, a newly developed laboratory exposure
 system can help to field verify the laboratory CETTP
 tests.

 Future Research and Refinements for In Situ
 Biomonitoring

 The most successful biomonitoring tests to date have
 been  conducted at the individual organism level. At higher
 levels of biological organization, the complexities of
 biological systems increase tremendously. Monitoring at
 the cellular and subcellular levels may simplify marine
 monitoring because, rather than predicting toxic effects
 on only one species, indicators at the cellular and
 subcellular levels might predict effects across many
 species. However, before this can be done, relationships
among cellular, tissue, individual, and population effects
 must be understood, to justify extrapolation from one level
to the next and to explain the mechanisms  behind whole
organism effects. Several promising techniques are
currently being investigated to expand the marine in situ
biomonitoring program.

For example, the slipper shell snail Crepidula fornicata is
under consideration as a likely candidate for in situ
                                                                                                          33

-------
biomonitoring. C. fornicata survive well in field cages and
reproduce frequently and prolifically with easily observable
reproductive behavior. They can, therefore, be used for
transplanting and for reproductive tests, directly indicating
the onsite population level effects of a marine discharge.

Biomarkers (sublethal biochemical measures of effect)
may serve as subcellular indicators  of exposure to
toxicants. In this context, biomarkers are defined as
"molecular biological techniques that may directly link
specific chemicals or classes of chemical  compounds to
observed biological effects." Two such biomarkers are
measurements of increased incidence of sister chromatid
exchange and increased synthesis of metallothionein.

Sister chromatid exchange is the  exchange of pieces of
DNA between the two arms of a chromosome. The rate
of this exchange has been shown to increase in the
presence of genotoxic agents (28).  Even chemicals that
do not directly affect DNA (nongenotoxic agents) may still
affect the complicated mechanism of sister chromatid
exchange, and, therefore, may  change the exchange rate.
This method will be explored further as an indicator of
pollution effects on invertebrate and fish species.

Metallothionein induction is another measure of toxic
exposure. Metallothionein is an intracellular metal-binding
protein. In mammals and fish, the binding  action induced
by this protein may be a mechanism for tolerance to trace
metals. Synthesis of the protein is induced by the
presence of metals such as copper and cadmium. This
and other "stress proteins" show promise as indicators of
exposure to specific chemicals (28,6).

The winter flounder Pseudopleuronectes americanus is
the first teleost organism studied  in this program, and
another candidate for population-level monitoring. This
benthic fish species returns annually to its nursery
grounds for spawning, making it a suitable indicator of
adverse population  effects due to pollutant exposure. In
winter flounder, simple morphometric measures may
indicate  exposure to certain pollutants. Moore and
Stegeman (40) have found liver tumors and severe
nonneoplastic changes in the liver, and suggest that
these are the result of exposure to aromatic hydrocarbons
and chlorinated pesticides.
Integrated Approach for Toxic Discharge
Control - Integration with Large-Scale Coastal
Monitoring

The integrated approach of combining short-term CETTP
tests with in situ biomonitoring not only promotes toxic
discharge control efforts, but also can determine
environmental trends. This integrated approach, therefore,
might be integrated with large-scale coastal monitoring
systems. A variety of Federally funded, regional
monitoring systems are compiling and integrating vast
amounts of chemical and biological environmental data to
better define present environments and document trends
that occur within them.  NOAA's Statusi and Trends
Program and  EPA's Environmental Monitoring and
Assessment Program (EMAP) are both designed for this
purpose.

The Integrated Approach - Field Tested
An example of the combined use of CETTP and in situ
biomonitoring tests is the New Bedford Harbor Pilot
Dredging Project (Section  5). In the late 1970s, the
Mussel Watch Program identified the harbor as a
"hotspot" of PCB contamination.  During the pilot project,
biomonitoring was employed to monitor potentially toxic
releases from the dredging and disposal operation.

The Superfund study also  demonstrated the effectiveness
of the  real-time monitoring approach. In routine monitoring
efforts, the time gap from data collection to analysis
hinders progress, or renders the data meaningless when
operations proceed before data analysis. In the New
Bedford Harbor study, decision criteria were developed to
help managers interpret and act on the monitoring data.
Immediate decisions were  made based on measurements
that were taken less than 24 hours earlier.

Finally, the New Bedford Harbor pilot study showed the
efficiency of the testing hierarchy - from mussel tissue
residues to in situ tests to  extensive lab and field
biomonitoring. The hierarchy ensured that resources were
conserved while the harbor characterization was
completed. The case study in New Bedford Harbor stands
as a model of the integrated approach for toxic discharge
control.
 34

-------
  7.     References
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      based permit limitations for toxic pollutants:
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   (2) U.S. EPA. 1985. Technical support document for
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   (3) U.S. EPA. 1987. Water Quality Act. PL100-4.

   (4) U.S. EPA. 1989. National Pollutant Discharge
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   (5) Mount, D.I., N. Thomas,  T. Norberg, M. Barbour,
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   (6) Phelps, D.K. 1988. Marine/Estuarine biomonitoring:
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   (7) Peltier, W.H. and C.I. Weber, eds.  1985. Methods
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   (9) Schimmel, S.C., ed. 1987. Users' guide to the
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(10)   Steele, R.L and G.B. Thursby. 1987. Draft
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(12)  Breder,  Charles M., Jr.  1948. Field Book of Marine
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(13)  Perricone, C. 1980. In: Symposium on Research
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(14)  Brandsma, M.G., L.R. Davis, R.C. Ayers, Jr., and
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(15)  ASTM. 1987. Standard guide for conducting life
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(16)  Grothe,  D.R. and R.A. Kimerle. 1985. Inter- and
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(17)  Morrison, G.E., E. Torello, R. Walsh, A. Kuhn, R.
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(19)  U.S.  EPA. 1987. Permit writer's guide to water
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                                                                                                          35

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(20)   U.S. EPA. 1985. Initial mixing characteristics of
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(21)   Abbott, R.T. 1974. American Seashells: The
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(22)   Phelps, O.K. and W.B. Galloway. 1980. A report on
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(24)   Nelson, W.Q., D. Black,  and D. Phelps. 1985.
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(25)   Conover, R.J. 1966. Assimilation of organic matter
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(27)   Nelson, W.G. 1988. Report on the comparison of
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(28)   Phelps, O.K., C.H. Katz, K.J. Scott, and B.H.
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(32)    Averett, D.E. and N.R. Francingues, Jr. 1988. A
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(33)   Allen, D.C.  and AJ.  Ikalainen. 1988. Selection and
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(34)   Otis, M. J. 1987. Pilot study of dredging and
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(35)   Otis, M.J. and D.E. Averett.  1988. Pilot study of
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(36)   U.S. EPA. 1988. Decision criteria: New Bedford
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(37)   U.S. EPA. 1988. New Bedford Harbor Pilot  Study;
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      Prepared by U.S. EPA Environmental  Research
36

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      Laboratory, Narragansett, Rl, for U.S. Army Corps
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(38)   Phelps, O.K., D.J. Hansen, K.H. Scott,  and A.S.
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(39)   Nelson et al. No date. Operational monitoring, the
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(40)   Moore, M.J. and J.J.  Stegeman. 1988.
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(42)   Schimmel, S.C., G.B. Thursby, M.A. Heber,  and
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      discharge: comparison of effluent and receiving
      water  toxicity. ASTM Proceedings, (in press)

(43)   Dettmann, E.H., J.F. Paul, J.S. Rosen, and C.J.
      Strobel. 1988. Transport, fate, and toxic effects of
      a sewage treatment plant effluent in a Rhode
      Island estuary. ERL-N Contribution No. 876.  EPA
      600/X-87/366.

(44)   U.S. EPA. 1988. Dilution study and toxicity testing,
      Bay County wastewater discharge: St. Andrew
      Bay, Panama City, Florida. U.S.  EPA
      Environmental Services Division, Athens, GA.
                                                                                                           37

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   Appendix A.
   Case  Studies
 CETTP Case Study I: Fernandia Beach, Florida

 Toxicity testing of a complex effluent discharged into the
 Amelia River north of Jacksonville, Florida, was a
 cooperative effort between EPA Region IV (Athens, GA);
 the EPA Environmental Research Laboratory 'in Duluth,
 Minnesota (ERL-D); and ERL-N. The focus of  the study
 was a point-source discharge from a pulp and  paper mill
 in Fernandia Beach, Florida. The mill discharged
 approximately 16 million gallons (60  million liters) of
 effluent each day into the Amelia River with the outgoing
 tide. The case study was a week-long analysis of the
 effluent and its effects on the surrounding ecosystem.
 The Amelia River, a high-salinity tidal basin, proved to be
 a good location for a field evaluation of the
 Marine/Estuarine CETTP test  methods. The goals of this
 study were:

   • To determine the absolute toxicity of the  pulp and
     paper mill effluent discharged at the site.

   • To determine the relationship between effluent and
     receiving water toxicity.

   • To generate data for deriving an NPDES  discharge
     permit for the mill.

 Dye studies, toxicity tests, and chemical analysis of the
 effluent were used to determine the source of the
 observed effects of the pulp-mill effluent.

 Effluent Tests

 Effluent samples were collected daily from the  pulp mill's
 effluent lagoon adjacent to the discharge point. The
 samples were composited over a 24-hour period and held
 on ice until they were used in  the toxicity tests. The
 following test methods were performed using the
 procedures specified in the EPA methods manual (8):

   • A 7-day red macroalga (Champia parvula)
     reproductive test.

   • A 7-day reproductive, growth, and survival test with
     the mysid Mysidopsis bahia.

   • Tests using the inland silverside (Menidia beryllina)
     and sheepshead minnow (Cyprinodon variegatus) to
     measure larval fish growth and survival (41).

The effluent was toxic to all test species at concentrations
higher than 32 percent; however, the receiving water
concentrations of effluent in the river did not exceed 2
 percent. The red macroalga and mysids were most
 sensitive to the effluent; the NOEC for the alga was 1
 percent effluent, and the mysid's NOEC was 3.2 percent
 effluent. Fish larval growth was not significantly affected
 at any effluent dilutions. Survival, however, was  reduced
 at 32 percent effluent for the sheepshead minnow, and at
 10 percent for the  inland silverside (see Table A-1).

 Ambient Receiving Water Tests

 Water samples were collected daily from 7 sampling
 stations in the river (except 1 day, when 21 stations were
 sampled; Figure A-1) during ebb tide, to coincide with the
 discharge. The following toxicity test methods were
 performed on the receiving waters (8):

   • 7- to 9-day reproductive tests (performed on all 21
     samples) with the red macroalga Champia  parvula.

   • A 7-day reproduction, growth, and survival  test with
     the mysid (Mysidopsis bahia).

   • A 7-day growth and survival test with the
     sheepshead minnow  (Cyprinodon variegatus).

 Using dye studies performed by EPA Region  IV,
 estimates were made of the effluent concentrations in the
 receiving water samples collected (see Table A-  2). The
 receiving water concentrations of effluent in the river did
 not exceed 2 percent effluent, as was predicted  by the
 dye studies. Only the red macroalga and mysids indicated
 toxicity at several monitoring stations. The alga was
 especially sensitive to several receiving  water stations,
 and cystocarp production was reduced to zero or slightly
 above  zero (indicating that the species was incapable of
 reproduction). The  mysid receiving water tests were
 inconclusive because receiving water toxicity  could not be
 correlated with estimated effluent toxicity based on the
 dye study. Finally, the sheepshead minnow showed no
 evidence of toxicity at concentrations as high  as  10
 percent effluent. The macroalga and sheepshead minnow
 tests showed a correlation  between laboratory and field
 results (42).

 Probable Causes  of Toxicity

 Using fractionation  methods, EPA scientists at ERL-
 Duluth  and Region  IV were able to identify ammonia as
the toxic constituent in the  discharge.  The freshwater
crustacean  Ceriodaphnia dubia showed  toxic  response to
                                                                                                           39

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          Table A-1.   Effects of Pulp Mill Effluent on Test Species (41)
Percent
Effluent
Control
0.5
1.0
2.5
3.2
5.0
10.0
32
50
Red Algae
Mean No.
Cystooarps
11+21
12 ± 3
13 ± 1
0.2'

0*
0*


Mysid
(%) Survival
100

100

88

41*
0*
0*
Inland silverside
(%) Survival
87

80

58"

33*
0*
0*
Sheepshead minnow
(%) Survival
98

98



93
11*
0*
           "Significantly different from control.
          "High variability between three replicates, but not significantly different from the control.
the ammonia in the effluent. Other tests, using the red
macroalga in effluent and ammonia mixtures containing^
the same concentration of the chemical, produced a very
similar toxic response (see Figure A-2). The calculated
concentrations of un-ionized ammonia (NHp) in the
effluent, along with the results from unpublished
laboratory studies also point to ammonia as the cause of
toxicity to the mysid and inland silverside (see Table A-3)
(42). Ammonia also exhibits a characteristic concentration
curve as ionization is affected by pH, temperature, and
salinity.

Setting Permit Limits
Numerical, nonbinding permit limits were calculated using
acute end-of-pipe toxicity data from the discharger's
records, along with the NOECs determined during the
study. Since the scientists were able to trace the toxicity
to a specific chemical, and the toxicity was found at the
very edge of the mixing zone set by the State of Florida,
it was relatively simple to direct the discharger on how to
reduce the effluent's toxicity. The State of Florida granted
the pulp and paper mill a 3-year variance to the water
quality standard, allowing them time to bring the effluent
within permit limits. The EPA Region IV office will follow
up on the toxicity testing  at the end of the 3-year waiver.

CETTP Case Study II: East Greenwich, Rhode
island
Greenwich Cove is an estuarine cove located in
Narragansett Bay  between East Greenwich, Rhode Island,
and Goddard State Park  in Rhode Island (see Figure A-3).
A Publicly Owned Treatment Works (POTW) in East
Greenwich discharges^an average of 0.7 million gallons
per day (MGD) into the cove. At the time of the first
phase of this case study, the POTW was designed to
treat  only 0.5 MGD. The Town of East Greenwich is
currently upgrading the plant, and after modification, the
plant will be capable of providing advanced secondary
treatment for 1.2 MGD of sewage. A variety of
marine/estuarine CETTP tests were used to determine
the effluent and receiving water toxicity for the cove
during four field studies performed over a 1-year time
period (December 1985, July, September,  and November
1986).

After the POTW is upgraded, additional toxicity tests will
be conducted, providing a "before and after" picture of
the effluent's toxicity and its effects on the cove.

Chemical and Physical Effluent Characterization

Chemical Characterization. The POTW effluent was
analyzed for dissolved and particulate trace metals
(cadmium, chromium, copper, iron, lead, manganese,
nickel, zinc);  organic compounds such as PCBs and
PAHs; and nutrients (e.g., ammonia,  nitrogen, and
phosphorous). Total suspended solids and total residual
chlorine also  were measured. Effluent samples were
collected for fractionation studies to determine what
class(es) of compounds caused the observed toxicity. As
part of the fractionation procedure, toxicity tests were
conducted. These tests showed that  chlorinated effluent
was more toxic than  unchlorinated, and that the  majority
of the toxicity could be attributed to a pompound in the
dissolved portion rather than one associated with the
particulates. When further separation of the effluent was
performed, it was determined that the toxicity was
probably due to moderately polar constituents (43).

Physical Characterization. Mixing in the cove was
determined by dye study techniques. The  dye studies
also provided information on the age  of the effluent at the
different sampling stations in the cove (see Figure A-4).
At the center of the boil area (the area above the
submerged outfall, or where the effluent actually enters
the water from the discharge pipe) effluent concentration
was measured at as high as 40 percent. The
concentration dropped to less than 1  percent, however, at
all other sampling stations. Old effluent (effluent remaining
 40

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 Figure A-1.   Amelia River, in Fernandina Beach, Florida (42).

in the cove after one tidal cycle) and new were found to
coexist in the cove - the mixture near the outfall was
primarily fresh effluent (Station 2); older effluent was
found near the mouth of the cove (Stations 4 and 5); and
the oldest effluent was found at the head, or
southernmost extent of the cove (Station 1). Figure A-5
shows the mixing characteristics as determined
studies.
using dye
No permanent (more than 2-day) stratification in
temperature, salinity, or dye concentrations was detected
in the study, although a salinity gradient was  measured
that showed increased salinity from the interior of the
                                                                                                               41

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     Table A-2.
Effect of Receiving Waters from the Amelia River, Jacksonville, Florida, on Test Species.  Effluent percents
were calculated based on a dye study (42).
                                                 Red Algae
                                            Mean No. of Cystocarps
                                                           Mysid
Station
Number
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
Percent
Effluent
2.0
control (0%)
- (no data)
1.5
-
0.05
1.4
0.08
1.4
0.5
0.2
0.9
0.7
0.5
0.5
-
0.6
0.5
0.4
0.4
0.5
May 17 May 18
0.5" 0.8*
14.7 19.0
1.0* 3.0*
1.0.4
2.3* 0.6*
1.1*
3.4
8.3 21.0
1.1*
15.2 13.5
12.8 19.9

7.2
5.3*
5.2*
7.9
11.7
9.7
10.4
10.2
9.6
(%) Females
(%) Survival with Eggs
90 64
93 71
88 77

98 47


93 14*"

93 66
93 31










Sheepshead Minnow
(5) Survival
98
98
100

100
(

95

100
' 100










      "Significantly different from control.
     "Significantly different from control, but not attributed to effluent toxicity.


cove (where the freshwater Maskerchugg River
contributes to the cove) to the outer cove, toward
Greenwich Bay. Table A-4 shows the flow rates in the
cove. Tidal range and currents, as well as the discharge
flow rate of both the POTW and the Maskerchugg River,
all affect the mixing and stratification characteristics in the
cove (43).

Complex Effluent Toxicity Tests
The chlorinated effluent samples were diluted to effluent
concentrations ranging from 32 to 3.2 percent with
relatively clean Narragansett Bay seawater, and the
salinity was adjusted to a standard of 30 parts per
thousand with Narragansett Bay  brine (8). The following
tests were performed on the effluent samples:

   • Sperm cell toxicity tests with the sea urchin Arbacia
     punctulata.

   • Reproductive effect tests on the red macroalga
     Champia parvula.

   • 7-day reproduction, survival, and growth tests using
     Mysidopsis bahia, the mysid.
                                             •  Tests with sheepshead minnows (Cyprinidon
                                                variegatus) and the inland silverside (Menidia
                                                beryllina) to indicate effects on fish  larvae growth
                                                and survival.

                                           The effluent toxicity varied from day to day. Averaging the
                                           toxicity over all of the test periods, a concentration of
                                           0.24 percent of the effluent affected sea  urchin
                                           reproductive capacity, while greater than 10 percent
                                           effluent concentration was toxic to the fish species tested
                                           (see Table A-5).

                                           Ambient Receiving Water Tests

                                           The scientists used the same methods to test the toxicity
                                           of the effluent as were used in the complex effluent
                                           analysis, and an additional test was performed, the kelp
                                           reproduction test. This test uses the sea kelp Laminaria
                                           saccharina (see Section 2) to evaluate the reproductive
                                           effects of the receiving  water.

                                           All samples from the boil station showed similar effects on
                                           the test species. The other stations gave variable results.
                                           Percent effluent at each station was determined using dye
                                           studies (Table A-6). The species' toxic responses differed
                                           from one test period to  the next, indicating a highly
 42

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       120 —i
                                   •  NH4CI

                                   D  Effluent Dilution
                          2345
                          % Effluent

 Figure A-2.  Number of cystocarps (as percent of the number
            on control-exposed red microalga) versus percent
            effluent. The effluent data are averaged over 2
            days. A stock solution of 26.4 mg NH4CI/100 mL
            was used to match the effluent with an assumed
            concentration of 70 mg NH4-N/L of effluent (42).

 variable effluent. The red macroalga Champia and the
 sea urchin were most sensitive to the contaminated
 receiving waters, (see Table  A-7).

 Conclusion

 The East Greenwich Cove Marine CETTP was generally
, successful in  its attempt to field-validate the laboratory
 effluent toxicity tests. The investigators  found that effluent
 "residence time" and toxicity half-life (or rate of toxicity
 reduction) are important factors  to build into estimations
, of lab to field  results. In this case, the variable receiving
 water toxicity  could be explained by the variability in the
 effluent coming from the POTW, and by toxicity reduction
 or degradation with time. Although the toxicity could not
 be traced to a specific chemical, it was  concluded that
 the total toxicity observed in the field generally could be
 attributed to the POTW discharge, although occasional
 toxicity observed in the more sensitive tests distant from
 the discharge could be from other sources (e.g., boats
 moored in the cove, runoff, and  contaminants in  the
 Maskerchugg  River, etc.).

 CETTP Case Study 111: Panama City, Florida

. This study in Saint Andrew Bay (see Figure A-6) focused
 on the effluent from the Bay County Wastewater
 Treatment Plant. The plant discharges about 30 million
 gallons of waste per day into  the bay. Scientists from
 EPA Region IV (Athens, Georgia) used dye studies,
 physical and chemical characterization of the effluent, and
 CETTP testing methods to determine the discharge's
 effect on the bay's ecosystem.

 A 25-hour dye tracer study was conducted to establish
 the mixing and dilution characteristics of the effluent. The
 release was continuous with the discharge and spanned
 one tidal  cycle. This initial measurement provided
 concentrations for the receiving water tests, and further
 measurements over five additional tidal cycles provided
 information on the effluent buildup in the bay. Table A-8
 shows the dilutions in the bay at the time of the study.
 Figure A-7 shows the dilutions averaged over six tidal
 cycles. The scientists concluded that those areas east of
 the boil receive effluent during flood tides, while those
 west of the boil receive maximum exposure during ebb
 tide (44).  They also found that the effluent's concentration
 was maximized in the upper 1 to 3 meters of the water
 column because  of its lower density.

 Effluent Characterization and Toxicity Testing

 Measures of biological oxygen demand  (BOD5), total
 suspended  solids (TSS), and fecal coliform counts were
 performed on the discharged wastewater, and the results
 were within the POTW's reported values.

 The toxicity of the freshwater effluent was determined
 using a larval survival and growth test with the freshwater
 species Pimephales promelas, or fathead minnow, and a
 survival and reproduction test using Ceriodaphnia dubia,
 a freshwater crustacean. The effluent was made saline
 (using commercial salt mix or hypersaline brine) for the
 following marine organism tests:

   • Static acute toxicity test using the mysid Mysidopsis
     bahia.

   • Sperm cell toxicity test with the sea urchin Arbacia
     punctulata.

   • Reproduction and toxicity test using the red
     macroalga Champia parvula.

 The results of these tests are summarized in Table A-9.
 The freshwater tests with 100 percent effluent showed no
 effect on daphnid or larval fathead minnow survival and
 growth. Reproduction in  the daphnid was affected,
 however, at 100 percent effluent. The No-Observed-
 Effect Concentration  (NOEC) in this case was 30 percent.
The effluent was acutely toxic to the mysids, with a LC50
concentration of 70.7 percent.  Sea urchin fertilization was
unaffected at 70 percent effluent, the highest
concentration tested. The NOEC for the red macroalgal
reproduction was  12 percent effluent.

Receiving Water Toxicity Tests

Receiving  water samples were collected in June and July,
1988. In June, four samples were collected at each
station (boil, 200E, and 200W)  in a consecutive  ebb,
                                                                                                             43

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     Table A-3.   Toxicity Comparisons of Calculated Un-ionized Ammonia in Effluent (Average Concentration was Calculated
                from Total Ammonia, which was 3.66 mg/L) and Separate Ammonia Tests (42)
Effluent Tests
Species
Champ/a parvula
(red macroalga)
Mysidopsis bahia
(mysid)
Menidia beryllina
(inland silverside)
Lowest Effect Concentration
(% Effluent)
2.5
3.2
10.0
Calculated Chronic Value"
for Un-ionized Ammonia (NH),
(mg/L)
0.06
0.07
0.20
Ammonia
Tests-
Chronic
Value
(mg/L)
0.04
0.12
0.06
     •Chronic value is the geometric mean of the highest no-effect concentration and the lowest effect concentration for 2-day tests (C.         ;
       parvula) and 7-day tests (M. bahia and M. beryllina).                                             ,
     -Unpublished ERL-N data; chronic numbers refer to calculated un-ionized ammonia concentrations from 2-day M. bahia tests, and 28-
       day M. beryilina tests.

                                                         A survey  of the  effluent's effects was performed on the
                                                         bay's seagrass community. That study is described in
                                                         Dilution Study and Toxicity Testing, Bay County
                                                         Wastewater Discharge (44).

                                                         In  Situ Biomonitoring Case Study: East
                                                         Greenwich, Rhode  Island
                                                         An in situ biomonitoring study was conducted in
                                                         Greenwich Cove in Greenwich Bay,  Rhode Island, during
                                                         October and November, 1986, to compare the results of
                                                         scope for growth tests on mussels (Mytilus edulis)
                                                         conducted over long and short time periods. The short-
                                                         term test (7 days) was conducted within the time period
                                                         of the long-term, 30-day test.  The observed toxicity could
                                                         be attributed to  the East Greenwich  POTW.

                                                         Method
                                                         Three mussel deployment stations were located around
                                                         the cove: one was in the cove near the outfall (Station 3);
                                                         the second was in the mixing  zone between the cove and
                                                         Greenwich Bay (Station 6); and the third was at the
                                                         mouth of Greenwich Bay (Station 10; see Figure A-10). A
                                                         reference station (REF) was chosen at the mouth of
                                                         Narragansett Bay. As described in Section 3,
                                                         polypropylene baskets containing mussels were anchored
                                                         at  each station, suspended 1  meter  above the bottom.
                                                         Clearance rate, respiration rate, and food assimilation
                                                         efficiency were  measured as described in Section 3 of
                                                         this document.  Average scope for growth was calculated
                                                         for each  group  of mussels, and the 7- and 30-day values
                                                         were compared for each station (27).

                                                         Results
                                                         There was a graded response in scope for growth of the
                                                         mussels  exposed for both 7 days and 30 days. Table A-
                                                         12 shows the SFG and individual parameter results at
                                                         each station, and compares the 7-day and 30-day
                                                         exposure times. The SFG of the mussels at the reference
Greenwich Cove in winter (ice in foreground).
slack low, flood, and slack high tide. Grab samples were
collected near the boil, Shell Island, and station 1E for the
July test period (Figure A-8). The same marine species
tests were performed as for the effluent tests. ERL-N
assisted Region IV, performing the sea urchin and algal
tests in the Narragansett lab. At the site, dissolved
oxygen, salinity, and temperature measurements were
taken, as well as light transmission readings. Figure A-9
shows the light transmission at four stations; Table A-10
gives the salinity, temperature, and dissolved oxygen
measurements for the same time period.

The mysids showed no response to the receiving water at
any of the stations. The red macroalga reproductive tests
were inconclusive; the varied results suggested that the
St. Andrew Bay waters are in themselves unable to
support Champia parvula reproduction (see Table A-11).
44

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                                    Maskerchugg
                         0  0.5  1.0   River
Figure A-3.  Greenwich Cove, Rhode Island (43).

station was almost unchanged from 7 to 30 days. SFG at
the remaining stations,  however, decreased moving
toward the East Greenwich POTW outfall with both time
periods. This trend in SFG was more pronounced after 30
days of exposure (27).  The more dramatic decrease
observed after 30 days may indicate continued
physiological decline with prolonged exposure to the
effluent, but since actual effluent exposure was not
measured concurrently, the physiological decline cannot
be tied conclusively to the effluent's presence in the
cove. Tissue residue analysis is being conducted, and will
provide more information about the conditions of the cove
waters during the exposure period.

The results of this study imply that the effects of 7-day
exposure are indicative of longer-term effects, and can be
used to predict biological response to effluent toxicity.
                                                                                                             45

-------
                                  East Greenwich POTW, viewed from Greenwich Cove (December
                                  1985).
                      41°40'N -
                   41 °38'45"N
                                     Boil Area
                                 East
                                 Greenwich
                                 POTW
Arbitrary Definition
of Greenwich Cove
Boundary
                                                                                500m
                          71°27'30"W                 71°26'30"W


Figure A-4.  Sampling stations in Greenwich Cove (43).
       71 °25'30"W
 46

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     December 9
       Low Tide
        (1100)
  December 9
Low Tide + 4 Hr
    (1500)
December 10
  Low Tide
   (1155)
     December 11
       Low Tide
        (1248)
                                     December 13
                                       Low Tide
                                        (1430)
Figure A-5. Dye distribution in Greenwich Cove during December,
           per billion (ppb) (43).
                  985. Numbers represent dye concentration isopleths in parts
                                                                                                                        47

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                                Table A-4.   Flow Rates into Greenwich Cove (43)

                                                                Flow Rate
Source
POTW
Maskerchugg R.
Mean Tidal Flow
(m3/day)
2,650
1,420 to 138,000
2,300,000
(million gallons/day)
0.70
0.38 to 36.46
607.70
                                Table A-5.   Rank Order of Species Sensitivity to Greenwich
                                            Cove POTW Effluent (43)
Study
Period
12/85




7/86



9/86



11/86



Species
Arbacia punctulata (sea urchin)
Mysidopsis bahia (mysid)
Champia parvula (red algae)
Cyprinodon variegatus (sheepshead)
Menidia beryll'tna (inland silverside)
Arbacia punctulata (sea urchin)
Mysidopsis bahia (mysid)
Champia parvula (red algae)
Menidia beryllina (inland silverside)
Arbacia punctulata (sea urchin)
Champia parvula (red algae)
Mysidopsis bahia (mysid)
Menidia beryllina (inland silverside)
Arbacia punctulata (sea urchin)
Champia parvula (red algae)
Mysidopsis bahia (mysid)
Menidia beryllina (inland silverside)
NOEC"
(% Effluent)
1.0
1.0
3.0
10.0
10.0
<0.3
3.2
5.4
10.0
<0.24
4.25
10.0
10.0
<0.6
2.5
3.2
10.0
                                "NOEC = No-Observed-Effect Concentration.
        Table A-6.    Mean Percent Effluent from  the  East  Greenwich POTW  Found in Surface Water Samples from
                     Greenwich Cove and Greenwich Bay, Rhode Island During Three 1986 Sampling Periods: July 16 to 22,
                     September 8 to 15, and November 9 to 18. All sampling was performed at slack low tide (43)

                                                         Receiving Water Stations                      '
Date
July
September
November
Overall
MEAN
Overall
MAXIMUM
Overall
MINIMUM
1
0.16
0.13
0.23
0.17
0.33
0.031
2
0.15
0.15
0.28
0.18
0.73 ,
0.010
outfall
26
29
17
24
40
6.3
3
0.28
0.30
0.38
0.33
0.66
0.082
4
0.35
0.30
0.24
0.30
0.68
0.14
6
0.12
0.12
0.27
0.16
0.86
0.039
7
0.064
; 0.054
0.083
0.063
0.14
0.013
48

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                                                     Bay Count'
                                                     Aerated Lagoon
Figure A-6.    St. Andrew Bay, Panama City, Florida (44).
                                                                                                                        49

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    Table A-7   Results of Toxicity Tests Conducted on Surface Waters from Greenwich Cove and Greenwich Bay,  Rhode
                Island, During Three 1986 Sampling Periods: July 16 to 22, September 9 to 15, and November 11 to 18. Toxicity
                is expressed in the numerator of each fraction as the number of occasions that the sample collected  from a
                specific station was significantly different from the control. The denominator denotes the number ol receiving
                water samples evaluated. Overall toxicity is the total  number toxic of all samples collected from the three test
                periods; the number in parentheses denotes the percent samples that were toxic of all tested (43).
         Species/Date
                                                            Receiving Water Stations
                                                       outfall
C. parvula
(red algae)
July
September
November
Overall Toxicity
A. punctulata
(sea urchin)
July
September
November
Overall Toxicity
M. bahia
(mysid)
July
September
November
Overall Toxicity
M. beryllina
(inland silverstde)
July
September
November
Overall Toxicity

0/6
0/5
1/1
1/12(8%)

1/7
4/6
2/6
7/19 (39%)


0/1
0/1
0/1
0/3 (0%)

0/1
0/1
1/1
1/3 (33%)

0/6
0/5
1/1
1/12 (8%)

5/7 .
3/6
1/6
9/19 (47%)


0/1
0/1
0/1
0/3 (0%)

0/1
0/1
0/1
0/3 (0%)

6/6
5/5
1/1
12/12 (100%)

7/7
6/6
6/6
19/19 (100%)


1/1
1/1
1/1
3/3 (100%)

1/1
1/1
1/1
3/3 (100%)

0/6
0/5
1/1
1/12 (8%)

3/7
3/6
2/6
8/19 (42%)


0/1
0/1
0/1
0/3(0%)

0/1
0/1
0/1
0/3(0%)

1/6
0/5
1/1
2/12(17%)

4/7
1/6
2/6
7/19 (37%)


0/1
0/1
0/1
0/3 (0%)

0/1
0/1
0/1
0/3 (0%)

1/6
0/5
0/0
1/12 (8%)

3/7
1/6
2/6
6/19 (32%)


0/1
0/1
0/1 .
0/3 (0%)

0/1 ;
0/1 ;
0/1
0/3 (0%)

1/6
0/5
1/1
2/12 (17%)

4/7
1/6
2/6
8/1 9 (42%)


0/1
0/1
0/1
0/3 (0%)

0/1
0/1
0/1
0/3 (0%)
50

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    Table A-8.   Dilution Summary, St. Andrew Bay, Florida, June,
                                         Low Tide
988 (49)
                                                                                High Tide
Station
10E
8E
6E
4E
2E
200E
Boil
200W
2W
4W
6W
8W
10W
12W
4S
Ultimate Tracer
Cone."
0.9
1.3
2.1
2.1
2.5
10.8
45.5
15.0
4.3
4.0
4.1
3.2
2.7
0.8
1.4
Wastewater
Dilution
800:1
550:1
340:1
340:1
285:1
66:1
16:1
48:1
165:1
180:1
175:1
225:1
265:1
900:1
515:1
Ultimate Tracer
Cone."
0.6
2.0
2.1
2.2
5.3
18.3
40.0
16:2
3.4
2.2
1.4
0.9
0.2
0.6
0.2
Wastewater
Dilution
1200:1
360:1
340:1
325:1
135:1
39:1
18:1
44:1
210:1
325:1
515:1
800:1
3600:1
1200:1
3600:1
— Average
Conditions
Wastewater Dilution
1000:1
455:1
340:1
335:1
210:1
53:1
17:1
46:1
190:1
255:1
345:1
510:1
1930:1
1050:1
2050:1
     *ppb
                 12W
                 11050:1
                                       North
Figure A-7.   Steady-state dilution, average conditions, June 1988 (49)
                                                                                                                             51

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    Table A-9.   Toxicity of Effluent from Bay County Wastewater Treatment Plant, Panama City, Florida (44)

                                   Daphnid
Percent
Effluent
0*
1
3
6
10
12
25
30
50
70
100
(%)
Survival
100
100
100

100


100


90
Mean No.
Offspring/Female
26.47
17.63
28.60

30.43


28.83


5.00*
Mysid (%)
Survival1
100
100


100

80

80

20*
Fathead
minnow (%) Sea urchin (%)
Survival Fertilization
100 95.5
100
100

100


100 !

89.6 i
100
Red Algae
Mean No.
Cystocarps
9.0


6.6

5.4*
2.4*

1.0*

0*
    'The calculated UC50 for Mysidopsis bahia was 70.7 percent.
    ^Control sample collected from Narragansett Bay.
    "Significantly different from control.
                                                                                                         North
                             Shell Island
Figure A-8.   St. Andrew Bay receiving water sampling stations (44).
 52

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              100
        H-

        o>
                                           8    10    12     14
                                                                   16    18    20    22    24

                                                            Dep'th (Feet)
                26    28
30
                                      __.__. Boi|        ._....(.— 2QOW


Figure A-9.  Light transmission, as measured by marine photometer (44).
 200E
                                      Table A-10.   Salinity, Terr perature,  and  Dissolved
                                                   Oxygen  at  the Boil, 200E,  and  200W
                                                   Stations in June, 1988 (44)

Boil
Depth
(ft.)
1
5
9
15
19
25
Sal.
(PFj>t)
32
32
32
33
34
35
3
9
8
6
9
0
Temp.
(•C)
26.9
27.6
27.2
26.6
26.8
26.6
D.O.
(ppm)
6.0
6.2
6.5
6.4
6.5
6.5
B-200E 1
5
9
15
19

B-200W 1
5
9
15
19
25

32
32
33
34
35

32
9
9
5
6
2

5
32.5
33.B
334
34.
34.

1
4

28.0
27.2
27.3
27.0
26.4
Avg. =
28.1
28.0
27.4
27.0
26.4
26.4
Avg. =
6.2
6.0
6:3
6.3
6.1
6.2
6.9
7.0
6.9
6.8
6.3
6.4
6.6
                                                                      Avg. =
6.4
                                                                                                                           53

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Table A-11.  Results of Mysidopsis bahia  (mysid), Arbacia punctulata  (see urchin)
             Champia parvula (red algae) Tests on  F^jceiying Wajers^rn St. ^ngrej

       Sample           Sample        Sample         (%)           (%)        Mean No.
         Date            Location        Time         Survival      Fertilization     Cystocarps
                                                                                                        and
6/14-15/88



7/15/88



6/15/88







7/19/88



Boil
SOW
100W
200W
Boil
SOW
50E
100E
Boil
50E
100E
200E
Boil
100W
200W
Control
Boil
Shell I.
1E
Control
1130 100
1325 100
1220 100
1305 90
2050
2110
2145
2130
0800
0815
0830
0840
1100
1110
1125
Treatment1
1321
1307
1347
Treatment1
92.8
89.4
87.9
88.8
81.3
86.9
84.5
86.5
86.0
82.8
93.5*
89.9
87.7
83.2
86.2
95.5
96.0
94.7
93.4
97.4
2.9
1.7*
4.5*
2.7











13.9 '
6.8
1.1*
3.3
9.0
                   "Significant difference from the boil based on Analysis of Variance and tests.
                   'Sample collected from Narragansett Bay.
                                       Figure A-10. Greenwich Cove, Rhode Island, with monitoring
                                                   stations (27).
54

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Table A-12.  Results of the Physiological Measurements and Scope for Growth Index
            of Mussels from 7- and 30-Day Exposure (Values in Parenthesis are
            Standard Errors) (27)

                                       7-Day Exposure
Station
Ref
10
6
3
Station
Ref
10
6
3
Clearance
Rate (L/h)
5.11 (0.22) AB
6.07 (0.23) A
4.48 (0.28) B
4.30 (0.39) B

Clearance
Rate (L/h)
5.88 (0.58) A
5.02 (0.46) AB
4.41 (0.42) AB
3.05 (0.54) B
Absorption
Efficiently %
85 (ik A
91 (0) B
89 (2)
86 (0)

Absorf
Efficien
84(5
95(0
96(1
89(1)
AB
AB
30-Day
ition
:y%
A
B
B
AB
Respiration
Rate (mL-02/h)
0.29 (0.03) A
0.48 (0.04) B
0.50 (0.02) B
0.50 (0.03) B
Exposure
Respiration
Rate (mL-02/h)
0.39 (0.04) A
0.53 (0.02) AB
0.68 (0.08) B
0.55 (0.06) AB
Scope for
Growth (J/h)
17.7(1.1) A
17.7 (0.9) A
13.0 (1.7) AB
11.4(1.3)6

Scope for
Growth (J/h)
17.2(2.4) A
15.0 (1.7) AB
10.7 (2.0) AB
6.5 (3.2) B
                                                                                                 55

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   Appendix B
   Glossary
 Absolute Toxicity. Laboratory-determined toxicity
 (usually determined using CETTP methods) of a complex
 effluent without comparison to receiving water toxicity.

 Acute. Occurring within a short period of time. In toxicity
 tests, a response observed within 96 hours or less
 typically is considered an acute  effect.

 Acute to  Chronic Ratio (ACR). The ratio of the acute
 toxicity of a substance (expressed as an LC50) to its
 chronic toxicity (expressed as a NOEC, LOEC, or EC50).
 ACRs can be used to estimate chronic toxicity on the
 basis of acute toxicity data, helping in the determination
 of NPDES permit limits.

 Ambient Toxicity. Toxicity produced by a sample of
 water taken from a larger water  body.

 Benthic Zone. The ocean floor.

 Bioaccumulation. The uptake and retention of
 substances by an organism from its surrounding  media
 and food.

 Bioavailability. The degree to which a substance is or
 becomes available to the tissues after exposure.  A
 substance's bioavailability determines  its effect on
 exposed organisms.

 Biochemical Oxygen Demand (BOD). The quantity of
 oxygen utilized in the biochemical oxidation of organic
 matter in a specified time and at a specified temperature.
 It is not related to the oxygen requirements in chemical
 combustion, being determined entirely by the availability
 of the material as a  biological food and by the amount of
 oxygen utilized by the microorganisms during oxidation.

 Biomarker. A molecular biological indicator that may
 directly link specific  chemicals or classes of chemicals to
 observed biological effects.

 Biomonitoring. Any method of testing that includes the
 use of aquatic plant  or animal species to measure acute
 or chronic toxicity, and any biological and/or chemical
 measure of bioaccumulation.

 Biosurvey. A method for determining the variety and
numbers of species  in a biological community.

Chronic. Occurring  after a prolonged period of time. A
chronic effect is one that is observed after long-term
 (usually low-level) exposure.
 Chronic Value (ChV). The geometric mean of the
 Lowest-Observed-Effect concentration (LOEC) and the
 No-Observed-Effect Concentration (NOEC).

 Clearance Rate (CR). The volume of water that a
 mussel can completely "clean" (remove particles larger
 than 3 microns) within a specific time period.

 Cystocarp. The spore case of the red marine alga
 Champia parvula. Cystocarps are evidence of sexual
 reproduction in the algae, and their formation is used as a
 toxicity test endpoint.

 Diversity. The number of species in a specific location.

 Direct Discharge. Release of a pollutant from a pipe or
 outfall directly into a water body.

 Effective Concentration (EC). The  effluent
 concentration that has an effect on a given percentage of
 test organisms.

 Effective Concentration/50 (EC50). The  effluent
 concentration that has an effect on 50 percent of the test
 organisms.

 Effluent. Any pollutant or mixture of  pollutants that is
 discharged into surface waters or treatment works.

 Effluent Limitation. A restriction imposed by State or
 Federal authorities on the quantities,  rates, and
 concentrations of pollutants that are discharged into
 surface waters.

 Estuary. The lower end of a river where freshwater
 meets and mixes with seawater.

 Fecundity. An organism's capacity for producing
 offspring.

 Fluorometry. An instrumental method for measuring the
 amount of fluorescence in a water sample. Used during
 dye studies to estimate the concentration of an effluent in
 the receiving  water.

 Fraotionation. A method for separating and identifying
the components in a chemical mixture and each part's
effects on test organisms. For example,  an effluent may
 be gravity filtered, and  the filtered solution used in a
toxicity test.

Gas Chromatography (GC). An instrumental method for
identifying the component parts (especially volatile
                                                                                                          57

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organics) in a chemical mixture. The solvent extract of the
aqueous sample is injected into a column containing a.
stationary phase (resin) and a carrier phase (gas). The
vaporized volatile components  leach off the column
according to their relative affinity for the stationary and
carrier phases, and show peaks on a chart proportional to
their presence in the mixture.

Hypersaline. A  solution whose salinity is greater than
normal seawater.

Larva. An immature form that is unlike the adult
organism.

Lowest-Observed-Effect Concentration (LOEC). The
lowest concentration of a substance at which an effect
was observed in test organisms.

No-Observed-Effect Concentration  (NOEC). The
highest concentration of a substance at which no effect
was observed in test organisms.

Nonpoint  Source. A pollution discharge source that is
not released from a distinct source, such as stormwater
runoff that  is collected and channeled  by humans.

National Pollutant Discharge Elimination System
(NPDES).  The National program for administering  permits
for the discharge of pollutants  into  the waters of the
United States under the Clean Water Act.

Outfall. The discharge point from any discernible
conveyance for effluent discharge  (i.e., a pipe, ditch, or
channel).

Phylogenetic.  Pertaining to ancestral development.
Point Source. A single source of effluent, such as a
manufacturing facility, that discharges into surface waters.

Precision. When repeated test results do not deviate
significantly among themselves. Compare this to
accuracy, in which  the test results do not vary
significantly from a standard.

Relative Toxicity.  The toxicity of an effluent after it is
mixed in a receiving water - determined  with either
CETTP  receiving water tests or in situ biomonitoring
methods.

Scope for Growth. A measure of the energy available to
an organism for tissue production, both  somatic and    ;
reproductive, after  routine metabolic  costs are accounted
for.

Superfund. Federal authority under  the Comprehensive
Environmental Response, Compensation, and  Liability Act
(CERCLA, ratified in 1980) to respond directly to releases
or threats of releases of hazardous substances.        ;
"Superfund"  refers to the Federal funds that are available
to help  pay for hazardous waste cleanups.

Teratogenicity. A substance's ability to produce
anomalies of formation or development.

Wasteload Allocation. The portion  of a receiving water's
total maximum allowable daily pollution load that is
allocated to one of its existing or future  point sources of
pollution.

Whole  Effluent Toxicity. The aggregate toxic effect of
an effluent (usually measured directly with a toxicity test).
 58

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vvEPA

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