Municipal Environmental Research
                         Laboratory
                         Cincinnati OH 45268
  EPA 600/9-79-034
          —„-,. Delation
          and Treatability of
          Specific Pollutants
-U.
                              UAiiii
                                     UUBUBE

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 US EPA-AWBERC LIBRARY
30701 100542890

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                               EPA-600/9-79-034
                               October 1979
 Biodegradation and Treatability
        of Specific Pollutants
                    by
         Edwin F. Barth and Robert L. Bunch
           Wastewater Research Division
                 AWBtRC. LIBRARY  U.S.
MUNICIPAL ENVIRONMENTAL RESEARCH LABORATORY
     OFFICE OF RESEARCH AND DEVELOPMENT
    U.S. ENVIRONMENTAL PROTECTION AGENCY
            CINCINNATI, OHIO 45268

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                       DISCLAIMER

  This report  has been reviewed by the Municipal Environmental
Research  Laboratory, U.S.  Environmental  Protection  Agency,
and  approved  for  publication.   Mention of  trade  names  or
commercial  products  does  not  constitute  endorsement  or
recommendation for use.

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                        FOREWORD

  The Environmental  Protection Agency was created because of
increasing public  and governmental  concern about  the dangers
of pollution to the health and welfare of the  American people.
The  complexity of the environment  and the interplay between
its  components  requires  a  concentrated and  integrated  attack
on the problem.
  Research and development is that necessary first step in problem
solving and it involves defining the problem,  measuring its impact,
and searching for solutions. The Municipal Environmental Research
Laboratory develops new  and improved  technology  and systems
for the prevention, treatment, and  management of  wastewater
and solid and hazardous waste  pollutant discharges from municipal
and  community sources. We are  committed to the  preservation
and treatment  of public drinking water supplies,  and to minimizing
the  adverse economic,  social, health,  and aesthetic  effects  of
pollution. This publication  is one of the products  of MERL research
and, as such,  is a most vital  communications  link between  the
researcher and the user community.
  This report describes biodegradation  and treatability studies that
have been conducted over  a period of  20 years by the Wastewater
Research Division.
                              Francis T. Mayo
                              Director
                              Municipal Environmental Research
                              Laboratory
                             in

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                         ABSTRACT

  This  report  discusses fundamental  and technical  considerations
relating to laboratory studies of the fate and effects of organics and
metals  in biological wastewater treatment processes.
  The  relationships  between  analytical  control tests  and biode-
gradation test methods are discussed.
  Various laboratory apparatuses for both  continuous flow and
static biodegradation methods are described.
  Both aerobic and anaerobic test results are included in the report.
                               IV

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                         CONTENTS

Foreword	iii

Abstract	iv

Introduction	vi
1. Biodegradation of Organic Compounds	1
     Fundamental Considerations
     Technical Considerations for Microbial Growth
     Analytical Control for Biological Treatability Studies
     Biodegradation Test Methods

2. Treatability of Specific Compounds	 19
     Anaerobic Studies
     Aerobic Studies

3. Treatability of Metals by Activated Sludge Systems	46
     Distribution of Metals through the Process
     Effects of Metals on Aerobic Process
     Effects on Anaerobic  Digestion
     Discussion

4. Bibliography	59
     General
     Treatability of Specific Organic Compounds
     Metals

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                      INTRODUCTION

  This  compilation  of biodegradation and  treatability  studies is
an overview of work  conducted by the staff of EPA's  Municipal
Environmental Research Laboratory over the last twenty years.
  The report is not intended to be detailed, but rather discusses
approaches,   considerations,  and  techniques  evaluated  during
laboratory studies of specific pollutants.
  Several protocols are presented with  a  summary of  the type
of results obtained for each.  There is  no  intention at  this time
of selecting one procedure as the best method for all environmental
decisions.
  We intend  that the  report serve as  a review  and summary of
past  investigations and provide the  user with  a  framework  for
deciding on directions  to pursue when undertaking similar studies.
                              John J. Convery
                              Director
                              Wastewater Research Division
                              Municipal Environmental Research
                                Laboratory
                              VI

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           Section 1. Biodegradation of
                 Organic Compounds
FUNDAMENTAL CONSIDERATIONS

  Biodegradation, as  a  theoretical concept, refers to the process
by  which an organic compound is converted to carbon dioxide,
water,  and  other inorganic constituents by the action  of  living
organisms. In practice, several  qualifying terms are used to modify
this concept.
  The  operational definition  of biodegradation can be divided
into three categories:
  Ultimate Degradation: Conversion  of the parent  compound  to
 carbon  dioxide, water,  and  inorganic  compounds (if  elements
 other than  carbon, hydrogen,  and  oxygen  are present  in the
 parent compound).
  Primary Degradation: A change in  the  parent compound such
 that it  no  longer responds to the analytical  measurement used
 for detection.
  Acceptable Degradation: Conversion of  the  parent compound
 to the extent that undesirable properties are no longer manifest.
  Because of the differences  in duration,  test procedures  used,
endpoint criteria, and analytical  methods selected, the demarcation
between  categories  is  not clear-cut.  Futher  complicating  this
situation  is the fact that  during biodegradation  a certain amount
of the constituents of  the parent compound will  be assimilated
into the active biomass. Thus,  it is impossible to do mass balances
for biodegradation studies on a routine basis.
  Most of the published work  on biodegradation deals with aerobic
systems.  However, anaerobic  degradation  can  be  an  important
environmental consideration  and  procedures  for  measurement
of this process are a necessary part  of a fully developed set  of
procedures  for  observing biodegradation.  This  is  an  area  that
requires considerable developmental work.
  The main  consideration in  biodegradability  is whether or not
an organism exists that can break down,  metabolize,  or oxidize
the  parent  compound,  either  on first exposure  to  it  or after a
suitable acclimation  period. This acclimation period is important
and should be accounted for in all biodegradation studies concerned
                             1

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with environmental acceptability. Acclimation can allow organisms
to produce an induced enzyme in response to the presence of the
unfamiliar substrate; or provide time for population shifts to favor
an increase in organisms utilizing the specific substrate.
  An  academic distinction  can be  made  depending on whether a
compound is degraded in a biological system  where it is the sole
source of  carbon;  or  whether degradation  takes place  in  the
presence   of  multiple  carbon  sources (co-metabolism). From  a
practical   point  of  view,  systems  employing co-metabolism  are
more similar to  environmental situations and  yield more realistic
estimates  of  biodegradation. Co-metabolism is also  important
to the treatment of diverse industrial wastes, which are produced
when the  manufacturer uses  a variety  of starting materials and
alternative reaction pathways. If  biological  treatment  is one  of
the possible means of  disposing of the unusable or unsalable wastes
from  an   operation,  the biodegradability of these mixed  wastes
may be an important factor in the  economics of  process selection.
  The above discussion  did not include  the factors of  time and
extent of  degradation.  In   real  world situations,  the  extent  of
degradation within  a fixed time limit  is a very important  con-
sideration  in  any  working  definition  of  biodegradability.  In
principle,   laboratory  testing  should show the  biodegradability
of the material tested as it would  occur  in the environment or in
the wastewater treatment  plant. Obviously, most  substances will
eventually be stabilized by  microbial action, given enough time and
the right  conditions.  We must, therefore,  consider the  biodegra-
dation process as a  rate process.  For practical  purposes  related
to wastewater treatment, the substance under consideration should
degrade at a rate equal  to  or greater than the degradation of the
usual constituents of domestic waste,  after  the organisms  have
become acclimated to  the substance. A control  test utilizing a
compound with  known biodegradation  characteristics, such  as
linear  alkylbenzene  sulfonate,  should  be  included  in  each
experiment.
  Common experience has  shown  that if a specific assay  is  used
to monitor the extent of biodegradation, 100 percent degradation
is seldom  achieved.  This is due  to the fact that  the  organisms
cannot  derive energy  by  oxidizing  compounds  at  very   low
concentrations.

TECHNICAL CONSIDERATIONS FOR MICROBIAL GROWTH

  The proper conditions must be maintained before any compound
can  be biodegraded.  Microorganisms cannot  grow,   and  hence
organic  compounds  cannot be degraded,  unless  a  number  of
conditions are fulfilled.  All necessary nutrients and growth  factors

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must  be present in the  appropriate chemical  and physical forms.
Solutions are more readily degraded than suspensions. An adequate
supply  of  water  and  sufficient  hydrogen  acceptors such  as
molecular oxygen,  nitrate, or  carbon  dioxide must  be present.
Because many diverse species of microorganisms are involved with
environmental degradation, their  growth can occur  over  a  wide
range of temperature and pH values; however, for any  one specific
case there  may  be optimal  values. In  general, the factors listed
below should be considered in aerobic testing.

PH

  The  culture medium  should  be buffered to maintain the pH
of the system between 6.5 and  8.5. If the medium contains nitro-
genous  components and  nitrification  occurs,  alkalinity can  be
depleted. During the conversion of one part of ammonium nitrogen
to nitrate nitrogen, 7.1 parts of calcium carbonate (CaC03) alkali-
nity will be destroyed. Sodium bicarbonate is an excellent buffering
reagent for pH control.

Dissolved Oxygen

  According to the culture method selected, oxygen can be supplied
by  shaking,  diffusion  from  the  atmosphere,  or aeration  with
compressed  air.  The  concentration  of  dissolved  oxygen  in the
culture medium should not be allowed to fall  below about 1  mg/l.

Nutrient Balance

  The proximate composition of protoplasm is C106H180045 N16Pi,
therefore, to ensure that synthesis of  new cellular matter is not
hindered, the culture medium  should  have these  elements  avail-
able.  In  the  process of degrading  organics, carbon and hydrogen
are supplied  by  the parent  compound and oxygen by substrates
such  as  water and air;  however,  nitrogen  and phosphorus  must
be externally supplemented.  For each 1 mg/l of parent compound
carbon  added  to  the  culture  medium, at  least  0.2 mg/l  of
ammonium nitrogen and 0.02 mg/l of  phosphorus should be pro-
vided.  Reagent grade phosphoric  acid and ammonium hydroxide
can be used to supply these two elements.
  Trace elements such as sulfur (S),  potassium  (K), calcium  (Ca),
and  magnesium  (Mg)  are required  in such  small amounts that
in all  but the most unusual cases these will be present  in sufficient
amounts as contaminants in the carriage water, chemicals used for
medium  preparation, or inoculum.  Preformed  growth  factors
such as thiamine or biotin  are usually provided by  the addition

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of yeast extract or domestic wastewater, depending on the medium
selected.

Temperature

  Initial screening tests for biodegradation  should  be conducted
at 20°C  ±  5°C.  This allows  experiments  to  be  conducted  at
ambient laboratory temperatures  and is a reasonable  temperature
for environmental  assessment. More  information  at 30°C and
5°C can be developed if the situation warrants such detail.

Inoculum

  Microorganisms to be used as the initial inoculum can be obtained
from  a  wide  variety  of  sources, depending  upon the  infor-
mation  desired,  the  culture  medium used,  and the choice  of
culture method. For assessment of environmental biodegradation,
natural inocula rather than pure culture systems are preferred.
  Natural  inocula can be  obtained from ambient air, forest  or
garden soil, activated  sludge  or  trickling filter  biomass, natural
waters, settled  wastewater or final effluent  from biological pro-
cesses.

Toxicity

  Controls  must  be included  in   biodegradation tests to  ensure
that  viable  inoculum  was  used and that absence of degradation
is not the result of the initial toxicity of the compound.
  If all the culture conditions cited above are controlled,  a known
degradable  compound, which is  included as a  control  in a test
system,  should show  the expected degree of degradation. If not,
either the inoculum or the medium must be the source of trouble.
  Since co-metabolism studies are recommended  for environmental
degradation tests, the  medium should contain readily degradable
materials  from  yeast  extract  or  wastewater.  Therefore,  slight
microbial  growth should  be  noted even  in  systems containing
non-degradable test compounds. If this growth  is absent, it is  an
indication the test compound is toxic. In that case, the test material
should be  evaluated at lower  concentrations and serial  transfers
should be  performed to accommodate acclimation of the micro-
organisms.


Medium Selection

  Four types of media are commonly used:
  Natural waters,

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   Mineral salts plus yeast extract,
   Mineral salts plus yeast extract and supplemental organic
    materials, and
   Settled domestic wastewater.
  The first two are used mainly for determination of biodegradation
in screening  tests. Because  of  their low organic content, oxygen
transfer  problems are minimized and  cell synthesis is minimal.
This latter point is  important  since  the low cell yield precludes
concern with sorption of the test  compounds on a massive surface.
These media are usually employed  in batch tests.
  The  last two media are used  for  treatability studies. The higher
organic  content  supports luxuriant  cell  growth.  Factors  such
as sorption  to cell mass  and varying sludge age can be studied
with these media. These media can  be utilized for  either batch
or continuous flow systems.

ANALYTICAL CONTROL FOR BIOLOGICAL
TREATABILITY STUDIES

  As in any chemical reaction, the process of biodegradation can be
followed  by  observing the disappearance of the  reactants  (organic
compounds and oxygen) or the appearance of products (cell matter,
carbon  dioxide,  and  inorganic  ions). The  disappearance of some
specific organic compounds  cannot, in  many cases, be proof that
the compounds  have  been  completely degraded. Most chemical
or physical  tests depend on a  particular functional group of the
organic compound. For example,  in testing for a surfactant of the
alkylbenzene sulfonate type, several  methods of assay could  be
used.  If  the  method were based on detection of the benzene ring,
then the  benzene  ring would have to be degraded for it not to be
detected. However, if the method  were based  on the  formation
of sulfate, then the molecule  would have to be so disintegrated
that the sulfonate group would split off the compound.

Specific Analyses

  Almost any specific physical or  chemical assay can be applied
to  treatability studies providing  the medium or metabolites  do
not interfere.  In many cases  preliminary clean-up or concentration
steps may be necessary before the final analysis.
  The  possible assays  cover  the  whole range of analytical tools
such as colorimetry, optical rotation, radionuclides, ultraviolet light
and  infrared  light absorption, liquid  and gas  chromatography,
and mass spectrograph.
  In order to gauge ultimate  degradation as compared with  primary
degradation,  it is necessary  to  use some  type of analytical test

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that  measures organic  matter in  conjunction  with  specific assay.
If  a  low concentration  of  total  organics is  measured after degra-
dation,  it  indicates that  partially  degraded  end-products  have
not accumulated.
Non-Specific Analyses

  For compounds or unknown wastes for which a specific analysis
of sufficient sensitivity and applicability in the biological  system
is not available, a  non-specific  analysis must  be used. The  most
commonly  used  methods have  been the  oxygen utilization  or
biochemical oxygen demand (BOD), the chemical oxygen demand
(COD), and the total organic carbon (TOO. Unlike the methods
that depend on  the observation of the disappearance of  some
specific organic compound,  these analyses  are based  on the dis-
appearance  of oxygen and organic matter as the degradation pro-
ceeds, which is a general function of aerobic systems.
  Of  the  three methods,  only  the carbon determination (TOC)
is a  definitive  measurement. The BOD and COD tests are empi-
rical tests that give  results in  terms of the  amount of  oxygen
utilized by matter under a prescribed analytical procedure. Because
they do not indicate  the quantity  of any  particular constituent,
it is  essential  that they  be conducted according to a standard
method.
The BOD Test -

  The  BOD of a  sample is the weight of oxygen consumed  by a
varying quantity  of bacteria  in contact with the sample at a spe-
cified  temperature  during a  stated time  period. The test  does
not  directly  measure the amount  of organic  material  present.
Phrases such as "BOD  is rapidly removed" and "the conversion
of BOD to  cellular material" should not  be used  since that  type
of thinking can lead  to incorrect assumptions about the amount
of  oxygen  required  to  stabilize organic  matter  under aerobic
conditions.
  The  test does  not  differentiate matter that is biologically oxi-
dized,  from  matter that  is first synthesized into protoplasm and
then  oxidized during  the  endogenous  respiration  of  cellular
material.  The  BOD  of  a  waste will always  be  less than the
theoretical ultimate BOD because the cellular material  is relatively
resistant to  microbial  oxidation. Whether or not the  BOD  includes
the oxygen  demand used in nitrification will depend on the sample,
inoculum,  duration  of  the  test,  and addition  of a nitrification
suppressor.

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  The  main limitation of the BOD test  is the inapplicability of
test  results  to  predicting what will take place when  the  waste  is
actually oxidized in a different environment. The BOD test  is a
measure of  avidity  for oxygen under specified conditions.  It does
not necessarily  follow that these same conditions exist in a waste
treatment plant or  a  stream.  In  spite of its imperfections,  the
biochemical oxygen demand  test  is useful  in  measuring easily
oxidized wastes in samples requiring more than 3 mg/l of oxygen.

The COD Test -

  The chemical  oxygen demand is the amount of oxygen consumed
by  a compound when it  is digested  with chromic  and sulfuric
acids. With  the proper catalyst,  all but a few organic compounds
are oxidized to carbon dioxide and  water under the conditions
of the standard test. The COD test does not measure the  oxygen
required to oxidize ammonia or organic nitrogen.
  The  main limitation of  the test is  that the chemical  oxidant
reacts  with  both  stable and  unstable  organic  matter. The  test
does  not  differentiate between  inert and biologically oxidizable
matter; neither  does it provide any  indication of the  rate at which
the  biologically active  material  would  be  stabilized  under  the
conditions of waste treatment or in streams.
  The  test  has  the  advantages of requiring only  a short time for
the determination  and not being  affected by toxic wastes. Used
in conjunction  with the BOD test, it will give an  indication of the
characteristics   of  wastes.  For example,  if  the  samples contain
substances  that are  difficult for microorganisms  to  oxidize,  the
COD value will be much greater than the BOD  value.
  This procedure is used principally for  samples  that have a COD
value above 10 mg/l.  It can  also be used on samples containing
less than this  amount to indicate  an  order  of magnitude of or-
ganic content.

The TOO Test -

  The most  commonly used total organic carbon  method  depends
on the rapid combustion of a  micro-sample in a stream of  oxygen
in a heated tube. The carbon dioxide produced is measured with an
infrared  analyzer  at  a  wavelength  specific  for  carbon  dioxide.
This method was designed  primarily for analysis of true solutions.
Unless the particles in samples are  reduced to about 50 microns by
blending, the needle used for injection will act  as a  filter. The usable
sample size  is  20  n\, so some difficulties may arise  in obtaining
a representative sample. For this reason, in biodegradation studies
the dissolved organic carbon  is usually  determined.  That is, the

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analysis is run on a filtered sample. The TOC apparatus is sentitive
to about 1 mg/l total carbon, but for quantitative determinations
it is  desirable  to have samples in the 2—3 mg/l range. In order to
measure only  organic carbon, the inorganic carbon must first be
removed from the sample.  In samples having a very small amount
of organic carbon, it is essential  to eliminate  completely any in-
organic carbon from the sample by first acidifying and then purging
with nitrogen. Some  of the newer instruments permit the separate
determination  of inorganic carbon. In that case, the decarbonation
step may be omitted as an external procedure.
  The advantages of the method are  that a definite substance is
measured  in a very short time, and the results  may be interpreted
in relation to the thoeretical  ultimate oxygen demand. The dis-
advantage  or limitation  of the method is that  all organic matter
is oxidized, which may or may not be the case  in biological treat-
ment systems or streams.  Many  things can be  oxidized at  high
temperatures  that would  not be oxidized in  a stream within a
reasonable time.  For example,  activated carbon  is usually  not
considered to  exert a dissolved oxygen demand on a stream, yet
it can be oxidized at high temperatures.

BIODEGRADATION TEST METHODS

  Over the years considerable research  has been done on evaluating
microbial  breakdown of  organic compounds.  Techniques  have
ranged from  Warburg respirometry  used by  microbiologists, to
BOD determinations  and laboratory activated sludge units used by
environmental engineers.
  The important  features  and limitations of various  methods of
testing biodegradability are discussed below.

River Die-A way --

  The river die-away method is very simple and has been used by
many investigators. The  compound to  be  tested is added to  a sam-
ple of  river  water at ambient room temperature.  Periodically,
the solution  is  analyzed  for  the test compound to establish a
die-away or disappearance curve. The curve usually shows a lag
period of  no degradation for several days and then a rapid removal.
The  advantage of the river die-away  test is that  it requires  very
little  equipment  and yet  can  indicate the  rate  of degradability
of the test material.  However, the rate  is only relative. If, after
the  disappearance of the compound,  an additional  amount  is
added, it  will  disappear in  less time. This can be repeated several
times, resulting in a die-away curve with a saw-toothed appearance
(Fig. 1).
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   100
LU
I-
LU
o
IT
    50
               10
                       20
                                30

                               DAYS
                                         40
50
         60
               Figure 1, River die-away pattern.
  A modification that can be used to  reduce the lag time and to
speed the process is to add a  small  amount  of activated  sludge
to the river water.
  The disadvantage of the test is that  all river waters are not the
same. Some have more microorganisms than others,  and the pH
and mineral constituents vary. To be able  to use this procedure,
it  is necessary to be able to analyze for the compound or  some
constituent of the molecule that is a likely degradation product.
The method of assay will detect the original compound, but per-
haps not a slight modification of the compound. Thus, intermediate
products may go undetected. In spite  of the inherent drawbacks
of the  river  die-away test,  it is useful in  obtaining preliminary
information on  the  ease  of biodegradation. A  large number of
compounds can be screened in a relatively short time.

Warburg Respirometer --

  The disappearance of oxygen as degradation proceeds is a general
characteristic of aerobic biological systems.  The Warburg respiro-
meter is one  of the many apparatuses  that permit oxygen uptake
rates  to be  determined for a  biological  system.  The Warburg
respirometer is popular because  it is commercially available, com-
pact, and permits a number of samples to be run simultaneously.

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  There are a  number of variables that can affect the results, but
the most important factor is the type and quantity of the micro-
organisms used. The seed (inoculum) used should be one that has
become  adapted  to  the test  compound  or  has at least  been
subjected to an adaptation period.
     Some  of the advantages and disadvantages are:
Advantages:
     1. Direct  measurement  of biological  oxidation  is  possible.
     2. This approach can  be  used  for  all compounds without
        using a specific analytical  test procedure. This is a decided
        advantage  in  cases where no reliable analytical method
        exists.
     3. The results are available within a reasonable time.
     4. Oxygen  consumption,  radioactive-carbon  dioxide  eval-
        uation,  and  respiration  assimilation balances can  serve
        as bases for measuring biodegradability.
     5. Continuous  observation of  multiple  samples  over periods
        of a few hours or several days is possible.
     6. Lag periods and rate changes are readily determined.
Disadvantages:
     1. Equipment  is  fairly  expensive  and requires a skilled  tech-
        nician,
     2. The small  sample size makes it difficult  to  obtain  rep-
        resentative samples of wastewater and industrial wastes.
     3. All  organic matter added to the system  is subject to oxida-
        tion.  Therefore, pure  organic   compounds  are needed
        for fundamental studies unless a  specific assay is included
         in the study.
     4. With  extended  runs, nitrification  can  occur,  giving  erro-
        neous high   results  of  oxygen  uptake.  Each  part of
        ammonium nitrogen converted to nitrate will consume 4.5
        parts of oxygen.
     5. Carbon dioxide evolved during respiration must be absorbed
        from  the gas phase.  It has been reported that complete
        absence  of  carbon dioxide  can inhibit  assimilation of
        organic matter by bacteria.
     6. Oxygen uptake results are  difficult to  interpret quanti-
        tatively unless supplemented by other analyses.

Biochemical Oxygen Demand Test --

  The biochemical oxygen demand test (BOD) is one  of the  older
methods for evaluating the  biodegradability of substances.  It was
first published as  a  standard  by the British  Royal  Commission
for Sewage Disposal in 1912. For many  years nothing better was
available for estimating the  strength and controlling the treatment

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of wastewater.  The sample is stored  in a filled bottle for five days
in the dark at 20°C, then the amount of dissolved oxygen utilized
is measured. Instead of a single final value, a curve showing oxygen
uptake as a function of time can be developed if replicates of  the
original sample are analyzed at different times.
  With the  development  of  reliable oxygen-sensing  probes, it is
now possible to determine the oxygen content of a single sample,
at intervals or continuously,  over  the entire test period.  Another
modification of the  test apparatus is  to determine  the  oxygen
uptake electronically based  on the electrical  current needed to
generate  the oxygen  required  to  maintain  the pressure at equili-
brium in  the closed flask.
  The BOD test has  numerous limitatioris  relating to the appli-
cability of the  test results to  predicting what will take place in a
waste treatment plant or a stream.  Only the limitations that  are
pertinent to  the  utilization of the  method for  determining bio-
degradability will be discussed.
  The advantages and disadvantages of the BOD test procedure are:
Advantages:
     I.The  apparatus and testing procedure are simple. The test
         does not require a highly trained technician.
     2. Extensive   BOD data  are  available on  compounds with
         known waste treatability.
     3. The  small  number of  bacterial  cells used and the organ-
         ic-free  BOD dilution water minimize interferences with
         specific chemical analyses either for the parent compound
         or intermediate breakdown products. •
     4. Carbon  dioxide  is  present  throughout  the  test  period
         and does not become limiting as is possible in the Warburg
         procedure.
     5. Problems  of  aeration   and oxygen transfer  are  not  en-
        countered  if suitable initial concentrations are selected,
         because all the oxygen required is present at the beginning
        of the test.
Disadvantages:
     1. Only a small amount of sample can be used because of  the
         limited solubility of oxygen in water.
     2. The  testing period is long. However,  intermediate  results
        can be obtained  in less time if oxygen depletion is meas-
         ured periodically.
     3. As with  the Warburg procedure, nitrification and extraneous
        organic matter can lead to erroneous conclusions.
     4. The  interpretation of the results has the same limitations
        as other respirometer methods in that the results are  not
         directly convertible  to a  percentage  of the compound
         degraded.
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Flask Test Method -
  Except for the activated sludge method, this is the most widely
used of all  the laboratory  procedures. There  are  many  versions
of  the  technique,  but most  use  a chemically defined medium.
The flasks may be aerated by continuous shaking on a mechanical
shaker,  agitated with  magnetic stirrers, aerated through a ceramic
filter, stirred with a paddle stirrer, or merely left  quiescent. The
test mixture can be analyzed each day or at the end of a specified
time. A variety  of inocula can be  selected  for  this procedure.
If degradation  is  not evident or toxicity  is suspected,  the test
compound  should be  run at a lower  concentration.  Parallel runs
with known degradable compounds are advisable to confirm that
the  conditions were  correct.  It is suggested  that a compound
of similar structure and class serve as a  reference standard.

  The popularity of the flask test  method is exemplified by the
fact that five leading countries (Austria, Canada, Germany,Sweden,
and England)  have adopted some version of it as  their screening
or  presumptive test method for determining the biodegradability
of anionic detergents.
  Some  of  the advantages  and disadvantages of  the flask test
method are:
Advantages:
     I.The  method does  not  require elaborate  or specialized
        laboratory equipment.
     2. Reproducibility is generally good.
     3. Sample size is not severely limited.
     4. The test compound does not have to be pure.
     5. Metabolic products  can  be identified  without too  much
        interference if mineral salts are used and if the test com-
        pound  is the major carbon source.
Disadvantages:
     1. The method requires a separate adapting process if adapta-
        tion is  not made a part of the  test procedure.
     2. Shaking  on a  mechanical  shaker  limits  the  number  of
        samples that can be  run.  If  a static  procedure  is used,
        sample  concentration  is limited  owing to poor oxygen
        transfer.
     3. The  test does  not simulate any waste treatment  process.
     4. A  chemical or  physical analytical  test method must  be
        available for each class of compounds tested.
     5. Simple  analytical  procedures  generally do not  measure
        partial  degradation residues.

                              12

-------
 Activated Sludge Method --
  The activated sludge method  is one of the most important waste-
water treatment processes. Laboratory scale models of the process
have  been used for many years to study the treatability of various
municipal and industrial  wastes. It was natural  that the method
should  become  a  major tool in biodegradability and treatability
testing.
  The laboratory methods based on the activated sludge principle
vary  in  feed,  residence time, and whether the feed  is continuous
or  batch  operation.  The  continuous activated  sludge system  is
preferred  because  it corresponds more nearly to actual conditions.
A system that approximates  the operation of  a plug flow aeration
tank  is known  as the fill-and-draw or  semicontinuous process.
In this  method, the inoculum,  the  test material, and the feed are
placed  in a vessel  and aerated. The feed is usually a chemically
defined medium with organic supplements. The mixture is aerated,
usually  for  23 hours. The air  supply is then turned off and the
mixture allowed  to  settle  (see Fig.  2).  A  portion  of the su-
pernatant is removed and replaced with new feed and  the  test
compound.  This sequence is then repeated for the duration of the
test.  If  the test lasts a  long time, it  is necessary to waste some
activated  sludge periodically  in  order to maintain a reasonable
level  of biomass. As  in all aerobic  biological  processes, there is a
net increase in bacterial cells.
  Although  each cycle in the semicontinuous operation constitutes
a batch run, the method  closely approaches plug flow conditions
in a conventional activated sludge plant. The aeration basin of the
activated sludge process is usually  long and narrow. The recycled
sludge and the wastewater enter at one end of the aeration tank.
Ideally, the mixture  moves  as a plug  along  the entire  length of
the tank.  The organisms are  subjected to all phases of the growth
curve from  the log phase to the endogenous stage. That is, at the
head  end of  the  tank, the  organisms have  an ample supply of
food. It is in  this  part of  the tank that the organisms grow most
rapidly, and consequently the  rate of  removal of organic matter
is the greatest. As  the plug moves along the tank, the food becomes
depleted,  and if the  retention  period  is correct,  the bacteria  will
start  to enter the  endogenous phase near the end of the aeration
basin.
  Various  designs  for continuous  flow activated  sludge  studies
have been reported in the literature. Figure 3 shows a continuous
flow,  complete  mix  reactor of  300  ml capacity developed by
Monsanto Chemical  Co.  Figure 4 shows a 6  liter complete mix,
continuous flow unit  developed by the U.S. Public Health Service.
                              13

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AERATION PERIOD (Al.
SETTLING PERIOD (B).
Figure 2. Four liter fill-and-draw reactors with two liters of test
         mixture, in both the aeration period (A), and the
         settling period (B).
                               14

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Figure 3.  Continuous flow 300 ml reactor.
                 15

-------
Figure  19 shows a 750 liter per day continuous flow, plug flow
reactor.
  Any material  used for constructing equipment for biodegradation
or  treatability  studies  on  very   low   concentrations  of  test
compounds should  be evaluated for sorption of the test compounds
on surfaces.
  Small  reactors  of only  several hundred milliliters  can be used
to establish  the biological characteristics  of a test system, such as
degradation  or sorption  on  the  sludge surface.  The  larger units
of several liters capacity are useful to collect additional information
on  the  physical characteristics  of  the  system, such  as  sludge
settleability.  Finally, the large continuous  flow  pilot  plants  of
several  hundred liters per day through-put capacity  are  necessary
to collect complete treatability and  engineering design data. The
large  pilot  units can  be used  to track  disproportionation of a
compound  through the various unit  processes and  give  guidance
on settler overflow rates, sludge production, aeration requirements
and detention time  for the unit processes.
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Separator made from a 10" section of 4"
Plexiglass tube cut in half longitudinally.
Bottom end cut at a 45°angle.  Part of the
lower end is closed by a 1/8" Plexiglass plate
so that the two compartments are connected
by a 1/2" x 4" opening.  Water proof "rubber
to metal" cement used to join Plexiglass to
glass.
               Figure 4.  Model activated sludge unit.
                                16

-------
  The test apparatus selected should not be so small that sampling
 would disturb  the system.  For biodegradability testing it is not
 necessary  to maintain  a  large unit.  Small units are  much  more
 desirable  because of space, cost,  and  labor of servicing several
 large units. In  scaling  down  to very small units, there is  some
 difficulty in designing settling  chambers. In most units the  settling
 chambers  are an  integral part  of  the  aeration tank  rather than
 being  separate  units. Most  of the  laboratory  units are the com-
 pletely mixed type rather than the plug flow type.
  A list of some of the  advantages and limitations of the activated
 sludge method follows:
 Advantages:
     I.The continuous and  the semicontinuous methods  using
        high concentrations of biomass simulate in the laboratory
        the treatment that wastewater normally  receives.
     2. There  is  considerable  information  on   the characteristics
        of these units since the method has been  used for many
        years.
     3. Reproducibility  is very good  if the unit is  operated until
        steady state is achieved.
     4. Sample size is not limited.
     5. The test compound  does not have to be pure. If mineral
        salts medium   is  used,  BOD or  COD   tests can be used
        to calculate an oxygen material balance.
 Disadvantages:
     1. Continuous operating  units  require money,  space,  and
        maintenance.
     2. Single batch  runs do not allow for acclimation unless the
        test period is unduly prolonged.
     3. Residence  time  of 23 hours  in  the fill-and-draw method
        is not  typical  of most treatment plants, and therefore
        relates only to extended aeration.
     4. Most investigators feel  that valid results cannot be obtained
        in less  than ten days. There  are always unexplainable
        fluctuations in the efficiency  of the  process that  must
        be averaged.
     5. A  chemical or physical analytical method must be available
        for the analysis of  each compound if  wastewater is used
        as feed.

Trickling Filter Method --

  Many small towns use trickling filters as the means of secondary
treatment. The  name is  a misnomer to the extent that the filter is
not a filter in the ordinary chemical engineering connotation. The
principle of the  method is that the supernatant from the primary
                              17

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settling  tank is  sprayed  on a  bed of crushed  stones about 6 ft
deep. Recently,  plastic  trickling filter  medium  has come  into
being. The  wastewater trickles through  the bed  to under-drains
that lead to a final settling tank. The action of the filter consists
in mechanically  removing the lighter solids not removed by sedi-
mentation and in the oxidation of the wastewater  by contact with
the microbial film on the filter medium.
  Experimental trickling filters have  never been a popular  device
for  measuring  biodegradability. Small  units  require  small size
media that increase operational difficulties as a result of ponding,
air circulation, and side wall effects. If recycling  is not  practiced,
the most important dimension is the depth of the filter bed because
that determines  the retention time.  The bed should be  at least 6
feet  deep.  The  main  limitation  is the time required to develop,
acclimate, and equilibrate the biological film. It can be a matter of
months  before steady  state is  reached. If a trickling filter is used
for biodegradability studies, the BOD or COD or TOC removals
also  should  be determined so that a comparison can be made be-
tween  the percentage removal  of  the other  constituents of  the
wastewater and the test compound.
                              18

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Section 2. Treatability of Specific Compounds
ANAEROBIC STUDIES

  Of all the biological  treatment  processes, anaerobic digestion  is
one of the most susceptible to poisoning by toxic substances, espec-
ially heavy metals and chlorinated organic compounds. Anaerobic
digestion  is two biological processes  rolled  into one. The first
group of bacteria degrade the starches,  fats, and proteins to mostly
short-chain  fatty  acids.  The  second  group, methane-producing
bacteria, continues the process of degradation to produce methane
as the end-product. This group is  much more susceptible to inhibi-
tion than the first. The  reduction of methane production signals
digester failure, which  results in offensive sludge that  is not well
digested. The toxic effect of chlorinated organic compounds at very
low concentrations is  remarkable, and provides a warning that a
close watch must be kept  on the disposal of the new chlorinated
hydrocarbons that come on the market. Toxic levels of a number of
organic substances are shown in Table 1.
  The effects  of  metals  on anaerobic digesters are dependent on
the amount of sulfides present. Sulfide precipitates heavy metals,
thus forming  insoluble metal sulfides that  are  not toxic. Most
digesters contain  enough  sulfides  that failures are not  usually the
result of metals  except for large slug dosages from spills. Some of
the more common metals  and their acceptable  concentrations in
digesters are shown in Table 2.
  Although the resilience of bacterial populations has  often been
stressed,  this  capability  is more commonly observed  in aerobic
processes  than in  anaerobic digestion. Once the performance of
an anaerobic digester has deteriorated,  it  is usually rather difficult
to reestablish.
  Anaerobic tests for  biodegradation should be managed so that
the test system has a hydrogen electrode (En) potential  of about
-0.4 volts, dissolved oxygen is excluded,  and methane  gas is gen-
erated.  Because  anaerobic  organisms have protracted  generation
periods, time studies should extend to at least thirty days. If any
change  is made   to  the  system,  the observations of  the  effects
should continue  for at least three  times  the hydraulic detention
time of the system.

                             19

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      TABLE 1.  TOXICITY TO ANAEROBIC DIGESTERS
                                        Concentration mg/l
          Organic compound           In wastewater   In sludge
Acrylonitrile                                -            5
Carbon tetrachloride                         -           10
Chloroform                                0.1
1, 2—Dichloroethane                         -          <1
Methylene chloride                         1
Di(5—chloro—2—hydroxypheny I (methane
 (dichlorophen)                            1
Pentachlorophenol                          0.4
1,1, 1—Trichloroethane                      -            1
Trichlorofluoromethane                     0.7
   TABLE 2. TOXICITY OF METAL IONS TO ANAEROBIC
	DIGESTION OF PRIMARY SLUDGE	

     Metal ion                Concentration in wastewater, mg/l

Cadmium                                    1
Copper                                    10
Dichromate (as Cr)                          50
Lead                                    50-70
Nickel                                     40
Tin(asSn)                                   9
Zinc                                       10
Sodium Salt of Nitrilotriacetic Acid

  In 1971, the sodium salt of nitrilotriacetic acid was suggested as
a replacement for the phosphates in detergent formulations. The
material  was  found to be aerobically  degradable, but reports of
anaerobic degradation  were conflicting.  To  resolve this  issue,
sodium nitrilotriacetic acid (NTA-Na3) was studied in thefill-and-
draw apparatus shown in Figure  5. At various times in the study
period, both  primary sludge and waste-activated sludge  served as
feed to the anaerobic digester.

                             20

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                                              ATMOSPHERIC VENT
                                                    PRESSURE
                                                    SQUEEZE BULB
                 Figure 5. Digester apparatus.
  The program  was divided into three  distinct study periods,  the
first of  which was a control period to  document the digester per-
formance.
  The second period concerned  the degradability of  NTA-Na3
in an anaerobic  system  receiving only primary sludge feed.  To
initiate this study, 30 mg of NTA-Na3  were added directly to  the
digester, yielding a concentration of 10  mg/l in the mixed contents.
For an interval  of one week, the primary sludge feed also contained
10 mg/l  of NTA-Na3. This was an acclimation period designed to
indicate  whether the  10 mg/l concentration would have any dele-
terious effect on the digestion process.  The gas production and pH
remained within  normal  limits.  The NTA'Na3  concentration in
the daily feed sludge was then increased to 20 mg/l,  and  maintained
at that   level for  three months.  Figure 6  shows  the  l\ITA'Na3
concentration in  the mixed  digester  contents  during this time
period.  On day zero the concentration  of  NTA-Na3  was abruptly
increased to slightly over  10 mg/l  as a result of the direct addition
of 30 mg of NTA-Na3 to the digester. The NTA-Na3  concentra-
tion  gradually increased to the  feed  concentration  20 mg/l over a
120-day  period. This would be predicted on the basis of the  hy-
draulic detention time of 30 days. Over the 120 days of observation,
no  appreciable degradation of the  NTA-Na3  was noted  in  the
primary  sludge  digestion process.  Table 3  shows that the digester
was  performing satisfactorily with regard to organic  loading,  gas
production,  and  volatile  solids  destruction during  this second
period.  The conclusion was that NTA-Na3 was not degraded in
an anaerobic process operated entirely on primary sludge feed.
                             21

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Figure 6. NTA-Na3 concentration of digester, day 10 through 205.
    TABLE 3. ANAEROBIC DIGESTER PERFORMANCE
          Parameter
                                            Study period
   1
 Volatile solids destroyed (%)
 Volatile solids loading (g/l/day)
 Gas produced from volatile
   destroyed (l/g)
 Gas produced from volatile
   added (l/g)
 Volatile solids in feed (%)
49       76       65
 0.71     0.80     0.57
 0.89
0.94
0.95
 0.43     0.72     0.62
82       69       83
  The third study period concerned  the  anaerobic digestion  of
primary and waste-activated  sludge.  The digester was switched  to
a daily feed of 80 percent primary  sludge and 20  percent waste-
activated sludge, on a volume basis. The overall concentration  of
NTA-Na3  was  20 mg/l. The waste-activated sludge was obtained
from an activated sludge plant that was acclimated to the degrada-
tion  of  NTA-Na3. The  digestion  process was monitored for two
weeks.   Operation was satisfactory,  and  the  concentration  of
NTA«Na3  in the digester contents  remained near 20  mg/l. The
percent  of waste-activated sludge in the daily feed was then  in-
creased  to 50 percent by  volume. Over a  two-month period, the
                             22

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 NTA-Na3  content  of  the  mixed digester  contents  gradually de-
 creased, until at the end of this period the concentration was near
 zero (see Fig. 6). To verify that the anaerobic system was biologic-
 ally  degrading  the   NTA-Na3,  a  direct addition of  NTA-Na3
 (to yield about 20 mg/l in the mixed digester contents) was added
 on day 190. The usual daily dose of 20 mg/l in the combined sludge
 feed was continued. Figure 6 shows the response of the system as
 measured  by the NTA*Na3  content  of the reactor contents.  In
 about  15 days (day 205),  the residual  NTA-Na3  was again near
 zero.
  Table 3 provides a summary of the digester's performance during
 each of the three study periods. Typical anaerobic digester perform-
 ance values were obtained in each period, a fact that  indicates
 good operational control of the reactor.
  The  results of this study support the conclusion  that primary
 sludge  does not contain a microbial population that can murtiply
 during anaerobic digestion  and  degrade  NTA-Na3.   However,
 once acclimated organisms are established in the mixed digester
 contents  by means  of  an inoculum of facultative organisms from
 waste-activated  sludge,  anaerobic degradation of  NTA-Na3 can
 proceed.

 AEROBIC STUDIES

  This  section summarizes a variety of aerobic biodegradation test
 methods that have been used with specific classes of compounds.


 Bacterial  Utilization of  Lignans

  Lignans  are a family of  methoxylated aromatic compounds of
known molecular composition that are structurally related to, but
 less complex than, lignins. Lignins are more complex methoxylated
polymers that are extremely  resistant  to microbial action. Alpha-
conidendrin  is  a  representative  of  the  lignan  type  molecule.
Organisms capable  of  degrading this material  were isolated  from
natural sources using adaptation techniques.
  A  synthetic  basal salts  medium  containing  0.1  to 1.0 percent
 a-conidendrin as the sole carbon source was used for  enrichment
 cultures.
  Since a-conidendrin  is insoluble, the analytical procedure used to
 detect degradation  was determination of methoxyl  groups on the
 residual solids  at various times. Alpha-conidendrin has a methoxyl
 content of  17.5 percent.  An adapted culture inoculated into an
 aerated culture medium containing 200  mg/l  a-conidendrin re-
 duced  this value to 0.8 percent in 20 days.  Figure 7 shows the
 daily progress of the degradation.
                              23

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                             PERCENT DEGRADATION OF
                             ALPHA - COWDENDRIN
                             METHOXYL CONTENT
                             RESIDUAL SOLIDS
                        INCUBATION TIME IN DAYS
   Figure 7.  Methoxyl content of residual solids and percentage
            degradation of a-conidendrin.
  The  organisms  adapted  to  the  utilization  of   a-conidendrin
dissimilated vanillic, p-hydroxybenzoic,  and protocatechuic acids
when grown on chemically defined media with each of these acids
as a sole carbon source.  Cultures freshly  isolated  from a natural
source by  streaking on  a-conidendrin agar when  incubated in  a
medium  with  either vanillic, p-hydroxybenzoic, or protocatechuic
acid as the only carbon source showed turbidimetric measurements
indicating a limited  amount of growth.  However, when the same
culture  was subjected  to many  successive  transfers  in a  liquid
a-conidendrin  medium,  and was then inoculated  into a medium
with any one of  the  above acids as  the sole carbon  source,  the
amount  of growth increased from  40-  to  60-fold  over that  ob-
tained in the same acids before the culture was adapted.
  Adaptation studies, together with  chromatographic and spectro-
photometric analyses of the degradation products, indicate that the
metabolic  pathway  in the  dissimulation   of  a-conidendrin  by
pseudomonad  strains may proceed from  a-conidendrin  —+ vanillic
acid —•» p-hydroxybenzoic acid — •»  protocatechuic acid —* keto-
adipic acid.
                              24

-------
 Degradation of Aromatic Compounds by Phenol-Adapted Cultures

  A culture acclimated to 300 mg/l phenol was tested in Warburg
 respirometers to determine its potential for degrading 104 aromatic
 compounds. The washed culture was added to 0.067 M phosphate
 buffer contained in the respirometers and the test compound was
 introduced from the side  arm to yield a  100 mg/l concentration.
 Oxygen uptake  was measured and chemical colorimetric analyses
 were done  for  the specific aromatic compounds.  Observations
 were carried out for about 200 minutes, with the oxygen  meas-
 urements made at 10 minute intervals.
  The various aromatic groups tested were phenols, benzyl alcohols,
 hetrocyclics, cyclics, benzoic and other acids, benzaldehydes and
 benzamides, and substituted benzenes.  The  results are shown in
 Figures 8, 9, 10,  11, 12, and  13.
  On the  basis of the results for these organic compounds,  there
 appeared  to be a  relationship between  molecular structure and
 resistance to bacterial degradation. The results indicated significant
 differences in  resistance to  oxidation within each of several well
 defined groups of compounds. The relationship between molecular
 structure and  ease of degradation was also apparently affected by
 the position of a group on  the ring,  the type of group, multiples
 of the same or different substituents,  and  the size and complexity
 of the substituent.  The  results indicated that there was a possible
 relationship  between molecular structure and  ease of degradation
 of some of the phenols. The presence of more than two hydroxyl
 groups on the ring  appeared to make  a compound highly resistant
 to oxidation, for example, phloroglucinol and pyrogallol. Dihydric
 phenols seemed  to  be oxidized to about  the  same, or possibly  a
 somewhat lesser degree than  phenol. The fact  that none of  the
 nitrophenols tested, except  o- and m-nitrophenol, and 2,4-dinitro-
 phenol, showed  any appreciable 02 uptake  indicated that adding
 a nitro group tended to increase resistance.
  Chloro  substitution showed  that  dichlorophenols were  more
 resistant to oxidation than monochlorophenols. The effect of the
 methyl group  was indicated  by the relatively  high level of activity
with cresols  and  with some dimethylphenols. The effect of position
of the substitution on the  ring was illustrated with the cresols and
dimethylphenols. With cresols, substitution of the methyl group in
the para position resulted  in a higher  initial O2 uptake rate,  but  a
 lower total  O2  uptake, than  was the case with o- and m-cresol.
The same  effect  for para-substituted compounds was observed with
dimethylphenols. High total 02 uptakes were noted with 2,4- and
3,4-dimethylphenol,  whereas  oxidative  activity was lower with
dimethylphenols having  methyl  groups  in   other  positions.  All
 monochloromethylphenols  had  a  measurable  02  uptake  while
                             25

-------
           SUBSTRATE CONCENTRAT ION = 100 ppm
           EXCEPT 2,4 DICHLOROPHENOL =60ppm
                         -2,6 DichloropHenolC'»JC'
                         _J	1   V  I
2,4,6 Tfi

2,4 Dic
                    60      90     120     150     ISO     210
                     DURATION OF WARBURG RUN, MIN
       Figure 8. Oxidation of hydroxy- and chlorophenols.
there was little activity with a dichloromethylphenol (4,6-dichloro-
m-cresol). This indicated that, as with the chlorophenols, dichloro
substitution   increased  the  resistance  of  chloromethylphenols.
The presence of a methoxyl or  phenoxyl group on  the ring  also
increased resistance to degradation.
  Differences in O2  uptake related  to molecular structure and  sub-
stitution were also  observed with  benzoic  acids.  Activity with
benzoic acid was relatively higher than with its hydroxyl derivatives.
The  latter  in turn  had  greater  activity  than the  trihydroxyl
derivatives.  In general,  the  hydroxybenzoic  acids were more  sus-
ceptible  to degradation than the other substituted benzoic acids.
The influence of position of substitution was  evident in the  case
of  methylbenzoic acids  because m-toluic acid was  more  readily
oxidized than the ortho and para  derivatives.  With methoxylated
                               26

-------
          SUBSTRATE CONCENTRATION = 100 ppm
          EXCEPT 2,4 DINITROPHENOL = 60 ppm

                              OH
    350
   O
    -250
                                  REPRESENT MAXIMUM ACTIVITY WITH
                                  ANY OF THE NITROPHENOLS TESTED
            30     6O     90      120     150     ISO     210
                   DURATION OF WARBURG RUN.MIN


        Figure 9. Oxidation of methyl- and nitrophenols.
benzoic  acids, increasing the number  of methoxyl groups on  the
ring apparently interfered with oxidation. Syringic acid  was more
resistant to degradation  than vanillic acid.  The  presence of two
carboxyl groups significantly  reduced the  rate  of O2 uptake by
the benzoic acids.  In  the case of hydroxybenzoic acids and  nitro-
benzoic  acids, the number  of  compounds tested appeared  to be
sufficient  to   provide  some  statistical  basis   for  tentatively
concluding that the former  were readily oxidized by the phenol-
adapted bacteria, whereas the latter were extremely resistant.
  A  high level of oxidation  was observed with benzaldehyde and
its  para  hydroxyl  derivative. Replacement of the hydroxyl by  a
nitro group blocked the activity. This  was demonstrated by  the
                               27

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   5 200 —
                                     SUBSTRATE CONCENTRATION --100 PD™
                                     EXCEPT 3,5 OINITROBENZOIC ACID = 60 pp
                                                        COOH

                                         ' 2,5 O'Mfoiybenzoic Acid  (n°H
                    120     ISO     0     60     120      ISO     24O
                     DURATION OF WARBURG RUN, WIN
              Figure 10.  Oxidation of benzoic acids.
resistance  of  m-  and p-nitrobenzaldehydes.  Benzamide,  a  com-
pound in which an NH2 group  replaces the hydrogen  in the CHO
group  of  benzaldehyde, is  also resistant.  A measurable level  of
activity was observed with vanillin.
  Results obtained with benzene and its chloro derivatives, regard-
less of the number of position of the chloro substitutions,  showed
that,  in  general, these compounds were resistant to degradation.
The  effect of a nitro group was significant, as indicated by such
compounds as nitrobenzene, and m- and p-dinitrobenzene, which
had a low level of activity.  In  particular, the activity of the mono-
nitro-substituted  benzene was  below the  endogenous level.  The
                               28

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             1       I      1      1

           SUBSTRATE CONCENTRATION = 100 pp
                   60     90     120     ISO     180
                    DURATION OF WARBURG RUN, MINUTES
Figure 11. Oxidation of selected benzoic and amino acids compared
          with that of tannic acid.
resistance of the nitrobenzenes to degradation seemed to decrease
as the  number  of nitro groups on the benzene  ring  increased.
Neither the methyl  nor amino  group appeared to interfere com-
pletely  with the oxidation of benzene because both aniline  and
toluene were  oxidized  to a  limited  extent. The position  of the
nitro substitution may be important because there was evidence
of some oxidation of m-nitroaniline but little, if any, activity with
p-nitroaniline.  Both  2,4-dinitrotoluene and  2,4,6-trinitrotoluene
(T.N.T.) appeared to be oxidized to some extent.
   Studies of this type are useful for the preliminary screening of a
large number of compounds. The  occurrence  of oxidation under
these conditions indicates that the compound would  be oxidized
                              29

-------
     500
     450 —
400 —
     360
   cfl
   CE
   o
   IE
     300 —
   X
   o
              A -  Benzamide  U  0=CH
                          v   /
              • - p-Nitrobenzoldehyde  (I
     250 —
      150 —
      100 —
              3O     60      9O     I2O      150     180     2IO

                   DURATION OF WARBURG RUN, WIN


     Figure 12.  Oxidation of benzaldehydes and benzamides.
in municipal biological  treatment plants. Failure  of a  compound
to be  oxidized  under these conditions is not  conclusive since in
actual practice microbial population selection could occur.


Organic Compounds for Replacement of Phosphates in Detergent


Formulations -


  The  major function of  phosphate in detergent formulations is
to prevent precipitation of  calcium  salts during laundering. As a
                              30

-------
      5OO
      450
      400
      350
    a:
    UJ
    t 300
    o
    a:
    u
      250
      200
       150
       100
       50
              SUBSTRATE CONCENTRATION = 100 ppm
                                          OH
                                r	Phi


                               -|_- J— ~~
                                   Phenol
   + 1,3,5 Tnchlorobenzene  Cl Cl
Representative of the Chlorobenzenes Tested
                30      60      90      120      ISO     180
                    DURATION OF WARBURG RUN.MIN

                 Figure 13. Oxidation of benzenes.
result  of the relationship between excessive phosphate discharges
and eutrophication, several organic compounds have been suggested
as substitutes for phosphates.
  In 1967, the compounds  shown in Figure 14 were  candidates
for consideration  as phosphate substitutes. These six materials were
tested  for biodegradation  using a basal salts medium  with yeast
extract and  an inoculum  of settled wastewater. The compounds
were  tested  at concentrations  of  5,  10, and 20 mg/l.  The flasks
                                31

-------
              NITRILOTRIACETIC ACID (NTA)
                                 0
                   0        CH2-C-OH
               HO-C-CH2-N
                            CH2-C-OH
                                 0
       HYDROXYETHYLIMINODIACETIC ACID (HEIDA)
                                 0
                   H        CH2-C-OH
               HO-C-CH2-N
                   H        CH2-C-OH
                                 11
                                 0
      DIETHYLENETRIAMINEPENTAACETIC ACID (DTPA)
    O
HO-C-CH,
                                   0
                                   11
                              CH2-C-OH
HO-C-CH2
    11
    0
.N-CH2-CH2-N-CH2-CH2-N
            i
            CH2
            i
            C-0
            OH
       Figure 14. Structural formulae of complexones.
                         32
CH2-C-OH
    0

-------
       ETHYLENEDIAMINETETRAACETIC ACID (EDTA)
           0

       HO-C-CH,
                                  0

                             CH,-C-OH
       HO-C-CH2'
           ii
           0
                    N-CH2-CH2-N
                            XCH2-C-OH

                                  0
N-HYDROXYETHYLETHYLENEDIAMINETRIACETIC ACID (HEDTA)

                                         0
    0

HO-C-CH,
                                     CHo-C-OH
       HO-C-CH2-
           11
           0
                     N-CH2-CH2-N
                                  H

                                  C
                                  i
                                  H
1, 2-DIAMINOCYCLOHEXAIME-N, N' -TETRAACETIC ACID (DCTA)
             0

         HO-C-CH2
                     N -
                      ,-N
                                0

                            CH,-C-OH
                                   CH2-C-OH
             0                          0

     Figure 14. (cont'd). Structural formulae of complexones.


                          33

-------
remained static and  weekly serial  transfers were  made for three
consecutive  weeks.  After  incubation  for seven days, each flask
was  analyzed  for  the  test compound to  detect  the  amount  of
degradation. The  chemical  method  used to  determine the  unde-
graded chelating  agent  was a complexation titration at  pH 2.5
with copper sulfate  using  PAN (1-(2-pyridylazo)-2-naphthol)  as
the indicator. Each  compound listed was run  on five different
occasions to eliminate  any variability of the inoculum. The bio-
degradable compounds showed biodegradability in every test.
  This method for testing for biodegradability provides an indica-
tion of the  time required for adaptation, as well as  a determination
of whether a compound is degraded during the test period.
  Only NTA (nitrilotriacetic acid) and HEIDA (hydroxyethylimin-
odiacetic acid) lost their chelating ability under the test conditions.
The  acclimation period for NTA was about two  weeks;  that for
HEIDA  was slightly  less. Both these compounds contain only one
nitrogen atom; the other four compounds, containing two  nitrogen
atoms, did not degrade under these test conditions.
  Although  the analytical  method  only determined whether the
compound  lost its chelating  ability,  it would  be fairly  safe  to
assume  that fragments  of  these molecules would be subject  to
complete degradation. No manometric testing was done.
  Another organic compound  under consideration  as  a phosphate
replacement  is carboxymethyl  tartronate,  more  commonly  re-
ferred to as GMT. This compound has the formula:

     (Trisodium 20-oxa-1,1,3-propane   Na02C     CO2 Na
      tricarboxylate)                    H-C-0-C-H
                                           I      I
                                      Na02C     H
  The biodegradation of this  compound at a concentration  of  20
mg/l was studied  in 300 ml continuous flow reactors as shown  in
Figure 9, using a mineral salts medium with yeast extract and sup-
plemental  organic  materials. The results are  shown in Figure  15.
The  remarkable feature is  that a period  of  about 12 weeks was
necessary for  acclimation.  Attempts  to shorten  the  acclimation
period by external  addition  of forest and  garden soil were not
successful.  However, once  acclimated, the culture could  degrade
shock CMT doses, and  could immediately  degrade CMT after a
substrate starvation period.
  During this study three analyses were used to monitor the CMT in
the reactor effluent: TOG, a non-specific colorimetric, and a specific
gas chromatographic  procedure.  After acclimation,  the  test reactor
and the control reactor  effluent had about the same TOG  concen-
tration. Figure 16  shows the comparison of the colorimetric and the
gas chromatographic assays during 18 weeks of the test period.
                             34

-------
 0

 Q
 z
 £
 t-~
 z
                                        TOC, EXPERIMENTAL
                                        REACTOR
                                        CMT,EXPERIMENTAL
                                        REACTOR (20mg/l)
                                        CMT, CONTROL
                                        REACTOR
50 mg/l CMT
        O
                  6   8   10  12   14   16   18   20  22 24
                    TIME, PROGRESSIVE WEEKS
Figure 15.  CMT and TOC of reactor effluents with synthetic feed.
  25-
  20-
  15-
  10-
   5-
                   SYNTHETIC WASTEWATER
                                       ^
-------
Synthetic Detergents --

  In the early  1960s great attention was focused on the biodegrada-
bility  of synthetic detergents because of the occurrence  of billows
of foam at wastewater  treatment plants and  along stretches  of
rivers.  Early  production of  detergents yielded compounds  with
branching  hydrocarbon  chains that were relatively  resistant  to
biodegradation.  Various  alternative  compounds  were  considered
for replacement.
  Using a static mineral  salts medium with  added yeast extract,
many  of  these  compounds  were screened for  biodegradability.
Provision for acclimation was a part of the protocol. Settled waste-
water was used to seed the medium. Tests on each compound  were
repeated on five different occasions  to  determine the  variability
of the seed. Table 4 gives the average results of these five replicate
tests for various synthetic detergents. Several  phenolic compounds
with  known degrees of degradability were included to  verify the
method. The compounds  were tested at a concentration of 10 mg/l.
  The  chemical  analysis used to  detect the  undegraded  alkyl
benzene  sulfonates  and alkyl sulfates  was  the methylene-blue
method.  The  cobalt-thiocyanate  test procedure was used for the
nonionic  surfactants.  Details of this  method can  be  found  in
Reference  10. Indirect determinations also were  made by judging
the foaming  tendencies  of the  surfactants.  The loss  of surface
activity or foam  does not  necessarily mean complete degradation
of the molecule. The  4-amino-antipyrene method was used for the
assay  of phenol,  o-cresol, and the chlorinated  phenols.  The  nitro
compounds were  adjusted  to  pH  10.0  and determined spectro-
photometrically.  As a further check on the phenolic compounds,
acid was added  to all the terminal flasks, ether extracted, and the
ether  examined  for the  benzene ring in the ultraviolet region of
the spectrum. No suitable solvent was  found that would extract
polyethylene   glycol  400  from  the  medium;  therefore,  it was
necessary first to dry the contents  of the flasks at 80° C before
using   the  cobalt-thiocyanate  procedure.  The  ether  extraction
step   was omitted  and  the complex  formed directly  with the
dried  residue.
  The  results  obtained  demonstrate  the general  applicability  of
this test.  In addition to determining whether a compound  is de-
graded during  the test period,  it provides an indication of the time
required  for adaptation.  This is very useful information  because it
is not enough  merely  to  know that a compound is degradable; the
rate  at which  it  is  degraded  is equally important where  water
quality depends  on it. Such  information is  helpful in  predicting
the behavior of a compound in a  waste treatment plant  and in sur-
face waters. For  example,  if microorganisms in a stream or waste

                              36

-------
TABLE 4. DEGRADABILITYOF SURFACTANTS

Percent
of compound degraded
Number of subculture
Name of compound
An ionic surfactants:
Linear alkyl (Ci2> benzene
sulfonate
Linear alkyl (Cio— Cis) ben-
zene sulfonate
Linear alkyl (Ci2— CH) ben-
zene sulfonate
Primary (Ci2) alcohol 3-mole
ethoxysulfate
Secondary (Cn— Cis ) alcohol
3-mole ethoxysulfate
Linear alkyl (Cu) benzene
sulfonate
Linear alkyl (Cn— CIB) phenol
5-mole ethoxysulfate
Branched alkyl (Cg) phenol
4-mole ethoxysulfate
N on ionic surfactants:
Linear primary (Cn) alcohol
8-mole ethoxylate
Linear primary (Ci2 ) alcohol
9-mole ethoxylate
Polyethylene glycol - 400
Branched alkyl (Cs) phenol
9-mole ethoxylate
Branched alkyl (Cg) phenol
1 5-mole ethoxylate
Branched (Cia) alcohol
9-mole ethoxylate
Phenol
o— Cresol
o— Nitrophenol
2,4— Dichlorophenol
m— Chlorophenol
2,4,6,— Trinitrophenol
Initial


84.5

90.5

93.5

99.5

96.0

61.0

39.5

9.5


98.5

99.5
52.0

0.0

12.5

1.5
99.5
99.5
81.0
8.0
0.0
0.0
1


98.5

90.5

97.0

99.5

97.0

92.5

12.5

9.0


99.0

99.5
42.0

0.0

6.5

0.0
99.5
99.5
84.5
47.0
0.0
0.0
2


99.0

92.5

97.0

99.5

97.0

96.0

11.5

3.5


99.0

99.5
63.5

0.0

0.0

2.0
99.5
99.5
93.5
91.5
0.0
0.0
3


99.0

92.0

97.0

99.5

97.5

96.0

12.0

7.0


99.0

99.5
61.0

0.0

0.0

2.0
99.5
99.5
98.5
98.5
0.0
0.0
                 37

-------
 treatment  plant were  exposed continuously to a  compound such
 as  2,4-dichlorophenoI,  the microorganisms  probably would adapt
 to  degrade  it.  If,  on the other hand, the exposure is intermittent,
 such as a spill, one would not expect the compound to be degraded
 in  a  waste treatment  plant. The compound might  persist in a
 stream  for  days  before the naturally  occurring microorganisms
 adapted to degrade it.
  As  a  result of the public concern about nondegradable deter-
 gents,  leading  manufacturers modified  their production processes
 to develop detergents with linear hydrocarbon chains. The market-
 ing of  these  new materials  began about  1965.  Five  treatment
 plants located around the  nation were  monitored to determine
 the  change in  influent wastewater  surfactant quality. Monthly
 raw wastewater samples were collected for  18 months. One liter
 samples were  aerated in 2-liter flasks for seven days after inocula-
 tion with 40 mg of volatile solids from  an activated sludge aeration
 tank. The determination of  the ratio of  hard  detergent (resistant
 to degradation) to  soft detergent (readily degraded) was calibrated
 by experimental procedures using various mixtures of known  lots of
 hard and soft detergents. Methylene-blue active substance (MBAS)
 was the analytical method used for determining degradation.
  The  data accumulated for one of  the  plants, in   Richmond,
 Indiana,  are given  in Figure  17.  Similar results were noted at the
 other four  sites.  It was concluded that a vigorous manufacturing
 and marketing strategy had been deployed and  the  quality of
 surfactants  in  influent wastewater  rapidly  changed  from  the
 undesirable  hard form to the acceptable soft form.
•Z. 80
LLJ
O
OC 70
(_• 60
z
111
O 50
or
LU
h- 40
111
D
Q 30
OC
                  J   J   A
                 	19 6 5
                          YEAR AND MONTH

 Figure 17.  Percent hard detergent in a raw wastewater; Richmond,
            Indiana.
                              38

-------
Steroid Hormones
  The increased medical use  of natural steroid hormones, coupled
with their recent use  as oral contraceptives  has  raised  concern
about their persistence in wastewater and the ability of conven-
tional  waste  treatment processes  to  remove these  compounds.
The normal concentration of steroids  in municipal wastewater in
the United States would  not be expected  to  be  more than a few
tenths of a milligram per liter. Nevertheless, since they are physio-
logically active in very small  amounts,  it  is important to determine
to what extent the steroids  are biodegraded in the normal course
of wastewater  treatment and in the receiving  bodies of water that
may eventually be used for water supplies.
  There  is  little information  on   the biodegradation of steroid
hormones.  The literature  is almost  void of studies that report ring
cleavage and complete oxidation of the  molecules. A  few studies
have been done on some of the natural steroids, but no biodegrada-
tion studies of the synthetic ovulation-inhibiting hormones have
been reported.
  Hormones representing different groups of steroids were selected
for the  biodegradability studies. The following is a list of represen-
tative hormone compounds  from  each of the classes of steroids
chosen for the study: (1)  Sterols — cholesterol; (2) Adrenocortical
hormones   —   progesterone  and  pregnanediol;   (3)   Estrogenic
hormones  —  estrone,  beta-estradiol, and estriol;  (4)  Androgenic
hormones — testosterone and androsterone;  (5)  Androgenic hor-
mone derivative — methyl  testosterone;  and  (6) 17-ketosteroids
— estrone and androsterone. The most commonly used synthetic
sex hormones were used for these studies. They were: (1) Estrogens
— ethynyl  estradiol  and mestranol  (ethynyl estradiol-3-methyl
ether)  and  (2) Progestins — norethynodrel  (17-hydroxy-19-nor-
17a -pregn-5(10)-en-20-yn-3-one), norethindrone  (17-hydroxy-19-
nor-17a-pregn-4-en-20-yn-3-one), dimethisterone  (6a,21-dimethyl
17-hydroxy-pregn-4-en-20-yn-3-one), etyhnodiol diacetate (19-nor
-17a-pregn-4-en-20-yne-3(3,17-diol),   norethindrone acetate,  me-
droxyprogesterone   acetate   (17-hydroxy-6a-methyl-pregn-4-ene-
3,20-dione  acetate)   and  chlormadinone  acetate  (6-chloro-17-
hydroxy-pregna-4,6-diene-3,20-dione).
  The compounds were tested at a  concentration of 20 mg/l in a
basal salts medium plus yeast extract, using activated sludge organ-
isms as  inoculum. The  flasks were  aerated by shaking during the
entire observation period. Since  these compounds are insoluble,
they were  suspended  in the  medium by introducing the hormone
dissolved in acetone and warming  the medium  to evaporate the
acetone.
                              39

-------
  Generic colorimetric assays for each group of steroids were used
to follow  the  progress of the degradation. The fate of the com-
pounds was also determined by  spectrophotometric  scanning to
determine  ring cleavage, and use  of ascending thin layer chroma-
tography   for  identification  of   residual  products.  Because the
materials  are  insoluble, both the supernatant and solids  portion
of the medium had to be extracted with  chloroform  to account
for the total  steroid content of  the  flask. Each  compound was
evaluated three times  in duplicate.  Provision for acclimation was
made  by transferring subcultures into fresh medium weekly. Table
5 shows the average percent degradation  for the three test runs
on each steroid.
   TABLE 5. AVERAGE PERCENT LOSS OF STEROIDS BY
   	WEEKLY SUBCULTURE ENRICHMENT	

                            Percent degradation of compound
Number
Steroid compound
Androsterone
Cholesterol
Testosterone
Progesterone
Estrone
Beta-estradiol
Estriol
Pregnanediol
Methyl testosterone
Ethynyl estradiol
Norethynodrel
Norethindrone
Norethindrone acetate
Dimethisterone
Medroxyprogesterone acetate
Chlormadinone acetate
Ethynodiol diacetate
Mestranol
Initial
96
96
95
93
91
90
82
79
77
72
71
70
68
65
56
55
52
47
1
100
100
100
100
98
97
96
95
91
89
88
87
84
82
80
78
72
73
of subcultures
2




100
100
100
100
97
97
97
96
95
93
92
91
90
88
3








100
100
100
100
100
99
98
97
94
89
4













100
100
100
96
90
                            40

-------
  The overall results of these studies show a variation in the relative
susceptibility of the natural  hormones and  the synthetic estrogen
and progestin components of oral contraceptives to biodegradation.
They  indicate that in  the microbial population  found in activated
sludge, there are  species capable of producing  enzymes that will
degrade both natural and synthetic steroid hormones if  conditions
for acclimation are provided.

Hazardous Organic Compounds

  Benzidine (1,1'-biphenyl)-4,4'-diamine) a bicyclic arylamine, has
been demonstrated to  be carcinogenic in experimental studies with
animals such as rats, mice, dogs, and hamsters. Literature citations
implicate many arylamines including benzidine  as causative agents
of bladder  carcinomas in  humans. Because  of this carcinogenic
property, benzidine has been included in the list of "14 Carcino-
gens" tabulated  in the Federal Register. The Occupational Safety
and Health Agency has  established standards to control  worker
exposure to these compounds, which  are used  in various process
industries.  Benzidine  is  used  in  hospitals,  research laboratories,
and dyestuff industries.
  Even though the use of materials such as benzidine is regulated, it
is  advisable to ascertain  the biodegradation  of this compound  in
conventional wastewater treatment, and to determine what protec-
tion a secondary facility could afford a receiving water in the event
that this material  is discharged into a sewer  system. Studies have
shown that the majority of carcinogenic compounds,  because of
their  strain-free  condensed  nuclear configuration, are resistant to
bio-oxidation by unacclimated microorganisms in activated sludge.
  The purpose of this study was to  find whether or not acclimation
to benzidine could  be established in  an activated sludge process
operated in a continuous feed mode within a specific range of feed
concentrations   of    benzidine   under   appropriate   operating
conditions.
  Continuous  flow reactors  of the type  shown in  Figure 4 were
used.  The  medium was settled municipal wastewater spiked with
selected concentrations of benzidine. The reactors were operated at
a hydraulic retention time of 24  hours and a  solids retention time
of 30 days. Phase I concerned benzidine concentrations of 0.5, 1,
and 5  mg/l. Phase  II studied benzidine concentrations  of 10,  20,
and 30 mg/l.  Biodegradation was  followed  by a  specific colori-
metric assay of benzidine, utilizing oxidation  with Chloroamine-T,
and residual chemical oxygen demand analysis.
  Since the biological  solids  were in the  range  of 4,000 mg/l,  the
sorption  of benzidine to these active surfaces had to be evaluated.
The  solids  for the 0.5  mg/l benzidine  reactor contained  0.003

                              41

-------
percent benzidine by weight. Solids from the 30 mg/l reactor con-
tained  0.3  percent sorbed benzidine.  On a mass through-put basis,
both these  determinations show that only about one percent of the
dosed material was sorbed to the biological floe.
  The color of the reactor sludges during Phase  II was remarkable.
The  control  unit mixed liquor was  light tan in color  during the
entire study. At the 6-week point the  10 mg/l, 20 mg/l, and 30 mg/l
dosed sludges were brown, dark brown, and brown-black, respec-
tively.  The  intermediate oxidation products and benzidine itself
are highly chromophoric compounds; and since these higher con-
centrations were not completely degraded,  this visual observation
confirms the analytical  results.
  Figure 18 presents the observations collected  during the 2 phases
of study.
    100
    90
  o 50
  0)
  cr
   ni(|

        O -	O  10 m
-------
  Results from the biodegradation studies on the several concentra-
tions of benzidine show that acclimated extended aeration sludges
can degrade continuous doses  of  1 mg/l benzidine. Less complete
degradation  occurs at higher dosage levels and intermediate oxida-
tion  products begin to  accumulate  in the system.  Even  at the
highest dose of benzidine, 30  mg/l, no marked interference  with
carbonaceous removal  efficiency  was  noted,  since the increased
COD of this effluent could be related to the presence of undegraded
benzidine.
  Concentrations  of  benzidine between  1  and  5 mg/l were not
studied, but because these studies employed an extended aeration
process and most secondary  municipal facilities have far shorter
hydraulic retention  times, it is recommended that  influent benzi-
dine concentrations  not  exceed 1 mg/l. This limit would ensure
that unacclimated systems would  not discharge significant concen-
trations of benzidine  into receiving waters.
  The work  also suggests that ambient concentrations of benzidine
below  1  mg/l will  be degraded by natural ecosystems, and the
material will not accumulate.
  Two  other compounds of environmental concern  are  acenaph-
thene (ANT) and dimethyl phthalate  (DIVIP).  Phthlate esters have
been recovered from  many environmental samples.  These  com-
pounds are  used  as plastisizers and industrial intermediates.  Some
studies have indicated  that these compounds  bioaccumulate in
plants and animals and subtle toxic effects have been  noted. Naph-
thalene  compounds  have also been noted in a wide variety of
environmental samples.  Recent studies have  shown  naphthalene
compounds  to  be toxic  to aquatic  invertebrates and fish.  Know-
ledge of  the biodegradation of these  two compounds would be
helpful  in   determining  the  efficiency of municipal  treatment
processes for control  of the substances.
  The compounds were  studied in  the continuous  flow  reactors
shown  in Figure  3. Settled wastewater was used as feed and  ANT
and DMP were added singly  and in combination. Concentration
levels of  0.1  and  1.0 mg/l were selected. It was possible to study
ANT and DMP in combination because the compounds could be
resolved using a Perkin-Elmer  # 900 gas chromatograph equipped
with a dual  flame ionization detector and a Varian Series Number 4
computer data programmer.
  The reactors were  started with  activated sludge and feed settled
wastewater  for one  week prior to the experimental run. Seven
reactors were operated at a hydraulic retention time of  6  hours
and a mixed liquor suspended solids concentration of 3,000  mg/l.
The ANT and DMP content of feeds and effluents was determined
by extraction of  the samples with  Freon-TF and evaporation of
the extract  by means  of  the Kuderna-Danish evaporation  pro-
cedure.  The gas  chromatograph  could detect 5  /ug of ANT or

                              43

-------
DMP. Table 6 presents the results.
  Reactors 1 and 2 were fed approximately 0.1 and 1.0 mg/l ANT,
respectively; Reactors 3 and 4, 0.1 and  1.0 mg/l  DMP; Reactors 5
and 6, 0.1 and 1.0 of both ANT and DMP; Reactor 7 was a control.
ANT is insoluble in water and had to be dissolved in ethyl alcohol.
The proper amount of 2 percent alcoholic stock solution was added
to the wastewater feed to achieve the desired concentration.
  Table 6 shows that after 48  hours of operation no ANT or DMP
could be detected in the final effluents of any of the seven reactors.
Trace amounts were  noted in the first  24 hour sample. The TOC
contents of the feeds to  Reactors 2 and 6 are elevated  above the
other reactors because of  the addition of the alcoholic solution to
achieve 1  mg/l ANT.
  The test procedure  was  replicated once each week for five weeks.
In no case was either ANT or DMP noted in the effluents collected
after 48 hours of operation.
      TABLE 6. BIODEGRADATION OF ANT AND DMP
Reactor number
Wastewater
feed, mg/l:
ANT
DMP
TOC
TKN
N03-N
Reactor effluent.
24 hr, mg/l:
ANT
DMP
TOC
TKN
N03-N
Reactor effluent,
48 hr, mg/l:
ANT
DMP
TOC
TKN
N03-N
1


0,13
-
19
13
0.3


0.00
-
3
1
10


0.00
-
5
0.8
10
2


0.97
-
35
13
0.1


0.02
-
7
1
9


0.00
-
7
1
8
3



0.08
18
12
0.1


-
0.00
3
1
11


-
0.00
3
1
10
4


-
0.89
17
14
0.1


-
0.01
5
2
13


-
0.00
3
0.8
13
5


0.10
0.10
20
12
0.1


0.00
0.00
6
1
10


0.00
0.00
4
0.9
10
6


0.93
0.85
38
11
0.1


0.01
0.01
5
0.09
9


0.00
0.00
7
0.8
8
Control


-
-
17
11
0.1


0.00
0.00
3
0.8
8


0.00
0.00
4
0.7
9
                            44

-------
  To ensure that the aeration solids did  not sorb the ANT or DMP,
after  one test  period the reactor  contents  were extracted  with
Freon-TF, Insignificant sorption  was noted. It is also unlikely that
the compounds were air-stripped from the reactors, on the basis of
the physical properties of ANT and DMP.
  Table 6 shows that ANT and  DMP singly  or  in combination at
concentrations of 0.1 or 1.0 mg/l do not interfere with the oxida-
tion efficiency of the microorganisms. The effluents contained low
residual TOC and nitrification was not inhibited.
                              45

-------
 Section  3. Treatability of Metals by Activated

                     Sludge Systems

  Extensive  treatability studies on the interaction of metals with
activated sludge systems have  been conducted at the U.S. Public
Health Service's Robert A.  Taft Engineering Center in Cincinnati.
  Continuous flow pilot plants with a flow capacity of 750 liters per
day were utilized. The design features of the pilot plants are shown
in Figure 19. This size pilot unit is sufficient to allow  study of
chemical distribution, biological response, and engineering design
features.
  These pilot units  proved to  be good  simulants of wastewater
treatment processes. They  were operated and maintained on a 24
hour, 7 day per week schedule,  with sustained  analytical super-
vision. Sufficient observations  were made to  establish statistically
valid  evidence  of performance  in  systems  with  metal input  and
metal withdrawal in general working balance. Over a 5 year period,
chromium,  copper,  nickel,  and  zinc were studied individually  and
in combination.
                               AIR HEADER
                                            CAPACITY: 7.9 gal
                                            DETENTION TIME: 2 hr
                                            SURFACE OVERFLOW RATE:
                                            102 gpd/ft2
        PRIMARY
        SETTLER
CAPACITY: 4.6 go!
DETENTION TIME: 1.2 hr
SURFACE OVERFLOW RATE:     SPIRAL-FLOW
142 9pd ft2                 AERATOR
                   CAPACITY: 23.6 gal
                   AERATION PERIOD: 6 hr
                   BOD LOADING: 0.5 TO 0.8 Ib 'day/lb
                   OF VOLATILE SOLIDS UNDER
                   AERATION
          Figure 19.  Plastic activated sludge pilot plant.
                              46

-------
  The wastewater feed  to the plant during the various studies was
either  a  weak supplemented  domestic wastewater or a  strong
nonsupplemented  domestic  wastewater. Both  feeds  give  results
indistinguishable by the usual analytic measures.

DISTRIBUTION OF METALS THROUGH THE  PROCESS

  Complete material balances of the metals were made during each
study. Table 7 summarizes these studies. The table is based  on the
amount of metal fed to a unit during a  compositing period.  Varia-
tion between compositing periods was common,  as indicated  by the
range of observations for the efficiency  of the process in removing
metals. The percent  metal unaccounted  for in Table 7 is not a firm
figure,  but represents the cumulative errors  involved  in sampling
sludges, flow measurements, and analytical methods.
  Metal balances were performed for each selected concentration of
the metals studied, not just those shown  in Table 7. Each metal was
studied in about five increments over the range  of 1 to 20 mg/l. In
addition,  four  metals were simultaneously traced during a combina-
tion study. Over the concentration ranges studied, no great  differ-
ence in the efficiency of the process in removing the metals was
noted. Zinc and copper, studied as the cyanide complexes, showed
the same overall removal as when studied as the sulfates.
  Chromium, introduced  to an activated sludge process as  hexa-
valent chromate, can show wide variation in concentrations  at the
various process outlets.  Reducing substances in the raw sewage can
cause precipitation of trivalent chromium with the primary sludge.
Also,  under anaerobic  conditions, the  organisms  in the  return
sludge  entering the  primary settler can  utilize  the oxygen  of the
chromate radical  and adsorb the  trivalent chromium  on the bio-
logical  floe.  Under these conditions chromium  removal  can reach
90 percent.
TABLE 7. DISTRIBUTION  OF METALS THROUGH ACTIVATED
       SLUDGE PROCESS WITH CONTINUOUS DOSAGE
% of metal fed

Outlet
Primary sludge
Excess activated sludge
Final effluent
Metal unaccounted for
Average efficiency of
process in removing
metal
Range of observations
Chromium (VI)
(15 mg/l)
2.4
27
56
15

44
18-58
Copper
(10 mg/l)
9
55
25
15

75
50-80
Nickel
(10 mg/l)
2.5
15
72
11

28
12-76
Zinc
(10 mg/l)
14
63
11
12

89
74-97
                             47

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  Table 7 shows that a considerable portion of the metal introduced
 is removed in the secondary sludge. The effects of the metals on the
 mixed liquor are apparent even in the 1 to 2 mg/l range. During 5
 years of study  no bulking was encountered in a metal-fed system.
 The floe in the final settler quickly settled. Control units frequently
 bulked. Table 8 shows the effects of  a combination of four metals
 on the  sludge density index and  volatile  solids  content of mixed
 liquor.
  With the exception of zinc, the conventional activated sludge pro-
 cess is not very efficient in the removal of metals from the influent
 stream.  The metal removed is concentrated at two points. In the
 primary sludge, a maximum concentration would occur if all the
 metal were removed with this sludge. Here the ratio of total flow
 volume to primary sludge volume  is a limiting  factor. Another
 point of concentration is in the secondary sludge. Since the volume
 of secondary sludge removed  from the process may be small com-
 pared to the  flow through the process, concentration  may be high
 at this point.
  There is  no net removal  of metal  if the primary and secondary
 sludges  containing the metals are not permanently removed from
 the line of flow.  For  instance, an extended aeration  plant passes
 all  the  metal to  the  receiving stream unless  secondary sludge  is
 removed.
  The copper, chromium,  and nickel discharged  with the final ef-
 fluent  from an activated  sludge  plant receiving these  metals are
 predominantly in a soluble form. At an influent concentration of
 10 mg/l, only a small amount of  zinc is discharged, and this is an
 insoluble zinc. At higher influent concentrations greater amounts
 of zinc were discharged as soluble zinc.

 EFFECTS OF METALS ON AEROBIC PROCESS

  Many  investigators of metal  toxicity have employed  batch opera-
tion or  direct dosing of the metal to the aeration chamber. Data
from individual studies (Table 9) show that primary settling has two
TABLE 8.  EFFECTS OF METALS ON MIXED LIQUOR SOLIDS
Analysis
Sludge density index
% Volatile solids

Control
unit
1.5
66.7
Mixed liquor from
Metal mixture
8.9 mg/l 4.9 mg/l 2.2 mg/l
3.2 3.4 2.4
57.9 61.8 63.8
                             48

-------
effects on the metals before entry into the aeration tank. First, the
total metal content of the primary effluent is less than that of the
influent because some  metal is removed with the primary sludge.
Second, the chemical and physical characteristics of the wastewater
alter the form of the soluble metal introduced. This was especially
true in the case of zinc where 90 percent of the added soluble zinc
was  converted to an insoluble form.  The differentiation between
soluble and insoluble metal in all  studies was  made by filtration of
the sample through an 0.45  /xn membrane filter and by acid diges-
tion of the filtrate before analysis.
  The procedure used to determine the concentration of metal in
the influent wastewater that would give a  barely detectable reduc-
tion in efficiency during the aeration phase of treatment is indicated
by Figure 20, which shows the results of a study of copper.
  During each run, data from an experimental pilot plant unit  and
a control  unit receiving no  metal were compared. The metal  was
added continuously to a constant  wastewater feed of the pilot plant
unit. Two weeks of acclimation  were  allowed before data on the
quality  of the final effluent were collected.  This time interval is
also required for the metal content of the activated sludge to build
up  to  a condition  of operating equilibrium.  Final effluents from
both units were assayed daily for  BOD, COD,  suspended solids,  and
turbidity. The run for any selected metal dosage was continued for
60 days to obtain sufficient data. The values for the two units were
then compared as frequency distribution curves. The parameter of
effluent quality in Figure 20 is COD;  this  is plotted as a frequency
distribution on arithmetic paper. As  shown  in the figure,  copper
present continuously at  0.4 mg/l did  not noticeably increase the
effluent COD of the experimental unit. A  copper concentration of
1.2 mg/l  however,  caused a significant increase in effluent COD.
From this and the other parameters measured, it is concluded that
copper present continuously at 1 mg/l in the influent is the thres-
hold dose for the aeration phase.

        TABLE 9.  METALS IN PRIMARY EFFLUENTS

                            Soluble metal
                            introduced in      Metal in primary
                           wastewater feed,    effluent, mg/l
Metal
Chromium (VI)
Copper
Nickel
Zinc
mg/l
50
10
2.5
10
Total
47
9
2.0
9
Soluble
38
3.0
1.0
0.6
                              49

-------
     200
      150 -
   ~3i  100 -
   Q
   O
   o
                                                         100
            10   20    30   40    50    60    70   80    90
             PERCENT OF OBSERVATIONS < STATED VALUE
Figure 20.  Effect of copper, fed continuously as copper cyanide
           complex, on COD of final effluents.
  It  is also useful to plot frequency distribution curves on proba-
bility paper. Readily available statistical measurements are given by
this  type of presentation. If a-straight line is obtained with arith-
metic probability paper, normal distribution of data is verified. The
50 percent point is  very close to the  true arithmetic mean of the
observations, and the slope of the line is a measure of the standard
deviation. Figure 21  is such  a plot of data collected during a study
of the effects of a mixture  of four metals on the activated sludge
process. The need for extensive sampling is shown here. The control
unit  had an average final effluent COD of 45 mg/l; however, contin-
uation of this point  to the experimental unit shows that 12 percent
of the time the experimental unit's final effluent had a COD of 45
mg/l or less.
  Copper and zinc are frequently used by the plating industry as
cyanide complexes. These two metals were studied in both the sol-
uble  cation form (as sulfate)  and as soluble cyanide complexes.
Results show that  once  the activated  sludge  acclimates  to the
continuous presence  of either form of the metal, there is no differ-
ence  in effects  on treatment efficiency. Figure 22A  shows that
where turbidity  of the final  effluent was used as the measure  of
treatment efficiency, after eight days the system receiving a 10 mg/l
                              50

-------
concentration of zinc as cyanide complex had acclimated to cyan-
ide and  was producing  effluent of stable turbidity. The cyanide
content  of  the  effluent followed a similar  pattern, with  almost
complete removal of cyanide at the end of seven days. Figure 22B,
from  a run with a 20 mg/l concentration of zinc as the sulfate,
showed  no  such acclimation.  Direct comparison  of the 10 mg/l
concentration of zinc  sulfate and a  10 mg/l  concentration of zinc
cyanide complex versus the same control unit is shown in Figure 23.
The BOD data collected after  two weeks of acclimation, showed
no significant difference between the two forms of zinc fed.
                METAL MIXTURE: Cr,Cu,
                Ni, Zn; TOTALING 8.9 mg/liter
 METAL
MIXTURE
                          ^0 30 40 50 60 70  80  90  95  98 99
              PERCENT OF OBSERVATIONS < STATED VALUE

                 Figure 21. COD of final effluents.
    IOO
    60
                Zn (CN)T
                COMPLEX, 10 mg/iiter
                                              —1	1	1	1

                                               B
                                               ZnS04, 20 mg,|,ler
      01  23456789  10 Ol  23456789  IO
                TIME, days                     TIME, days

 Figure 22. Comparison of acclimation to complexed zinc (A) and
           zinc sulfate (B).
                              51

-------
  Q
  O
  CD
                                  I	I
                           30  40  50  60
              PERCENT OF OBSERVATIONS < STATED VALUE
Figure 23.  Cumulative frequency data on quality of final effluents
           with zinc concentration of 10 mg/l in  wastewater feed.
  The reaction pattern of the activated sludge process was the same
for each of the metals studied. A small dose of metal gives a signifi-
cant reduction in treatment efficiency, but substantially larger doses
do not further decrease the efficiency greatly. Figure 24 graphically
illustrates this situation. One to two milligrams per liter yield a de-
tectable  response after which  a plateau is reached, and complete
failure does not  occur until one to two orders  of  magnitude are
reached.
  Table 10 lists the concentrations  of metals that give a significant
increase in the usual parameters applied  to judging treatment effic-
iency. These may be considered threshold concentrations. It should
be borne  in mind, however, that these  limits were obtained under
carefully controlled laboratory operation. The significance of Figure
24 is that the  threshold concentration is  mainly of academic, rather
than practical  interest and actual plant situations are concerned with
the plateau region of metal dosage and response.
                              52

-------
  I-
  LU
  o
  DC
  o
  z
  UJ
  u.
  UJ
100
     40
  LU
  or
20
  in
  tn
  O
            CONCENTRATION OF METAL IN INFLUENT SEWAGE

         Figure 24. Response of system to metal dosage.
TABLE 10.  CONTINUOUS DOSE OF METAL THAT Wl LL GIVE
    SIGNIFICANT REDUCTION IN AEROBIC TREATMENT
                        EFFICIENCY
     Metal
                                  Concentration in
                                 influent wastewater,
                                        mg/l
Chromium (VI)
Copper
Nickel
Zinc
                                         10
                                          1
                                        1-2.5
                                        5-10
  The results of these studies showed that the aeration phase of bio-
logical treatment can tolerate in the influent wastewater, chromium,
copper, nickel, and zinc up to a total heavy metal concentration of
10 mg/l,  either singly or  in combination, with about a 5 percent
reduction in overall plant efficiency.
  Slug doses of metals to the activated sludge process were also
studied. The concentration of  metal that constitutes a harmful slug
dose is determined by the waste volume, the volume and character-
istics of the dilution water, the specific  form of the metal, and the
usage of the stream below the point of effluent discharge. For conven-
ience, only a single measure of effluent  quality, such as an increase
in organic  material  passing through the plant, has been used to
                             53

-------
judge a harmful slug dose. As an example, in Figure 21, the control
unit has a  COD of 70 mg/l or  less 98 percent of the time; then a
harmful  slug dose can be defined as that concentration of metal
that will yield an effluent COD  in excess of this value for the sub-
sequent 24 hours of performance.  The effects of slug doses were
observed on 4-hr metal doses to the influent wastewater.
  Table 11  gives the results obtained. To fix  these concentrations
more accurately would require  an  inordinate  amount of time and
money. Table 11, therefore, is the best estimate of what concentra-
tion of metals causes  an exceptional displacement of treatment
plant performance as the result of a slug dose.
  Not reported in the table are the results  of slug studies in which
the metals  were added as cyanide  complexes. In  these cases, the
cyanide toxicity completely obscured the toxic effect of the metal.
In  general, acclimation of the  system to low concentrations of
metals or cyanide did not offer protection from slug doses.
  The inhibition of nitrification by heavy metals has been previously
studied with regard  to individual metals. A pilot plant that received
a combination of four metals also showed inhibition of nitrification.
There was no evidence of acclimation of the nitrifying organisms to
the metals.  The oxygen requirement of this metal-loaded sludge was
less than that of the control unit because oxygen for the biological
transformation of ammonia to  nitrate was not utilized. Figure 25
shows  the nitrate content of the final effluents of a control and
metal-fed unit. Inhibition of nitrification is  regarded as an important
effect of metal toxicity. A plant so affected would  discharge all the
influent nitrogen in  excess of that needed for  synthesis, predomin-
antly in the form of ammonia. Such an effluent would require con-
siderable chlorine if downstream breakpoint chlorination were used,
and nitrification  in the receiving stream would  use large amounts of
oxygen.
 TABLE 11. METAL CONCENTRATION IN 4-HR SLUG DOSE
   THAT WILL PRODUCE  HARMFUL SLUG, AS MEASURED
 _	BY COD	

                                     Concentration
    Metal                        influent wastewater, mg/l
Chromium (VI)                            >500
Copper                                       75
Nickel                                   >50<200
Zinc                                        160
                             54

-------
                            FINAL EFFLUENT OF PILOT PLANT
                            CONTROL UNI T
      FINAL EFFLUENT OF PILOT PLANT
      UNIT RECEIVING METAL MIXTURE?
                                    DO OF FINAL EFFLUENT
                                    OF PILOT PLANT UNIT
                                    RECEIVING METAL MIXTURE
                                I*-DO OF FINAL EFFLUENT OF
                                ', PILOT PLANT CONTROL UNIT
                                                           o
                                                           Q
          10    20    30    40    50    60    70    8O
                          TIME, DAYS

          Figure 25.  Nitrate nitrogen in final effluents.
EFFECTS ON ANAEROBIC DIGESTION

  The metal-bearing sludges  produced by the pilot plants were di-
gested in single stage non-mixed digesters.  Organic loading was for
non-mixed operation. A small circulating pump was used once each
day to obtain representative samples of sludge for material balances.
In each metal study both primary sludge and combined primary and
secondary sludges were digested. The metal contents of the sludges
fed to the digesters during several of the runs are given in Table 12.
The primary sludges were about 2 percent solids, and the secondary
sludges, about 0.5 percent solids during these studies. On a percentage
-of-solids basis, the metals in the secondary sludge are concentrated
to a much greater extent than in the primary sludge.
  A digester receiving combined sludges will contain more metal on
a percentage-of-solids basis than a digester receiving primary sludge,
when operated at the same  influent  metal concentration.  Digester
failure as a  result of heavy metals occurs at a lower influent metal
concentration  in  a combined sludge digester than  in  a  primary
sludge digester. The maximum continuous influent sewage metal
concentrations for  satisfactory  anaerobic  digestion  are given  in
Table 13.
                               55

-------
      TABLE 12. METAL CONTENT OF SLUDGES FED TO
                        DIGESTERS


Metal
Chromium
(VI)
Copper
Nickel
Zinc
Continuous dose in
influent wastewater
mg/l

50
10
10
10
Excess activated
Primary sludge
mg/l

330
280
62
375
sludge
mg/l

530
160
89
328
   TABLE 13. HIGHEST DOSE OF METAL IN CONTINUOUS
        DOSAGE THAT Wl LL ALLOW SATISFACTORY
           ANAEROBIC DIGESTION OF SLUDGES

                                    Concentrations in
                                 influent wastewater, mg/l
Metal
Chromium (VI)
Copper
Nickel
Zinc
Primary sludge
digestion
>50
10
>40
10
Combined sludge
digestion
>50a
5
>10a
10
a Higher dose not studied
  The response of the anaerobic system to metal dosage does not ex-
 hibit a plateau region as does the aeration phase; it is an all-or-none
 reaction. Digestion  either proceeds  normally  or ceases entirely.
 This may be more apparent than real, however, because  the analy-
 tical measures used to assess digester  performance are not as direct
 as those for the aeration phase.
  The  results of these metal studies show  that  in the cases of
 chromium,  nickel, and  zinc an influent metal concentration of 10
 mg/l, either singly or combined, will  not affect digestion. Copper
 continuously present at 10 mg/l causes failure of combined sludge
 digestion.
  The  prevailing  conditions of anaerobic digestion are such  that
 soluble metal introduced  with the feed sludges is efficiently  con-
 verted  to an insoluble form. This is shown in Table 14.
                            56

-------
  A few slug doses to anaerobic digesters were studied. The slugs to
the digesters were in conjunction with the aeration slugs. The sludges
produced by the activated sludge process during a metal slug were
collected and fed to satisfactorily operating digesters. In no case was
there  any  interruption  of  digestion  caused  by the  metal-bearing
sludges. Concentrations of  metals in the influent during these slug
studies are given in Table 15.
  More detailed studies were not conducted because the logistics of
digester operation  make it unlikely that an operating digester would
be upset by the sludges produced during a slug period. This belief is
based  on the facts that a digester is not on the main flow stream and
only a small part of the total flow through the plant reaches it, and
the daily additions to a digester are only  a fraction of the total
digester volume.
    TABLE  14. SOLUBLE METAL CONTENT OF SLUDGES
       COMPARED WITH TOTAL METAL CONTENT OF
                     DIGESTED SLUDGE
Soluble metal


Metal
Chromium (VI)
Copper
Nickel
Zinc
Concentration
in influent
waste water,
mg/l
50
10
10
10
Feed

Primary,
mg/l
38
2
10
0.3
sludges

Excess
activated,
mg/l
32
0.5
9
0.1

Digested
combined,
mg/l
3
0.7
1.6
0.1
Total metal

Digested
combined,
mg/l
420
196
70
341
      TABLE 15. DIGESTERS FED COMBINED SLUDGES
           PRODUCED DURING METAL SLUG TO
                ACTIVATED SLUDGE PLANT

Metal
Chromium (VI)
Copper
Nickel
Concentration of metal in
wastewater feed, mg/l
500
410
200
Effect on
digestion
None
None
None
                             57

-------
DISCUSSION

  The results of these studies show that for each phase of treatment,
aerobic or anaerobic, there are different bases for judging the con-
centrations of metals that are acceptable in the influent wastewater.
  The plateau-type response of the aeration phase shows that concen-
trations of metal many times higher than the threshold concentration
can be received  without greatly reducing efficiency. In a situation
in which  removal of organic matter is not critical, the most sensitive
performance criterion may be the ability of the digester to handle
the sludge  produced.  Generally,  biological  secondary treatment
processes can tolerate up to 5 mg/l of the inorganic toxicants without
noticeable impairment of treatment efficiency. The composition of
municipal wastewater and  the chemistry of the inorganic toxics is
such that disproportion  of the inorganics occurs during treatment
and the materials are conservative in nature. Thus, if the inorganics
enter the treatment facilities at concentrations of 1 to 5 mg/l in the
raw water and removal occurs in the system, to yield low effluent
residuals, the inorganics will be found  in concentrated side streams
such as primary sludge, waste  biological sludge,  digested sludge,
digester  supernatant,  or lagoon  bottom  sediment. Mainly,  the
inorganics will exist as  insoluble products in these side streams or
sludge  deposits.  Assessment  of  technology to enhance inorganic
toxics control must be based on  the overall environmental trade-off
of low final  effluent residuals versus concentration of the inorganics
in the sludge or sludge handling  operations of municipal treatment
systems and their subsequent fate during ultimate disposal practices.
  In other cases the amounts of metals passing through the plant to
the receiving stream may be the factor that determines the concen-
tration of metals permissible in the plant influent. Metals are  not
destroyed in wastewater treatment processes, so management prac-
tices must be based on their distribution in various process streams.
  A survey of four municipal treatment plants (Ref. 18, Chap.  10)
concerning the receipt of heavy metals, distribution of metals in the
process outlets, and effects of metals on treatment efficiency,  has
shown a pattern of response similar to pilot plant studies.
                              58

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               Section 4. Bibliography
GENERAL

  1.  Standard Methods Committee — Subcommittee on Biodegrad-
     ability. 1967.  Required  characteristics and measurement of
     biodegradability. Jour. Water Poll. Control Fed. 39(7): 1232-
     1235.
  2.  Bunch, R.  L.  1977. Criteria and assessment of waste treat-
     ability. Proc.  5th  U.S./Japan  Conf. on  Sewage Treatment
     Technology, Tokyo. 17 pp.
  3.  Ludzack, F. J. 1960. Laboratory model activated sludge unit.
     Jour. Water Poll. Control Fed. 32(6): 605-609.
  4.  Bunch, R.  L.,  and C. W. Chambers. 1967. A biodegradability
     test for organic  compounds. Jour.  Water Poll. Control Fed.
     39(2): 181-187.

TREATABILITY OF SPECIFIC ORGANIC COMPOUNDS

  5.  Bunch, R.  L.  1976. Effects and removal  of toxic substances
     on/by  conventional  biological treatment systems. EPA Toxic
     Substances Seminar, Washington, D.C., October 1976. 10pp.
  6.  Moore, L., and E. F. Barth. 1976. Degradation of NTA acid
     during anaerobic digestion. Jour. Water Poll. Control Fed.
     48 (10): 2406-2409.
  7.  Tabak, H. H., C. W. Chambers, and  P. W.  Kabler. 1964.
     Microbial  metabolism  of aromatic compounds.  I.  Decompo-
     sition  of phenolic compounds and aromatic hydrocarbons
     by phenol-adapted bacteria. Jour. Bacteriol. 87(4):910-919.
  8.  Chambers,  C. W., H. H. Tabak, and P. W. Kabler.  1963. De-
     gradation of aromatic compounds by phenol-adapted bacteria.
     Jour. Water Poll.  Control Fed. 35(12):  1517-1528.
  9.  Tabak,  H.  H., C. W.  Chambers, and  P. W.  Kabler. 1959.
     Bacterial utilization of lignans. I. Metabolism of  a-conidendrin.
     Jour. Bacteriol. 78(4): 469-476.
 10.  Bunch, R.  L.,  and M.  B. Ettinger. 1968. Biodegradability of
     potential  organic substitutes  for  phosphates.  Proc.  22nd
     Ind. Waste  Conf., Purdue  Univ.,  Ext. Ser.  No.  129:393.
                            59

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 11.  Barth, E. F.,  and M. B. Ettinger. 1967. Anionic detergents
      in  wastewater  received  by  municipal  treatment  plants.
      Jour. Water Poll. Control Fed. 39(5): 815-822.
 12.  Tabak, H.  H.,  R. N.  Bloomhuff, and  R. L. Bunch. 1972.
      Coprostanol:  a  positive tracer of fecal pollution. Develop.
      Ind. Microbiol. 13: 296-307.
 13.  Tabak, H.  H.,  and  E.  F.  Barth.  1978. Biodegradability of
      benzidine in aerobic suspended growth reactors. Jour. Water
      Poll. Control Fed. 50: 552-558.
 14.  Tabak, H. H., and R. L. Bunch.  1970.  Steroid hormones as
      water pollutants.  I.  Metabolism  of  natural  and synthetic
      ovulation-inhibiting  hormones  by  microorganisms  of  ac-
      tivated sludge  and  primary settled  sewage.  Develop.  Ind.
      Microbiol. 11: 367-376.
 15.  Barth, E. F., et  al. 1978. Biodegradation studies of carboxy-
      methyl  tartronate.  Municipal  Environmental  Res.  Lab.,
      Cincinnati, Ohio, EPA Rept. No. EPA-600/2-78-115,
 16.  Bunch,  R.  L,  and C.  W, Chambers.  1967. A biodegrad-
      ability test  for organic compounds. Jour. Water Poll.  Control
      Fed., 39(2): 181-187.
 17.  Tabak, H. H., and E. F. Barth. 1979. The microbial degrada-
      tion of naphthalene and phthalate ester compounds. In press.

METALS

 18.  U.  S. Department of Health, Education, and Welfare, Public
      Health Service,  Water Supply and Pollution Control. 1965.
      Interaction  of heavy metals and biological sewage treatment
      processes. Rept. 999-WP-22, Cincinnati, Ohio.
 19.  Barth, E. F,,  M. B.  Ettinger, B.  V.  Salotto, and G. N. Mc-
      Dermott. 1965. Summary  report on the effects of heavy
      metals  on  the  biological treatment  processes.  Jour. Water
      Poll. Control Fed. 37:86.
 20.  Mulbarger,  M. C., and J.  A. Castelli. 1966. A  versatile ac-
      tivated sludge pilot plant—its design, construction and opera-
      tion.  Proc. 21st  Ind. Waste  Conf., Purdue Univ., Ext. Ser. No.
      121:322.
                             60

                            T U.S. GOVERNMENT PRINTING OFFICE: 1979—657-060.

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