PROCEEDINGS
     1998 WATER AND WATERSHEDS
            PROGRAM REVIEW
               28-29 January 1998
                Corvallis, Oregon

Sponsored by the EPA/NSF Partnership for Environmental Research

          \  NATIONAL SCIENCE FOUNDATION
      ^60 ST4t.
            ENVIRONMENTAL PROTECTION AGENCY


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Cover Photo:  Aerial photograph of the White Clay Creek watershed near the Stroud Water
Research Center, Chester County, Pennsylvania. The photo, taken after a snowstorm, shows patches
of forest (dark areas) interspersed among deforested meadow or cropland (white) areas.  This region
of southeastern Pennsylvania contains many of the study streams for Sweeney et al.'s study entitled
"Streamside Reforestation: An Analysis of Ecological Benefits and Societal Perceptions" (see page
38).  Photograph from Stroud Center archives; R.L. Vannote.

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                PROCEEDINGS
     1998 WATER AND WATERSHEDS
           PROGRAM REVIEW
               28-29 January 1998
                Corvallis, Oregon
Sponsored by the EPA/NSF Partnership for Environmental Research
          ENVIRONMENTAL PROTECTION AGENCY
          NATIONAL SCIENCE FOUNDATION

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                                          Table of Contents

Introduction      ....        ...           .                         .    .          .        .      vii

Section 1.  Projects Initiated With Fiscal Year 1997 Support

Community Values and the Long-Term Ecological Integrity of Rapidly Urbanizing Watersheds           .        2
        M. Bruce Beck, T.C.  Rasmussen, B.C. Patten, K.G. Porter, B.C. Norton, A. Shepherd

Connecting Ecological and Social Systems:  Watershed Research Relating Ecosystem Structure
        and Function to Human Values and Socioeconomic Behaviors                  .             ...  3
        Gaboury Benoit, S. Kellert, M. Ashton, P. Barten, L. Bennett, D. Skelly

Social and  Ecological Transferabiliry of Integrated Ecological Assessment Models              ...           .4
        Linda A. Deegan, James Kremer, Thomas Webler

From Landscapes to Waterscapes:  Integrating Framework for Urbanizing Watersheds               .            6
        PanosDiplas,  W.E. Cox,  D.F. Kibler, V.K. Lohani, R.G. Greene, D.J. Bosch, L.A. Shabman,
        K. Stephenson, E.F. Benfield,  P.S. Nagarkatti, S. Mostaghimi, D.J. Orth

An Integrated Ecological and Socioeconomic Approach To Evaluating and Reducing Agricultural
        Impacts on Upper Mississippi  River Watersheds                                            .        8
        Prasanna H. Gowda, R.J. Haroa, A.D. Ward, T.L. Napier

Integrated Ecological-Economic Modeling of Watersheds and Estuaries at Multiple Scales                     10
        Charles Hopkinson, Edward Rastetter, Joseph Vallino

Linking Watershed-Scale Indicators of Changes in Atmospheric  Deposition to Regional Response Patterns       11
        Jeffrey S. Kahl, I. Fernandez, D. Mageean, S. Ballard, S. Norton, J. Cosby, P. Ludwig, L. Rustad

Comprehensive Watershed Management: A Spatial Water Quality Assessment System                        12
        C. Gregory Knight, Robert P.  Brooks, Barry M. Evans, James M. Hamlett, Archie J. McDonnell,
        Gary W. Petersen,  Todor N. Hristov

Development and Implementation of Decision Support Systems for Predicting Economic and Ecologic
        Impacts of Alternative Land and Water Management Policies in Urbanizing Regions                  14
        Daniel P. Loucks, Tammo S. Steenhuis, Mark B. Bain,  Warren Brown, Walter R. Lynn

Effects of Natural and Anthropogenic Processes on Tillamook Bay and Its Watershed: An Integrated
        Process Study and Land Use Perspective                                                         16
        James McManus, Paul Komar, M.S.  Jesse Ford, Courtland Smith, Debbie L. Colbert, Greg Bostrom

Ecological  Risks, Stakeholder Values, and River Basins: Testing Management Alternatives for the
        Illinois River             . .                              .    .                                 17
        Mark Meo, Baxter Vieux,  James Sipes, Edward T. Sankowski, Robert Lynch, Will Focht,
        Keith Willett, Lowell  Caneday

Risk-Based Urban Watershed Management:  Integration of Water Quality and Flood  Control Objectives         19
        Vladimir Novotny, Robert Griffin, David Clark, Douglas Booth

Impact of Social Systems on Ecology and Hydrology in Urban-Rural Watersheds: Integration for
        Restoration                                                                                   20
        Steward T.A. Pickett

Section 2.  Projects Initiated With Fiscal Year 1996 Support

Strategic Renewal of Large Floodplain Rivers: A Preliminary Status Report        ...         .         .25
        John B. Braden, Misganaw Demissie, Eric DeVuyst, Paromita Mitra, Daniel Schneider,
        Richard E. Sparks, David C. White, Renjie Xia
                                                   in

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                                   Table of Contents (continued)

 Integrating Modeling and Management of Agriculturally Impacted Watersheds:  Issues of Spatial
        and Temporal Scale    .        .  .         	      .  .    .        .   .     26
        Patrick L. Brezonik, K. William Easter, David Mulla, James A. Perry

 Urban Stream Rehabilitation in the Pacific Northwest:  Physical, Biological, and Social Considerations  ...   28
        Stephen J. Surges, Derek B. Booth, Sally Schauman, James R. Karr

 Influences of Forest Fragmentation on Watershed Functions in Northern Vietnam—Preliminary
        Field Results     	             . .  29
        Jeffrey Fox, T. Giambelluca, A.T. Rambo

 Geochemical, Biological, and Economic Effects of Arsenic and Other Oxyanions on a Mining
        Impacted Watershed  . .        	     ....        .        	        	  31
        Glenn C. Miller, Watkins W. Miller, Scott Tyler, Douglass Shaw, Ron Hershey, Lambis Papellis,
        Susan Anderson

 Effectiveness of Regulatory Incentives for Sediment Pollution Prevention: Evaluation Through Policy
        Analysis and Biomonitoring         	            ...           .   .                    32
        Seth R. Reice, Richard N. Andrews

 Watershed Protection in Agricultural Environments: Integrated Social, Geomorphological, and
        Ecological Research To Support Ecosystem-Based Stream Management                       . .       33
        Bruce L. Rhoads, Edwin E. Herricks, David Wilson

 Towards an Integrated Regional Model of River Basins of the Western Pacific Rim .                  ...  34
        Jeffrey E. Richey

 An Integrated Approach To Assessing Water Management Options in a Major Watershed:  Extending
        a Hydrodynamic-Water Quality Model To  Include Biological and Politico-Economic Components       35
        Paul Sabatier, Loo Botsford, Mike Johnson, Jay Lund, Peter Moyle, Gerald Orlob, James Quinn,
        Peter Richerson, Tom Suchanek, Marca Weinberg

 Modeling Effects  of Alternative Landscape Design and Management on Water Quality and Biodiversity
        in Midwest Agricultural Watersheds . .            	       	        . .        .           36
        Mary Santelmann, K. Freemark, D. White, S.  Polasky, G. Matzke, J. Eilers, J. Bemert,
        B. Danielson, R. Cruse, J.  Nassauer, S.  Galatowitsch

 Streamside Reforestation: An Analysis of Ecological Benefits and Societal  Perceptions    .        .            38
        Bernard  Sweeney, Thomas Bott, John Jackson, Louis Kaplan, J. Denis Newbold, Laurel Standley,
        Richard Horwitz, W. Cully Hession, Janet  Johnson, James Finley, Caren Glotfelty, Cecilia Ferreri

 Integrated Urban Watershed Analysis: The Los Angeles Basin and Coastal Environment    ...            .40
        Richard P. Turco

 Section 3.  Projects Initiated With Fiscal Year 1995 Support

 Development and Application of Spectroscopic Probes for Measurement of Microbial Activity in
        Aquatic Ecosystems    ....     . .      ...     . .    . .       .         .            	  45
        Carol Arnosti, Neil V.  Blough

Watersheds and Wetlands:  Large-Scale Disturbances and Small-Scale Responses        .      .           .47
        Charles Andrew Cole,  Robert P. Brooks, Denice Heller Wardrop

Integrated  Ecological Economic Modeling and Valuation of Watersheds  ....       ...         .     .49
        Robert Costanza, RoelofBoumans,  Tom Maxwell, Ferdinando Villa, Alexey Voinov,
        Helena Voinov, Lisa Wainger
                                                   IV

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                                   Table of Contents (continued)

Oyster Reefs as Structural and Functional Components of Tidal Creeks:  An Ongoing Ecosystem
        Experiment	      ...                 .  .                            ...  51
        Richard F. Dame, E. Koepfler, L. Gregory, T. Prins, D. Allen, D. Bushek,  C. Corbett, D. Edwards,
        B. Kjerfve, A. Lewitus, J. Schubauer-Berigan

Tracing the Fate of Nitrogen Inputs From Watersheds to Estuaries                             .             53
        Linda A. Deegan, Bruce J. Peterson

Probing the Relationship Between Fulvic Acid Aggregation, Metal Ion Complexation, and the Binding
        of Organic Compounds     ...          	          	                55
        A. Dixon, W.R. Carper,  C.K. Larive

Diffusional Rate Limitations hi Heterogeneous Porous Media: Model Structure, Scale, and Geologic
        Characterization	      	     	         	      56
        David L. Freyberg,  Paul V. Roberts

Integrating Planning, Forecasting, and Watershed Level Ecological Risk Assessment Techniques:
        A Test in the Eastern Cornbelt Plains Ecoregion                                                  57
        Steven I. Gordon, Andy Ward, Dale White

Development of Geomorphological Artificial Neural Networks (GANNs) for Modeling Watershed Runoff ...  59
        Rao S. Govindaraju

Physicochemical Mechanisms Governing Virus Filtration               ...                               60
        Stanley B. Grant,  Terese M. Olson, Mary K.  Estes

Watershed Impacts on Sediment Pollution History and the Viability of the Zooplankton Egg Bank               61
        Nelson G. Hairston, Jr., Colleen M. Kearns, Charles T. Driscoll

The Role of Colloidal Particles in the Transport of Chemicals Through an Agricultural Watershed   .  .          62
        George M. Hornberger, Janet S. Herman, James E. Saiers

Geomorphic, Hydrologic, and Ecological Connectivity in Columbia River Watersheds:  Implications for
        Endangered Salmonids                .               ...              ....          .          64
        Hiram W. Li, Bruce A. Mclntosh, J. Boone Kauffman, Judith L. Li, Robert L. Beschta,
        Patricia McDowell

Resistance of Communities to Chronic Haloaromatic Contamination From Biogenic and Anthropogenic
        Sources  .   .       ....                     	              . .             . .         65
        David E. Lincoln, Sarah A.  Woodin, Charles R. Lovell, V. Pernell Lewis

Influences of Watershed Land Use on Stream Ecosystem Structure and Function                              66
        Judith L. Meyer, E.A.  Kramer, M.J. Paul, W.K. Taulbee, C.A. Couch

The Role of Hg (II) Reduction and Chemical Speciation in Controlling the  Concentration of Mercury
        and Its Methylation in  Natural Waters         	           .       .         .  .  68
        Franfois MM. Morel

Formation and Propagation of Large-Scale Sediment Waves in Periodically Disturbed Mountain
        Watersheds  ...           ....                .                                          69
        Gary Parker

Multiscale Statistical  Approach  to Critical-Area Analysis and Modeling of Watersheds and Landscapes    ...    70
        G.P Patil, W.L. Myers

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                                   Table of Contents (continued)

 Contemporary Water and Constituent Balances for the Pan-Arctic Drainage System: Continent to
         Coastal Ocean Fluxes	          .                          .      .     .   .     72
         Bruce Peterson,  Charles Vorosmarty, Richard Lammers

 Modeling Temporal Rainfall via a Fractal Geometric Approach          .             .                     .74
         Carlos E. Puente

 Effects of Food Web Structure and Nutrient Loading on Lake Productivity and Gas Exchange With
         the Atmosphere	      . • •      -  •                  •  • • •        •  •     .  .  75
         Daniel E. Schindler,  Stephen R. Carpenter, James F. Kitchell, Jonathan J. Cole, Michael L. Pace

 A Comparative Institutional Analysis of Conjunctive Management Practices Among Three Southwestern
         States  .               	                                           ......  76
         Edella Schlager

 Water and Sustainable Development in the Binational Lower Rio  Grande/Rio Bravo Basin         .             77
         Jurgen Schmandt

 Environmental Change and Adaptive Resource Markets:  Computer-Assisted Markets for Resource
         Allocation       .                                           .                                .79
         Vernon Smith, S. Rassenti, E. Hoffman, R. Howitt, A. Dinar

 Detecting Fecal Contamination and  Its Sources in Water and Watersheds                   .         .       .80
         MarkD. Sobsey

 Ecoregion-Specific Comparison of Stream  Community Responses to Nutrient Gradients Using Both
         Survey and Experimental Approaches  .        .                                      .81
         R. Jan Stevenson, Mike Wiley, Joe Holomuzki

 Holocene Floodplain Development as a Function of Climate Change and Human Activities:
         The Arroux and  Loire Rivers, Burgundy, France                                   .               82
         Eric C. Straffin,  Michael D. Blum

 Response and Compensation to a Bivalve Invasion by an Aquatic  Ecosystem                                  83
         David L. Strayer, Nina  Caraco, Jonathan J. Cole, Stuart Findlay, Michael L. Pace

 Scaling Up Spatially Distributed Hydrologic Models of Semi-Arid Watersheds    .                        .     84
         David G. Tarboton, Christopher M. U. Neale, Keith R. Cooley, Gerald N.  Flerchinger, Clayton L. Hanson,
         Charles W. Slaughter, Mark S.  Seyfried, Rajiv Prasad, Charlie Luce, Greg Crosby, Changyi Sun

 Traveling Wave Behavior During Subsurface Transport of Biologically  Reactive Contaminants:
         Implications for In Situ  Bioremediation                           .                           .      85
         Albert J. Valocchi

 Carbon Exchange Dynamics in a Temperate Forested Watershed (Northern Michigan):  A Laboratory
         and Field Multidisciplinary Study        .                       .                .                .87
         LynnM. Walter,  L.M. Abriola,  J.M. Budai, G.W. Kling, P.A. Meyers, J.A. Teeri, D.R. Zak

 A Comparison of Agricultural vs. Forested Basins:  Carbon and Nutrient Cycling Within the
         Hyporheic Ecotone of Streams                                                                   89
        David S.  White, Susan P. Hendricks,  Timothy C. Johnston, George Kipphut, William E. Spencer

In Situ Assessment of the Transport  and Microbial Consumption of Oxygen in Groundwater                  .  99
         Tadashi Yoshinari, R.L. Smith, J.K. Bohlke, K. Revest

Index of Authors .  .                                                                                   91
                                                   VI

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                                       Introduction
The  U.S. Environmental Protection  Agency / National  Science Foundation (EPA/NSF) Water  and
Watersheds competition  is one of three  special extramural  awards  competitions  supported  by the
EPA and the NSF under a partnership for environmental research initiated in 1994.

The  competition emphasizes interdisciplinary  research taking a systems approach  to issues of water
and watersheds. Its goal is to 1) develop an improved understanding of the natural and anthropogenic
processes that  govern the quantity, quality, and availability of water resources in natural and human-
dominated systems, and 2) improve understanding of the structure,  function,  and dynamics  of the
terrestrial and  aquatic ecosystems that comprise watersheds.

The  1995  Water and Watersheds competition  reviewed  656  proposals requesting ~$335M and made
36 awards. In  1996, the competition announcement was narrowed to  focus more on interdisciplinary
research—as well as to limit the number of proposals. The 1996  competition reviewed 249 proposals
requesting ~$168M and 12 awards were made. The agencies  further  narrowed  the  announcement in
1997 primarily in  response to concerns that the competition had too low a success rate. Proposals
were required  to integrate physical, ecological and social science  research and, for  the first time, the
investigators were  encouraged  to take  a community-based approach. The  1997  competition,  with an
emphasis on urban/suburban research, reviewed 128 proposals  requesting ~$77M and made 13 awards.

The  abstracts  in this volume are organized alphabetically, within three  sections that correspond to
the year of award.  The most recent awards (FY97) are first. Many of these projects were just  getting
underway  as this publication went to press  and the  abstracts indicate  goals and  plans. The  FY96
cohort of projects  appear next. These abstracts  report early  findings  and plans for  future years. The
projects that were initiated with FY95 support are in the third section. These abstracts  report results
based on several years of research.

The  competition is currently  entering  its fourth  year,  and the U.S. Department of Agriculture
(USDA) lias joined the two original agencies in supporting the effort.  The theme for the fourth year
competition is rehabilitation of watersheds. Program reviews  such as this one will allow investigators
to interact  with one another,  and to discuss progress and findings with EPA, NSF, USDA and other
federal officials who are interested in the program.

Any opinions, findings, conclusions, or recommendations expressed in this  report are  those  of the
investigators that participated  in the research  and the Program  Review meeting, and not necessarily
those of the  NSF or the  EPA.  For further information on the  EPA/NSF Water and Watersheds
competition   please   contact   the   Program   Officers:   Ms.    Barbara   Levinson,    EPA,
levinson.barbara@epamail.epa.gov or  Dr. Penelope Firth, NSF, pfirth@nsf.gov.
                                          vn

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               Section 1.
Projects Initiated With Fiscal Year 1997 Support

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 Community Values and the Long-Term
 Ecological Integrity of Rapidly  Urbanizing Watersheds
 M. Bruce Beck and T. C. Rasmussen
 Daniel B. Warnell School of Forest Resources, University of Georgia, Athens, GA
 B.C. Patten and K.G. Porter
 Institute of Ecology, University of Georgia, Athens, GA
 B.G. Norton
 School of Public Policy,  Georgia Institute of Technology, Atlanta, GA
 A. Shepherd
 City Planning Program,  College of Architecture,  Georgia Institute of Technology, Atlanta, GA
                Untoward Disturbances
               Controls/Policy
                                         THE LAKE/WATERSHED

                                             Natural Science Model
                                             (Hydrology; Ecology)
                                                                    Short-Term Response
                                                                    Long-Term Response
                                        Desired Short-
                                        Term Response
                          Potential Changes of
                           Model Structure
                                            THE COMMUNITY

                                              Belief Networks and
                                            Plurality of Mental Models
                                            (Social/Policy Sciences)
             .Mismatch & Dissonance
                                                                                              Desired Long
                                                                                              Term Response
 Figure 1. Schematic representation of the interaction between natural and social/policy science models in which the tasks are to identify: (a) on
         which critical, key unknowns in the natural science base may hinge the reachability of feared/desired futures; and (b) how community
         and stakeholder concerns for the future change in the light of changing information about the hydrological and ecological status of the
         lake and its watershed.
       Watersheds surrounding metropolitan Atlanta are
 expected to experience substantial development during
 the next two decades. Indeed, Atlanta's continued rapid
 economic development could be constrained by pro-
 blems of access to sufficient water supplies from the
 relatively small headwater catchments  to  its north.
 Within these watersheds lies Lake Lanier, which is used
 for hydroelectric power generation, water supply, flood
 protection, and most significantly, recreation.
       The investigators  are developing a new approach
 in  which both community interests  and  a complex
 (mathematical)  representation of the lake's ecosystem
 can be engaged in exploring how shorter term individual
 preferences  can  be  reconciled with  longer  term
 community values regarding the maintenance of the
 integrity  of  an environmental  system. The primary
 hypothesis is  that citizens'  value  commitments  are
 dependent upon scale (both in time and place) and that
 these scales have their counterparts in the range of time-
 constants  typifying the behavior of the physical system:
 the short  term, spanning 0-5 years; and the long term,
 more than 20-100 years.  The secondary hypotheses are
 that:  (1)  radical  shifts of behavior,  as gauged by
 qualitatively different patterns of model outputs (such as
populations of  fish),  may be a function of  slowly
evolving  changes in the  values  of the model's  co-
efficients (i.e., growth, mortality, and predation rates);
and (2) longer term change, and hence maintenance of
the integrity of a system, is primarily a function of such
coefficient variations.
      Adaptive community learning will be facilitated
by working  with small groups  of stakeholders. The
procedure will comprise several iterations around the
cycle of  eliciting community values, allowing  their
modification, encoding stakeholder concerns about the
long-term future (as target features to be matched by the
model's simulation results), identifying consensus  on
policy  options  and/or  reconciling discord between
longer and shorter term aspirations, and generating in-
sight into  the attainability of the target futures through
the scientific model (see Figure 1). The model will be
developed from past studies of Lanier and its watershed
and will incorporate new material for description of the
microbial  food web  and sediment biochemistry, the
latter being supported by modest field and laboratory
work. The model will be used principally within the
computational  framework  of a sensitivity analysis
employing Monte Carlo simulation.
      In addition to developing and testing a prototype
approach to facilitating adaptive  community learning,
with special reference  to preserving the attainability of
goals on an intergenerational time  scale, this project will
develop a new model for the dynamics  of an ecosystem
in a southeastern impoundment and policy guidance on
options for maintaining the longer term integrity of such
a system.

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Connecting Ecological and Social Systems:
Watershed Research Relating Ecosystem Structure
and  Function to Human Values and Socioeconomic Behaviors
Gaboury Benoit, S. Kellert, M. Ashton, P. Barten, L. Bennett, and D. Shelly
Yale Environmental Studies, New Haven, CT
                                   Ecosystem
                                    Health
                                Primary Productivity
                               Organic Matter Cycling
                               Biogeochemical Cycling
                               Hydrologic Regulation
                                 Energy Processing
                                   Biodiversity
           Human
      Environmental Values
     Aesthetic    Dominionistic ^
     Humanistic   Naturalistic
     Negativistic   Moralistic   I
    v Scientific    Symbolic   /
          Utilitarian
                                              Socioeconomic Benefits
                                              Property Values
                                              Material Production
                                              Consumption Patterns
                                              Outdoor Recreation
                                              Water and Land Use
                                              Intellectual Skills
                                              Sense of Place
                                              Community Stability
    Figure 1. Hypothesized feedback loop linking ecosystem structure and function, environmental values, and Socioeconomic benefits.
       The goal of this project is to examine how eco-
 logical and social systems influence each other through
 either positive or negative feedbacks within watersheds.
 The central hypothesis is that the relative health and in-
 tegrity of ecosystems cause, and in turn, are caused by
 enhanced human performance and productivity. Con-
 versely,  damaged and degraded ecosystems cause, and
 in turn, are caused by diminished human performance
 and productivity.
       The  current  research will use  observational
 techniques integrated by  standard statistical methods to
 measure the quantifiable linkages between biophysical
 and social systems. This investigation also will lay the
 groundwork for a field experiment to test whether  re-
 storing a degraded watershed can significantly enhance
 social interactions in associated human communities.
 State-of-the-art  analytical methods  will be  used to
 characterize approximately 30 sub-watersheds in terms
 of key hydrological, chemical, biological, economic,
 and social parameters. This large number of biophysi-
 cal and  social measures will  be  integrated within a
 quantitative context that allows  us  to test our pre-
 dictions and to generate an overall index of watershed
 quality.
       The first step has been to select study sites from
 among an initial set of more than 60 possible candidates.
 The investigators see objective selection of appropriate
 sites as crucial to the  long-term success  of this project.
 The final set of sub-watersheds need to fulfill several
 criteria:  (1) relatively homogeneous  land use or land
cover in each;  (2) all large enough to include a sufficient
number of residents to yield robust statistics on social
science surveys,  even in areas  having the minimum
regional  population  density; (3) adequate in size to
ensure year-round streamflow; (4) all sites within  +/-
50 percent of the mean area; (5) representative of the
entire range of development from rural to urban; and (6)
covering the full  spectrum of environmental quality
from nearly pristine to badly degraded.  Because these
last two characteristics are not entirely correlated, a set
of sites are being  sought that includes representatives
having: (1) a high  level of development with good
environmental quality  and (2) low development with
poor environmental quality as well as the more common
converses of these two.
      Every sub-watershed within our three river basins
fulfilling criteria 1 through 4 have been delineated on
24,000 scale USGS topographic maps and all ambiguous
boundaries clarified  through field visits.  Preliminary
land cover and land  use have been calculated for each
sub-watershed, allowing them to be ranked as a test of
criterion 5. Final land use characteristics will be de-
rived from an existing  detailed geographic information
systems' database. Rapid assessments of water quality
and biological characteristics are being conducted to
evaluate criterion 6.
      Also, we  are  in the process of designing  and
testing surveys to evaluate human attitudes, values,
knowledge, and behavior towards nature and economic
conditions within the  watershed.  The process of select-
ing sub-watershed study sites  also has  required us to
begin examining in detail how to reconcile differences
between physical watershed boundaries and political or
other human social boundaries.

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Social and Ecological Transferability
of Integrated Ecological Assessment Models
Linda A. Deegan
The Ecosystems Center,  The Marine Biological Laboratory, Woods Hole, MA
James Kremer
University of Connecticut, Avery Point, CT
Thomas Webler
Social and Environmental Research Institute, Leverett, MA
      The coastal zone, near-shore waters and adjacent
uplands, is heavily used for a variety of purposes—not
all of which are compatible.  In past years, urban and
suburban development in the uplands have been ac-
companied by  profound changes in ecosystems with
declines in fish and shellfish harvests.  However, people
are reluctant to believe mat what occurs  in their back-
yards can affect the health of the sea.  This project will
develop  a general  integrated ecological assessment
model for coastal watersheds. This model will be used
to determine how the level of involvement with the
model by managers and planners affects their acceptance
of it  as  a useful management tool.  Our project  will
determine how best to make the model  relevant  and
useful to local citizens in their decisions regarding land
use management in  the coastal zone.  Two interrelated
issues will be addressed: (1)  the ecological transfer-
ability of the model, the model's ability to capture the
dynamics of a previously unconsidered ecosystem; and
(2)  the   social transferability  of  the  model,  the
willingness of local managers  and planners to use the
integrated planning model.
      An existing watershed-loading model will be used,
and an estuary model will be extended to include a new
and socially important management endpoint—fish and
shellfish. This new integrated assessment model will be
used  in  a series of experiments  designed to  test
simultaneously the ecological and social transferability of
the model in  a variety of new  settings.   Empirical
relationships will be developed between predicted N-load
and  endpoint  responses,  using  new field data  and
literature information. This project's research involves
using replicated experiments to determine the effect of
familiarity with  and involvement in the  model  on its
acceptance and usefulness by the  planning community.
The  effect of  four treatment conditions on  model
acceptance will be tested (see Figure 1): (1) explanation
of the model by an "expert," (2) participation  in a
learning workshop ("Wkshp I"); (3) a learning workshop
supplemented with intensive dialogue ("Wkshp II"); and
(4) a learning workshop supplemented with a citizen
field-data collection program ("Wkshp + Field").
      There  is  an urgent  need  for an  estuarine
ecological model of broad application, incorporating
components that  are of direct concern to the public.  It
is not possible to build "from scratch" an integrated
ecological model for each  community using local re-
searchers and local data.   Most  communities do not
have  the local researchers nor the data  needed,  and
society cannot afford the money nor the  time to fund
de  novo  model  development for each  local case.
Therefore, planners must  rely on models developed
elsewhere to  be  applied to their  local situation.   It is
common knowledge,  however,  that  people distrust
policy recommendations for their community when
they are based on data gathered elsewhere. One of the
most  interesting  results will be to explain how much
gain  in model   acceptance is achieved  by differing
levels of training and familiarity building.   Conclusions
will  be  drawn  about  whether  the  existing  models
provide information that community policymakers  find
useful and relevant.  This research project is designed
to benefit urban/suburban coastal  communities interest-
ed in protecting estuarine  ecosystems from nitrogen
loading.  Integrated  ecological assessment  models have
the potential to  improve  the competence of  local
communities in making policy decisions.  This research
will  result in an integrated ecological model of the
consequences of coastal land use change  on estuarine
systems and,  perhaps  more importantly,  better infor-
mation on how  to apply that  model to new environ-
mental and social settings.

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 CD
 0
 c
 CL
 0)
 O
 O
•u
 o
           Expert      Wkshp I     Wkshp II    Wkshp +  Field

                             increasing interaction


                                Treatment
    Figure 1. Hypothetical response of subjects to increasing intensity of interaction with the model. Dependent variable is tine

          degree of acceptance of the model as a useful planning tool.

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From Landscapes to Waterscapes:
An Integrating Framework for Urbanizing Watersheds
Panos Diplas,  W.E. Cox, D.F. Kibler, V.K. Lohani, and R.G. Greene'; D.J. Bosch, L.A. Shabman, and K.
Stephenson 2; E.F. Benfield, and P.S. Nagarkatti3; S. Mostaghimi *; and D.J. Orth 5
' Department of Civil Engineering;2 Department of Agricultural and Applied Economics;3 Department of
Biology;4 Department of Biological Systems Engineering;5 Department of Fisheries and Wildlife Sciences,
Virginia Polytechnic Institute and State University, Blacksburg, VA
      A generally adverse relationship between urban-
ization  and water resources  conditions is  normally
assumed.   However, the variability of impacts  from
alternative landscape scenarios on water supply, flood-
ing, and the ecological status of waters  within a water-
shed subject to urbanization/suburbanization is not well
understood.  Approaches and  methods for analyzing
cause-and-effect relationships  from  a  comprehensive
perspective are not widely available.
      Land settlements  within urbanizing areas take
place in a  socioeconomic  framework within which
landowners attempt to maximize returns  associated with
land consistent with public  policies on such issues as
taxation, density of development, wastewater disposal,
and drainage.  These constraints have increasingly been
applied on  the  basis  of hydrologic  units  such  as
watersheds.  However, watershed management has  tend-
ed to consist of a collection of disparate measures that
focus on specific aspects of development in relative
isolation and sometimes without adequate evaluation of
unintended  side  effects.  For  example, measures to
protect surface waters from erosion and sedimentation
may result in additional aquifer recharge and an increase
in chemical content of runoff, which ultimately may
affect not only use of groundwater but also use of the
surface  water  into which the groundwater discharges.
This  situation creates  the  potential  for ineffective
management actions and wasted resources as individual
control measures are implemented without evaluation of
their interactions  or cumulative effects.  The lack of
methodology for a comprehensive assessment of water-
shed management strategies is a major obstacle  to
assessment of policy effectiveness and  adoption of a
more holistic approach to watershed management.
      The goal of this  project is to develop pro-
cedures for integrated assessment  of the hydrologic,
ecological, and economic consequences of alternative
landscape  scenarios  that occur during the urban-
ization/suburbanization processes.   More specific
objectives include:  (1) development of an integrated
hydrologic  framework  for  assessing impacts  of
alternative land-scapes on  surface  and subsurface
water flows  and movement  of sediments and pol-
lutants;  (2) development of procedures to predict the
response of fish and macroinvertebrate communities
to urbanization-induced  changes in water quantity,
water quality, and other biological conditions; and (3)
identification and assessment of policy and economic
conditions consistent with  alternative  landscape
scenarios as well  as estimation  of  the  effects of
alternative landscapes  on land  values  and fiscal
consequences for local governments.
      A case study focusing  on the Upper Roanoke
River Watershed will be employed to test the operation
of the system of linked models to be developed.  This
study area is shown in Figure 1.  This project will use
a  stakeholders'  panel from the case study  area to
ensure  relevancy  to actual  community  watershed
management.
      The project will produce a set of linked models
incorporated  into a  general methodology for holistic
watershed  assessment.  Figure   2   illustrates  the
methodology that will be used as  the basis for this
assessment. The creation of this analytical capability
will facilitate the development of policy likely to be
effective in achieving social objectives by allowing an
evaluation  of  consequences during  policy devel-
opment.

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                                                                                      Washington, D C.
            Figure 1.  Upper Roanoke River watershed study area.
                   Initial Conditions in Watershed Study Area:
                   Alternative Landscape Scenarios Modeled
                   Land Use Change

          Urban                       Rural
    Change In Population          Change in Land Cover
Change in Settlement Patterns     Change in BMP Practices
  Change in BMP Practices      Change in Animal Density
                                                          Policy & Economic
                                                      Conditions Consistent With
                                                           Land Use Change
                                                         Policy, Preferences, &
    Changes in Hydrologic Parameters
    Hydrology & Hydraulic Submodels
        Output Indicators Display:  User Friendly Computer Interface
    Aquatic Conditions
Biolocial Impact Assessments
    Biomarker Species
                                       Flood Hazards
                                    Change in Flood Risk
                                       Stream Quality
Economic Conditions
Land Price Changes
 Net Fiscal Returns
              Figure 2.  Study components and integration plan.

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An Integrated Ecological and Socioeconomic Approach To Evaluating
and Reducing Agricultural Impacts on Upper Mississippi River Watersheds
Prasanna H. Gowda and R.J. Haroa
University of Wisconsin-La Crosse, La Crosse, WI
A.D. Ward and T.L. Napier
The Ohio State University, Columbus, OH
      Nutrient-enriched waters from the Mississippi
River are one of the main causes of hypoxic zones in the
Gulf  of  Mexico  and  adversely  impact  aquatic
ecosystems within the basin. Nutrient loadings in the
Mississippi  River  have been associated  with  the
production of a substantial portion of the Nation's corn,
soybean,  and  wheat, but the specifics of these links
remain unclear.  Our research will:  (1) use a process
model to predict agricultural  discharges  from two
watersheds in  the Upper Mississippi River basin (see
Figure 1); (2)  use observed and predicted runoff, sed-
iment, and nutrient loadings to estimate the contribution
of urban areas, wastewater treatment plants, and point
sources to the total loading; (3) evaluate potential water
quality  benefits  associated with   the  adoption of
alternative management strategies on these watersheds;
(4)  develop  regional-scale  predictive  models  of
ecosystem health, biodiversity, and  sustainability by
relating information on biota and ecosystem  processes
to current and potential landscape composition and
structure; and (5) identify factors that affect the adoption
of conservation production systems  among landowners
in the two watersheds.  Our research focuses  on the
Lower Minnesota River watershed in eastern Minnesota
and the Maquoketa River watershed in northeastern
Iowa  (see Figure  2).  A  spatial-process  watershed
modeling approach that incorporates  a field scale model
will be used for predicting daily hydrologic and water
quality responses.  Landsat  Thematic Mapper data,
Natural Resources Conservation Service's  soils data-
bases,  published  farming   system   information,
topographic data, and historic climatic data will be used
in conjunction with Geographic Information Systems
(GIS) software to delineate  watershed and hydrologic
units to be used as inputs to the model. Available water
quality data will be used to determine flow attributes
associated with point discharges and nonagricultural
activities.
      Potential alternative farming  systems will be
developed based on interactions with stakeholders in
each watershed. Process-model estimates will be made
of  potential  reductions   in  environmental  effects
associated with  the  adoption of alternative farming
systems.  Data on macroinvertebrate communities in the
study watersheds will be collected and analyzed in the
laboratory. Statistical relationships between the habitat,
biotic community structure, and land use activities will
be developed  and used to predict expected regional
attainment for  management plans. Finally, the project
will evaluate socioeconomic factors that have influenced
the success of past soil and water protection  initiatives
within the watersheds.  Socioeconomic characteristics
of the farm operator and the farm enterprise will be
used  to  develop  statistical  models  to predict the
following:  (1) adoption of  specific  soil and  water
conservation production systems; (2) agricultural pro-
ductivity; and  (3)  propensities  to  adopt innovative
conservation farming systems. A  survey will  interview
750  landowner-operators in  each  watershed. Socio-
economic, ecological, and  water quality data will be
analyzed by using a combination  of parametric  and
nonparametric statistics. It is  anticipated that factors
affecting the adoption of conservation behaviors at the
farm level will be identified.  This will make it possible
to implement  more effective  technology  transfer
programs within the region and provide a tool to  assess
the environmental  benefits  of  adopting alternative
management practices.

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 Figure 1.   An infrared aerial photograph (acquired on September 12, 1989) showing movement of sediments from Maquoketa
           River watershed into Pool 13 of the Upper Mississippi River.
                                                 b.  Lower Minnesota River watershed in Minnesota
              a.  Upper Mississippi River Basin       c.  Maquoketa River watershed in Iowa
Figure 2.  A map showing the location of Lower Minnesota River and Maquoketa River watersheds in the Upper Mississippi
          River basin.

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Integrated Ecological-Economic Modeling
of Watersheds and Estuaries at Multiple Scales
Charles Hopkinson, Edward Rastetter, and Joseph Vallino
The Ecosystems Center, Marine Biological Laboratory,  Woods Hole, MA
C.J. Cleveland
Center for Energy and Environmental Studies, Boston University, Boston, MA
                                           IPSWICH RIVER SYSTEM
                                          & ASSOCIATED WETLANDS
                                                           IMMOBILIZATION
                                                           SEDIMENTATION
                                                           DENITRIFICATION
                                                           RIPARIAN FOREST
 Figure 1. Annual budget of N loading, uptake, and export for the Ipswich River drainage basin (metric tonnes). Objectives of the research
         program are to develop spatially explicit models of water and nutrient transport and fate in the Ipswich River watershed.
       The  consequences  of  large-scale,  long-term
 changes within watersheds  on estuaries  are  poorly
 understood.  Integrated assessments of ecological im-
 pacts of economic development in coastal watersheds
 will require interdisciplinary approaches.   Ecological-
 economic  models  must  be integrated  into  biogeo-
 chemical and landscape  models to realistically assess
 human impacts on watersheds and  estuaries.   The
 objectives of this project are to:  (1) integrate socio-
 economic models  with spatially explicit land use, bio-
 geochemistry,  and  watershed hydrology models  to
 provide comprehensive assessments of human activities
 on nutrient, sediment, and water fluxes in  a -500 km2
 coastal watershed;  and  (2) to develop  aggregation
 schemes that will allow the linked models to be applied
 to other watersheds.
       The  Ipswich River  watershed  in the  Boston
 metropolitan area will serve  as the experimental water-
 shed.  Rapid economic  and land  use changes in this
 region, plus ongoing research  and an extensive data-
 base, make the Ipswich River drainage basin particularly
 appropriate for this research project (see Figure 1).
       To assess human impacts  on  watershed and
 estuarine dynamics, four types of models will be used:
 (1) socioeconomic models, (2) spatial land use change
 models,  (3) terrestrial  hydrologic models, and  (4)
 stream hydrologic/biogeochemical models.  A goal  of
 our research is to  develop  a means of assessing the
 impacts  of urbanization and land  use  change  on
 watersheds and estuaries  in general, not just a  specific
 assessment for the Ipswich River.   Pending successful
completion  of Ipswich  River watershed modeling,
aggregation schemes will be developed that will allow
for applying our linked models to larger watersheds or
watersheds with less intensive input data than ours.
Aggregation procedures can ensure that our approach is
transferable.   Our approach will enable us to better
assess  human impacts on watersheds and to evaluate
management  strategies  for  rninirnizing  or  reversing
watershed, riverine, and estuarine degradation.
      The following  scenarios  will  be  evaluated
following the development and linkage of models: (1)
How do the magnitudes and temporal patterns of water,
sediment, organic matter, and nutrient loadings differ
among subcatchments with different land use patterns,
and how are loadings processed within the river before
being discharged to the coastal zone?  (2) What effects
do the spatial patterns of land use and river habitats
have on magnitudes and temporal patterns of loadings
to the mouth? How did historical land use and economic
activities affect the magnitudes and temporal patterns of
estuarine loadings? (3) What changes in watershed land
use can be  expected based on projected  economic
activity, and what will be the impact on water yield and
material loadings?
      Stakeholder involvement is an integral part of our
research. Our research will be implemented with the
involvement of three  stakeholder groups in the Ipswich
River watershed. Collaboration with these groups will
facilitate the collection of data critical  to parameterizing
hydrologic   and  biogeochemical   models  of  the
watershed.
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Linking Watershed-Scale Indicators of Changes in
Atmospheric  Deposition to Regional Response Patterns
Jeffrey S. Kahl \ 1. Fernandez 2, D. Mageean 3, S. Ballard 3, S. Norton 4, J. Cosby 5, P. Ludwig 6, and
L. Rustad 2 7
1 Water Research Institute; 2 Department of Applied Ecology and Environmental Sciences; 3 Smith Center
for Public Policy; 4 Department of Geological Sciences, University of Maine, Orono, ME
5 Department of Environmental Sciences, University of Virginia, Charlottesville, VA
6 Champion International, Inc., Bucksport, ME
7 U. S. Forest Service, Durham, NH
                    o
                    z
                         1986   1987   1988   1989   1990   1991   1992   1993   1994   1995

                          Figure 1. Bear Brook Watershed in Maine (BBWM) stream time series.
       The possibility of negative impacts from acidic
 deposition on aquatic ecosystems was widely recognized
 in the United States less than 20 years ago. Methods to
 assess the status of aquatic systems have been available
 for only about 10 years, and the models to evaluate this
 assessment are still  being modified and tested.  In this
 project, a specific ecosystem response to experimental
 watershed acidification is being determined, and this
 knowledge   is  being  scaled  to a  regional level to
 determine the extent of acidification and N-saturation in
 a  sensitive  subpopulation  of  high-elevation lakes.
 Concurrently,  mechanisms  are being developed to
 involve state, federal, and industry resource managers
 in the process of using these results in management and
 policy.
       The project activities are hierarchical  from site
 specific to regional.  First, the indicators of, and model
 predictions  for,  acidification and  N-saturation at the
 experimental site will be examined at the Bear Brook
 Watershed in Maine (BBWM).  This site has had  an 8-
 year experimental treatment of dry ammonium  sulfate.
 The  model  of acidification of  groundwater in catch-
 ments, and its nitrogen sequels, will predict acidification
 and responses as well as determine where  BBWM is
 located on the response continuum.  From new  and on-
 going data  collection on soils,  stream chemistry, and
 forest growth, the response indicators will be assessed
 at this site.   Second, the site-intensive information will
 be scaled to the region using  data from high elevation
 lakes in Maine and  their watersheds. These lakes had
 the highest concentrations of nitrate of any known lake
 population  in  the northeastern  United  States in the
 1980's.  Their acidification status was similar to  lakes
 in the Adirondacks.  Resampling of these lakes will
occur in 1997 and 1998 to determine lake-chemistry
changes during the past decade.  Using  mechanisms
defined at BBWM, parallels will be determined between
Bear Brook and the  high elevation lakes.  Then the
focus will be  on transferring new information about
changes in water quality and forest productivity to the
state, federal, and industry contacts.  A key component
is  the  early  and  ongoing  involvement of  these
stakeholders in a two-way communication process.
      At BBWM,  dry  ammonium sulfate has  been
applied  bimonthly since 1989 to the 10.2 ha. experi-
mental catchment, with an adjacent 10.9 ha. catchment
serving  as an untreated  reference.  This experimental
loading is 1,800 eq/ha/yr, vs. 600 eq/ha/yr ambient wet
+ dry N deposition.  In the experimental watershed,
nitrate concentrations increased from a volume-weighted
mean of 26 ^eq/L in 1989 to 78  ^eq/L by 1996 (see
Figure 1).  The yearly maximum increased from 40 to
106  jueq/L.  Nitrate  concentrations  in the reference
stream decreased from an average of 23 to 3 /ieq/L, and
the yearly maximum decreased from 43 to 11  fteq/L,
reflecting a regional decrease in nitrate concentrations
and flux.
      This information will be used by the EPA to meet
the  congressional  mandate in  the Clean Air  Act
Amendments of ascertaining trends  in ecological re-
sponse and determining the effectiveness of the Act in
influencing these trends. Site-specific data from Bear
Brook  scaled to the  regional high elevation lakes also
will provide  a template  for the  recognition  and
understanding of possible N-saturation and base cation
depletion. This information will be evaluated for use in
management and policy decisions by industry, and at the
local, state, and federal  levels.
                                                    11

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Comprehensive Watershed Management:
A Spatial Water Quality Assessment System
C. Gregory Knight
Department of Geography and Center for Integrated Regional Assessment, Pennsylvania State University, University
Park, PA
Robert P. Brooks, Barry M. Evans, James M.  Hamlett, Archie J. McDonnell, and Gary W. Petersen
Environmental Resources Research Institute, Pennsylvania State University, University Park,  PA
Todor N.  Hristov
Institute of Water Problems, Bulgarian Academy of Sciences, Sofia, Bulgaria
      The geography of natural  resources, land use,
and human economic activity influence the nature and
spatial distribution of water quality conditions and the
health of many aquatic  ecosystems.  The goal of this
project is to create an approach to achieve multiple goals
for water quality improvement from both physical and
biological viewpoints in river reaches that are subject to
a variety of geographical, social, and economic factors.
Basic to achieving  this  complex  goal is the ability to
answer two kinds of questions:  (1) if we invest in pollu-
tion control at one location, where and to what degree
will the stream quality goals  be achieved?  and  (2)
vice-versa, to attain particular quality standards for a
given reach for a basin, what alternative strategies could
be implemented under various stream-flow scenarios?
Thus, integrated approaches to water quality improve-
ment must be linked to available technological interven-
tions, to spatial and temporal patterns of implementation
and to social and economic dimensions of community
decisionmaking.
      This  project  is  a  collaboration  between  the
Institute of Water Problems in the Bulgarian Academy
of Sciences  and the Pennsylvania State University,
Environmental Resource Research Institute and Center
for Integrated Regional  Assessment, using the Yantra
River Basin (see Figure 1)  as a study site.  This basin,
the location  of Bulgaria's  first river basin planning
council,  is among the major Bulgarian contributors to
Danube  pollution  as a  result of its  long history  of
industry and its farming and livestock production.
      This work will bring together geographic infor-
mation  systems  (GIS),  water  quality  models,  and
decision-support software for simulating the impacts of
community decisions about improving water quality.
GIS will assist in the development of model input for
estimating  nonpoint source  pollution  where directly
measured data are not available and will bring together
such key variables contributing to water pollution and
remediation as population centers, settlements, industry,
soil types, land use, water use, stream hydrology, treat-
ment facilities, and biochemical and aquatic indicator
measures.  GIS also will help decisionmakers visualize
the impacts of alternative community decisions about
priorities for pollution abatement.
      This project's approach has been  to link new and
existing water quality model components in a holistic, and
transferable, GIS-based water quality assessment system.
Bulgarian colleagues will contribute important components
of  the  research  beyond   empirical  data  for  model
calibration,  including work  with aquatic   indicators,
experience in modeling multireservoir hydrologic systems,
multicriteria decision modeling, and interaction with local
resource managers.  The initial  review of decision support
software  suggests  that  a  basin  managment  system
developed at the International Institute for Applied Systems
Analysis  (Austria) can be coupled with  existing  models
used in the United States and other new approaches.  This
project will develop scenarios based on a finite combination
of  economic   and   climatic   conditions,  which  are
representative of past and future conditions in the basin.
                                                   12

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 /\,Vratsa  O <-^-  }•
S     s—fy     I.ovech
                                                         YANTRA RIVER BASIN

                                                        BULGARIA
               Figure 1. The Yantra River Basin, a Danube sub-basin in Bulgaria
                                   13

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Development and Implementation of Decision Support
Systems for Predicting Economic and Ecologic Impacts of
Alternative Land and Water Management Policies in Urbanizing Regions
Daniel P. Loucks ', Tammo S. Steenhuis 2, Mark B. Bain 3, Warren Brown 4, and Walter R. Lynn 5
'Civil and Environmental Engineering; 2Agricultural and Biological Engineering;3Natural Resources;
4'Institute for Social and Economic Research; sCenterfor the Environment, Cornell University, Ithaca, NY
      Community planning and environmental boards
often evaluate proposed changes in land and water use.
The aim of this research is to develop and then provide
government agencies with an interactive computer-based
technology for predicting the economic and ecological
impacts of decisions on water and land use policies.  To
do this,  this project must identify and quantify how
various land use policies impact surface and ground-
water quantity and quality, and how flows and pollutant
concentrations affect  aquatic  and adjacent terrestrial
systems.  Also, the investigators must attempt to predict
how changes in land and water use policies influence
socioeconomic activities, local economic development,
and tax revenues.
      Our immediate goals are to improve the ability to
predict the interdependent socioeconomic,  terrestrial,
hydrological, and ecological processes that define the
dynamics of land use changes and their effects on water
and associated biota.   This capability will be incor-
porated  into a  data-driven,  interactive,  microcom-
puter-based  decision support  system (see Figure 1).
This research is being conducted by a multidisciplinary
team that includes a  regional  economist, three agri-
cultural-environmental  engineers, and  an ecologist.
Work is being conducted closely with town and county
planners, and the methods will be tested on  a local
watershed. The approach and models being developed
will be applicable to a variety of watersheds similar to
those found in the Northeastern United States.
      In the few weeks since this project began, work
on  developing  the  required   surface  groundwater
simulation models for water quantity and quality  has
been conducted.  This will be an expanded Interactive
River-Aquifer Simulation (IRAS) model that has been
developed steadily. To assess economic impacts of land
and water use policies and to predict likely scenarios for
future land use changes,  this land use and land cover
information  is being gathered  for  the  five-county
economic region in which  this study watershed  is
located.  Data from the  Census Bureau's Survey of
Population and Housing for 1970, 1980, and 1990; the
New York State Office of Real Property Services' tax
assessment for every land parcel; and USGS' GIRAS
files on land use and land cover are being obtained.
The  initial objectives include  building a model  that
accounts for  land converted to residential uses since
1970, and then using it to identify areas in the watershed
having a high probability for future residential develop-
ment under various  economic growth scenarios.   The
structure of the ecological components of the simulation
model  are  being developed by  expanding a simple
system relating macroinvertebrate and fish species to
predicted habitat and water quality status.  New quan-
titative information from the surface groundwater sim-
ulation model will  permit much more sophisticated
predictions on aquatic communities than were previously
available.
     This project's goal is to develop a model system
that will simulate decision options, policy ramifications,
and environmental enhancement strategies.  The model
should be valuable for uses such as designing the spatial
distribution of conservation  incentives, selecting re-
quirements for development mitigation measures, com-
paring  zoning and  siting approaches, and planning
reserves, greenways, and other forms of natural area
designations.
                                                  14

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          MODULE 1
          Social &
          Economic
  MODULE 2
  Rainfall -
   Runoff
   MODULE 5:
     Decision
   \Support

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Effects of Natural and Anthropogenic Processes  on Tillamook Bay
and Its Watershed:  An Integrated Process Study and Land Use Perspective
James McManus ', Paul Komar', M.S. Jesse Ford2, Courtland Smith ', Debbie L. Colbert', and
Greg Bostrom 3
1 College of Oceanic and Atmospheric Sciences; 2 Department of Fisheries and Wildlife;3 Department of
Anthropology, Oregon State University, Corvallis, OR
                                    Figure 1.  Tillamook Bay and its watershed.
       Human  activities can  significantly  alter  the
 physical, chemical,  and biological nature of aquatic
 environments. Furthermore, community perceptions of
 environmental hazards can lead to mitigative actions that
 may themselves  have their own detrimental impacts  on
 ecosystems.  There is an interplay between social per-
 ceptions and societal decisions that can influence the
 physical environment, chemical cycles, and biological
 communities. An understanding of the problems as-
 sociated with the interactions between social and natural
 systems requires a multidisciplinary approach.
       This  project is  employing  such techniques  in
 addressing research problems associated with the inter-
 play between natural and anthropogenic processes within
 the Tillamook Bay  watershed.   Tillamook  Bay is a
 drowned-river estuary,  formed  initially  about  9,000
 years ago when rising sea level  at the end of the Quater-
 nary ice age inundated the lower reaches of the Trask,
 Wilson, Tillamook, Kilchis, and Miami Rivers, which
 currently drain into the bay  (see Figure 1). The Bay is
 located in the Pacific Northwest, roughly  100 km west
 of Portland, Oregon,  and approximately 80 km south of
 the Columbia River.
       Land use  practices vary significantly among the
 different riverine watersheds.  The Trask  runs through
 the urban center of Tillamook, whereas the Tillamook
 River to the south is heavily  influenced by dairy farming
 activities.  The northern-most three rivers experience
progressively  less urban  and  agricultural  activities
within their lower  watersheds and  increase  in  the
relative proportion of forested lands. Within the frame-
work of examining the relative influence of different
watershed activities on processes occurring within the
Tillamook basin, four specific hypotheses will be tested:
(1)  Land  use practices have  significantly altered  the
sedimentary budget of the Tillamook Bay watershed.
(2)  Carbon, nutrient,  and  trace metal cycles are sig-
nificantly different among the five major river systems
feeding  the Tillamook estuary because of the different
land use practices. (3) Differences in land use practices
between the Tillamook and  Kilchis River sub-basins
have led to quantifiable differences in aquatic ecosystem
health and biotic  integrity in the  riverine and  stream
environments.  (4) Local knowledge of watershed act-
ivities and processes  are  influenced  by  many  social
factors that  may result in perceptions quite different
from scientific measures  of sedimentation, nutrient
cycles, and aquatic health.
      The above  hypotheses will be  approached with
field and analytical programs designed to yield infor-
mation  on spatial and temporal  patterns within the
watersheds.  This project will attempt to explain the fac-
tors influencing watershed quality in such a way that
plans can be developed to  enable  residents of the five
river basins to improve the  environmental quality of
Tillamook Bay and the basin in which they live.
                                                   16

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Ecological Risks, Stakeholder Values, and River
Basins:  Testing Management Alternatives for the Illinois River
Mark Meo
Science and Public Policy Program, University of Oklahoma, Norman, OK
Baxter Vieux
Civil Engineering and Environmental Science, College of Engineering, University of Oklahoma, Norman, OK
James Sipes
Division of Landscape Architecture, College of Architecture, University of Oklahoma, Norman, OK
Edward T. Sankowski
Department of Philosophy, College of Arts and Sciences, University of Oklahoma, Norman, OK
Robert Lynch
Department of Occupational and Environmental Health, College of Public Health, University of Oklahoma Health
Sciences Center, Oklahoma City, OK
Will Focht
Department of Political Science, College of Arts and Sciences, Oklahoma State University,  Stillwater, OK
Keith Willett
Department of Economics,  College of Business Administration,  Oklahoma State University, Stillwater, OK
Lowell Caneday
College of Education, Oklahoma State University, Stillwater, OK
      The Illinois River, one of the most scenic rivers
in Oklahoma, has been the center of political contro-
versy about private property rights and environmental
protection for more than 25 years.  The Illinois River
has provided multiple social benefits to the citizens of
Oklahoma through its use  for recreation, water and
power supply, flood control, and nutrient removal. Yet,
the inability  of different interests to reach agreement on
how to protect the Illinois River watershed has placed its
hydrologic  resources at increased  risk of long-term
degradation.  With the absence of a unique environmen-
tal  issue  or feature to  catalyze political  support for
policy change, the Illinois River basin exemplifies the
challenge to sustainable  river basin management.
      This  3-year interdisciplinary  research project
demonstrates how different environmental and social
values held by river basin stakeholders can be identified
and compared so that  more effective environmental
protection strategies can be determined and adopted by
local land and water use interests and state agencies.
The investigators  will develop and test an integrated
impact assessment management protocol for the Illinois
River watershed by linking together the ecological,
economic, hydrological,  social, and political aspects of
the  watershed  in  an interdisciplinary approach  that
provides a more realistic framework for  calculating,
communicating, and negotiating environmental  risks and
competing social values (see Figure  1).  In the first 2
years of the project, the research team will:  (1) identify
stakeholder  views  on  the  Illinois  River  basin; (2)
determine the economic effects of alternative  land and
water uses for three study  sites in the river basin; (3)
calculate the ecological risks associated with different
intensities of  resource  use;  (4)  develop hydrologic
models  using Geographic  Information Systems (GIS)
that incorporate water quality aspects of alternative land
use practices; and (5) develop computer-generated im-
agery of each of the three sites  that will enable stake-
holders  to  visualize more  easily  the  implications of
different management options for  the river basin's re-
sources.  At the same time, members  of the research
team will be investigating stakeholders' perspectives of
natural,  economic, and sociopolitical impacts through
interviews and focus group sessions.  These groups  will
include  technical experts, lay stakeholders, and policy-
makers.   Background data will be drawn from the
investigators' prior studies of  the Illinois  River water-
shed, its carrying capacity, and comprehensive land use
plans.
      In the third year of the project, stakeholder groups
will be  engaged in a policy dialogue and a test of the
effectiveness of integrated computer models to facilitate
the risk communication of  complex  environmental
management issues.  Visual simulations developed from
CIS-based hydrological models will be shown to stake-
holders  in conjunction with focus group sessions to
ascertain management preferences and the overall legiti-
macy of negotiated agreements. Negotiation workshops
will be held to develop a consensus about land use prac-
tices that afford an  adequate level of protection to the
basin. The entire process will be tested to determine the
degree to which the process is viewed by experts and lay
stakeholders as efficient, effective, and legitimate, and
therefore acceptable.
                                                   17

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Year 1: Baseline Assessment
 Year 2: Alternatives Evaluation
                          Modeling N1A
                          regional policy
Modeling EIA
regional policy

alternatives
1

\

Year 3: Policy Recommendation

xU


Prepare
visualizations
\
/
Meeting of lay
stakeholders and
researchers

\

/
Refine policv
alternatives
alternatives
1
\

^

                                                                                       Modeling of EIA
                                                                                          basin wide
                                                                                      policy alternatives
                                                                                        Phone survey for
                                                                                        larger evaluation
      Figure 1. Illinois River Project: Research Plan.  Ecological risks, stakeholder values, and river basins: testing management
                alternatives for the Illinois River.
                                                              18

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Risk-Based Urban Watershed
Management:  Integration of Water Quality and Flood  Control Objectives
Vladimir Novotny, Robert Griffin, David Clark, and Douglas Booth
Marquette University, Milwaukee, WI
Figure 1.  The Milwaukee River is being restored. The picture shows a site that was previously an urban reservoir with very poor water quality.
         The dam that has created the reservoir is being removed, contaminated exposed sediments are either removed or capped and vegetated,
         stream banks are being reinforced by bioengineering, and the channel is restored to provide a natural habitat to fish, including
         migrating salmon from Lake Michigan.
       Urbanization is  known to  increase  flooding
potential and to impair the ecological integrity of water
bodies. In the past, however, most urban drainage pro-
jects focused almost solely on flood protection and were
conveyance oriented.  This project will develop and test
methodology for watershed management  in urban and
urbanizing watersheds.  Management is based on quan-
titative risk assessment of urbanization  to the watershed
ecosystem, including people residing in the watershed
and affected  by  the  receiving  water  body and biota
residing in receiving water bodies.  An evaluation of the
flood and ecological risks will help to  set  priorities for
management, find an optimum solution considering both
risks, and provide necessary information to agencies for
funding decisions.
       The objectives of this project are to: (1) develop
statistical  flow,  loading, and  water  quality  models
applicable to risk  assessment; (2) develop objective and
quantitative risk assessment procedures for estimating
ecological risks of stormwater and subsurface discharges
from urban and  suburban watersheds; (3) develop a
methodology for  the assessment of  flood control and
water  quality benefits  and resolving conflicts  between
flood  control and  ecological preservation-restoration
objectives; (4) develop benefit/cost models for urban
watershed  management to optimize both flood control
and receiving system ecological integrity; (5)  research
innovative  financing of urban watershed management,
identify key players, and assess the willingness  to pay
for  different types  of  benefits; and  (6)  examine
homeowners' risk/benefit perceptions, values, effective
responses to the risk,  subjective norms, sociocultural
backgrounds, and use of communication in the willing-
ness to pay for these different types of benefits.
      Figure  1 shows the concept of optimization of
flooding and ecological risks in watershed management.
Ecological risk assessment quantitatively enumerates the
hazards of pollution discharges and physical alteration
of urban water bodies and their hydrology/hydraulics to
the biota residing in the water body and riparian eco-
systems. A similar notion is applied to flood risk that
denotes a probability of damage to the water body itself,
to properties in the flood plain,  and hazards to human
health and well being.
      Presently, no ecological standards are available
for urban drainage and water body restoration projects,
and most projects are  driven by flood  protection ob-
jectives, leading to conveyance-oriented modification of
the urban (suburban) water bodies. Conveyance  solu-
tions (such  as channel  lining and straightening)  have
adverse ecological  consequences and commonly are
economically  inefficient.  Today,  such  projects  and
solutions may not  be  acceptable.  Incorporating eco-
logical integrity objectives  may lead to storage-oriented
approaches  and restoration of  ecological integrity of
urban receiving water bodies.
      The methodologies developed in this project will
be tested on  two pilot watersheds in Milwaukee County.
One watershed is fully urbanized and is found in the
City of Milwaukee.  The  other pilot watershed is in a
suburban part of the County that is undergoing  rapid
urbanization.
                                                    19

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Impact of Social Systems on Ecology and Hydrology
in Urban-Rural Watersheds;  Integration for Restoration
Steward T.A. Pickett
Institute for Ecosystem Studies, Millbrook, NY
      Policymakers, planners, and managers need to
understand the biophysical and social processes con-
trolling water  yield and  quality as  well as aquatic
biodiversity in human-dominated landscapes.   This
requires an integrated watershed model, which  can
address three types of social interventions for ameliorat-
ing water quality:  Type  1— reducing point sources of
pollution at the "end of the pipe;" Type 2—regulating
land  use  to reduce nonpoint inputs; and  Type  3—
modifying sociocultural processes within different types
of land uses  to minimize the  polluting behavior of
individuals, groups, and institutions. Types 2 and 3 are
underutilized.
      This research, to be initiated in late 1997, will
develop a model to test whether the sociocultural factors
involved  in Types  2  and 3 control  water  quality,
watershed  dynamics, and stream biota of watersheds
that  extend across  urban through  rural land uses.
Specifically,   the  model  will  test  to  what  degree
sociocultural  factors affect water quality, watershed
dynamics,  and stream biota  indirectly by modifying
ecological processes, or directly by modifying people's
actions. Indirect sociocultural effects include clearing,
planting, or managing forest patches that can affect the
extent, distribution, and structure of vegetation cover.
Direct sociocultural effects on  ecological processes
include inputs of solids and nutrients from dumping or
runoff from fertilizers and pesticides. The research will
be conducted in the 17,150 ha Gywnns Falls watershed
in the Baltimore region of the Chesapeake Bay drain-
age.
      An integrated approach  is needed  for  two
reasons. First, the U.S. EPA has mandated a 40 percent
reduction in nitrogen loading to  the Chesapeake  Bay.
Controlling point sources of pollution (Type 1) has not
met this mandate.  Second, forest conversion and sub-
urbanization are proceeding rapidly (Type 2), leaving
few options to mitigate water quality. Thus, a model of
the interaction  between the  ecological and physical
patterns  and processes in  the landscape with socio-
cultural patterns and processes (Type 3) is a potential
decisionmaking tool to identify and evaluate social and
ecological interventions for improving water quality in
the Chesapeake region.
                                                  20

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                      LAND COVER-ECOLOGICAL
                               HIERARCHY

                              Land Cover Type
                             Patch Configuration
                                    Soil
                             Permeability Class
    SOCIO-POLITICAL
        HIERARCHY

        Political Units
           Land Use
        Neighborhood
          Household
     WATERSHED
HYDRO-ECOLOGICAL
     HIERARCHY

        Watershed
     Sub-Watershed
     Sub-Catchment
         HUlslope
        Floodplain
         Channel
Figure 1.  The three hierarchies of patch or land units to be integrated in the proposed research. Sociopolitical units are those defined by
       political or administrative entities. Watershed hydroecological units are those delimited by decomposing large watersheds into smaller
       units that contribute runoff. The land cover-ecological hierarchy describes the specific surface features that govern the flow rate and
       pathway of runoff and the degree of saturation.
                                      21

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               Section 2.
Projects Initiated With Fiscal Year 1996 Support

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Strategic Renewal of Large
Floodplain Rivers;   A Preliminary Status Report
John B. Braden ', Misganaw Demissie 2, Eric DeVuyst', Paromita Mitra, Daniel Schneider ''3,
Richard E. Sparks '-3, David C. White ', and Renjie Xia 2
'University of Illinois at Urbana-Champaign, Urbana, IL
2 Illinois State Water Survey, Champaign, IL
3 Illinois State Natural History  Survey, Champaign, IL
                        136
                      O
                     _Q
                      rt  134
                                       Pre-dam (1887)
                                                       natural dry
                                                        season
                                          100           200
                                                 Day of Ycnr

                                      Figure 1. Effects of Illinois River dams.
                                                                    300
       This project  is investigating the potential for
 selective restoration of the hydrology and ecology of
 floodplains.   Interrelated ecological, hydrologic,  and
 economic models are being developed to compare the
 effects of  current  floodplain  management  versus
 ecosystem-based alternatives.   Specific objectives are:
 (1) to estimate thresholds at which floodplain restoration
 measures can have significant impacts; (2) to identify
 combinations  of structural  changes  and floodplain
 restoration efforts that enhance sustainability; and (3) to
 identify institutional  obstacles to policies that  might
 otherwise balance the costs and benefits of floodplain
 restoration. Our project uses the LaGrange Reach of
 the Illinois River as a study site.  Figure  1 illustrates the
 impact that current river management has had on river
 and floodplain hydrology.
       The hydrology of the Illinois River varies on time
 scales  of days, seasons, and decades.  The role of varia-
 bility is being  examined at each of these time scales on
 the ecological processes in  the  floodplain and  the
 resulting  economic  activities.  Ways to  increase  the
 overall economic value of the river using ecological and
 hydrologic restoration are being sought.
       Project hydrologists are combining one- and
 two-dimensional hydraulic models to  compute flow
 fluctuations in  three critical  time scales: (1) short-
 term changes related to navigation needs, (2) seasonal
 patterns, and (3) long-term extremes (both floods and
 droughts).  For selected restoration sites, intensive
 modeling using  the two-dimensional model will allow
 prediction of lateral-flow velocities and sedimentation
 processes that influence  biotic regimes.  To date, the
 one-dimensional model  has been run under selected
high flow conditions to test its  sensitivity and  to
 simulate the removal of levees.  The analysis will be
extended to low flow conditions. The model results
appear highly sensitive to Manning's roughness co-
efficient.
      Project ecologists are creating dynamic models of
floodplain vegetation, for both forested and nonforested
regions.   A critical feature of the models has been to
incorporate daily hydrological variation rather than the
average annual conditions used in existing floodplain
forest models.  These dynamic ecosystem simulators
interact with complex water regimes provided by the
hydraulic  model and, in turn, generate input for  static
habitat suitability models for floodplain animal species.
More than a century of river-stage records, floodplain
forest descriptions from early 19th century surveyor
notes,  and information from present-day  vegetation
surveys  have been used to calibrate the forest model.
The model has demonstrated that hydrological variation
at long time scales is necessary to maintain floodplain
forest diversity.  The next steps include incorporating
the nonforest vegetation model and linking the dynamic
models to  habitat suitability.
      Project  economists  are  developing a model to
estimate the impacts of restoration on the local econ-
omy.  This includes collecting, organizing, and assess-
ing data of the various economic elements (e.g., farm
production budgets, crop insurance, recreation values,
flood damage records and estimates,  transportation
values, institutional influences, etc.). A limited land use
allocation  model has been developed and used to assess
how agricultural land use might change as levee heights
are  reduced.  Agricultural land use  values will be
supplemented with survey data on recreational values to
permit a  more  complete  analysis of  social values
associated with floodplain land uses.  This project has
emphasized improving the information flows between
disciplines and model components, perhaps through
expanded  use of geographic information systems.
                                                    25

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Integrating Modeling and Management of Agriculturally
Impacted Watersheds:  Issues of Spatial and Temporal Scale
Patrick L. Brezonik', K. William Easter2, David Mulla3, and James A. Perry 4
1 Water Resources Center and Department of Civil Engineering;2 Department of Agriculture and Applied
Economics; 3 Department of Soil, Water and Climate;4 Department of Forest Resources, University of
Minnesota, St. Paul,  MN
      Scientists  and managers concerned with water-
shed management are faced with a dilemma: Scientists
are developing models that explicitly address cumulative
effects, large spatial scales, and longer timeframes, but
societal policy shifts are forcing managers  to  make
decisions on an increasingly more local basis.  Effective
implementation  of watershed management principles
requires models that resolve this dilemma (e.g., models
that incorporate knowledge about  effects at  longer
temporal scales and larger spatial scales into more local-
ized decisionmaking). The goals of this project are to
develop  a  more quantitative  understanding of  the
implications that this dilemma poses to aquatic science
and watershed management and to develop  solutions for
resolving the dilemma.
      This  project  focuses  on the Minnesota  River
basin, which occupies much of southern Minnesota. The
basin  presents  great  opportunities for  integrative
research to advance watershed science  as  well  as
contribute to solving a major environmental problem.
The basin drains almost 40,000 km2, including 34,000
km2 of intensively fanned land, and it is considered one
of  the  most  polluted rivers  in  the country.  High
concentrations  of nitrogen, phosphorus, pesticides,
bacteria, and sediment, mostly from diffuse sources,
cause degraded  conditions throughout  the river and
many of its tributaries.  In turn, the river  acts as a major
source for these pollutants to the Mississippi River, into
which it flows at the Twin Cities.
      A state-initiated  program, the Minnesota  River
Assessment Program, studied water quality in  the basin
and set improvement objectives during the period of
1989-1993.   This  program  led to a phase in which
comprehensive basin plans  are being developed and
implemented.  These plans include Best Management
Practices (BMPs) for farm management, wildlife and
habitat protection, sewage treatment facilities, riparian
zone  and wetland restoration,  and pesticide/nutrient
management.   However,  these plans rest on an in-
complete knowledge base. Barriers to implementation
include incomplete  knowledge about sources and the
effectiveness of proposed  BMPs.  This  project is ad-
dressing these needs with one overarching objective:  to
improve  the understanding  of how biophysical and
socioeconomic variables interact in agricultural water-
sheds of varying scales, landscape  conditions, and land
use management practices to affect  the  export of nu-
trients and their impacts on instream biological com-
munities, and in turn, to assess the role  of knowledge
about those effects in decisionmaking processes on local
level land use.
      We are conducting  the research in three coor-
dinated phases,  respectively  dealing  with landscape
processes, aquatic processes, and socioeconomic issues.
Biophysical phases  of the project encompass spatial
scales ranging from the whole drainage basin to several
scales of watersheds. In particular, we are evaluating:
(1) ways to account for scale effects  in hydrologic and
nutrient transport modeling within watersheds; (2) the
effectiveness of agro-ecosystems as  complementary
landscape units (to watersheds) for analysis of nutrient
export from the land (see  Figure  1); (3) the scales at
which aquatic ecosystems  process and retain nutrients
and how they influence rates of stream recovery when
land management practices  are improved; and (4) ways
to identify priorities  for controlling diffuse sources of
sediments and phosphorus based on technical, social,
and economic feasibility. All of these are critical for the
protection and enhancement of aquatic ecosystems by
integrated watershed management.
                                                  26

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                                                                              Steeper Cropped Moraine






                                                                              Steeper Urban Moraine







                                                                              Flatter Moraine






                                                                                       ,J QuUtnli






                                                                              Olivia Till






                                                                              Diyer Blue Earth Till






                                                                              Wetlei Blue Earth Till






                                                                              Poorly Drained Wetter Silts and Clays






                                                                              Poorly Drained Dryer Silts and Clays
Figure 1.  Map of Minnesota River Basin delineating its 12 major watersheds and 10 agro-ecoregions.
                                                   27

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 Urban Stream Rehabilitation in the Pacific
 Northwest:  Physical, Biological, and Social Considerations
 Stephen J. Surges, Derek B. Booth, Sally Schauman, and James R. Kan-
 University of Washington, Seattle, WA
                                 Figure 1.  McAllister Creek, Thurston County, WA.
       Society is eager to rehabilitate urban streams and
 to limit future damage as new areas are urbanized.  Any
 framework  for  rehabilitation  or protection must be
 grounded in understanding the specific factors respon-
 sible for degradation, their consequences, and the causal
 pathways linking them to human activities.  With an im-
 proved understanding of what determines stream condi-
 tion in urban areas, we can better evaluate prospective
 rehabilitation candidates;  define realistic rehabilitation
 goals; guide the design of successful rehabilitation pro-
jects;  and limit damage to high-quality aquatic systems
 through improved planning, design, construction, and
 management in urban areas.
      One of the goals of this project is to document the
consequences of urban development on the physical and
biological condition of urban streams and to demonstrate
specific rehabilitation strategies likely  to restore valued
properties of those  systems (see Figure 1).  Another
goal is to develop successful rehabilitation methods for
urban areas and to test public attitudes and preferences
for rehabilitation  measures.  The  questions  for  this
project  are  framed in a  process-based,  watershed
context:  How do landscape processes translate to pat-
terns  in channel form  and  process, and how  does
urbanization  affect spatial and temporal  pattern of
landscape processes?   Since  April 1997, we have ac-
complished several important tasks: (1) selected sites for
examining processes,  elemental changes, and cultural
contexts  of  stream  degradation,  benthic  macroin-
vertebrates, channel forms, bed sediments,  and cultural
landscapes—we have  considered 18  sites  that span a
gradient from low to high urbanization;  (2) evaluated
visual preferences for rehabilitation design and human
behaviors exhibited toward urban  riparian areas and
rehabilitation designs; and  (3)  assessed rehabilitation
project  costs  and outcomes.  These  efforts are being
expanded as  we  continue the development  of our
project.
                                                  28

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Influences of Forest Fragmentation on Watershed
Functions in Northern Vietnam—Preliminary Field Results
Jeffrey Fox,1 T. Giambelluca,2 and A.T. Rambo1
1 Program on Environment, East-West Center, Honolulu, HI
2 Department of Geography, University of Hawaii, Honolulu, HI
      It  is likely that the atmospheric and hydrologic
effects  of deforestation  depend  on  the  degree of
fragmentation of the remaining forest.  As fragment size
is reduced, a greater fraction of the remaining forest is
in close proximity to a forest edge,  where humidity, air
temperature,  and  wind  speed  may  enhance   the
transpiration rate.  Forest clearing and subsequent land
use can have major effects on soil hydrologic properties,
including hydraulic conductivity.  A fragmented land-
scape will translate into high spatial variability in  the
saturated hydraulic conductivity of the  soil known as
Ks.  As  a result, overland flow may be less  likely to
reach the stream channel or to attain velocities capable
of initiating erosion or transporting sediment  as far as
the stream channel.  Together, fragmentation effects on
evaporation and overland flow can alter the hydrologic
process differently than monolithic clearing. Thus far,
one of several planned intensive field experiments has
been conducted to  investigate edge effects on trans-
piration.   During the first  experiment, a transect  was
instrumented,  crossing  both upwind and downwind
forest  fragment edges.   Along  the transect,  four  re-
cording microclimate stations were created.  Also,  two
arrays of Granier-type sap flow sensors were installed,
monitoring 10 trees near the forest edge and 5 trees in
the interior of the patch.  Preliminary analysis of the sap
flow   measurements  indicate  significantly  higher
transpiration in near-edge trees as compared with trees
of the same species located in the patch interior.
      To scale up the hydrologic effects from a single
forest fragment to a larger area, knowledge of the larger
area's land cover is needed.  Preliminary analysis of the
research  area's landscape suggested that the relatively
homogeneous land cover, dominated by forest, present
in 1952  has become much more  heterogeneous  and
fragmented over time.  To test  this  hypothesis, 1995
Landsat  TM imagery  was acquired, classified,  and
compared with a 1952 air photo interpretation.  Because
of the different  resolutions of the two  data  sets,
intensive  field  work  was  carried out to adequately
ground truth the Landsat TM image.  More than 200
ground truth points were  collected as randomly as
possible.  A method was devised for using differential
global  positioning  systems or  GPS  ( + /-  7  meter
accuracy), and the satellite imagery was registered to the
field data with  a resulting root mean  square or RMS
error of 0.74 pixels  (22.2  meters).  A "supervised
classification" (whereby the image analyst "supervises"
the process by choosing the information categories on
classes  desired and then selecting training areas  that
represent  each  category) of the imagery  resulted in
seven land cover classes. Two accuracy assessments of
the classification were done. The first showed 79 per-
cent of 155 check points correctly classified without
attempting to account for GPS surveying and  regis-
tration error.  The second attempted to account for GPS
surveying and  registration  error  and resulted  in 95
percent of the check points being correctly classified.
A comparison of the Landsat TM classification with the
1952 air photo interpretation suggests that the research
area's land cover is more fragmented and heterogeneous
than in  1952.  Today, trees still make up  a relatively
large  part of the land cover (27  percent),  but it is
significantly more fragmented than in 1952  and the
average tree cover fragment size  is  two-thirds of  a
hectare  as compared with the air photo in 1952.
      Preliminary   socioeconomic  research  results
indicate  that  the  local  community  manages  the
fragmented landscape for different types of bamboo.
The land  cover analysis also corroborates this.   The
largest land cover  type  is dominated  by bamboo (32
percent), and the average fragment  size is 2 hectares.
      Future efforts will include continued field work
to investigate edge effects on transpiration in dry and
wet seasons, further ecological surveys to determine the
major tree species  that compose the vegetative cover,
and  additional  socioeconomic  studies  to  determine
factors that cause and maintain forest fragmentation.
                                                   29

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Figure 1.    Forest fragment where we established four recording microclimate stations, Ban Tat, Vietnam.
                                                30

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Geochemical, Biological and Economic Effects  of
Arsenic and Other Oxyanions on a Mining Impacted Watershed
Glenn C. Miller, Watkins W. Miller, Scott Tyler, and Douglass Shaw
University of Nevada, Reno, NV
Ron Hershey and Lambis Papellis
Desert Research Institute, Reno, NV
Susan Anderson
University of California at Davis-Bodega Marine Laboratory, Davis, CA
                              Rain/Snow
              Evaporation
                 ~
             Ground Water
                      \
               Flow In
                                           Turnover
         Wall Rock  ,
         Dissolution
                   Pit Wall
                   Sloughing
  Chemical
Precipitation
Pit Lake Water
  '.	\  Flow Out
                                         Newly Oxidized Rock
                                         (from air penetration of ground
                                         water cone of depression)

                                Figure 1.  Processes affecting pit lake chemistry.
      More than one-half of the annual production of
gold is  mined in the Humboldt River Basin, between
Elko and Winnemucca, Nevada. More than 15 major
precious metals  mines are located in this watershed,
including the Nation's two largest mines, which together
produce more than 5 million ounces of gold per year.
The expansion of precious metals mining in the past 15
years has been made possible, in part, by relatively new
techniques  of cyanide extraction  that have allowed
profitable extraction of gold at very low concentrations.
This expansion of mining has introduced environmental
problems that are poorly understood, both in terms of
their immediate environmental impact and in our ability
to predict long-term changes that may occur as mines
begin to close. Many of these large  mines penetrate the
groundwater  table.   The  required pumping of large
amounts of water significantly impacts the groundwater
systems.  Arsenic (As)  is  often associated with  gold
deposits, and release of As in soils  and surface  waters
has become an increasing concern.
      This research  is focusing on the geochemical,
biological, and economic impacts of As released  during
mining  in the Humboldt River watershed  in northern
Nevada. The objectives are to examine four aspects of
precious metals mining: (1) studies  on the evolution of
precious metals pit lakes, (2) long-term drainage from
cyanidization heaps, (3) biological effects of arsenic and
antimony, and (4) economic effects of long-term alter-
ation of the watershed.
      The  pit lake  studies are using a variety  of
geochemical models (see Figure 1) to develop  a pre-
        dictive understanding of an existing 106 meter deep pit
        lake in Yerington.   The water quality in the pit lake
        appears  to be significantly affected by dissolution of
        wall rock because the equilibrium models underpredict
        the contaminant concentrations in the lake.  Trace met-
        als are particularly difficult to model.  Initial laboratory
        studies on leached heap material show a large array of
        constituents being released from the rock. As expected,
        As concentrations in the drainage water are controlled
        by sorption to the heap material. Development of math-
        ematical models are considering both  equilibrium dis-
        solution effects  for heap  drainage water quality  and
        chemical reactions on the heap material.
              Biological research on  the effects of arsenite on
        water flea (Ceriodaphnia) indicate that brood sizes are
        decreased from 9.4 +/- 1.8 to 1.5 +/- 1.1 individuals
        when two generations are exposed to arsenite at 1.5
        mg/L. These multigenerational studies indicate that ar-
        senite may cause long-term genetic damage at sublethal
        doses.
              Studies on the economic impacts of mining on
        the Humboldt River system have  been initiated using
        five  focus  groups with  interests in  the Humboldt
        River watershed.   Groups consisting of represen-
        tatives of the mining industry,  the  agricultural in-
        dustry, the conservation community, and the govern-
        ment have provided views on the potential impacts.
        Water quantity  issues, rather than water quality,
        appear  to be a dominant concern, and  the  focus
        groups have reflected strongly held views of water
        usage in the watershed.
                                                  31

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Effectiveness of Regulatory Incentives for Sediment Pollution
Prevention:  Evaluation Through Policy Analysis and Biomonitoring
Seth R. Reice and Richard N. Andrews
University of North Carolina at Chapel Hill, Chapel Hill, NC
                             Minimum Disturbed
                                Area Requiring
                                 Erosion Plan
    # Field
     Staff
Total Area
 (Miles 2)
# Active
Projects
Orange County
Wake County
District 4*
(16 Counties)
0.5 Acres
1 .0 Acres
1 .0 Acres
3
13
5
400
858
8,116
-60
-500
-1000
 * District 4 oversees all construction projects in all 16 counties without a Local Erosion and Sediment Control
 Program. It covers all governmental construction in the District 4 area, including Orange and Wake Counties.  So,
 a single stream can have adjacent construction sites administered by different regulatory agencies.

                            Figure 1. Characteristics of the regulatory jurisdictions studied.
      A  critical problem in  American  rivers  and
 streams is sedimentation.  Sedimentation degrades water
 quality, alters habitat for fish and macro invertebrates,
 limits the ecosystem functions and services, and reduces
 the aesthetic and economic value of rivers and streams.
 Many  regulations and  policy  incentives  have  been
 devised to control sediment pollution of our rivers. Yet,
 there has  rarely been  an attempt to reconnect the
 policies with the ecology of the  rivers. That is the goal
 of this research.  The overall objective is to study me
 effectiveness  of different policy incentives  in the
 reduction of die ecological risks and consequences of
 sedimentation.  The  aim is to create more effective
 management  strategies  to  provide   environmentally
 sustainable social and economic development in U.S.
 watersheds.  This work will  integrate the social and
 regulatory theory behind  sediment  ordinances  and
 policies as well as the resultant ecological impacts of
 sedimentation on the rivers and  streams. This project's
 goal is to discover which policies and regulations really
 work to enhance stream biota and ecosystem health and
 the reasons why they  are or are not effective.
      A comparative watershed  approach is being used
 to contrast the ecological effects of different intensities
 of sediment control standards and enforcement. Similar
 watersheds  are being  selected from three political
 jurisdictions  that  differ in the  stringency  of their
 sediment  and erosion  control  requirements,  their
 staffing, and enforcement (see  Figure 1).  Field bio-
monitoring  data,  water  chemistry,  and  leaf  litter
decomposition rates are being used to document changes
in stream ecosystems above and below construction
projects under these different regulatory regimes.
      North Carolina has one of the strongest state laws
in the Nation for erosion and sedimentation control, and
at least 38 of its  county and municipal governments have
enacted even more stringent control ordinances.   The
differences among these  regulatory regimes, applied to
similar   types   of  socioeconomic   activities  (e.g.,
commercial and housing  construction, road building) on
otherwise similar watersheds, provide opportunities to
draw broader inferences on a comparative basis.   An
unusually detailed evaluation  of the  North Carolina
erosion and sedimentation program and a close working
relationship  with the North Carolina Division of Land
Quality provide substantial baseline  and supporting
information  for this research. This project focuses on
the regulations governing controls on sedimentation into
rivers and streams  to determine their effectiveness and
ecological consequences.  A direct comparison of dif-
ferences in  the programs'  regulatory stringency with
their ecological  outcomes in the  streams will expose
what actually results in the protection of stream  eco-
systems.  This will determine the actual effectiveness of
these regulations  in maintaining  stream  ecosystem
health, functioning, and biodiversity.   In the end, the
answer to the question,  "What really works?"  will be-
come clear.
                                                   32

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Watershed Protection in Agricultural Environments:
Integrated Social,  Geomorphological, and Ecological
Research To Support Ecosystem-Based Stream Management
Bruce L. Rhoads, Edwin E. Herricks, and David Wilson
University of Illinois at Urbana-Champaign, Urbana,  IL
                                          Historical Social/Cultural Context
                                               T        T
                                        Values/Attitudes Towards the Environment   I
                                                   t
/' Predominant
/ Ethic
Legal
/ Economic
/ Available i
\
Context
Context
r Technology
/ Interaction with Environment 1-
/ Scientific
/ Research
Perceptual
Assessment
/ /' Environmental Response I ~~

Scientific Knowledge |


                                                                   Nonscientists
                                                                 Local Knowledge I

 Figure 1.  Conceptual model of interaction between scientists and nonscientists m the social  negotiations of community-based resource
         management.
       The overall goals of this project are to provide an
 improved understanding of the roles of scientists  and
 scientific information  in  community-based  environ-
 mental decisionmaking (see Figure 1) and to advance
 knowledge  of the  connections  between  stream geo-
 morphology and aquatic  ecology in human-modified
 agricultural stream systems. The proposed project  has
 a four-pronged research design: (1) historical analysis of
 the attitudes and values of rural stakeholders toward
 water and  watersheds in the agricultural Midwest,
 (2) social analysis of the mechanisms that facilitate  and
 impede infusion of new scientific knowledge into local
 decisions about  stream management in  agricultural
 watersheds, (3) GIS-  and  field-based analysis of inter-
 relations  between  geomorphological  and ecological
 dynamics  of  human-modified  agricultural  stream
 systems at the watershed and reach scales, and (4) dis-
 semination  of scientific  information  from the geo-
 morphological and  ecological  research to  farmers  and
 other local stakeholders.
       First-year  data collection  activities have con-
 centrated mainly on the social component of the project.
 Fifty-seven in-depth, open-ended  interviews have been
 conducted   with  drainage  district  commissioners,
 farmers,  and associated   stakeholders in the  rural
 communities of East  Central Illinois.  The interviews
 have focused on three  issues: (1) farmer and stakeholder
 perception of watersheds  as  an  agricultural,  cultural,
 and aesthetic  resource; (2) current  usage and main-
 tenance of watersheds; and (3) perceived best and  rel-
 evant strategies for future usage and maintenance. Pre-
 liminary findings indicate that farmers are a surprisingly
heterogeneous group in outlook and perspective and that
important commonalities can be identified.  Farmers
value streams not only for the purpose of land drainage,
but also perceive themselves as fundamental stewards of
the land.   Future  work will explore the interplay of
these contrasting values. Additional open-ended inter-
views will be conducted after scientific/technical infor-
mation is imparted to farmers and stakeholders to gauge
their responses.
      Work on the geomorphological component of the
project  has emphasized the development  of  a geo-
referenced grid of control points for digitizing stream-
channel traces and for analyzing changes in these traces
through time.  Preliminary analysis indicates that over
the last several decades, humans have been the dominant
agents of geomorphic change in the fluvial systems of
East Central Illinois.  Field activities have involved the
selection, preparation, and mapping of seven study sites
for detailed investigations of the fluvial  dynamics of
human-modified agricultural streams. During the next
year, geomorphological  research will  focus on  GIS
analysis of stream-channel change and on data collection
at the field sites.
      A fish-species data  matrix containing  critical
habitat  and life history  information for Illinois  fish
species  is under development. The matrix will be used
to assist in interpreting habitat  fisheries  relationships
from field data  and will provide a means of specifying
critical habitat conditions when management programs
are being developed. A systematic fisheries collection
program has been  initiated at  the  field sites  in co-
ordination with geomorphic data  collection activities.
Preliminary sampling suggests that fish-species diversity
is tied strongly to physical habitat diversity.
                                                   33

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 Towards an Integrated Regional Model
 of River Basins of the Western Pacific Rim
 Jeffrey E. Rickey
 University of Washington, Seattle, WA
                     Evapotranspiration
                   Soil Texture
                            Vegetation
               Precipitation
                                                                            Temperature
                                                                    Population
                                                 Runoff

                       Figure 1. Schematic of integrated regional model for the Western Pacific Rim.
       The  extraordinary  pace  of development and
 population growth along the Western Pacific Rim has
 placed dramatically increasing pressure on river basins
 and their downstream coastal ecosystems.  Although
 decisions  about the usage  and allocation of  water
 resources are generally made according to economic and
 political criteria, the sound management and optimiza-
 tion, and hence sustainable use of these resources, will
 require  increasingly sophisticated information on the
 functioning of the biophysical systems and how they are
 affected by socioeconomic and political institutions. To
 address  these issues, the investigators are building an
 "integrated regional model" of the river basins of the
 Pacific Rim,  with a focus on Southeast  Asia.   The
 model  has the  objective of coupling  hydrological
 pathways  and  biogeochemical  indicators  with the
 interactions and effects of human impact to describe
 how materials are mobilized and transported from the
 land surface  to the coastal zone.  The  project is ad-
 dressing two  interrelated questions: (1) The  science
 question is, "What are the effects of changing land use
 (and  climate) on  the  mobilization of water  and its
 dissolved and paniculate load from the land surface into
 the coastal zone at local to regional scales?"  (2) The
 practical question is, "How is information and training
 in  a  geographically  and  politically  diverse region
 mobilized and focused on the problem?"
      The  project  approach  is to link  large  scale
 attributes of  drainage  basins to  fundamental  bio-
geochemical processes and fluxes through  rivers  via a
regional-scale  routing model (see Figure 1). This ap-
proach is derived from the extensive experience of this
 group on the Amazon River.  To address the inherent
 space and time scaling assumptions that must be made,
 the  investigators have developed a "drainage basin
 element" model (DBE), expressed within the modeling
 environment of a global biogeochemical model. Each
 DBE is georeferenced within a drainage network and
 linked  to  databases  of  basin characteristics  (e.g.,
 elevation,  soil texture, and vegetation types).  The
 immediate requirements are to translate precipitation
 into runoff with a water balance model for each DBE in
 a manner that  allows estimates of residence time and
 flow through each  element (to "mobilize"  the chem-
 istry). Selection of  the size and temporal resolution of
 each DBE is a "scaling" problem, where tradeoffs must
 be made between expectations and data availability.
      The current  expression of the model for the
 region is  1 degree  (about  100 km),  based on readily
 available data sets and summed to a monthly composite.
 At  this scale, the hydrographs of such rivers as the
 Mekong and Yangtze are readily reproduced.  However,
 it  is  not  possible  to  resolve  the dynamics of,  for
 example, the myriad small rivers of the coastal areas so
 typical of the region, nor to address the typical scales of
 land use and cover change. Hence, the next phase of the
 work will  be to establish the model for 1 km elements,
 taking advantage of newly available data.  An integral
part of this project will be to  establish a network of
 cooperating institutions across the  region, including
 science, policy, and  economic organizations. Based on
preliminary feedback, this project will contribute to
useable assessments  of water resources along the Pacific
Rim.
                                                   34

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An Integrated Approach To Assessing Water Management
Options in a Major Watershed:  Extending a Hydrodynamic-Water
Quality Model To Include Biological and Politico-Economic Components
Paul Sabatier, Loo Botsford, Mike Johnson, Jay Lund, Peter Moyle, Gerald Orlob,
James Quinn, Peter Richerson, Tom  Suchanek, and Marca Weinberg
University of California at Davis, Davis, CA
      The primary objective of this project is to develop
and  demonstrate  an  integrated methodology for  as-
sessing  management  alternatives  for watersheds that
support aquatic species at risk.  Such a methodology
requires an  interrelated set of deterministic and sta-
tistical models designed to characterize quantitatively the
responses  of sensitive species to hydrodynamic, water
quality, and ecological influences within riverine and
estuarine reaches of the watershed under various mana-
gement  alternatives.  Alternatives may include restor-
ation of riparian habitat,  regulation of stream flows,
curtailment or rescheduling of diversions,  and control of
point and  nonpoint sources of pollution.  The project
links not only hydrologic, water quality, sedimentation,
and  fishery  models,  but  also  the impacts of various
management alternatives on urban and agricultural water
supplies.  Finally, the  conditions under which such
modeling  capability  can  produce consensus on  the
perceived impacts and feasibility of various management
alternatives  in situations  of political conflict will be
assessed.    The models are being developed for  the
Sacramento  River from  Shasta Dam   through  the
Sacramento-San Joaquin  Delta, with  the  principal
fisheries being the winter run chinook salmon, striped
bass, and Delta smelt.
      In  the  12 months  since funding  began,  the
project has focused on: (1) developing the hydrologic
and water quality (i.e., temperature, salinity, and toxic
particle)  models  for  1984, 1992,  and  1993;  (2)
integrating models of particle and fish movement into
the hydro model, and then estimating winter run and
striped bass survival;  (3) continuing  long-standing
work on the Clear Lake and Cache Creek watersheds,
particularly with respect to sedimentation and mercury;
(4) gathering the  data to build the models relating
water quantity and quality to urban and agricultural
water  use; and  (5) conducting  a mail survey  of
approximately 1,300 policy participants in Bay-Delta-
Central Valley water policy.
      The number of water  years was  increased from
three to six to include a wider variety of flow and tempera-
ture conditions. Project members also have obtained ad-
ditional support to study mercury contamination and urban
water management in the context of this research.
                                                 35

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Modeling Effects of Alternative Landscape Design and Management
on Water Quality and Biodiversity in Midwest Agricultural Watersheds
Mary Santelmann, K. Freemark, D. White, S. Polasky, and G. Matzke
Oregon State University, Corvallis, OR
J. Filers, and J. Bernert
E & S Environmental Chemistry, Inc., Corvallis, OR
B. Danielson, and R.  Cruse
Iowa State University, Ames, IA
J. Nassauer
University of Michigan, Ann Arbor, MI
S. Galatowitsch
University of Minnesota, St. Paul,  MN
      Serious concerns exist about ecological degrad-
ation from modern agriculture.  An integrated, multi-
disciplinary approach  linking multiple  purposes of
watershed could inform and improve our management
of agricultural watersheds and water resources.  The
goal of this  project is to develop  alternative  future
landscape design and  management scenarios and to
evaluate their effects on agricultural production, water
quality, and native biodiversity in watersheds charac-
teristic of the midwestern United States.
      This project involves the following:  (1) iterative
design of alternative future landscape and management
scenarios;  (2) development  of computer  models to
estimate  effects  of  landscape change  on agricultural
production, water quality,  and biodiversity (i.e., aquatic
organisms, terrestrial vertebrates,  plants); and  (3) a
farm planning exercise to incorporate input from local
farmers and decisionmakers and to explore how human
attitudes as well as practical  and economic constraints
are translated into land use and management decisions
and  the spatial implications  of these decisions at the
watershed  scale.  This  project  is  focusing on two
agricultural watersheds (5,130 and 8,790 ha) in central
Iowa.   Databases exist  on  land  cover,  avian  bio-
diversity, and aquatic biodiversity for these watersheds
as a  result of the  USEPA/USDA Midwest Agrichemical
Surface/Subsurface  Transport and  Effects Research
(MASTER) program.
      For each watershed, 3 alternative scenarios for 25
years into  the future  are being  developed using a
normative expert informed design process (see Figure
1).   The  scenarios reflect a set of alternative national
priorities with an emphasis on: (1) production of agri-
cultural commodities such  as corn,  soybeans,  and
livestock;  (2) conservation of native biodiversity and
associated ecosystem  functions; (3) improvements in
water quality; and (4) a compromise among all three.
Digital  maps of  each scenario  may  incorporate the
effects  of heterogeneous  implementation  of  each
watershed design to reflect situations in which farmers
and/or landowners  may  decide  not to  participate in
programs  designed to achieve improvements in water
quality  and biodiversity.  Therefore, the manner in
which   patterns of implementation may influence the
success of these efforts can then be evaluated.  Digital
coverages among  scenarios can also be recombined to
evaluate effects on ecological and reorganized model
results.
      Existing spatially explicit computer models for
terrestrial  vertebrates  (multiple species and individual
species  populations) are being adapted for the present
watersheds.  Parallel models for terrestrial vertebrates
will crossvalidate  results.  New models  for plant bio-
diversity (particularly in wetlands) are being developed
from existing databases in central Iowa and southern
Minnesota.  Currently, only  rudimentary accounting
models are being developed for agricultural production
and water  quality.
      This project  is pioneering a landscape design
approach  for  agricultural  watersheds  based  on  a
normative  process using experts. Evaluation efforts will
contribute to the  development  and  comparison  of
spatially explicit computer models for individual versus
multiple species for a variety of taxa. The longer term
significance lies in the ability of the project to inform
landowners  and  policymakers (for  example,  those
crafting Farm Bill 2000) of  effects of  land use  and
management choices  on  water resources,  ecosystem
function, and human social systems in the Western Corn
Belt Region.
      The next steps  for this project  will involve the
completion of digital maps of future scenarios as well as
the construction  and  parameterization  of computer
models.
                                                  36

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\
\—
Design
Scenarios ,-,•

«•
/
X
Design
Scenarios
2


Examine

CON

Design
Scenarios
3
C E P T U A L

Define

5 P E C 1

Design
Scenarios
4
F 1 C 1 TY
Design
Scenarios
U^ for Modeling and
Field Testing
^^^^^—^^^^^^^•k
^•^^•t
PORTRAY    COMPARE     PORTRAY
                                       COMPARE
                                                         PORTRAY
                                                                          , COMPARE
                                                                                                PORTRAY
                                                                                                                     . COMPARE
                                                                                                                                           PORTRAY
                                             Figure 1. Agricultural watersheds—A normative, expert design process.

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Streamside Reforestation:  An Analysis of
Ecological  Benefits and Societal  Perceptions
Bernard Sweeney,  Thomas Bott, John Jackson, Louis Kaplan, J. Denis Newbold, and Laurel Standley
Stroud Water Research Center, Academy of Natural Sciences, Philadelphia, PA
Richard Horwitz and W. Cully Hession
Patrick Center for Environmental Research, Academy of Natural Sciences,  Philadelphia, PA
Janet Johnson
University of Delaware, Newark, DE
James Finley, Caren Glotfelty, and Cecilia Ferreri
Penn State University,  University Park, PA
      Streamside forests are recommended as a land use
practice to protect aquatic ecosystems from nonpoint
source pollution. To date, the scientific basis for the
benefits of these forest buffers has stressed their role in
removing watershed inputs of nutrients, sediments,  and
toxic contaminants.  In this project, forests along small
streams in  the  Piedmont  region  of Eastern North
America are viewed as the primary regulators of stream
width.  Preliminary  measurements indicate that small
forested streams  are consistently wider than contiguous
meadow reaches.  The alteration of stream width  that
results from deforestation  profoundly influences  the
stream  ecosystem,  both locally and in  downstream
rivers  and estuaries, through effects on habitat  and
water quality.
      Public policies to restore Streamside forests  will
require documentation of ecological benefits and an un-
derstanding of societal factors affecting their implemen-
tation.  The principal goals of this project are to: (1) un-
derstand the relationship between forest buffers, stream
width, and the condition of stream ecosystems; (2) de-
velop databases concerning stream organisms and their
activities  that can guide the development of policies
concerning  restoration  of  Streamside  forests;   and
(3) identify  and quantify social and economic issues
affecting Streamside forest restoration.
      This  project  includes four interrelated  com-
ponents:  (1) field  measurements  in 15-18 different
streams to  ensure  general applicability  of results;
(2)  supplemental  experiments in  controlled  model
streams; (3) mathematical modeling to assess watershed-
wide water quality implications; and (4) interviews  and
surveys to assess landowner  attitudes and  land  use
practices.  The ultimate goal is to bring both the natural
and  social   science  components   into  focus  for
policymakers and regulatory agencies.
      Field work began in March 1997. Longitudinal
profiles  and detailed  cross-sectional measurements
have  been performed on  18  stream reaches (i.e.,
paired meadow-woodland  reaches on 9  streams) to
estimate  the geomorphological  characteristics of each
stream reach (e.g., slope, cross-sectional area, width,
depth,  sinuosity, pool  and  riffle  characteristics,
floodplain   height    and   width,   and   habitat
heterogeneity).   Preliminary  results  indicate  that
woodland reaches are 1.6 to  3.4 times  wider than
meadow  reaches.  Community metabolism studies have
been conducted on 5 streams (10 reaches) using open-
system measurements  of dissolved oxygen change.
Macroinvertebrate communities in the riffle and pool
sections  of 9  streams (18  reaches)  were  sampled
quantitatively in March, May, July, and November of
1997.  Fish community analysis  was performed by
electroshocking studies of 6 streams (12 reaches; 3-4
passes per reach) in the fall,  including  length  and
weight analysis of individual  fish of each species.
Degradation/decomposition studies of natural organic
matter   (i.e.,  humic  substances,   amino   acids,
carbohydrates, and total dissolved organic carbon) and
pesticides (i.e., atrazine, dursban, linuron, and meth-
oxychlor) in meadow and forest reaches was initiated
in 1997.  Preliminary  studies involving bromide and
chloride  releases  in 5 streams (10  reaches) have been
completed  in  1997 in preparation for nitrogen and
phosphorus spiraling work.
      In the anthropological work, investigators de-
veloped,  pretested, and refined an interview protocol
and objective statements describing Streamside forests.
Target watersheds and streams in central and southeast-
ern Pennsylvania,  and northern Delaware and Maryland
were randomly selected, and the interview process with
landowners was started in 1997 (see Figure 1).
                                                   38

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                   f\/ Watersheds
                    A/R°ads
                    /\ / Rivers and Streams
Land Use/ Land Cover
     Water
     Low Intensity Development
     High Intensity Residential
     High Intensity Industrial/Coi
     Hay/Pasture
     Row Crops
     Lawns/Parks/Golf Courses
:.vV: Forest
    I Wetlands
                                                                                                    N
Figure 1. Contemporary land use map near rural Chester County, Pennsylvania, showing the location of study streams (blue
          lines) and their watersheds  (red boundaries) where stream ecosystem structure and function are being evaluated in
          response to the presence/absence of a streamside forest.  Each dot represents either the upstream or downstream limit
          of a given reach of stream; each study stream has two experimental reaches—one forested, one deforested. Map was
          derived from EPA Region Ill's 30 m Landsat Thematic Mapper Land Cover Data Set Version 3alb by Cully Hession,
          Linda Misiura, Maeve McBnde, Academy of Natural Sciences of Philadelphia.

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Integrated Urban Watershed Analysis:
The Los Angeles Basin and Coastal Environment
Richard P. Turco
University of California, Los Angeles,  CA
      This project is investigating the processes  that
control water availability and quality  in a major urban
watershed—the Los Angeles basin in southern California
(see Figure 1). The study consists of a synthesis of data
and models that bear on numerous aspects of the water
resource problem, including: regional meteorology and
climatology; basin hydrology, vegetation and land use;
human  water consumption  and disposition;  runoff
sources of sediments, toxics and nutrients; air pollutant
transport, transformation and surface deposition; down-
stream wetlands ecology and impacts; and coastal water
circulation, biogeochemistry,  and sediments. This pro-
ject consists of  a  broad  synthesis  of observational
data—in situ measurements, remotely sensed data, and
local geographical information—and model simulations
of  material  flow,   deposition,  transformation,  and
bioassimilation.
      Regional simulations of storms over the  Los
Angeles basin were  carried out with the  MM5 meso-
scale dynamics model. This project focuses on a record
rainfall event that  occurred in  1993. Model calculations
of precipitation compare favorably with  meteorological
observations. The corollary aspects of extreme regional
drought are  under  investigation through  a tree ring
analysis, which  correlates moisture  availability with
seasonal growth rates and wood density.   A 500-year
reconstruction indicates an extended  period of severe
drought in southern California in the late 1500's, within
the epoch of the "Little  Ice Age."
      A  coastal  ocean model for the Los Angeles
watershed is  close to completion. To provide realistic
large  scale  forcing of the  circulation,  the  model
incorporates a nesting of spatial scales from that of the
Pacific basin, to the California Bight, to Santa Monica
Bay. Moreover, initial results were obtained from a bio-
geochemistry code  that is being integrated with the
coastal dynamics  model to study the  "red tides." The
coastal modeling also is being  connected with the runoff
component  of the project,  for  which  a GIS-based
analysis has been developed to quantify the chemical
composition (nutrients, toxics) in key regional Los Angeles
outflows.  The runoff model is being calibrated using
streams in the region, particularly those heavily impacted
by  urbanization.  The runoff component  uses  a com-
prehensive hydrological  database and modeling system
developed for several of the Los Angeles watersheds.
      New research was conducted on the response of
vegetation in a major southland estuary to anthropogenic
nitrogen and phosphorus. Measurements taken along an
upstream-to-downstream transect through the Newport
Bay  estuary  indicate   that  macroalgae  efficiently
scavenge  nutrients  from the freshwater inflow.  The
levels of nitrogen in the salt marsh sediments were quite
low, however, and the  nitrogen/phosphorus ratios in
these sediments were unusually low.
      An analysis of  the  contribution of  airborne
particulates  to the  southern California  coastal  ocean
surface microlayer showed that the concentrations of a
suite of trace metals (i.e., Cd, Cu, Ni, Pb, Zn, Cr, Mn,
Fe,  and Ag)  are  more  strongly  correlated  with a
planktonic source than with either an aerosol  or bulk
water source.  The  plankton appear to bioconcentrate
these metals effectively, although the  mechanism by
which  this  occurs  remains  uncertain.    Hence,  the
situation of southern California is quite different from
that in  the  Great Lakes,  where  the  trace  metal
composition of  the surface layer, bulk  water, and
aerosols are strongly correlated.  In other work, a novel
analysis of DDT in offshore sediments succeeded in
separating the contributions due to sewage outflows and
to industrial dumping. The sewage source is identified
through a close  relationship that exists between DDT
and another class of stable organic compounds used in
detergents. Thus, coastal  regions with heavy industrial
DDT pollution have been identified, which will help in
formulating remediation schemes.
                                                   40

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:MalibuufjF
 CreelifW
 II
 Santa
Monica  Balloha
  Bay   Wetlands
III
                            Los Angeles

                                Basin-
                             I
                            Newport
                              Bay

                 Figure 1. Los Angeles Regional Watershed.
                          41

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               Section 3.
Projects Initiated With Fiscal Year 1995 Support

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Development and Application of Spectroscopic Probes
for Measurement of Microbial Activity in Aquatic Ecosystems
Carol Arnosti
University of North Carolina-Chapel Hill, Chapel Hill, NC
Neil V. Blough
University of Maryland-College Park,  College Park, MD
      The microbial degradation of organic matter is a
key part of carbon cycling in aquatic systems. Bacteria
are responsible for a  high organic-carbon turnover in
water  and  sediments,  hydrolyzing   organic  macro-
molecules to smaller  pieces that can be remineralized
and interconverting organic structures through a variety
of reactions. Measuring the net degradative capabilities
of a complex community of microorganisms is a major
challenge in understanding the dynamics  of carbon
cycling because we lack the  means  to  measure ac-
curately  the  rates  at  which bacteria  hydrolyze  large
macromolecules to  smaller pieces, which can be further
transformed or remineralized.   In this project, we are
developing a new  generation of spectroscopic probes
that  can  be used  to measure  extracellular enzymatic
hydrolysis rates of  organic macromolecules in the water
column and sediments.  This  approach  is  based  on
efficient intramolecular energy transfer between donor
and acceptor fluorophores covalently attached to single
macromolecules, so hydrolysis rates can be  measured
using simple fluorescence techniques. For this study, the
fluorophores are covalently linked to polysaccharides,
which comprise a significant proportion of total organic
matter. The sensitivity of these probes should permit
measurement of hydrolysis rates at picomolar levels;
ease of analysis means that a large number of samples
can be surveyed rapidly.
      Two different polysaccharides as well as lower
molecular weight oligo-saccharides with different sets of
fluorophores have been covalently labeled to determine
optimal probe characteristics and coupling strategies.
Based on initial laboratory investigations, we are focus-
ing on two fluorophore pairs. Laboratory investigations
demonstrate  that  the  double-labeled  probes  exhibit
fluorescence energy transfer. Donor quantum yield is
significantly higher in single-labeled  as compared to
double-labeled probes.  Field tests of these probes are
being conducted; the addition of fluorescein/Texas Red-
xylan  to  porewater  from  Delaware  Bay  sediments
demonstrated an increase in the ratios of donor to
acceptor emission with incubation time (see left side of
Figure 1). Concurrent measurements using gel perme-
ation  chromatography  showed  changes in substrate
molecular weight (hydrolysis of xylan; see right side of
Figure 1) consistent with the  fluorescence measure-
ments. Changes in fluorescence characteristics were
observable on much shorter time  scales than changes in
substrate chromatography;  the  fluorescence analyses
also required much  less time than chromatographic
analyses (< 4 min. vs. > 1 hr).  Our technique already
allows us to screen rapidly large numbers of samples to
evaluate enzymatic activity and select samples  for fur-
ther investigation.
      This project focuses on using purified enzymes to
calibrate  the  fluorescence responses  of the  double-
labeled probes  in the laboratory.  The synthesis  of
several new double-labeled probes of different chemical
structures are being planned to test for  a wider range of
enzyme activities. This expanded suite of probes will
then be used to make larger scale field measurements of
enzyme activities in the water column and sediments.
Preliminary studies have indicated that we may find
distinct differences in relative  activities  of  specific
enzymes  in sediment pore waters as compared to the
water column,  and an expanded suite of probes will
enable us to investigate this possibility in detail. These
fluorescent-labeled probes will provide the means of
making rapid, high-resolution measurements of specific
enzymatic activities with minimal  disturbance to the
naturally complex microbial community, providing a
more realistic picture of the transformations that actually
take place.
                                                   45

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Watersheds and Wetlands: Large
Scale Disturbances and Small Scale Responses
Charles Andrew Cole, Robert P. Brooks, and Denies Heller Wardrop
Penn State Cooperative Wetlands Center,  University Park, PA
      As pressures for development increase and more
wetlands are permitted for modification or destruction,
our understanding of the watershed-level impacts of
these activities remains poor.  In addition, although
wetland restorations usually try to provide a replacement
that  looks and  operates  like a natural wetland, the
complexity of these systems may limit success. The
goals of this project are to: (1) assess characteristics of
natural  reference  wetlands  and created wetlands,  by
watershed, disturbance, and hydrogeomorphic (HGM)
category, to determine improved design characteristics
for  created  wetlands  implemented  for  mitigation
purposes;  (2) assess  succession in reference wetlands
using seed banks and soil dating; (3) assess character-
istics of created wetlands of different ages to begin
determining successional pathways; (4) compare created
wetlands with the reference sites to see if the  created
wetlands  are  successful  in  any  sense; and  (5) use
disturbance theory  to evaluate the  impacts  of the
surrounding landscape on both reference and  created
wetlands and their successional trajectories.
      The Penn State Cooperative Wetlands  Center
(CWC)  has  been assessing wetland  structure  and
function by classifying a wetlands' position within the
watershed  (e.g., headwater) and by its source of water
(e.g., groundwater).   Research  indicates that sur-
rounding disturbance impacts wetland function so that
events occurring at the watershed scale will likely be
revealed by changes  in function at the wetland scale.
The  CWC is  studying  63  reference wetlands  by
measuring  soil  characteristics, sedimentation  rates,
hydrology,  basin  morphometry,  plant  community
composition,  biomass  and production, seed  banks,
macroinvertebrates,  and wildlife habitat. The investi-
gators have found that soil characteristics, sedimentation
rates,  hydrology,  and  plant  communities  all  are
predictable (to some  degree) based on position and
sourcewater for the wetland.
      This  project  involves  studying   12  created
wetlands and  looking  at  the  same  parameters.  Our
research found that created wetlands  are not at all like
natural wetlands in Pennsylvania. They have different
physical  structure, different soils, markedly different
hydrology, and  different  plant communities.  A  pre-
liminary time series analysis shows that created sites are
not converging in structure or function towards natural
wetlands.
      In addition, this project is considering the historic
use of each of several watersheds to  determine if past
land use practices resulted in any discernable changes in
wetland plant communities.  A subset of the reference
wetlands is being examined by sampling the soil column
and dating  soil layers using Cs137. Once the soil layers
are dated,  the seeds in  the  seed bank will then be
germinated or otherwise  identified to develop a plant
community history.   A comparison of land  use changes
since the 1930's for a 1 km radius around each site will
be made to see if land use has been reflected in the seed
bank (i.e.,  the plant  community).  To date, successful
dating of the soil layers  has been accomplished, and the
process of collecting seed bank samples is  under way.
      Our  findings  are  helping  to  develop data on
wetland function by HGM subclass. HGM  will soon be
the primary   wetland  classification  and  assessment
protocol within the United States, and  we will be able to
provide real data towards assessing functions. Also, we
are adding to basic ecological knowledge on succession
by looking  at historic seed banks in reference wetlands
and at chronological sequences in created wetlands.  This
will help to define permit expectations and requirements
as well  as  aid in streamlining an already  contentious
mitigation process.  We are just now beginning to ex-
plore  the  impacts of watershed-level  activities on
individual wetland function. This data will allow for a
better understanding of large  scale decisions on  site-
specific processes.
                                                    47

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Water Authority "AS":  1994-1996
  McCall Darn "AS": 1995-1997
        •dr
H'H«! H« J|«Hs
                                                                                                                        Bald Eagle Creek "C720": 1996-1997
  100
  75

1 I


I -
G -50
  -75
                                                                                                                           BESPPEM"AS": 1994-1996
                                                   Figure 1.  Examples of wetland hydrographs by HGM subclass.

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Integrated Ecological Economic
Modeling and Valuation of Watersheds
Robert Costanza, Roelof Boumans, Tom Maxwell, Ferdinando Villa,
Alexey Voinov, Helena Voinov, and Lisa Wainger
Institute/or Ecological Economics, University of Maryland, Solomons,  MD
      This  integrated modeling effort is intended to
provide tools to manage water quality and ecosystem
responses by synthesizing data from intensive experi-
mental  studies on small scales  and using  dynamic
models to extrapolate that information to large drainage
basins.  A grid-cell, process-based model developed for
the 2,500 km2 Patuxent River watershed in Maryland
integrates information over areas of spatial, temporal,
and complexity scales.  A unit model calculates stocks
and flows  for water, nutrients, plants,  and animal
populations within each cell, and cells are linked to
simulate water and material fluxes across the landscape
(see Figure 1). An economic submodel estimates land
development patterns and effects on human decisions
from site  characteristics, ecosystem properties, and
regulatory paradigms. Patterns of development are used
in the ecological model  to estimate effects of human
activities.  The software we have developed links geo-
referenced and other databases to the model for input
and calibration. We have developed methods and tools
to simplify and partially automate model calibration and
result analysis.
      Preliminary model runs  show good agreement
with measured data for  components of the model at
several scales and for several years.  Streamflow  in the
hydrologic submodule was compared with 2 years of
gage data after the initial calibration year.  The model
predicted overall  surface and groundwater flow across
the watershed  generally within  10 percent of daily
values,  although some large  flood peaks deviated to a
larger extent  from  measured values.   Several  sub-
watersheds were used to test model behavior at smaller
spatial extents  (23 km2 to 940  km2).   This modular
approach to calibration worked well for the hydrologic
component, with little fine tuning required when switch-
ing scales.  Forest biomass calculated by the model
matched boundary conditions established from calibra-
tion data.  Temporal and spatial calibration  data on
annual increments  to forest biomass were developed
using species-specific  tree ring records and spatial
distributions in the watershed.  Seasonal and long-
itudinal dynamic records for phosphorus and nitrogen
concentrations in the river were compiled from numer-
ous sources for calibration.  Unit model estimates were
constrained to boundary conditions for nutrients and
deposited matter.
      The investigators developed methods to examine
the effect  of spatial pattern of land use on ecological
indicators developed from model variables. Preliminary
results from an adaptation of the hydrologic model for
a suburban watershed suggest that sensitivity to  land use
proportions and spatial pattern decrease sharply above
60 percent high-density residential and commercial uses.
Stream buffers in particular lose the ability to  mitigate
storm peak flows at this level of development. These
effects will be tested in the full landscape model.
      Scenarios are being  developed to investigate the
implications of changing land management policies on both
the ecological and economic components of the system.
These include: (1) clustered vs. sprawl for agricultural and
urban areas development; (2) implementation of nutrient
and water management practices for agricultural and
urban areas; (3) development with varying proportions
of forest,  agricultural,  and urban uses; and (4) spatial
arrangement of land uses. Dynamic models examine the
numerous  competing effects of human and naturally
imposed variation to suggest potential long-term impacts
to system function and resiliency.  Model results can
help guide land use policy and help to develop goals for
ecosystem quality indicators.
                                                   49

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Figure 1.  Elevation of the Patuxent Watershed. Grid cells shown are 3 km.  Model grid cells range from 0.2-1 km.
                                                  50

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Oyster Reefs as Structural and Functional
Components of Tidal Creeks;  An Ongoing Ecosystem Experiment
Richard F. Dame, E. Koepfler, L. Gregory, T. Prins
Coastal Carolina University, Conway, SC
D. Allen, D. Bushek, C. Corbett, D. Edwards, B. Kjerfve, A. Lewitus, and J. Schubauer-Berigan
Baruch Marine Laboratory, Georgetown, SC
                                      ATMOSPHERE
                                         WATER
       PARTICULATE
                          FILTRATION
     SEDIMENTATION
                  DISSOLVED
METABOLITES
                                     MINERALIZATION
                    RESUSPENSION

                                  RELEASE
                                   AEROBIC SEDIMENTS
             DON
                                                            DEITRIFICATION
           MOBILE PHQSpHATBpO
                                                            METHANOGENESISCH
                       Figure 1. Ecological processes coupling oyster reefs to tidal creek waters.
      Shallow tidal estuaries are highly productive and
ecologically diverse systems because the complexity of
the physical environment  generates  a multitude of
habitats.  These estuaries are threatened by environ-
mental changes such as pollution, overharvesting of
fisheries, and sea level  rise. The investigators in this
project are conducting an ecosystem-level experiment to
examine the structural and functional role of oyster reefs
in tidal creeks (see Figure 1). Eight similar tidal creeks
were chosen to standardize for oyster dry body biomass
to creek water volume  at full bank conditions.  Each
creek has  been  topographically and bathymetrically
surveyed, and these data have been used to determine
area and volume relationships for each creek basin.
      Preliminary findings  from the first, premanipu-
lation year include: (1) less mature creeks with upland
interfaces have lower nutrient concentrations than more
mature creeks nearer  the ocean;  (2) oyster growth
appears to be negatively correlated with distance from
the creek mouth; (3) grazing by bivalves and micro-
zooplankton is important in controlling phytoplankton
year round; (4) nutrient limitation effects are only found
during the late fall to early spring period (see Figure 2);
(5) tide pools within creeks accumulate nutrients and serve
as refuges  for nekton; and (6) differences  in nekton
assemblages among creeks appear to be determined by
creek geomorphological  and flow pattern variations.
      In the manipulation year, which begins in March
1998, oyster  reefs will be removed from four creeks,
and their structure and function will be compared to
four  manipulated creeks  for 1  year.  The  replicated
experimental design will further the basic understanding
of the role of oyster reefs in sustaining the ecosystems
that they dominate.
                                               51

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                                  Growth response to
                                  grazer reduction
                              Growth response
                                 toNH
     Spring
Summer        Fall
         Season
Winter
Figure 2. Predicted phytoplankton responses in creeks without oyster reefs compared to creeks with oyster reefs.
                            52

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Tracing the Fate of Nitrogen
Inputs From Watersheds to Estuaries
Linda A. Deegan and Bruce J. Peterson
The Ecosystems Center, The Marine Biological Laboratory, Woods Hole, MA
      This project's  goal is to illuminate the trophic
pathways by  which  river-borne dissolved inorganic
nitrogen is incorporated into and processed by  food
webs in the upper, tidal freshwater-oligohaline reaches
of coastal plain estuaries.    Specific research  goals
include determining: (1) the key species in pelagic and
benthic food webs; (2) the strength of linkages between
benthic and pelagic food webs; and (3) the importance
of watershed nitrogen inputs  to consumer organisms.
To meet  these goals, we conducted the first whole-
ecosystem stable isotope tracer amendment to be carried
out in an estuary. First, 15N-enriched nitrate was added
to the upper reach of the Parker River, Massachusetts,
during  the summer of  1996 and,  through intensive
sampling, the progress of the isotopic label through the
water column, organisms,  and sediments was tracked
during the amendment  and for several months there-
after.
      Three major sources  of organic matter  fuel the
consumers of the upper Parker River estuary: (1) most-
ly microscopic pelagic  (water column) producers, (2)
benthic  (sedimentary)  primary  producers,  and (3)
detritus derived predominantly from the tidal freshwater
marsh (see Figure 1).  The pelagic diatom, Actinocydus
normanii, was identified as a critical determinant of the
fate of watershed-derived nitrogen in the Parker River
estuary during  the  summer  period  when biological
activity reaches its maximum.  A bloom of this single-
celled alga rapidly assimilated watershed-derived nitrate
entering the upper estuary and was the primary vector
of this  nitrogen to benthic  and pelagic  food webs.
Among those species feeding directly on the diatom, or
on an herbivorous intermediary, up to one-half of their
assimilated nitrogen was derived from this source, and
thus from new nitrogen entering from the watershed.
On the basis of 15N tracer incorporation, 25-50 percent
of the dietary  nitrogen assimilated by planktivorous
fishes, such as juvenile alewife and white perch,  was
derived originally  from A. normanii by way of their
copepod food.  Surprisingly, we found that the tracer
was  rapidly assimilated by major crustacean benthos
(i.e., mud crabs, amphipods, and grass shrimp), indicat-
ing an unexpectedly strong linkage of pelagic primary
production with benthic secondary production.  It  is
estimated that 20-40 percent of their assimilated nitro-
gen derived from the pelagic diatom.  A. normanii was
incorporated into surficial  sediments, providing both a
food  reserve  for  benthos, and  a probable  nitrogen
source for locally important benthic primary producers
that fed small, productive  intertidal fauna. Other spe-
cies  were largely  uncoupled  from  A. normanii  and
depended more on unlabeled food sources, derived most
likely  from  freshwater   marsh  detritus.  Abundant
benthic-feeding fishes  (e.g., white sucker, mumrnichog)
assimilated the tracer minimally by feeding primarily in
a detritally based food web.
      This project is among the first to measure directly
the relative trophic importance of in situ primary pro-
ducers and detrital organic matter in an intact ecosys-
tem, and it is the first to do so by using a stable isotope
tracer  in an estuary.   The research has made a major
contribution to understanding the fate of watershed-
derived nitrogen in the critical zone where rivers inter-
face with estuaries, a region that has received relatively
little study despite its potential to control nutrient trans-
formation and transport from uplands to coastal waters.
                                                    53

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        Trophic Pathways of Nitrogen Flow in the Upper Parker River Estuary
    §

    I
    v>
          grass shrimp
           mud crab
           amphipods
           Nereis
                                                pelagic copepods
                                                  E. affinis
                          tidal freshwater
                          marsh detritus
                                  mummichog
                                 white sucker
     deposited detritus
                                        benthivores
                                                                                  planktivores
    alewife
white perch (Juv-)
                                  ^^Tnjcrophytobenthos, bacteria
                                 N
                         remifieralfeation
sediments
striped bass
                    Figure 1. Conceptual model of nitrogen flow through biota of the upper Parker River estuary, showing trophic linkages supported
                           by isotopic evidence, feeding ecology, and habitat use.

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Probing the Relationship Between Fulvic Acid Aggregation,
Metal  Ion Complexation, and the Binding  of Organic Compounds
A. Dixon ', W.R. Carper2, and C.K. Larive '
' Department of Chemistry, University of Kansas, Lawrence, KS
2 Department of Chemistry, Wichita State University,  Wichita, KS
                                                                                     D Region  1
                                                                                     13 Region2
                                                                                     H Aromatic
                                                                                     • Average
                Sm Angle      BPPLED     Spin Echo-   Relaxation-
                  X-Ray                       BPPLED      BPPLED
 Figure 1. This figure shows the results of NMR experiments for the analysis of the average molecular size of fulvic acid solutions.  These results
         are compared with the reported radius of gyration for this sample using small-angle X-ray scattering.  Region 1 refers to the portion
         of the proton NMR spectrum containing CH3, CH2, and CH groups adjacent to aromatic, carbonyl and carboxyl groups.  The standard
         NMR experiment, the Bipolar Pulse Pair Longitudinal Encode-decode pulse sequence (BPPLED), produces a radius of gyration that
         is the average over all fulvic acid molecules and is directly comparable to the small-angle X-ray scattering results. We have used two
         spectral editing methods based on spin-spin coupling (spin echo) and T, relaxation to modify the standard BPPLED experiment to detect
         fractions of die sample that are either lower or higher in molecular weight than the average. All three NMR experiments are being
         used to see how the average size of each fration changes as a result of cadmium complexation.
       Soluble organic acids (humic substances) leach
 into ground and surface waters from soil and plant
 organic matter.  Humics play a key role in water quality
 because they  are the major constituents of dissolved
 organic carbon and they are important in controlling the
 bioavailability and transport of toxic metals and organic
 pollution. The goal of this project is to develop a better
 understanding of the physical and chemical properties of
 natural aquatic organic materials such as fulvic acids.
 Our approach makes use of the high specificity of nuclear
 magnetic resonance  spectroscopy (NMR). NMR pro-
 vides information about the local chemical environment
 of a molecule or chemical group. For example, binding
 constants of NMR active metal ions (such as cadmium)
 can  be  measured by monitoring the  concentration-
 dependence of the NMR chemical shift, hi addition, the
 NMR chemical  shift also yields information about the
 local chemical environment of the metal.  In this way,
 it is possible to determine which functional groups (such
 as carboxylate or amine) form the complex.
      NMR relaxation   measurements and  detailed
molecular mechanic calculations also are being used to
further elucidate the nature of the  metal-fulvic acid
complex. One  interesting facet  of this  project is the
relationship between metal complexation and fulvic acid
aggregation. NMR experiments have been modified for
the measurement of diffusion coefficients to make them
more specific for different classes of compounds.  In this
way, the extent to which metal ion complexation induces
aggregation in a fulvic acid sample can be examined.
The  high  specificity of NMR can provide additional
information about the  fulvic acid aggregates when
compared  with  that provided  by  other analytical
methods, which could be used for this purpose, such as
small  angle X-ray  scattering  and dynamic  light
scattering.  The ultimate goal of this project  is to use
NMR and specially labeled compounds to examine the
extent of metal binding  to a fulvic acid solution,  how
metal ion  binding affects the aggregation of the fulvic
acid sample, and how metal-induced aggregation affects
the binding of organic  pollutants  by the fulvic  acid
aggregates.  The molecular level specificity of our an-
alytical  methods will increase our  knowledge  of the
molecular-level behavior of aquatic  fulvic  acids.  This
information will be useful  to  develop a better under-
standing of how the chemistry of fulvic acids contributes
to the availability and transport of pollutants in aquatic
ecosystems.
                                                    55

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 Diffusional Rate Limitations in Heterogeneous
 Porous Media:  Model Structure, Scale, and  Geologic Characterization
 David L. Freyberg and Paul V. Roberts
 Department of Civil Engineering, Stanford University, Stanford, CA
                                                                       (B)
                                                           80'^
                                                           60
                                                                 20
                                                                                80
                                                                      40    60
                                                                       [cm]
 Figure 1. Regions of low groundwater velocity within a point bar deposit. (A) Natural log of the hydraulic conductivity (cm/s) for a vertical
         section. (B) Regions of low velocity (dominant flow direction is left to right): mean velocity magnitude=4.TxlO^cm/s; dark regions
         are those with velocity magnitudes less than 4.8xlO"5cm/s.
       The goal of this project is to enhance our ability
 to predict the transport and fate of hazardous chemicals
 in subsurface waters by increasing our understanding of
 the nature and significance of diffusion during transport
 in saturated, heterogeneous porous media.  The objec-
 tive is to develop the understanding  and tools necessary
 to  evaluate  and  predict the relative  importance of
 diffusion, or diffusion-like  processes, over a range of
 spatial scales (from grains to regionals).  The approach
 rests on a program of  numerical experimentation and
 analysis examining: (1) the mathematics of multiscale
 diffusional rate limitations  on  sorption; (2) the inter-
 actions  between  geologic  structure,  heterogeneity,
 upscaling, and real and  apparent transport process rep-
 resentation in mathematical  models; and (3) physically
 based aggregation schemes that are consistent  with
 different model formulations of transport processes and
 multiscale diffusional rate limitations.  The investigators
 are  using  both  random-field  models  and  a  very
 fine-scale,  quasideterministic, numerical model to  re-
 present spatial heterogeneity and uncertainty.
      The researchers  have developed and applied a
 model to analyze a set of laboratory column experiments
 conducted  to determine the rate at which trichloro-
 ethylene (TCE)  can be removed from natural soils.  A
 wide range of  diffusional  time  scales  for the TCE
 sorption-desorption process,  spanning up to 15 orders of
 magnitude, has been found.  To investigate  the effects
 of a distribution of contaminant sorption time scales in
 macroscopically homogeneous  aquifers, the temporal
 moments of a contaminant  pulse were  examined.  A
 distribution  of  sorption time scales was found to not
 affect the first two temporal moments,  but can affect
 strongly and even dominate the third moment.  In ex-
 tending the analysis to heterogeneous  aquifers,  the
 statistics of the arrival times of a contaminant are being
examined after  transport through  a hydraulic  con-
ductivity field. Preliminary results suggest that we will
be able to distinguish among scenarios in which:  (1) the
heterogeneity of the  aquifer controls the spread  of
arrival times,  while  sorption can be treated as  an
equilibrium process; (2) the kinetic sorption controls the
spread of arrival times, and the aquifer can be treated as
macroscopically homogeneous; and (3)  both hetero-
geneity and kinetic sorption contribute significantly to
the spread of arrival times.
       Pure advection  through spatially heterogeneous
groundwater velocity  fields itself can lead to skewed
breakthrough curves.  In addition, velocities in some
regions of a heterogeneous velocity field may be so low
that solute transport is dominated by diffusion, even
though the mean velocity is relatively high. To inves-
tigate the nature of low-velocity regions, the investi-
gators began by exploring  simple geometries such as
spherical  inclusions.   Subsequently, this project has
examined simulated flow through a  highly  resolved
representation of a point-bar deposit. Preliminary results
show that regions of very low velocity do exist, and that
the spatial  structure of these regions is complex and a
strong function of the conductivity heterogeneity as well
as of the mean head gradient (see Figure 1).
      The results from this research will  contribute to
a fundamental  understanding of  the  impact  of dif-
fusionally  rate-limited  sorption   and   diffusionally
controlled slow release from zones of low velocity on
transport, with applications to contaminant remediation
design, site characterization, and the interpretation of
field monitoring and laboratory experimental data.
      Continuing   work  is  focused  on  exploring
heterogeneity-kinetic sorption interactions in  different
heterogeneous settings; characterizing  low-velocity re-
gions and their relationship to geologic structure and
flow as  well as transport boundary  conditions; and
exploring the roles of aggregation, upscaling, model
formulation, and spatial structure in predictive modeling
of transport.
                                                    56

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Integrating Planning, Forecasting,  and Watershed Level Ecological  Risk
Assessment Techniques: A Test in the Eastern Cornbelt Plains Ecoregion
Steven I. Gordon and Andy Ward
The Ohio State University, Columbus, OH
Dale White
Ohio Environmental Protection Agency,  Columbus, OH
      The objectives of this research are: (1) to test the
relationships between biological conditions of streams
and the nature and distribution of human activities on
the watershed; (2) to demonstrate methods for linking
physical models of urban and agricultural  impacts on
runoff volume and runoff quality; (3) to define  the
relationships between physical model forecasts and the
biological conditions of streams; and (4) to integrate all
of the findings into an expert system to  be used by
planners.
      During the  past year, we have been working
simultaneously in three areas: (1) assembly of a com-
prehensive regional database and testing of watershed
scale relationships between biological measures of water
quality  and watershed characteristics; (2)  testing  the
impacts of different scale soils data on our predictive
models   of  hydrologic  response; and (3) finalizing
methodologies and data  for  preparing and analyzing
riparian zone and  detailed  watershed data aimed  at
testing models at larger scales.
      The   USEPA    RF3    stream   Geographic
Information System (GIS) files were used  as the basis
to divide the 4 Eastern Cornbelt Plains region into two
distinct  sets of watersheds—25 coarse scale  watersheds
and  137 fine scale watersheds.   For the  region as a
whole, Landsat imagery was  used to define the major
land use  categories.   A number of other data were
compiled using the  same geographic base, including
the  IBI  (Index  of Biologic  Integrity),   the  major
dependent variable in our regional scale analyses.  The
data were collected at 522 locations  throughout the
watersheds.
      At this regional scale,  a number  of empirical
relationships between the IBI as a measure of biological
quality and some proxy variables for point and nonpoint
water pollution stresseshave been tested. More detailed
modeling at the  watershed scale has proceeded  by
testing the impact of soils data at  two scales on the
results of our sediment model.   A test of the model
behavior using two soils data at two different scales has
been completed. Finally, work at the riparian zone level
has led  to a  strategy for extracting relatively detailed
vegetation-type assemblages and patterns in the riparian
zone using Landsat imagery.  A new set of techniques
will be used in the  coming year to derive  the most
detailed data sets and to  test empirical models at  this
scale against our regional scale and watershed scale
empirical and physical models.
      A set of  empirical models for  the  Eastern
Cornbelt Plains Ecoregion has been  defined, which
explains significant amount of the variation in IBI levels
across watersheds.  Table  1 compares the results for the
larger  scale  (more  detailed)  and   smaller  scale
watersheds.
      The significant variables  are the percentages of
dense  urban settlement,  and  several measures  of
watershed condition and location,  including  stream
order, substrate type, and pool  and riffle  run quality.
Adding an   estimate of chemical  stressors  using
STORET  data  produced significant  improvement in
model results.  IBI levels also are strongly influenced
by upstream and downstream IBI.
      Model runs made for the  Big Darby Creek
basin with the  two alternative  sources of soils  data
lead to different conclusions concerning scale changes.
The predicted  annual  flow,  sediment,  and  nitrate
losses for the analysis with conventional tillage and no
subsurface drainage are presented in Table 2.  There
is no significant difference between the mean annual
flow and sediment values predicted by using data of
finer vs.  coarser resolution.  However, the  mean
annual nitrate loss prediction based on the two soils
databases  are  significantly  different  due  to  the
approximately  10 percent difference  in  the loss  for
1992. There are considerable  differences  in the flow,
sediment,  and  nitrate prediction  for  individual  map
units of fine vs. coarse resolution (not presented).  The
results suggest that coarser resolution data can be used
to model hydrological responses at a watershed scale
but  should not be used at a field scale, or if the goal
is  to predict responses  for  short time  periods. It
should be noted that in this study,  the model  was not
calibrated and the results  only provide an indicator of
the   relative responses   associated  with   different
agricultural  practices and  soil characteristics. It is
probable that sediment losses are underpredicted  and
some  of  the  model  input  variables need  to  be
adjusted.
                                                   57

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                Table 1.  Regression models for mean IBI at the watershed scale.
Coarse Watersheds (25)
R2 Values
R2 Values
R2 Values
.384
.494
.694
Fine Watersheds (137)
.409
.525
.703
Significant
Independent Variables
Dense urban; substrate;
stream order;
Pool/Riffle
Adding chemical
conditions based on
ranking of segment by
STORET readings
Adding upstream and
downstream IBI to
account for spatial
autocorrelation
Table 2. Comparison of annual flow depths, sediment losses, and nitrate losses for conventional tillage
        without subsurface drainage.
   Precipitation
                       Flow Depth (mm)     I  Sediment Loss (ton/ha)   I    Nitrate Loss (kg/ha)
                   STATSGO     MUIR     STATSGO     MUIR      STATSGO    MUIR
1991
1992
1993
1994
1995
Mean
718
792
931
696
937
815
197
211
264
155
317
229
198
226
270
160
318
235
3.0
3.7
3.7
3.9
6.8
4.2
2.8
3.9
3.8
3.9
6.9
4.3
9.3
16.2
9.9
6.9
10.7
10.6
9.3
17.8
10.2
6.8
11.0
11.0
                                          58

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Development of Geomorphological Artificial
Neural Networks (GANNs) for Modeling Watershed Runoff
Rao S. Govindaraju
School of Civil Engineering, Purdue University, West Lafayette, IN
     Input
                                                                             Weighted
                                                                             Summation
                                                                                                  Output
                              Figure 1. A schematic of modular artificial neural network.
       Predicting streamflow  is a  fundamental com-
 ponent of a wide range of watershed-modeling efforts.
 In this project,  geomorphological  artificial  neural
 network  techniques  have  been  applied  to  predict
 watershed response to precipitation and temperature.
 These  networks can be used to resolve watersheds at
 different spatial and temporal scales through a massively
 interconnected network of  processing units.  System
 architecture is determined by the topology  of  the
 watershed.   Network development  is  characterized
 through  an   iterative  training  procedure.   Model
 calibration involves the selection of the optimal values
 of connection strengths and processing-unit thresholds,
 so that the network best reproduces historical stream-
 flow records for the watershed. This  requires a large
 number of training examples,  in terms of historical
 precipitation and streamflow records for a watershed.
 Once network training has been accomplished, it can be
 used to forecast streamflows for the watershed based on
 rainfall and temperature inputs. The model preserves
 some geomorphological and probabilistic characteristics
 of watershed  response as represented  by historical
 records.   Data from several Kansas watersheds  are
 being used to test the  performance of the model.   To
 avoid  problems  associated  with  over-training,   an
 additional cross-validation  data set has  been used to
 determine optimal training and network architecture.
 The  data set is partitioned into three portions: (1) a
 training set, (2) a cross-validation data  set, and (3) a
 testing data set.  Historic  records have been used to
 determine  the time  lags for watershed response to
precipitation  and  have  been  incorporated  in  our
modeling effort.  A cross-correlation analysis between
the average monthly precipitation and watershed runoff
revealed that the past 3 months  of rainfall data would
have a significant impact on the runoff for some Kansas
watersheds.
      Singular   neural   networks   predict   average
streamflow characteristics satisfactorily, but they have
not been successful in predicting high  and  low runoff
events.  A number of singular networks were arranged
in a layered structure to form a  modular network, and
stream discharges  were partitioned into three classes,
each using a singular network (see Figure 1).  Each
singular network or module covered a particular range
of the dataset, and a final module assigned weights to
the prediction of the individual modules.  Each module
was developed based on the data  in a class, but the final
network was based on the entire training dataset. This
kind of network resulted  in an improved prediction of
streamflows for three Kansas watersheds.  Inclusion of
rainfall and temperature from previous months allows
the model to better represent watershed conditions.
      The preliminary success of modular networks
suggests that different rules are required to represent
extreme and average events.  The architecture of the
modular network can be  adapted to represent a stream
network  and  includes  the  geomorphology  of  a
watershed.   Future efforts are  planned to extend our
applications  of  geomorphological  artificial neural
networks and consider  the  influence of  spatial  and
temporal resolution on watershed-runoff computations.
                                                    59

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Physicochemical
Mechanisms Governing Virus Filtration
Stanley B. Grant and Terese M. Olson
Department of Civil and Environmental Engineering, University of California, Irvine, CA
Mary K.  Estes
Division  of Molecular Virology, Baylor College of Medicine, Houston, TX
                                                          3.00
                                                          2.00
                                                           1.00
                        ,23.4  r»M
                                                                               11.7  HM
                                                                              "0.0  nw
                                                                  NanoScope
                                                                  Scan size
                                                                  Setpoint
                                                                  Scan rate
                                Contact  AFM
                                       3.000  JJM
                                      -2.025  U
                                       1.001  Hz
                                                                  Number of  sanples
                                          256
Figure 1. Atomic force microscope image of Norwalk virus particles absorbed to a mica surface. This adsorption experiment was conducted
        in an aqueous solution at pH 5 and lOOmM NaCl. Under these conditions, the virus is positively charged and the mica is negatively
        charged.  Consequently, there is no electrostatic barrier to virus adsorption.
      The use of reclaimed sewage for groundwater
 recharge raises  a number of important human health
 issues.  Foremost among these is the possibility that
 human enteric viruses present in the reclaimed sewage
 may be transported to production wells and, ultimately,
 end up in a water distribution system. Laboratory-scale
 experiments  are being conducted that are aimed  at
 identifying the geochemical features of a recharge basin
 influencing the removal of viruses from groundwater by
 physicochemical filtration (see Figure 1). Initial experi-
 ments of this project examined the influence of water
 pH on the surface charge of recombinant Norwalk virus-
 like particles  and their filtration  rates in packed beds  of
 quartz sand.  Many  outbreaks of gastroenteritis in the
 United States are caused by waterborne or foodborne
 transmission of Norwalk virus.  The initial  results from
 this project  suggested that pore water pH may be the
most important factor in deterrnining the capacity of
groundwater systems to provide natural disinfection by
physicochemical filtration.
      Further experiments have used  a total of four
different viruses, including three coliphage (a somatic
phage, and two  different male-specific phages) and
Norwalk virus particles  generated using a baculovirus
expression system.   In  all cases, we found that pore
water chemistry dramatically influences the removal of
these viruses in packed beds of quartz sand.  When the
filtration  results are  combined with microelectro-
phoresis measurements of the viruses' surface electrical
potential, the results are generally consistent with the
predictions of  classical theory—that is,  electrostatic
interactions between the viruses  and quartz sand ul-
timately  determine a filter's ability to  remove viruses
from water.
                                                   60

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Watershed Impacts on Sediment Pollution
History and the Viability of the Zooplankton Egg Bank
Nelson G. Hairston, Jr., and Colleen M. Kearns
Section of Ecology and Systematics, Cornell University, Ithaca, NY
Charles T. Driscoll
Department of Civil and Environmental Engineering, Syracuse University, Syracuse, NY

                                            log Qig Hg/g Sediment)
                                                     10  -10
  E
  u
  Q.
  »
  Q

  O
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              1950
              1930
              1890
              1850
              1810
              1770
              1730.
Onondaga   1690'
   Lake      1650
                                                             10
                                                     100   0

j %Hatch
i

f
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> Lake

1990-
1970-
1950-

1930-

1890
1850
1810-
1770-
1730-
                              20
40
60
80
100
                                          % Hatch of Diapausing Eggs


 Figure 1.  Comparison of the mercury concentrations (logarithmic scale) and hatching success of zooplankton diapausing eggs in the sediments
         of two lakes in central New York State. Oneida Lake has only received metals from atmospheric deposition on the watershed.
         Onondaga Lake has experienced direct inputs from industry effluent.  Sediment dates determined by 210-Pb analysis.  Note that
         analysis of hatching success in Oneida Lake is only partially complete.
      The zooplankton  community of lakes and its
 response to environmental change may be substantially
 impacted by the diversity, viability, and population
 dynamics of long-lived diapausing eggs buried in lake
 sediments.  The history of environmental change in
 lakes is  manifested in the pollutants that  enter their
 watersheds and are transported to the lake, where they
 ultimately accumulate in the sediments.
      This project is  investigating the impact of this
 pollution on diapausing egg viability.  The two major
 sources  of  heavy  metals  in  lake sediments in the
 Northeastern United States are:  (1)  direct inputs of
 industrial effluent to  the watershed or lake and (2)
 atmospheric inputs that are deposited within the wate-
 rshed and wash into the lake. Replicate sediment cores
 taken at  two lakes in  central  New York State reflect
 their distinct histories  of industrial impacts.  In both
 lakes,  cores  were  sectioned,  dated by 210-Pb,  and
 analyzed for concentrations of metals, including  lead
 (Pb) and mercury (Hg).  In both lakes, fluxes of these
metals to the sediments increased from 1850  to the
mid-to-late 1900's.  The increase was initially due to at-
mospheric  deposition  to  the  watersheds, but in
                  Onondaga Lake, major inputs from chemical and steel
                  industries on the shore occurred during the 1960's,
                  1970's, and 1980's.
                        Sediment  enrichment  factors  (modern  metal
                  deposition/metal deposition in 1850) were on the order
                  of 2 to 3 for Pb and Hg in Oneida Lake, but between 30
                  and 120 for these metals  in Onondaga Lake.  These
                  deposition  patterns  will be compared with  those for
                  remote  lakes in  the Adirondack region of New York
                  State.  Diapausing eggs of crustacean and rotifer zoo-
                  plankton are found in abundance in the sediments of
                  both lakes (see Figure  1). Both show substantial hatch-
                  ing for eggs deposited 40 to  100 years in the  past.
                  There is no evidence for a reduction in egg viability in
                  Onondaga  Lake, as compared  with Oneida Lake, or
                  within Onondaga Lake as a function of sediment metal
                  concentration. Apparently, the  '"ametabolic" state of
                  zooplankton eggs in diapause protects them from metal
                  toxicity in the sediments.  These eggs may thus repre-
                  sent a source for the recovery of lake zooplankton
                  communities from past environmental pollution.
                                                   61

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The Role of Colloidal Particles in the
Transport of Chemicals Through an Agricultural Watershed
George M. Hornberger and Janet S. Herman
University of Virginia, Charlottesville, VA
James E. Saiers
Florida International University, Miami, FL
      Organic contaminants exhibit complex chemical
patterns as they are transported through watersheds to
streams.  This project has examined the chemistry of the
widely used herbicide,  atrazine, as it passes through
hydrologic systems—from its initial occurrence in the
unsaturated soil zone in agricultural areas through the
groundwater of a bedrock water-supply aquifer to its
appearance in a  stream draining the watershed.  The
goal has been to expand our understanding of herbicide
movement through the entire soil-bedrock-stream system
in an agricultural  watershed.   This information will
permit informed  management decisions about land use
and its impact on a sustainable water supply.
       Work is  being  conducted  on Muddy  Creek,
which drains  an  agricultural catchment in the Shenan-
doah Valley,  Virginia.  The research addresses three
primary questions:  (1) What factors control the varia-
bility in the concentrations of colloids and agricultural
chemicals in the aquifer-stream system at Muddy Creek?
(2) How do aquifer materials and  colloidal particles
affect the movement of  agricultural  chemicals such as
atrazine and hydroxyatrazine? (3) Do laboratory-based
observations  of  colloid-facilitated  transport of agri-
cultural chemicals provide a reliable estimate of trans-
port at the field scale?
      In this  project,  the  forms  and transport  of
herbicides  in the  groundwater flow  system  will  be
reported.  To evaluate in situ soil-water composition,
samplers were installed to collect  water held  under
different tensions in the unsaturated zone at three depths
at three sites in the catchment. Tension lysimeters were
installed in 8 cm diameter holes at depths of 15, 45, and
90 cm.   Zero-tension (pan) lysimeters, which  sample
water  flowing  through macropores  in the soil, were
installed by excavating a trench and fitting three pans at
the same  depths as the tension lysimeters. Soil-water
samples were collected, and water volume was recorded
from both types of lysimeters approximately every 4-6
weeks from April 1996 through June 1997. Samples
were analyzed  for atrazine, nitrate,  dissolved organic
carbon, and colloid concentrations.  In the clay-silt loam
at our field site, significantly higher concentrations of
atrazine were identified in zero-tension  lysimeters at
three depths (15, 45, and 90 cm) than in corresponding
tension lysimeters. Atrazine concentrations in the zero-
tension lysimeter were as high as 21 ppb, whereas the
corresponding concentrations in tension lysimeters were
only ~2 ppb (see Figure 1). Colloid concentrations in
the zero-tension lysimeters  were as high as  650 mg/L.
Although the differences between the particle-associated
atrazine and  the  dissolved atrazine  in the  pan
lysimeters are small, they are significant statistically
(p  =  0.027).  Concentrations  of nitrate are highly
variable and not significantly higher in  the pan
lysimeters than the tension lysimeters, indicating that
different  processes  may  control  the transport of
nitrate relative to atrazine.
      Our results indicate that: (1) transport of atrazine
is largely via macropores; (2) colloids are mobilized and
transported over distances  of at least 1  meter in  the
vadose zone  at our site; and (3) nitrate is not a good
surrogate  for judging atrazine transport.  Macropore
sampling is required to  estimate transport of herbicides
in the field, and colloid-facilitated transport of strongly
sorbing constituents  is  of potential  importance  at
agricultural sites such as Muddy Creek.
                                                   62

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 Figure 1.  Atrazine concentrations in (a) pan and tension lysimeters and (b) raw and filtered fractions from pan lysimeters.
                                                     63

-------
 Geomorphic,  Hydrologic, and Ecological Connectivity in
 Columbia River Watersheds:  Implications for Endangered Salmonids
 Hiram W. Li, Bruce A. Mclntosh, J.Boone Kauffman, Judith L. Li, and Robert L. Beschta
 Oregon State University, Corvallis, OR
 Patricia McDowell
 University of Oregon, Eugene,  OR
                    100n
                           Fishes
                   -o-Warmwater spp.
                       redside shiner
                       speckled dace
                       northern squawfish
                       largescale sucker

                    -iV'Coolwater spp.
                       mt whitefish
                       torrent sculpin
                       bridgelip sucker

                    -a- Coldwater spp.
                       chinook salmon
                       rainbow trout
                       ABODE   FGHI   JKLM
                      Warm   Cool            Cool           Warm    Cool
                                        Geomorphic reach
         Figure 1. Patterns of fish assemblage structure corresponding to temperature patches among stream reaches.
       Coldwater  fish  communities  throughout the
 Columbia River basin face high risks of extinction due
 to degraded freshwater habitats.   The  survival and
 persistence of these fish communities  will  depend on
 our understanding of the processes controlling stream
 temperatures and fish habitats. This project is investi-
 gating the interplay among geomorphic, hydrologic, and
 riparian-community  features  and  the  dynamics  of
 groundwater input and hyporheic flow in pristine and
 badly  disturbed streams  in  the  high desert of the
 Columbia River. Images captured using Forward Look-
 ing  Infrared (FLIR) videography  revealed  that the
 temperature  profile in a  watershed  disturbed by
 intensive livestock grazing, the Middle Fork John Day
 (MFJD), varies by as much as 50° C.   However,  a
 stream draining the wilderness basin,  the North  Fork
 John Day (NFJD), exhibits the normative temperature
 patterns. Patterns are similar throughout the year, with
 consistent magnitudes between stream differences in all
 seasons.
       Presently, there is no clear, single explanation for
 the temperature patches.  Ground truthing  has estab-
 lished that it is not an artifact. Intensive stream gaging
 at selected locations, dye release tests, and measures of
 vertical hydraulic gradient do not indicate that they are
 caused by hyporheic groundwater exchange, which is
 relatively minor in comparison to that documented in
 streams of the northern  Rocky Mountains.  Likewise,
 analyses of bed materials  suggest that hydraulic  con-
 ductivity may be low, although this will be decided with
 further analysis by ground penetrating radar to  map
potential pathways of hyporheic  flow  by  measuring
depth to bedrock and  defining alluvial  stratigraphy.
These studies were conducted in the MFJD and adjacent
drainages with similar geomorphologies.  Biogeochem-
 ical relationships between subsurface flows, sols, and
 vegetation in the riparian zone were characterized by
 measuring annual dynamics of water-table depth, soil
 redox potential, and chemistry of subsurface and stream
 waters.  In contrast to instream measurements,  pre-
 liminary results show some degree of lateral hyporheic
 exchange between the stream and riparian communities
 as documented by water temperature, redox potential,
 and dissolved organic carbon concentrations.  Livestock
 grazing appears to affect groundwater exchange.  Un-
 grazed riparian reaches were found to have greater  root
 bio-mass, soil organic matter, water infiltration rates,
 pore space, and lower soil bulk densities.
      Fish surveys in the two watersheds reveal  that
 longitudinal fish community patterns strongly reflect the
 temperature profiles.  There is a gradual shift from  cold
 water to warm water communities  in the wilderness
 stream system (NFJD); whereas, the relative composi-
 tion of community types in the disturbed stream (MFJD)
 follows the  distribution of temperature  patches  (see
 Figure 1). Subsurface flow within reaches was found to
 be very slow and poorly oxygenated, suggesting  that
 high stream  temperatures  may  force  organisms to
 choose between microhabitats  of cooler temperatures,
 but with  low dissolved oxygen versus  mainstream
 habitats that are  warmer,  but higher  in  dissolved
 oxygen. Water chemistry, primary production, macro-
phytic  and algal  biomasses, macro invertebrate abun-
 dances,  and fish diets at  12  sites along the MFJD were
to examine the effects of water temperature and possible
hyporheic up-welling/down-welling on the trophic  net-
work.  This project's analyses do not support the hy-
porheic corridor  concept  as an  explanation  for the
distribution  of  organisms,  but an  evaluation of the
factors dictating temperature conditions will be made.
                                                  64

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Resistance of Communities to Chronic Haloaromatic
Contamination From Biogenic and Anthropogenic Sources
David E. Lincoln, Sarah A. Woodin,  Charles R. Lovell, and V. Pernell Lewis
University of South Carolina, Columbia, SC
      Halogenated aromatic compounds are important
pollutants characterized by their toxicity, persistence,
and  accumulation in  the environment,  and abundant
production and frequent use in a variety of industrial
processes. This project examines the extent to which the
capacity  to  degrade naturally occurring halogenated
aromatic compounds determines the biological impact of
anthropogenic halophenol pollutants on marine benthic
communities.  Dehalogenase enzyme activities and ge-
netic probes are being used to assess the potential for
dehalogenation among the polychaete  worms,  which
dominate  these  sedimentary ecosystems,   and  the
worm-associated  bacteria,  as  well as other principal
community  members.  The enzyme  assays and DNA
probes plus induction experiments will enable us to
determine if dehalogenation potential is an important
determinant of organism  survival and persistence in
biogenically and anthropogenically  contaminated lo-
cations.
      Our findings to date highlight the abundance and
broad  distribution  of  benthic  macro-  and micro-
organisms capable of production  and/or degradation of
halometabolites.  A large  proportion of invertebrate
taxa, which are numerically dominant in coastal marine
sediments,  are  halometabolite  producers  and  have
enzymatic dehalogenation activity.  The native  worm
species—the most abundant intertidal benthic macro-
fauna—appear to be segregated among sites with regard
to their dehalogenation capacity, and this may be related
to the presence  of halophenols in their environment.
Twelve  reductively dechlorinating anaerobic  bacterial
strains have been  isolated from the  burrows  of  a
bromphenol-producing  and a nonproducing  species.
These   strains  reductively   dechlorinate  2,4,6-tri-
chlorophenol.
      A  survey  of  polychaetes  at an undisturbed
estuarine site showed that 40 percent of the organisms
contained halogenated metabolites. Additional surveys
indicated that native species appear to be segregated
among  sites  with  regard  to  their halgenation and
dehalogenation capacity, and this may be related to the
presence of halophenols in their environment.
      Twelve  reductively  dechlorinating  anaerobic
bacterial strains have been isolated from polychaete and
hemichordate burrow lining materials.  Dechlorination
of 2,4,6-trichlorophenol, with phenol as the apparent
terminal product, was initially rapid but declined after
successive subculturing of all strains.  A new method
for analysis of phospholipid fatty acid data, allowing a
more sensitive  detection of changes in community
composition, also was developed as part of this project.
      The investigators have initiated a series of species
transplant and halophenol contamination experiments in
the field to  assess  the  ability  of  worms that do  not
produce halogenated aromatic compounds to  survive in
their presence and to determine whether dehalogenation
activity  can be induced. Experiments creating refuge
from predators also are in progress to determine the role
of predators in controlling the representation of fauna
with halometabolites.
      Additional  near-term  activities  include   the
completion of purification and  characterization of the
bacterial dehalogenase enzymes and the development of
probes to characterize the genetic potential for dehalo-
genation by bacteria associated with the macrofauna.
Dehalogenation activities against brominated (common
biogenic) and chlorinated (common anthropogenic) halo-
aromatics  will be tested. We  have developed a new
method, based on neural  networks, for analyzing phos-
pholipid fatty  acid profiles  from sediment bacterial
communities. The new method is more sensitive than
conventional cluster  and principle components analyses
and yields  greater resolution of community composition
than previous approaches.
                                                   65

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Influences of Watershed Land Use on
Stream Ecosystem  Structure and Function
Judith L.  Meyer, E.A. Kramer, M.J. Paul, W.K. Taulbee
Institute of Ecology, University of Georgia, Athens,  GA
C.A. Couch
U. S. Geological Survey, Norcross,  GA
      Because natural  resource management is more
frequently conducted at the  landscape and ecosystem
levels, it must be determined whether the species  and
habitat-based approaches currently used by water-quality
monitoring programs provide data relevant to  assessing
interventions at the scale of the landscape or ecosystem.
This project has  two objectives: (1) to determine how
differing patterns of land  cover  and land  use  in  a
watershed alter stream ecosystem structure and function;
and  (2)  to compare measures of stream ecosystem
function (rates of community metabolism and transport
and cycling of carbon  and nutrients)  with traditional
chemistry-based, water-quality assessment and organ-
ism-based bioassessment.   The project approach  has
been to measure ecosystem  structure and  function in
streams draining  eight watersheds in the Chattahoochee
River basin; two streams drain watersheds in each of
four land use categories (i.e., forest, agriculture, urban,
and suburban).  Urban and suburban watersheds are
within metropolitan Atlanta.   Poultry is  the primary
commodity produced in the two agricultural watersheds.
The eight watersheds studied range in size from 47
200 km2; human population density ranges from 190
8,200 persons/km2.  The progress in four areas of re-
search will be  reported,  including: (1) watershed
characterization, (2) patterns of material transport, (3)
organic matter dynamics,  and (4) nutrient spiraling.
      Land cover and land use has been characterized
in the watersheds, and impervious surface area has been
estimated in different land use classes.   Percent  im-
pervious surface increases with distance from the stream
in urban and suburban watersheds.  An Index of Biotic
Integrity (ffll) based  on  fishes was lowest in watersheds
with much of the impervious  surface area in com-
mercial, industrial, and transportation land uses. There
apparently is a threshold at about 30 percent impervious
surface area; when percent  impervious  surface rises
above that, IBI scores decrease sharply.  Correlations
between IBI scores  and forest cover in riparian areas
(50-meter buffer) were  similar to correlations between
IBI scores and forest cover in whole watersheds, even
though  riparian  zones had 20  percent  greater forest
coverage. This  analysis suggests that in urban water-
sheds, the success of riparian restoration projects may
be limited by the influence of land use elsewhere in the
watershed.
      Export of sediments, nitrate, total phosphorus,
and paniculate organic carbon were highest in agri-
cultural watersheds (see Figure 1).  In contrast, dis-
solved organic carbon  export was greatest in urban
watersheds,  presumably a result of wastewater inputs
and runoff from impervious surfaces.  These data, when
combined with standing crop and respiration data, will
provide  valuable  comparative  measures of organic
carbon turnover in urban, agricultural,  and forested
watersheds in the same physiographic area.
      Organic matter inputs, retention,  storage, and
metabolism  were  measured  seasonally  in  the  eight
streams.  The amount of leaf  litter falling directly into
streams was  greatest in agricultural streams and lowest
in urban streams.  However, lateral input of litter was
greatest in urban streams. Retention of organic matter,
measured as uptake lengths of fine particles (fluore-
scently  labeled yeast),  and standing crop of  benthic
organic matter was lowest in urban streams. Leaf de-
composition  rates were highest in urban and agricultural
streams and lowest in  forested streams.   All streams
were  heterotrophic,  and  variance  in net ecosystem
metabolism was best explained by the standing crop of
benthic organic matter.
      Uptake  lengths  of nutrients were  shortest
(200-400 meters) in forested streams  and longest
(> 800 meters) in the urban streams. Uptake is best
explained by hydrologic properties of each channel,
especially the size of the transient storage zone and
its exchange rate. If human activities decrease tran-
sient  storage zone size, then nutrient uptake  length
will increase, thus indicating a reduction in the ability
of the stream to  remove nutrients from the water
column.
                                                   66

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                                                                        Sediment

                                                                        Total  P

                                                                        Nitrate +  nitrite
      Forest      Agriculture    Suburban       Urban
                   Watershed  Type
Forest       Agriculture     Suburban
                Watershed  Type
                                                                           DOC

                                                                           POC
                                                        Urban
Figure 1. Average daily export of suspended sediment (metric tons km2 d"'), total phosphorus (kg km"2 d"1), nitrate and nitrite
        (kg km2 d"1), dissolved organic carbon (kg km"2 d"1), and particulate organic carbon (kg km"2 d"1). Values shown are
               standard deviations for the first year of the study.
        means
                                            67

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 The Role of Hg (II) Reduction and
 Chemical Speciation in Controlling the
 Concentration of Mercury and Its Methylation hi Natural Waters
 Franfois M.M. Morel
 Department of Geosciences, Princeton University, Princeton, NJ
                 (A) Filtered river water
 c
 .0
 "2
 H—'
 c
 o
 o
 c
 o
» Hg(0)

i Hg total

• Hg(ll)
                  (B) Unfiltered river water with KCI added
                 9 T       :
                                                               M chloride added
                                              Time (min)
Figure 1. Time series of dissolved gaseous Hg°, total Hg and Hg(II) concentrations in (A) filtered river water and (B) unfiltered river water
        with 0.5 M chloride. At the onset, 8.3 nM of Hg° was added. Hg(II) was obtained by the difference between Hg° and total Hg.
      High concentrations of mercury (Hg) that bioac-
cumulate in fish are a major environmental and human
health concern.  Research has shown that there is a poor
correlation between the extent of Hg contamination and
the Hg content  of  fish  in the  environment.   The
objective of this project  is to elucidate the conditions
and processes that lead to a high rate of formation of
methyl  mercury,  the organic mercuric  species  that
accumulates in the food chain. This project's research,
which is based on experimental process studies in the
laboratory  and in the field, has followed two principal
axes:  (1) the  determination of the chemical conditions
that  lead   to  a high rate  of  bacterial  uptake  and
methylation of Hg; and  (2) the quantification of the
transformation processes—dissolution, reduction, oxida-
tion—that control the concentration of the various forms
of Hg in aquatic systems.
      In natural waters,  Hg is  methylated chiefly by
sulfate-reducing bacteria  that live in anoxic waters or
sediments and respire by reducing sulfate to  sulfide.
This project has tested the hypothesis that the presence
of polysulfides (formed by the reaction of sulfide with
elemental  sulfur) might  enhance the rate of bacterial
methylation of Hg via the formation of mercury poly-
sulfide complexes, which would keep the Hg in solution
and diffuse rapidly through bacterial membranes.  This
project  found  that  mercury poly sulfide  complexes
indeed form and enhance the solubility of Hg in sulfidic
waters and that  the addition of elemental Hg  (in the
form of polysulfides or particulate sulfur) to cultures of
             sulfate reducers enhances markedly their rate  of Hg
             methylation.  This result is complicated, however, by
             the changing concentrations of sulfur species  in the
             cultures,  and control experiments are under way.
                    Besides  precipitation of mercuric sulfide in
             sediments, the major pathway of elimination of Hg from
             aquatic systems is by reduction of ionic Hg to  elemental
             Hg that,  as a  gas,  is then volatilized into the atmo-
             sphere.  Much attention has been paid to the  chemical,
             photochemical, and microbial processes  that reduce
             ionic Hg, and it has generally been assumed that, once
             formed, all the elemental Hg eventually escapes to the
             atmosphere. This  study has demonstrated that  this is
             not the case:  elemental  Hg can be  oxidized back to
             ionic Hg in oxygenated water.  This process,  which
             competes with volatilization and decreases the rate of
             elimination of Hg from aquatic systems, depends both
             on the chloride concentration in the medium and the
             presence  of appropriate particles. This is illustrated in
             Figure 1, which depicts oxidation of elemental Hg in
             unfiltered freshwater after the addition of chloride ions,
             whereas little oxidation was observed in unsalted filtered
             freshwater.
                   These results modify radically our understanding
             of the processes that control the concentration and the
             methylation of Hg in aquatic systems.  Once confirmed
             and extended, the results should provide a fundamental
             understanding of the chemical and biological parameters
             that determine  the extent of bioaccumulation of Hg in
             various ecosystems.
                                                  68

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 Formation and Propagation of Large-Scale
 Sediment Waves in Periodically Disturbed Mountain Watersheds
 Gary Parker
 St. Anthony Falls Laboratory,  Department of Civil Engineering, University of Minnesota, Minneapolis, MN
                                                                       water surface showing
                                                                       a weak hydraulic jump
                      original channel bed
                                            imposed sediment wave
   a new delta composed of
   finer sediment develops  in
   the ponded area upstream
   of the sediment wave
                                sediment wave
                                some time later
                       Figure 1. Evolution of a sediment pulse from a sample numerical run.
      Human-induced disturbances in watersheds such
 as mining or timber harvesting, combined with certain
 hydrologic conditions, often result in the formation of
 large-scale waves of sediment that are moved (pro-
 pagated) downstream.  These waves have deleterious
 effects on aquatic and riparian habitat and can result in
 bridge and pipeline failures at crossings as well as the
 loss of infrastructure such as roads adjacent to the river.
      The goal of this research is to develop a single,
 global numerical model encompassing backwater ef-
 fects, abrasion, gravel sorting, transport and deposition
 of fines, point and distributed sources of water  and
 sediment, and input hydrographs to predict the evolution
 of sediment waves in rivers.   A combination of field
 work, laboratory experiments,  and numerical experi-
 ments is employed.  The purpose of the field work and
 laboratory experiments is to provide verification of the
 numerical model.
      A  preliminary  numerical model has been  de-
veloped and verified against two small-scale laboratory
experiments. The results of a sample run are shown in
Figure 1. Also, Figure 1 shows successful modeling of
the ponding of water  upstream, the formation of a
hydraulic jump downstream,  the  overall upstream
propagation of the deposit, and the formation of a new
downstream-propagating  delta composed  of  fine
sediment at the upstream of the ponded region.
      It is easy to suppose that sediment waves in rivers
should propagate downstream.  However, results from
the numerical  model  reveal  that dispersion  always
dominates the process.  That is, the apex of a sediment
wave  usually does not  move too far from its original
position. For gravel bed rivers, which usually have
steep  slopes, the position of the apex is more likely to
move  upstream than downstream.  This  finding is im-
portant in determining the fate of landslides in rivers.
      The next steps of this research include: (1) per-
forming more experiments to verify the findings of the
numerical model and (2) fine-tuning the model for an
actual  river (i.e.,  Redwood  Creek in California).
Corresponding field data for this  case  have been
collected. The model will be modified to a user-friendly
form so that interested parties can use it as a predictive
tool.
                                                 69

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Multiscale Statistical Approach to Critical-Area
Analysis and Modeling of Watersheds and Landscapes
G.P. Patil
Center for Statistical Ecology and Environmental Statistics, Department of Statistics, Pennsylvania State
University, University Park, PA
W.L. Myers
School of Forest Resources and Environmental Resources Research Institute, Pennsylvania State University,
University Park, PA
      Public agencies  and corporate landholders are
developing and maintaining major environmental data-
bases for computerized mapping and analysis. The cost
of such  databases  increases rapidly  with  increasing
spatial detail. This project undertakes the development
of objective and  efficient methods  for determining
patterns  of spatial variation from such databases—
particularly relating to water resources and landscape
ecology—and for making pattern comparisons between
databases having different levels of detail. A compari-
son of patterns emerging from data at different levels of
detail will  enable  managers to select the appropriate
spatial resolution  necessary  for particular purposes.
Cost efficiency also will be increased by the ability to
indicate likely zones of spatial uncertainty and to detect
areas of disagreement  between alternate hydrologic
models, so that detailed data acquisition can be better
targeted and modeling will be more precise.  Applica-
tion of these approaches will provide  a better basis for
districting  watersheds,  thereby reducing the cost of
monitoring and detecting the need for remediation.
      The general  nature of the approach to multiscale
characterization of  landscapes in this project is through
analytically induced pattern  deterioration.  The basic
idea is akin to survival of the  fittest in natural selection,
whereby  strong elements persist and weaker ones are
progressively vanquished. Although spatial pattern is a
complex of properties,  one can think  in terms of large
coherent patches that stand in contrast to their neighbors
as being the stronger elements, whereas small internally
variable patches having relatively similar neighbors are
the weaker ones. As with a large field of contestants in
a marathon, the weaker will drop out early while the
stronger will withstand considerable stress before being
subordinated.  Multiscale character is thus reflected in
the rate and extent of deterioration for pattern elements
under the influence of a progressive degenerative pro-
cess.
      The project considers three kinds of spatial data
layers: (1) surface data such as elevations or concen-
trations; (2) signal data  such as  remotely sensed
reflected energy; and (3) categorical data such as land
cover, soil, or geology.  Analytical methods appro-
priate to each kind of data have been conceptualized,
and software capability for their application is being
developed. Echelon hierarchies of hillforms organized
as dendrogram trees  apply  to surface data,   with
pruning  of the tree being the degenerative  process.
Patchwork representation by our "PHASE" version of
statistical clustering is appropriate for signal data, with
information capacity and patch size constraints serving
to induce pattern deterioration.  Random  and modal
filters bring about progressive deterioration of spatial
pattern in categorical data.
      Progressive filtering of categorical data  has been
studied  intensively,  and entropy-based  measures  of
patch structure are used to capture the  trajectory  of
pattern deterioration across scales.  Transition matrix
models  have been formulated that exhibit pattern
deterioration similar to land cover maps of selected
watersheds in Pennsylvania.  Model parameters are
currently being  estimated for  104 major watersheds in
Pennsylvania.  The fitted transition matrices will be
studied for their ability to characterize and differentiate
among watersheds.
      PHASE  formulation  not only provides for
multiscale exploration of signal data, but also offers an
alternative to conventional methods of handling and
analyzing digital image data in remote  sensing.  It
compresses multiband image data into a single layer of
patch-type identifiers with accompanying  table(s)  of
characteristics for patch types.  PHASE compressed
image data is a value-added product that is beyond the
scope of usual copyrights on image data, thus permit-
ting unlimited distribution (see Figure 1).  The  com-
pression is sufficient to accommodate 10 scenes  of
Lands at  Thematic Mapper  formulations  covering
Pennsylvania on a single CD-ROM, along with  a
variety of supplemental vector geographic information
systems (GIS) data.
                                                   70

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Figure 1. Central Pennsylvania sample PHASE from Landsat Thematic Mapper.
                                 71

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Contemporary Water and Constituent Balances for
the Pan-Arctic Drainage System:  Continent to Coastal Ocean Fluxes
Bruce Peterson
Marine Biological Laboratory, Woods Hole, MA
Charles Vorosmarty and Richard Lammers
University of New Hampshire, Durham, NH
      Current predictions  of global  climate change
indicate  a greater  sensitivity in the  Arctic  than in
temperate or tropical regions.  During the next century,
average  temperatures  could  be warmer  by several
degrees  Celsius,  resulting in changes  to snow  ac-
cumulation  and melting,  depth  of soil  thaw,  and
precipitation.  Changing plant communities could alter
evapotranspiration rates and therefore the water balance
across the entire pan-Arctic.  These changes, in turn,
influence the Arctic Ocean through feedbacks between
freshwater discharge, sea ice, and deepwater formation.
The primary  goal  of  this research is  to provide  a
quantitative,  contemporary estimate of  spatially  dis-
tributed water balance, discharge through  rivers,  and
associated fluxes  of constituents from the entire  ter-
restrial drainage system to the Arctic Ocean.
      Our approach combines a data-rich  geographic
information   system (GIS)  with  redundant  models
applied to establish the  pan-Arctic  freshwater  and
material balances.  Aerological estimates  of water flux
conergence and precipitation minus evaporation (P-E)
are calculated using rawinsonde (a balloon-borne atmos-
pheric sounding instrument) archives.  This has been
performed over the Mackenzie, Ob, Yenesei, and Lena
drainage basins. In parallel,  ongoing  development of
a  permafrost water balance  model  (P/WBM)  has
yielded a  first set of estimates of water budgets  at  0.5
degrees (latitude x longitude) resolution.   Freshwater
runoff from  each of these cells is then cascaded down
the river network and discharged into the Arctic Ocean.
A database of river discharge gauges (n =  3,500)  has
been assembled covering land mass contributions from
the United States, Canada,  and Russia.  Additionally,
an  improved  digital   stream  network  has  been
established to provide the best possible routing of water
and constituents downstream to the  Arctic  Ocean.
About 555 gauges from large catchments, greater than
15,000 km2,  were used to validate the network.  Figure
1  shows  the  improvement  in the comparison  of  ob-
served and simulated drainage areas using  these gauges.
The new  discharge  data  set  from the  large  basins
represents an increase of data resolution by an order of
magnitude over previous data  sets (from  89 to 646
gauges).
      Successful  simulations   using  P/WBM  were
carried  out for Imnavait Creek, Alaska.  Upscaling of
the model is currently taking place with applications to
Boreal  zone catchments  and  the  entire  pan-Arctic
region.  Preliminary simulations have taken place that
span  the  full  30-year  timeframe for  this  project
(1960-1990).  High year-to-year variability in many
locations  potentially masks any  progressive  climate
change.
      Preliminary  results  from  rawinsonde   data
(1974-1992) show high interregional variability in P-E
for the Eurasian watersheds. This variability is driven
by an occasional large  summer negative  flux con-
vergence with southerly outflow of water vapor from the
basins.   In some years,  this  negative  flux  is  large
enough  to cause annual P-E to turn negative.  Winter
balances appear to be realistic.  The summer problem
appears to stem from the inability of the rawinsonde
network to "capture" the high topography at the water-
shed divides, an effect that is currently being remedied.
      Simulations  using  P/WBM over   the  entire
pan-Arctic region show large errors when compared
with observed river discharge records.  A  significant
source of this error is in precipitation, particularly over
mountainous regions due to: (1) bias in the locations of
the precipitation gauges (typically at low elevations) and
(2) the sparse network of meteorological stations in the
northern regions giving interpolated fields a high level
of uncertainty. Modifications to these interpolated fields
are under way  and combine  existing elevation and
rawinsonde data to improve temperature and precipi-
tation estimates.
      There continues to be a need to better identify
the sources  of error in the components of the pan-
Arctic water balance. This issue will be pursued in the
context  of looking at the  consistency between  rain-
sonde,  water  balance  model  results,  and discharge
station records.
                                                  72

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     Drainage Area (sq. km) - Before Edit
 CO
    1e6
    1e5
    1e4
   1e3
            •   •••-"     - '.x.r
       1e4
         .  .
"•'           "
          1e5
                                  1e6
                    Observed
      Drainage Area (sq. km) - After Edit
   1e6
O

CO ie5
   1e4
   1e3 -
      1e4
         1e5
1e6
                   Observed
   Figure 1. Contemporary water and constituent balances for the pan-Arctic drainage system.
                         73

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 Modeling Temporal Rainfall
 via a Fractal Geometric Approach
 Carlos E. Puente
 Hydrologic Science Program, University of California, Davis, CA
                                                         ox,
 Figure 1. Alternative derived measures (DY's) arising from the same fractal interpolating function / using alternative parent multifractal measures
         (DX's).
       Accurate rainfall modeling is of vital importance
 for the  proper  management  of  our  environment.
 Rainfall descriptions  are  required, among others, to
 model pollution migration, address issues related to cli-
 mate change (i.e., global circulation), estimate extreme
 weather events, and manage our watersheds. Although
 several sophisticated (stochastic) rainfall  models exist,
 they do not capture the variability observed at a fixed
 location when a storm passes by.  They approximate the
 irregular and intermittent rain patterns by superimpos-
 ing, for example, randomly arriving rectangular pulses
 and consequently preserve only some statistical features
 of the rainfall series (e.g., mean, variance, correlations,
 etc.).   Because these representations  are  typically
 limited by their analytical tractability and because there
 has been a recognition of chaotic and multifractal effects
 in rainfall,  a new class of  rainfall models has been
 developed by our group.  The basis for these new mo-
 dels is the belief that predictability could only be im-
 proved  when the  overall trends and observed  details
 present in rainfall events are considered explicitly.
      The goal of this project is to develop a better
 understanding of temporal and spatial rainfall patterns
 within the State of California under alternative climatic
 conditions.  Parsimonious  representations of  rainfall
 records under  alternative  climatic conditions will be
derived  so that a classification of observed rainfall pat-
terns may be elucidated. This task will be accomplished
by employing the fractal-multifractal representation to
 encode rainfall records throughout the State  of Cali-
 fornia as suitable fractal transformations of appropriate
 (turbulence-related) multifractal measures.  Once the
 time series is represented by this approach, a classifica-
 tion of rainfall events will be attempted via surrogate
 parameters that define the fractal transformation and the
 multifractal measure.  This research should lay a firm
 foundation for a new approach towards hydrologic dy-
 namics in terms of surrogate geometric information.
 This methodology should be viable because it concen-
 trates on capturing what is observed (i.e., the geometry
 of rainfall series).
       During this  year,  studying  the  nature of the
 derived measures that  are generated via the  fractal-
 multifractal representation has continued under a variety
 of alternative scenarios.  Simple modifications to the
 original procedure could be made so that data may be
 generated, similarly  to data obtained in rainfall  records,
 transforming a variety of Cantorian measures (measures
 with holes) via the same fractal interpolating function.
 The existence of these extensions is most welcome
because they are quite parsimonious, requiring only five
 surrogate parameters.   Such extensions  are being em-
ployed to encode actual data sets in California, solving
an inverse optimization problem via genetic algorithms
that minimizes squared differences of real and predicted
records.  A study of the encodings obtained and their
relationship to climatic conditions will  be performed
during the next year.
                                                    74

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Effects  of Food Web Structure and Nutrient  Loading
on Lake Productivity and Gas Exchange With the Atmosphere
Daniel E. Schindler, Stephen R. Carpenter, and James F. Kitchell
Center for Limnology, University of Wisconsin, Madison, WI
Jonathan J. Cole and Michael L. Pace
Institute for Ecosystem Studies, Gary Arboretum, Millbrook, NY
g T
o ™

20
Q- O)


E rl
       2.8
       2.6
       2.4
       2.2
2.0
              A
 ^ ^
minnow.
 __ _ A
 bass
    0        1.0      2.0      3.0
          P input  (mg C rrV2d'1)
                                                        160

                                                          80
                                                       x
                                                       13 ^
                                                       *=  E      0
                                                       O
                                                       O
                                                                -80
                                                              -160
                                                                      B
                                                                  A   O
                                                                    0        400      800      1200
                                                                  Primary production (mg C m"2d"1)
Figure la.  Effect of P input rate on primary production in four
lakes with contrasting food web structures. Each symbol represents
a summer mean for 1 lake year combination from 1991 to 1995.
Lakes characterized by high planktivory and low-grazing rates are
shown by circles.  Triangles represent lakes with piscivores and
high-grazing rates. Lakes that were experimentally fertilized are
denoted by  filled symbols: gray symbols are lake years before
enrichment; black symbols are during enrichment.
                                                  Figure Ib. Relation  between calculated carbon dioxide flux
                                                  between lakes and the atmosphere and the estimated primary
                                                  production rate in 1992-1995 (correlation -0.84).  Positive values
                                                  of carbon dioxide flux represent net flow out of lakes, and negative
                                                  values represent flow into lakes.  The dashed line represents con-
                                                  ditions in equilibrium  with the atmosphere. Symbols are as de-
                                                  scribed for (la).
      When lakes are  viewed  in  a landscape per-
spective, lake ecosystem dynamics are viewed  as the
outcome of interactions between drivers at the landscape
scale (e.g., water,  nutrients, and organic carbon) and
internal processes (e.g.,  nutrient  cycling and food web
interactions).  In this project, whole-lake experiments
were performed to evaluate the  interactions between
food web structure and nutrient loading in controlling
primary productivity of lakes. Food webs were manipu-
lated by changing  the  dominant fish species,  which
resulted in a restructuring of the zooplankton grazer
communities.  Lakes with piscivorous fishes (bass) have
large-bodied zooplankton that are very effective algae
grazers.   Lakes  with planktivorous fishes (minnows)
have small-bodied zooplankton  that  are much  less
effective at suppressing  algae growth. Experimental
additions of inorganic nitrogen and phosphorus resulted
in substantial  increases in algae growth (primary pro-
duction) in all lakes.  However, primary production in
lakes with piscivorous fishes was  much less responsive
to nutrient addition than in lakes with planktivorous
fishes.  This effect of food web  structure on primary
                                                production was sustained over phosphorus loading rates
                                                that ranged from less than 0.2 mg P/m2/d to greater than
                                                3 mg P/m2/d (see Figure la).
                                                      Food web effects on primary productivity also had
                                                implications for carbon dioxide exchange between lakes
                                                and the atmosphere.  Prior to nutrient enrichment, all
                                                study  lakes were net sources of carbon dioxide to the
                                                atmosphere.  Nutrient enrichment increased algae de-
                                                mand  for carbon dioxide, causing lakes to become net
                                                sinks for atmospheric carbon. At identical nutrient load-
                                                ing, atmospheric carbon invasion was greater in a lake
                                                with planktivorous fishes and low grazing than in a lake
                                                with piscivorous fishes and high grazing (see Figure  Ib).
                                                Carbon  stable-isotope  distributions corroborated the
                                                drawdown of lake carbon dioxide and traced atmospheric
                                                carbon transfer from algae to top predators. Thus, top
                                                predators altered ecosystem carbon fixation and linkages
                                                to the atmosphere.  Our study reinforces the idea that
                                                exploitation of top  predators  and the introduction of
                                                exotic species that cause changes in trophic structure are
                                                an aspect of environmental change that have important
                                                implications for structure and function of ecosystems.
                                                    75

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A Comparative Institutional Analysis of Conjunctive
Management Practices Among Three Southwestern States
Edella Schlager
University of Arizona, Tucson, AZ
      Among the  more  popular contemporary
recommendations for improved watershed use and
protection is conjunctive use of surface and under-
ground water resources. Conjunctive use involves
the coordination of surface  water supplies  and
storage with groundwater supplies and storage for
purposes of sustainable watershed use and enhanced
watershed protection. Conjunctive use in a water-
shed  requires a  great deal of joint effort among
human beings, the prospects for which will be af-
fected strongly,  though not exclusively, by insti-
tutional  arrangements  that define organizational
forms and jurisdictions  and provide incentives and
disincentives to individuals.  Using a comparison of
Arizona, Colorado, and California, this  project
seeks to advance the  theoretical  and empirical
understanding of the relationships between insti-
tutional arrangements governing the allocation, use,
and protection of water  resources and the develop-
ment,  implementation,  and performance  of  con-
junctive use programs.
      The  approach taken in this project is one of
institutional analysis. Institutional analysis is based
on in-depth  comparative  analyses  of  the  per-
formance of diverse institutional arrangements. The
technique of institutional analysis was developed
during the last  quarter-century  at  Indiana  Uni-
versity, primarily by Elinor and Vincent Ostrom
and colleagues at the Workshop in Political Theory
and Policy  Analysis (a complete description of the
approach is available  in E. Ostrom, 1990).   In
essence, institutional analysis involves: (1) the use
of a systematic  framework for classifying insti-
tutional arrangements (conceived as sets of rules)
and levels of action; (2) viewing the development of
institutional arrangements as  processes of inten-
tional (individual or collective) human choice and
action constrained by physical circumstances and
community context, recognizing that human beings
are capable of multiple levels of action (e.g., as
rule  makers and rule followers,  as  organization
designers as well  as organization members);  and (3)
assessing the effects of institutional  arrangements
on human choice and action as well as evaluating
the effects  and  effectiveness  of institutional ar-
rangements relative to their intentions.
      Given  a  well-developed  and systematic
framework, institutional analysis allows for com-
parisons  across situations on  a variety of dimen-
sions. For instance, community characteristics and
physical settings may be held constant, while rules
are varied,  permitting an  understanding  of  how
different configurations of rules promote  various
outcomes.
      Although Arizona, California, and Colorado
face similar physical problems of water distribution
in the same geographic region, the three states have
devised markedly different approaches  to con-
junctive water  management. Arizona has  taken a
more state-centered approach. The primary entities
engaged in conjunctive water management are the
Central  Arizona  Water  Conservation   District
(CAWCD),  the operator of the Central  Arizona
Project, and the Arizona Water Banking Authority
(AWBA). By far, the dominant conjunctive water
management approach is the exchange of surface
water for groundwater credits, or in lieu recharge.
CAWCD  makes available to  CAP  water  for
agriculture in  exchange for agriculture  ground-
water.  CAWCD uses the groundwater for drought
protection and  to  enhance the reliability of  the
CAP. If a pumping station or a portion of the canal
becomes inoperable, CAWCD can deliver ground-
water to its customers.  Although California uses
conjunctive  water  management  primarily  as  in-
surance against water shortages, it is primarily local
level jurisdictions,  and not state  entities,  that
engage  in  it.  The dominant  conjunctive water
management  approach  is  constructed recharge
projects whereby surface water is  stored under-
ground  through  injection  wells and spreading
basins. Due to  recent changes in state law, in lieu
recharge is beginning to be used in California. Like
California,  Colorado's  conjunctive  management
activities  occur  primarily among local  level
jurisdictions. Unlike both Arizona and California,
however, conjunctive water management  is used
primarily to protect the rights of senior  surface
water users  under the state's prior appropriation
doctrine, and not for drought protection. Colorado
jurisdictions use a combination of in lieu recharge
and constructed recharge projects.
      Institutional arrangements govern the type,
development,   and  uses  of  conjunctive water
management  projects.  Data collection  for  this
project is in its  preliminary stages. As further data
are collected, issues of institutional performance,
the allocation of costs and benefits, and  the  en-
vironmental impacts of conjunctive water manage-
ment projects will be explored.
                                               76

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Water and Sustainable Development
in the Binational Lower  Rio Grande/Rio Bravo Basin
Jurgen Schmandt
The Houston Advanced Research Center (HARC), The Woodlands, TX, and The Instituto Technologico y de
Estudios Superiores de Monterrey (ITESM), Monterrey, Nuevo Leon, Mexico
      In  its  1994  Regional Assessment  of  Water
Quality in the Rio Grande Basin, the Texas Natural
Resource Conservation Commission made two recom-
mendations for further research and policy develop-
ment:  (1) an improved understanding of water issues
as they relate to the binational border dynamic and (2)
development of lasting links across  the  U.S.-Mexico
border.   The  goals of this  project  are  to:  develop
reliable watershed-based data sets from both Mexican
and U.S. sources to analyze the whole binational region;
analyze water resource issues as critical  factors in the
region's long-run planning and sustainable development;
and  engage researchers, policymakers,  and the civic
community to help answer the question, "What actions
can be taken to achieve sustainable development in this
rapidly growing, drought-prone, environmentally fragile
watershed?"
      Mexican and U.S. researchers are paired in teams
to perform data compilation and  analysis in the follow-
ing  areas: (1) water supply and  demand,  (2) water
quality, (3) population and socioeconomic conditions,
(4) ecology, (5) water management and institutions, and
(6) geographic information systems (GIS).  The teams
produce  an integrated  "baseline  report" of current
conditions  in the region, which then provides the basis
for subsequent analysis of alternative future  scenarios.
Scenarios are  developed by combining regional  demo-
graphic projections to the year 2030 with alternatives for
future water  availability,  irrigation technologies, and
management practices.
      Involvement of the  community is a key element
of the project, and mechanisms for stakeholder input
and consultation are incorporated into the  research pro-
cess. Researchers and graduate  students  from Mexico
and the United States (as  part of a joint University of
Texas-ITESM Policy Research Project) have conducted
a survey among stakeholders and community leaders.
In addition,  workshops  are  scheduled involving  re-
searchers  and stakeholders  from both  sides of  the
border.  These outreach components gather insights into
the  community's  concerns,  perceived  threats,  and
opportunities related to water and development in  the
watershed.
      The preliminary findings of the study include  the
following:  (1) The population will double by the year
2030, while surface water supply  will stay  at current
levels  or decrease.  (2) The single source of surface
water  for  2 million people and  intensive  irrigated
agriculture in the  Lower  Rio Grande—the combined
Amistad-Falcon international reservoir system—is at its
lowest level since coming online 25 years ago.   (3) The
current drought in  the upstream parts of the watershed
began  in 1994 and continues unabated.  Mexican far-
mers lost two harvests, and several irrigation districts in
Texas went dry in 1996.  (4) The adequacy of the exist-
ing water supply system under drought conditions is
being tested for  the first time.  During the drought of
record in the 1950's, only 500,000 people lived in the
same area where 2 million live today.  Amistad reser-
voir did not yet exist, and Falcon reservoir came on line
in the middle of the drought.  (5) Significant changes in
fauna appear to be correlated with decreasing stream-
flows, the proliferation of exotic species, and chemical
pollution.  (6) GIS analysis indicates that irrigated land
area in the basin  is significantly larger than  official
records indicate.  (7) Rapid growth and development in
the  San Juan and  Conchos River basins in Mexico could
significantly reduce the amount of water these tributaries
contribute to the main stem of the Lower Rio  Grande/
Bravo. (8)  Initial runs of the agricultural model suggest
that significant long-term  reductions in water supply
would at most have a very small impact on the  regional
economy because new irrigation technology and crop-
ping patterns would be adopted.  (9) Transfers of agri-
cultural water to  municipal use occur in Texas through
the  use of  water markets and joint  infrastructure im-
provements on the part of cities and irrigation districts.
      The  significance  of these findings  is  that the
system is  flexible—small  increases in irrigation ef-
ficiency can release significant  amounts of water for
other uses,  hi addition, a tremendous opportunity exists
for  "win-win" arrangements in which municipal water
suppliers agree to  finance  improvement in irrigation
efficiency in return for the rights to the water saved.
Furthermore, the existing regional water market holds
great potential for more efficient use of water.
      The research teams plan to meet in January 1998
to discuss the scenario analysis portion of this project.
A community workshop  involving researchers and
stakeholders  is  scheduled for  May  1998,  and the
integration  of  scenario analysis is planned  for spring
1998.
                                                   77

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           '"51
        Basin Boundary -
                                          United States
                                           of America
                          tad Reservoi
                                         Falco'n. Reservoir

                                                 "1
                                                    McAllen

                 '*>
 Estados Unidos
    Mexicanos
Shaded area represents primary study region
TAMAUL1PAS
                      Figure 1. Lower Rio Grande/Rio Bravo Basin.
                                78

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Environmental Change and Adaptive Resource
Markets:  Computer-Assisted Markets for Resource Allocation
Vernon Smith, S. Rassenti,  E. Hoffman, R. Howitt, A. Dinar
University of Arizona, Tucson, AZ
      Water delivery systems have traditionally been
operated by  large centralized authorities due to the
natural monopoly nature of the industry and the quasi-
public good nature of a reliable and clean water supply.
In most countries, the era of meeting new demands by
increased supply development has been terminated by
economic  and environmental  costs,  environmental
change, and  the  shortage of suitable sites.  Like the
energy and  natural gas industries,  the  water supply
industry will have to look to market mechanisms to
reallocate existing water supplies and increase efficiency
to meet future demands.
      This project's goal is to develop a formal mech-
anism  for evaluating alternative market structures for
integrated water systems. This goal will be achieved by
the  following objectives:  (1) developing a structural
market model for the California water  industry, (2)
deriving  value and  cost  functions  for  the different
components  in the structural model, (3) developing an
experimental design that uses the water market model,
(4)  running experiments on alternative market options
using economically motivated subjects in a replicable
design, and  (5)  developing  a  remote (Internet) ex-
perimental ability that allows extension of the market
experiments  to market participants in the field.
      A prototype of a  smart  water market for the
central California water industry has been  designed and
tested at the  Economic Science Laboratory in Tucson,
Arizona.  The input needed for the player at each node
is in the form of a step function  representing  their
willingness to pay for different quantities  and qualities
of water. Experiments on market price convergence and
market efficiency were conducted using students whose
earnings were based on their market success. Players
represented   agricultural,  urban  and  environmental
interests, and were given initial endowments of money
and water rights reflecting the current allocation in the
state. After initial training, the experiments were run for
15  trading  periods  in  four experimental  sessions.
Student participants were paid between $2 and $70 as a
result of their trading actions.
      The first set of experiments with the simplified
model obtained realistic results. A "smart" computer-
assisted market with a double-blind auction structure
was used. Under this structure, buyers and sellers do
not have  to reveal their true valuation for the goods to
achieve an outcome that  is close  to the theoretical
optimum under perfect competition. However,  the
relative share  of  their defined supply  or demand
schedule  that the players were  able to realize differed
significantly among players  and groups.   Alternative
allocations of water transport facilities  made a signi-
ficant difference in the outcome between players.  The
outcomes of  the initial experiments  shows that  it is
possible to reproduce price efficiency and convergence
results in natural resource markets.  The results were
obtained using players that included both users and an
environmental agent, who was  only interested in the
level  of flows through a certain sensitive node.  The
initial results show  that the  novel concept of market
interactions between resource users and environmental
interests can be modeled in a replicable manner using
this approach.
      The next steps are:  (1) extending the model to a
larger number of participants in each sector of the water
economy; (2) incorporating a water quality component
into the supplies, demands, and the smart market soft-
ware; (3) developing Windows NT-based Internet soft-
ware to enable use by outside participants in the exper-
iments ; and (4) extending the smart market analysis to
water allocation problems in  a developing country.
Through step (3), the performance of participants  who
are actually working in the industry  can be compared
with the student-based results. This latter work will have
benefits as a  validation of  the formal  results  from
student-based experiments and also will make the smart
market a practical tool for developing market skills by
members of the industry.
                                                   79

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Detecting Fecal Contamination
and Its Sources in Water and Watersheds
Mark D. Sobsey
University of North Carolina at Chapel Hill, Chapel Hill, NC
                                                         MS2
                                                         SW1
                                                         SW6
                                                         SW11
                                                         SW16
                                                         SW21
                                                         SW26
           GA
           SW2
           SW7
           SW12
           SW17
           SW22
           SW27
 QP
SW3
SW8
SW13
SW18
SW23
SW28
 FI
SW4
SW9
SW14
SW19
SW24
SW29
SP
SW5
SW10
SW15
SW20
SW25
SW30
         Probe nA                  Probe HB
Figure 1. Lysis zone hybridization for identification of 30 F RNA coliphage isolates from surface water by oligoprobes IIA and nB. Positive
        controls consisting of 5 prototype F RNA coliphages (MS2, GA, Q(3, FT and SP) are shown on the top. Thirty field isolates (SW1-SW30)
        are arranged on the key.
      Current methods to detect, quantify, and predict
water  and watershed  quality with respect to fecal
contamination are inadequate and unreliable. Because
so little is actually known about the levels and sources
of  fecal  contamination  in water,  there  are  great
uncertainties about the human and environmental health
risks from the pathogens associated with contaminated
water.  The objectives of this research are  to further
develop, evaluate, and apply new and improved mea-
sures of human and animal sources  of fecal contami-
nation in water and watersheds.  Two main criteria will
be  used to judge  the value  and reliability of  the
candidate indicators: (1) their ability to  predict the pre-
sence and concentrations of the pathogens relative to
the sources of the organisms and (2) their ability to
predict the presence, persistence, transport, and fate of
the pathogens  in response to natural environmental
processes.
      The investigators  have  developed reliable
methods for the simultaneous concentration of viruses,
bacteria, and  protozoans from water using disposable,
hollow fiber ultrafilters. Recoveries of seeded organ-
isms from surface  water are 50-75 percent and  the
coefficients of variation (CVs) were relatively small.
      The new  methods  previously  developed   to
concentrate human gastroenteritis viruses from water
were field tested on a groundwater supply of drinking
water that was implicated in  an  outbreak of viral
gastroenteritis.  The virus isolated from the  water was
genetically identical to the virus independently isolated
from stool specimens of ill persons by another lab-
oratory.  This is  the first time such a virus was suc-
cessfully recovered and detected in an incriminated
water supply, and the work  was recently  published
(Bellar et al.,  JAMA 278(7):563-568).
      New and improved methods to  detect indicator
viruses known as coliphages using "gene probes" were
improved and field tested (see  Figure 1).   Also,  the
researchers found that it was  possible to distinguish
fecal from nonfecal indicator viruses in water on the
basis of bacterial host range and growth temperature.
Coliphages of fecal origin grow well at temperatures of
42-45° C, and they have a narrow host range.  Coli-
phages of possible  nonfecal, environmental origin will
grow well at temperatures of 37° C or lower and not at
42-45° C.  Some of these environmental coliphages have
a wider  host  range and  will grow on  some other
bacteria.  In field samples of surface waters impacted by
known sources of human and animal fecal contamination,
levels of various fecal indicator organisms generally
increased when fecal waste sources were nearby. Even in
areas  where  animal wastes were managed by land
application, adjacent surface waters showed evidence of
fecal contamination. Levels of indicators were 10-1,000
times higher than background levels.
      Coliphages in the water of  stations near animal
waste sources were  primarily animal groups, and
coliphages in the water of stations impacted by muni-
cipal wastewater  discharges or nonpoint source human
wastes were primarily human groups. At stations where
both sources were impacting water  quality, both groups
were detected.  These results indicate that sources of
fecal contamination can be reliably  identified and traced
by  identifying and grouping the  coliphages that are
detected using gene probes.
      Studies on detecting and quantifying microbial
pathogens  and indicators of  fecal contamination in
waters and watersheds impacted by human and animal
fecal contamination will be continued in  the watersheds
now being monitored.  An additional watershed will be
added during the next 12-month period.  Also, the new-
ly developed methods to detect human pathogens of
fecal origin will be  applied to these waters to determine
if the known  human and  animal waste  sources are
important contributors of these protozoan and bacterial
pathogens.  The reliability of the various microbial
indicators in predicting the presence and concentrations
of these pathogens  also will be examined.
                                                  80

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 Ecoregion-Specific  Comparison of Stream Community Responses to
 Nutrient Gradients Using Both Survey and Experimental Approaches
 R. Jan Stevenson
 University of Louisville, Louisville, AT
 Mike WUey
 University of Michigan, Ann Arbor, MI
 Joe Holomuzki
 Transylvania University, Lexington, KY
Landscape Development

       I
     Nutrients
           Algal mat &
           filaments
Flow Stability
                             	^.  Scraping
                              , - ' Herbivores
  Collector/
  Gathers
                                                 Predators
       Most evidence that exists for regulatory purposes
 comes  from  toxicological studies,  which are often
 conducted in  laboratory experiments.  This project is
 aimed at providing a credible scientific basis for setting
 regulatory  criteria   relating   to  changes  in  stream
 ecosystems along  an  environmental gradient.
       Nutrient loading commonly causes an increase in
 primary productivity and a replacement of clean-water
 ecosystem algal-invertebrate communities with a new
 suite of organisms that results  in very different stream
 structure and function.   Quantitatively assessing the
 relationships between nutrient concentrations and algal-
 invertebrate communities in streams has been challeng-
 ing because  stream  ecosystems  are so  physically
 variable  and  biologically  dynamic.  This  project is
 designed to develop a better predictive understanding of
 nutrient  effects on stream communities in  ecoregions
 with different hydrological stability. In this project, the
 investigators are using stream surveys with large within-
 ecoregion sample  sizes, experiments,  and analyses of
 large preexisting databases.
       Initial  results  indicate  that chances for  high
 accrual  of algae with increasing nutrients is greater in
 the hydrologically  variable streams of Kentucky than in
 the stable  streams of Michigan.   Invertebrate  abun-
 dances,  much lower in Kentucky than in Michigan, do
 not respond to nutrients  in Kentucky but  do show a
 positive response to nutrients in Michigan.
       Three  groups  of experiments were conducted
 during the summer  of 1997 to complement  previous
 work. In one experiment, the hypothesis that thresholds
 in  nutrients occur where algae can grow  faster than
 grazers  can consume them  was tested in experimental
Figure 1. Positive and negative effects on invertebrates in streams.
Nutrient  effects on algae and  invertebrates in streams are highly
dependent upon climate, geology, and flow stability of streams.
Nutrients stimulate growth rates of algae.  If unregulated by grazers,
nutrients stimulate the change of algae from thin mats to thick mats
and filaments.  High flow stability allows invertebrate densities to
increase and regulate accumulation of algal mats and some filamentous
algae. Low flow stability, either from scouring spates or drought,
constrain invertebrate densities so that  algae can easily overgrow
substrata when nutrients are abundant. When these algae overgrow
substrata, changes in invertebrate communities due to the positive
effect of the abundant  algae on some invertebrates are observed.
Little evidence has been  found that overgrowths of algae have negative
effects on invertebrates in streams (solid lines indicate positive effects;
dashed lines indicate negative effects).

streams with 96 experimental reaches.  Eight different
nutrient concentrations, 2 different grazer treatments,
and 6 replicates of  each treatment were manipulated in
the 96 reaches.  In a second  experiment, nitrate and
phosphate concentrations were varied in 72  different
experimental streams to  determine  the thresholds  in
nutrient concentrations that saturate algal growth rates
when biomass is low and the concentrations that cause
peak algal accrual on substrata (nuisance algal growths).
Nine different nutrient levels of nitrate and phosphate
were used with 4 replicates of each treatment.  The third
set of experiments  assessed the relationship between
invertebrates and  chariging habitat conditions when
algae accumulate.   Preliminary results indicated over-
growths of  streams by long,  filamentous green algae
(Cladophora) can have a positive effect on some stream
invertebrates,  particularly  hydropsychid  caddisflies.
These experiments showed that these invertebrates
prefer rocks with  Cladophora as compared  to  bare
rocks, and stoneflies preyed more successfully on these
caddisflies on bare rocks than on rocks covered  with
Cladophora.
      Future work will develop stronger quantitative
predictions  of the effects  of specific concentrations  of
nutrients on algae and invertebrates in streams in  diff-
erent ecoregions. This will be accomplished by evalu-
ating data from 2 years of algae and invertebrate data
from 130  streams in Kentucky and Michigan and from
two sets of experiments in which nutrient concentrations
were manipulated.   In addition,  relations between in-
dices of  biotic integrity,  indices of environmental
stressors,  and nutrient concentrations will be assessed
with data from  the  130 streams that were in the study.
                                                      81

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Holocene Floodplain Development as a Function of Climate Change
and Human Activities: The Arroux and Loire Rivers,  Burgundy, France
Eric C. Straffin and Michael D. Blum
Department of Geosciences, University of Nebraska-Lincoln, Lincoln, NE
      Holocene age floodplains of the Arroux and Loire
Rivers contain a regionally extensive record of fluvial
activity,  which can be interpreted within established
archaeological,  paleoenvironmental, and historically
documented climatological frameworks.   River mor-
phology and associated deposits record the magnitude
and frequency of floods, associated with  climate regimes
driven  by  global  atmospheric  circulation.  The
reconstruction of past  fluvial activity  and correlated
discharge regimes facilitates the prediction  of  fluvial
adjustments  to  climatic and anthropogenic changes
expected in the near future.
      The southern  Burgundy  region of France  is
situated at the boundary between maritime, continental,
and Mediterranean climates.  During the period  of his-
torical monitoring, the position and dominance of these
climatic regimes has shifted as a function of changes in
atmospheric circulation. Discharge regimes have fluctu-
ated as well, but details differ regionally due to geologic
and antecedent conditions, and the latitudinal position
and style of storm tracks. Historic analog models sug-
gest that the dominance and persistence of a  particular
climate  regime should result  in  regionally circum-
scribed,  but isochronous   alluvial fills containing
differing styles of floodplain construction. For example,
the Little Ice Age in Europe (ca. 1300-1850 A.D.) was
marked by episodic cooling, resulting  in large  floods
and flood variability  between basins, while  Medieval
times  (ca.700-1300 A.D.) were marked by warmer con-
ditions, and more steady flood regimes,  although the
magnitude of floods varied between western and eastern
drainage basins.
      Throughout  Europe,  enhanced  fluvial activity
occurred  during the  period  1250  to 1550 A.D., es-
pecially during transitions from dominantly  cold sub-
periods to warmer conditions. The Loire and Arroux
valleys follow  the same  regional trend,  with pre-
dominantly overbank deposition of sands  and silts
during  the Neoglacial  period,  which  resulted  in
straightened, chute-cut-off channel patterns  and the
burial of older alluvial surfaces. This style of fluvial
activity began ca. 1050 B.C., contrasting sharply with
earlier gravelly, laterally accreting, meandering rivers
characteristic of the mid-Holocene. This morphological
and sedimentological change predates significant human
impacts on the landscape,  is regional  in extent, and
when combined with proxy evidence supports the notion
that climate change has been the dominant mechanism
in driving fluvial adjustments.
      The separation of climatic  from human influences
on the response of rivers is often difficult to determine,
but there is  increasing recognition that at regional
scales, climate has been the dominant mechanism for
changes in fluvial dynamics. Human  activities have
conditioned these responses locally, and at regional
scales have served to amplify existing climatic in-
fluences. In southern Burgundy,  Roman land use prac-
tices were introduced ca. 50 B.C., and noticeable hu-
man impacts  in upper  portions  of the Loire basin
occurred even later.   Roman and Medieval land use
practices may  have increased the  overall thickness  of
overbank deposits but were most likely not responsible
for the overbank discharges permitting such deposition
at regional scales.
      Land use changes  in this rural region have
changed little over the last several hundred years, and
if we can  use the past response of the  Loire and
Arroux  Rivers as analogs to what may happen to this
region with future climatic change, we may  expect
that with decreased temperatures  and increasingly
meridional circulation, variability in flood frequency
will increase, resulting in flood-prone valley bottoms.
Moderate increases  in temperature and  increased
zonal circulation may produce less variable discharge
regimes,  but  more  regular  inundation  of lower
floodplain positions.
                                                  82

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 Response and Compensation to a
 Bivalve Invasion by an Aquatic Ecosystem
 David L. Strayer, Nina Caraco, Jonathan J. Cole, Stuart Findlay, and Michael L. Pace
 Institute of Ecosystem Studies, Millbrook, NY
                                            % change due to zebra mussel
                                                                           o
                                                                           o
                            microzoopl.
                         phytoplankton -
                           macrozoopl. -
                              unionids
                         macrobenthos
                           susp. solids
                                  DIN
                                secchi
                              bacteria
                                  SRP
 Figure 1. Variation in the response of North American freshwater ecosystems to the zebra mussel invasion. All ecosystems have estimated zebra
         mussel filtration rates of 70-125 percent of the water column per day. The solid line shows the response of the Hudson River, while
         the points show responses from Lake Erie, Lake St. Clair, Saginaw Bay, and Oneida Lake (from data of Holland [1993], Fahnenstiel
         et al. [1995], Fanslow et al. [1995], Holland et al. [1995], Johengen et al. [1995], Maclsaac et al. [1995], Mellina et al. [1995], and
         Horgan [1996]). DIN=dissolved inorganic nitrogen; SRP=soluble reactive phosphorus.
       Bivalves are so abundant in many shallow aquatic
 ecosystems that their filter-feeding regulates pelagic and
 benthic variables.  Humans cause sudden changes in bi-
 valve  populations through  the  introduction of alien
 species, water pollution, and overharvesting, which may
 have large effects throughout the ecosystem.  The in-
 vestigators postulate that the response of an ecosystem
 to changes in bivalve populations  will depend on the
 characteristics  of  the ecosystem  as  well as on  the
 numbers and biological  characteristics of the bivalve.
 Thus,  two  ecosystems might respond very differently to
 the same bivalve population. Specifically, the project
 hypothesis  is that the ecosystem response will depend on
 some characteristics that are fixed (e.g., morphometry,
 hydrology) as well as  flexible compensatory pathways
 (e.g.,  functional substitution of species, environmental
 feedbacks, and induced defenses).
       The investigators have been looking at ecosystem
 responses to bivalve populations by studying the response
 of the Hudson River to the ongoing zebra mussel invasion
 and comparing its  response to that of other ecosystems,
 where large bivalve populations appeared or disappeared.
 This project combines long-term field studies, models,
 small-scale experiments, and cross-system comparisons.
 Zebra mussels first appeared in the Hudson in 1991 and
 became the dominant consumer in the ecosystem by the
 end of 1992.  Estimated clearance  times for the entire
 volume of water in the freshwater  tidal Hudson River
have been in the range of 1-4 days. Consequently, large
changes have emerged through the ecosystem.  Densities
of phytoplankton  and small zooplankton, which are
consumed by zebra mussels, fell by 80-90 percent, and
the composition of the remaining phytoplankton shifted
markedly from preinvasion communities. Populations
of native bivalves  fell by 50-70 percent in response to
the loss of their phytoplankton food. Bacterial densities
doubled, perhaps because of losses of small zooplankton
that feed on bacteria.  Water clarity and dissolved re-
active phosphorus  both rose as phytoplankton densities
fell.  The density  and extent of rooted plant beds may
have increased as the Hudson's water cleared.  Densities
of other sediment-dwelling animals fell in deep waters
but rose in shallow waters, probably in response to
changing patterns of primary production.
      Generally,   other  systems exposed  to bivalve
grazers show qualitatively  similar  responses  to the
Hudson, but with very large quantitative  differences
across systems (see Figure 1).  These differences pro-
bably reflect the extent to which ecosystem character-
istics constrain the strength of the interaction pathways
that connect the bivalve with the rest of the ecosystem.
Specifically,  cross-system comparisons support the idea
that ecosystem responses depend on vertical mixing of
the water column; the size, structure, and specific com-
position of the plankton;  the  nature  of factors  that
otherwise limit growth rates of plankton (i.e., nutrients,
light, advection, etc.); the  diet breadth of consumers;
the magnitude of allochthonous inputs that can  substitute
as food for phytoplankton; inputs of physical energy to
the system; and system morphometry.
                                                    83

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 Scaling Up Spatially Distributed
 Hydrologic Models of Semi-Arid Watersheds
 David G.  Tarboton \ Christopher M. U. Neale 1,  Keith R. Cooley 2, Gerald N. Flerchinger2,
 Clayton L. Hanson 2, Charles W.  Slaughter2, Mark S. Seyfried2,  Rajiv Prasad 1,  Charlie Luce ',
 Greg Crosby 1, and Changyi Sun 1
 1  Utah State University, Logan, UT
 2  USDA Northwest Watershed Research Center, Boise,  ID
                       3 a
                           OcU.92
                                    Dec 1.92
                                               Feb1.93
                                                        Apr 1,93
                                                                  Jim 1.93
                                                                            Aug1,93
 Figure 1. Modeled basin average snow water equivalent in Upper Sheep Creek, 1992/93, using a fully distributed model on a 30-meter grid and
         lumped model with snow-covered area fraction parameterized using a depletion curve. The good comparison indicates that the depletion
         curve approach is promising for scaling up the modeling of snow accumulation and melt, allowing small basin size areas (26 Ha) to be
         modeled as a single node in a distributed model with larger elements.
       Semi-arid rangeland  and forested watersheds
 comprise a large portion of the Western United States.
 The quality, quantity, and timing of runoff from these
 watersheds is  crucial  for  water  supply  and affects
 agriculture, fisheries, recreation, and hydropower.  The
 purpose of this project is to understand interacting
 watershed processes over  a range of scales in the
 Reynolds Creek Experimental Watershed (RCEW) in
 southwest Idaho.
       The  investigators  are developing  a  spatially
 distributed  modeling framework  that  accounts for
 spatial variability in topography, vegetation, and soils
 to quantify the  complete water balance at a range of
 spatial scales.   This  will provide a framework within
 which to test hypotheses regarding the hydrology and
 water balance  at Reynolds Creek.  As such, it is a
 working model with alternative modules being switched
 in and out for comparison against data.   The model
 development is proceeding  in parallel with the acquisi-
 tion and processing of the remotely sensed data.  This
 includes data from six aircraft  overflights as well as
 Landsat data.  Field data being used consist of stream-
 flow,   meteorological   data,  soil   moisture   and
 groundwater data, and evapotranspiration flux data.
      Parameterization of snow subgrid variability has
 focused on causes for this variability and how to model
 it. The results to date indicate that the subgrid varia-
bility due to drifting was equally or more important than
subgrid variability in solar radiation for estimating the
quantity and timing  of surface water  inputs  from
snowmelt.   A parameterization linking snow covered
area to basin average  snow  water equivalent  shows
promise as a tool in scaling the energy balance up model
to larger model elements.  Figure  1 shows a comparison
of the time evolution of basin snow water equivalent
calculated using a 255-point distributed model and the
spatially integrated snowmelt  model.  The close com-
parison is indicative of the potential of this approach.
      Leaf  Area Index (LAI) measurements to  date
indicate that: (1) there is approximately a twofold range
in LAI within the major plant communities in the
watershed;  (2) the maximum  LAI occurs early in the
growing season, shortly after snowmelt; and (3) the LAI
of all the plant communities (i.e., at all sites) decreases
steadily during the summer to  an apparent minimum in
the fall. Different approaches for using remotely sensed
imagery to map the  spatial distribution of the critical
vegetative  communities  within the  watershed  are
currently being tested. This will allow the incorporation
of vegetative dynamics  into  the surface energy  flux
modeling.
      This project will lead to a better understanding of
the  spatial  variability   and   scale   dependence  of
hydrologic  processes in RCEW.  Because this work is
aimed at gaining a better understanding of the physical
processes  and  their  interactions,  results  will be
generalizable to other watersheds in the semiarid mount-
ainous Western United States.
                                                    84

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Traveling Wave Behavior During Subsurface Transport of
Biologically Reactive Contaminants:   Implications for In  Situ Bioremediation
Albert J.  Valocchi
University of Illinois at Urbana-Champaign,  Urbana,  IL
      Contamination of  soil  and  groundwater  by
hazardous chemicals is widely recognized as one of the
major environmental problems faced by modern society.
Although there is ongoing debate regarding the extent
and severity of contamination, there is no disagreement
over  the  staggering  economic  costs  incurred  in
monitoring  and rehabilitating  polluted groundwater
supplies.  In-the-ground (in situ) remediation of con-
tamination costs a fraction of other approaches. The
objectives of this project are to investigate how  transport
and mixing processes affect the overall performance of
engineered in situ bioremediation. Although these pro-
cesses play a key role in the ultimate success  of actual
remediation projects,  their significance cannot be ascer-
tained through typical laboratory-scale studies.
      Mathematical models are  being  analyzed for a
typical bioremediation scenario  in which a uniformly
distributed organic contaminant is  degraded by in-
digenous soil microbes that are stimulated by an injected
material  (e.g., an  electron acceptor  such as oxygen).
Our analysis starts with simple, one-dimensional homo-
geneous  systems  and  progresses to  more  realistic
mutlidimensional heterogeneous  aquifers.
      The following cases have been  studied:  (1)
one-dimensional  uniform  flow  in  a  homogeneous
aquifer;  (2) ideal radial flow from an injection  well  in a
homogeneous aquifer; and (3) two-dimensional flow in
a stratified aquifer.  For a wide variety of cases, the
system evolves to form traveling  waves;  that is, the
spatial profiles of the organic pollutant, electron ac-
ceptor, and biomass attain constant shapes that travel in
unison.  These traveling  waves  form because of  a
balanced interaction  between solute  mixing processes
and localized biodegradation processes.  This results in
a very localized reaction zone where the pollutant and
electron acceptor mix together, as can be seen in Figure
1, which is for a two-dimensional stratified aquifer.
      For conditions when traveling waves exist, simple
formulas have been derived to calculate the long-term
rate of pollutant removal due to biodegradation.  The
removal rate expressions are similar for all the different
flow systems that we have examined. The pollutant re-
moval rate depends upon transport properties of the
aquifer, but it does not depend upon the initial biomass
concentration or upon the rate at  which the biomass can
degrade the pollutant. Results from the simple formulas
were  verified by comparison with numerical simula-
tions.
      The traveling wave framework is a useful  sim-
plifying tool for approximating  the complexity of bio-
remediation modeling.  Use of the analytical formulas
reduces significantly the computational burden of es-
timating the efficiency of a remedial design.  The results
are significant  because they  indicate  conditions  for
which the long-term pollutant  removal rate does not
depend  upon the laboratory-determined rate of  bio-
degradation.  This helps explain the common observa-
tion that laboratory-determined degradation rates over-
estimate the degree of biodegradation attained in many
field projects.
      In the future, the analysis will be  extended  to
more realistic patterns of aquifer heterogeneity where
the soil permeability changes randomly in all directions.
Detailed numerical simulations of the cases  studied to
date  indicate  that the traveling  wave behavior is not
established until a certain time elapses.  The inves-
tigators plan to study this initial time period because the
biodegradation rate,  which is often much larger than the
long-term rate, which  is attained  after the  traveling
waves form.
                                                   85

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    1.0-1
N  0.5-
N 0.5
N 0.5-
                                                                                              20.0
                                                                                              20.0
                                                                                              20.0
                                  T
                     0.0         0.2         0.4-         0.6         0.8         1.0
                                Normalized concentration scale
 Figure 1.  Normalized concentration profiles of the organic substrate (pollutant), electron acceptor, and biomass characterizing
           enhanced in situ bioremediation in a stratified aquifier. The aquifier consists of two layers, with the more permeable
           layer on the bottom.  The electron acceptor is injected from the left-hand boundary.  The profiles maintain their
           shapes as they migrate downstream (to the right) as traveling waves.
                                                   86

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Carbon Exchange Dynamics  in a Temperate Forested Watershed
(Northern Michigan):  A Laboratory and Field Multidisciplinary Study
Lynn M. Walter ', L.M. Abriola 2, J.M. Budai \ G.W. Kling 3, P.A. Meyers  ', J.A. Teeri 3,  and D.R. Zak 4
'Department of Geological Sciences;2 Department of Chemistry and Environmental Engineering;3 Department
of Biology; •* School of Natural Resources; The University of Michigan, Ann Arbor, MI
      Beginning in the mid-1800's, humans started an
uncontrolled experiment with carbon on earth.  It is now
apparent that the effects of increasing levels of atmos-
pheric CO2  resulting  from  this experiment will alter
carbon dynamics and the functioning of terrestrial and
aquatic ecosystems in ways that are not entirely under-
stood.  The goal of this project is to determine the fate
of organic carbon produced in temperate forests.  Such
forests constitute major potential sinks for anthropogenic
CO2.   Importantly, studies of carbon  allocation in
forests  under  enhanced and ambient  CO2  growth
conditions have shown that above and  below ground
carbon storage as well as root and microbial respiration
processes all increase with elevated CO2.  The question
driving this research effort is, "How does the increased
rate of carbon fixation and processing in forest stands
and  soils  affect  the  overall  carbon budget at the
watershed scale?"   Is the  additional organic carbon
merely recycled via respiration  and returned to the
atmosphere, or is it transformed and transported from
the  soil zone to the regional  groundwater system for
longer term storage?
      The project is taking place in a hydrologically and
physiographically constrained portion of the Cheboygan
watershed. Within  the  watershed confines are two well-
studied forest stands and an established  elevated CO2
experiment in which aspens and sugar maples are grown
in mesocosms.  Soil compositions and water chemistry
are characterized in each experimental mesocosm and in
instrumented natural forest stands (aspens vs. sugar
maples) to determine how root respiration and microbial
processing of organic  matter may be linked to carbon
transport out of the rooted zone. As a direct measure of
mineral weathering effects  on  dissolved  inorganic
carbon  transport,  prepared  experimental arrays  of
feldspar and carbonate minerals have been implanted in
natural forests and  in experimental mesocosms.
      The  first year of the project was devoted to
hydrologic characterization of the watershed system,
establishment  of soil  water sampling  arrays,  and
general chemical  characterization  of soil  waters,
surface waters, and groundwaters in the study area.
Soil waters exhibit large vertical chemical variations,
generally  grading from dilute,  dissolved  organic
carbon-rich solutions  in  the  upper  20  cm  into
mineralized solutions chemically similar to regional
groundwaters by  4 meters  in depth.  Mass balance
among dissolved carbon species suggests that dissolved
organic carbon (DOC)  originating from reactions in
the upper rooted zone  is transformed to dissolved
inorganic carbon (DIG) via respiration and coupled
mineral solubilization reactions.   Dissolved silica and
aluminum increase rapidly  in soil waters suggesting
that aluminosilicate minerals, as  well as  carbonate
minerals, are dissolving as  DOC (especially organic
acid anions) is transformed to DIG (see  Figures la and
Ib).   The  significant solute acquisition and carbon
transformation evident  in soil water profiles suggest
that organic processes active in the upper soil horizons
are closely linked to mineral dissolution mechanisms
and the overall rate of solute transport out of the soil
zone.
      The next 2 years of the project will involve
generating an overall carbon budget for the watershed
and modeling carbon exchange rates among the main
carbon reservoirs. The  experimental mesocosms will
yield   more information on links among  growth
conditions,  soil  water  chemistry,  and  DOC/DIG
fluxes as the second summer of tree growth begins in
1998.  Regional groundwater flow modeling together
with more  detailed groundwater sampling transects
will better define the  horizontal transport effects
along flow paths relative to the vertical effects in soil
water chemical profiles.
                                                  87

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                      Figure la.
                                                              Figure Ib.
Figures la and Ib.
As shown in scanning electron phomicrographs,  "Minerals present in glacial drift soil horizons, such as
carbonates (a) or aluminosilicates (b), undergo dissolution and enhanced solubilization in organic acid and
CO2-rich soil waters."

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 A Comparison of Agricultural vs. Forested Basins:  Carbon
 and_Nutrient Cycling Within the Hyporheic Ecotone of Streams
 David S. White, Susan P. Hendricks, Timothy C. Johnston,  George Kipphut, and William E. Spencer
 Center for Reservoir Research and Hancock Biological Station, Murray State University, Murray, KY
                   Figure 1. Installing gravel bar wells at the Ledbetter Creek, Kentucky, hyporheic study site.
       Processes and patterns within the hyporheic zone,
 the  interface between groundwater and surface water
 beneath streams, may help explain differences in stream
 functions  between forested and agricultural  basins.
 Land use effects on subsurface processes mediated by
 increased  nutrient,  carbon, and sediment loads are
 largely unknown.  This  project's  goal  is to examine
 differences in  the  function  of hyporheic ecotones of
 third order streams in pristine and agricultural water-
 sheds.  The hyporheic ecotone functions as a  biogeo-
 chemically active   interface  where  biogeochemical
 storage  and processing  are  directly  linked  to the
 longterm health  and productivity  of surface  waters.
 Anthropogenic alterations within watersheds (such as
 increased  sedimentation  or runoff) potentially  alter
 hyporheic  function and therefore the biotic integrity of
 stream  ecosystems.  Ledbetter  Creek (LC)  is  an
 agricultural watershed  located in  Galloway County,
 Kentucky;   Panther  Creek  (PC) is a  forested basin
 located in  Stewart County,  Tennessee.  The two study
 streams both empty  into Kentucky Lake. Hyporheic
 zones within specific third order  sites on both streams
 have been mapped and monitored  with wells, mini-
 piezometers  and  seepage   meters  (see Figure  1).
 Monitoring provides baseline data for specific investiga-
 tions and  manipulations  under  base  flow and  high
 discharge conditions.
      At base flow, LC surface discharge contains more
 nitrogen but less phosphorus than PC.  Also, LC carries
 twice the suspended solids and four times the suspended
 organic matter load.  Fine sediment deposition on the
bed  surface in LC (at approximately 10 times the rate
 for PC) appears to decrease surface flow infiltration into
the bed and through  gravel bars. Decreased surface-
subsurface  interaction  is  demonstrated by   several
measurements.  Vertical hydraulic gradients (VHG) are
minimal or slightly positive throughout the  LC  site.
 Conversely, VHG at PC are more distinct,  indicating
 extensive upwelling and downwelling areas through bed
 sediments. Conservative tracer studies using bromide
 and a one-dimensional transport model indicate a much
 higher degree of surface-subsurface connectivity at PC.
 Several  gravel  bar  wells  at  PC  have  shown rapid
 hydrologic exchange with surface water (100 percent),
 while similar wells at LC have shown little hydrologic
 exchange within a 9-hour period. There is less oxygen
 in the LC hyporheic ecotone, commensurate with higher
 concentrations  of phosphate,  ammonium,  iron,  and
 manganese as  well as methane, particularly within the
 gravel bars.   In general at base flow, the  hyporheic
 ecotone at LC is much more isolated from  surface water
 than at PC.
      A 1-hour, half-inch  rainfall  in the  LC basin
 produces a steep hydrograph (0.03 to 2.0 m3 sec"1) within
 2  hours  but little measurable change in PC  discharge
 because of basin retention differences.  With  increasing
 discharge, LC surface water contains exponentially more
 suspended solids, nitrogen, and phosphorus, presumably
 derived from runoff but potentially from hyporheic storage
 as well. At PC, increased discharge causes a slight increase
 in suspended solids, but surface water nitrogen and phos-
 phorus concentrations  decrease, demonstrating surface
 water dilution.  Storm flows in LC tend to remove settled
 fine sediments,  increase subsurface  flow,  and decrease
 hyporheic concentrations of ammonium and phosphate.
 Whether the changes in subsurface chemistry  represent flux
 into surface  water or  redox  changes in the subsurface
 environment is now being investigated.  Concurrent studies
 in PC and LC hyporheic ecotones are now being conducted
 on  microbial diversity and  production, benthic algal
production, and  sediment fauna (hyporheos)  secondary
production.  Initial results  indicate  that PC  is more
biologically diverse with generally higher rates of second-
ary production.

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In Situ  Assessment of the Transport and
Microbial Consumption of Oxygen in Groundwater
Tadashi Yoshinari
Wadsworth Center, New York State Department of Health and SUNY, Albany, NY
R.L.  Smith
U.S.  Geological Survey, Boulder, CO
J.K.  Bohlke and K. Rtvesz
U. S.  Geological Survey, Reston, VA
    50
 = 40

 8.
 o
oo
    30
    20
\AfelllD

 473; E = 5.4
 26-11; E = 10.0
 343 top; E = 5.9
 343bot;E = 1.r
 13-16; E = 5.2
 62-12; E = 3.6
 593; E = 2.8
                                             air
                                           saturated
                                         water (10 oq
      -10123
             In O2 concentration in mg/L

       Levels  of dissolved oxygen  in groundwater
 influence the  rates and extents of biodegradation of
 organic pollutants within an aquifer.  Because microbial
 respiration is the most important mechanism of oxygen
 consumption  in this environment and  thus  is  a  key
 determinant of aquifer geochemistry,  the investigators
 have been conducting a  study  to  quantify aerobic
 respiration within a plume of dilute treated sewage in a
 sand and gravel aquifer in Cape Cod,  Massachusetts.
       The  approaches being  used to assess aerobic
 respiration  include:  (1) quantifying  rates of oxygen
 consumption using  in  situ tracer tests and laboratory
 incubations  with  sediment  core  and  groundwater
 samples and  (2)  analyzing the concentrations  and
 isotopic composition of O2 in various geochemical zones
 within  the  aquifer  and  determining  the  relative
 importance of dispersion and respiration in controlling
 oxygen levels within these zones.
       Rates of oxygen consumption were determined by
 three methods:  (1) A natural-gradient tracer test was
 conducted to directly measure in situ rate of oxygen
 demand.  (2)  Oxygen  consumption was measured in
 laboratory  incubations at in   situ  temperatures  and
 oxygen concentrations with sediment core slurries using
 a gas chromatographic (GC) technique. (3) An assay of
 bacterial electron transport system (ETS) activity was
 conducted in parallel with the GC analysis by amending
Figure 1. Variations in the concentration and isotopic composition of
dissolved oxygen near  the upper  boundary of a contaminated
groundwater plume at Cape Cod. Epsilon (e), the apparent isotope
fractionation factor, is defined by a version of the Rayleigh distillation
equation; l,000*ln(R/Ro) = -e*ln(C/C0), where R = 18O/16O, C =
concentration, and RQ and C0 denote the initial condition (before
reaction). The 618O value on the y-axis,  a measure of the relative
enrichment of 18O/'6O in a sample,  is defined by: 618O = 1,000*
(R/RSTANDARD -1) ^d is approximately equal to l,000*ln(R/R°). Thus,
the e value is a measure of the rate of change of 618O during decline
in the dissolved O2 concentration. Here it is assumed that groundwater
had initial values around O2 = 340 ^M and 618O = +24.3 per mil (in
equilibrium  with air at  10  • 12°C).   The  concentrations of O2
decreased while values of 6I8O-O2 increased from +24 to at least
+45 per mil, indicating that  the O2 consumption by microbial
processes is  taking place in the plume water to varying degrees.
                 sediment core  slurries with the tetrazolium salt  INT,
                 which acts as an artificial electron acceptor.
                       Rates  of oxygen  consumption in the sediment
                 core incubations, as measured by GC, were dramatically
                 higher than rates calculated from the in situ tracer test.
                 Conversely, rates of oxygen consumption from the INT
                 assay were significantly lower than the in situ rates.
                 The data  suggest that incubation  of aquifer  samples
                 within bottles in the laboratory appreciably stimulates
                 microbial respiration, even when in situ temperature and
                 oxygen levels are maintained and that INT is toxic.
                       The main  factors  that determine  the  relative
                 enrichment of 18O/16O in O2 in an aquifer are: (1) microbial
                 reduction  of O2 that causes  an increase  in the  ratio  of
                 18Q/16Q  ^ ^  remaining fraction of O2  due to kinetic
                 isotope fractionation and (2) dispersion of O2, which could
                 result in concentration gradients without  major  isotopic
                 variation.   Figure 1 shows an increase of  18O  (see the
                 definition   in the  figure  caption)  with   decreasing O2
                 concentration through  a vertical O2 gradient  at  seven
                 locations  in the  aquifer,  clearly   indicating   that O2
                 consumption has occurred at these sites. The low apparent
                 e values suggest that the vertical O2 concentration gradient
                 within the aquifer  is caused by a  combination of O2
                 consumption and dispersion in varying degree. Numerical
                 models  are currently being applied  to better define the
                 relationship between O2 consumption and dispersion.
                                                     90

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                                    Index of Authors
Abriola, L.M., 87
Allen, D., 51
Anderson, S., 31
Andrews, R.N., 32
Arnosti, C., 45
Ashton, M., 3
Bain, M.B., 14
Ballard, S., 11
Barten, P.,  3
Beck, M.B., 2
Benfield, E.F., 6
Bennett, L., 3
Benoit, G., 3
Bernert, J., 36
Beschta, R.L., 64
Blough, N.V.,45
Blum, M.D., 82
Bohlke, J.K., 90
Booth, D.,  19
Booth, D.B., 28
Bosch, D.J., 6
Bostrom, G., 16
Botsford, L., 35
Bott, T., 38
Boumans, R., 49
Braden, J.B., 25
Brezonik, P.L., 26
Brooks, R.P., 12,47
Brown, W., 14
Budai, J.M., 87
Surges, S.J., 28
Bushek, D., 51
Caneday, L., 17
Caraco, N., 83
Carpenter, S.R., 75
Carper, W.R., 55
Clark, D., 19
Cleveland, C.J.,  10
Colbert, D.L., 16
Cole, J.J., 75, 83
Cole, C.A.,47
Cooley, K.R., 84
Corbett, C., 51
Cosby, J., 11
Costanza, R., 49
Couch, C.A., 66
Cox, W.E., 6
Crosby, G., 84
Cruse, R., 36
Dame, R.F., 51
Danielson, B., 36
Deegan, L.A., 4, 53
Demissie, M., 25
DeVuyst, E.,  25
Dinar, A., 79
Diplas, P., 6
Dixon, A., 55
Driscoll, C.T., 61
Easter, K.W., 26
Edwards, D., 51
Eilers, J., 36
Estes, M.K.,  60
Evans, B.M., 12
Fernandez, I., 11
Ferreri, C., 38
Findlay, S., 83
Finley, J., 38
Flerchinger, G.N., 84
Focht, W., 17
Ford, M.S.J., 16
Fox, J., 29
Freemark, K., 36
Freyberg, D.L., 56
Galatowitsch, S., 36
Giambelluca,  T., 29
Glotfelty, C., 38
Gordon, S.I., 57
Govindaraju,  R.S., 59
Gowda, P.H., 8
Grant, S.B.,  60
Greene, R.G., 6
Gregory, L.,  51
Griffin, R., 19
Hairston, Jr., N.G., 61
Hamlett, J.M., 12
Hanson, C.L., 84
Haroa, R.J., 8
Hendricks, S.P.,  89
Herman, J.S., 62
Herricks, E.E., 33
Hershey, R., 31
Hession, W.C., 38
Hoffman, E., 79
Holomuzki, J., 81
Hopkinson, C., 10
Hornberger, G.M., 62
Horwitz, R., 38
Howitt,  R., 79
Hristov, T.N.,  12
Jackson, J., 38
Johnson, J., 38
Johnson, M., 35
Johnston, T.C., 89
Kahl, J., 11
Kaplan,  L., 38
Karr, J.R.,28
Kauffman, J.B., 64
Kearns,  C.M.,  61
Kellert,  S., 3
Kibler, D.F., 6
Kipphut, G., 89
Kitchell, J.F., 75
Kjerfve, B., 51
Kling, G.W., 87
Knight,  C.G., 12
Koepfler, E., 51
Komar,  P., 16
Kramer, E.A.,  66
Kremer, J., 4
Lammers,  R., 72
Larive, C.K., 55
Lewis, V.P., 65
Lewitus, A., 51
Li, H.W., 64
Li, J.L.,64
Lincoln, D.E., 65
Lohani,  V.K.,  6
Loucks, D.P.,  14
Lovell, C.R., 65
Luce, C., 84
Ludwig, P., 11
Lund, J., 35
Lynch, R., 17
Lynn, W.R., 14
Mageean, D., 11
                                             91

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                            Index of Authors (continued)
Matzke, G., 36
Maxwell, T., 49
McDonnell, A.J.,  12
McDowell, P., 64
Mclntosh, B.A., 64
McManus, J., 16
Meo, M., 17
Meyer, J.L., 66
Meyers, P.A., 87
Miller, W W., 31
Miller, G.C., 31
Mitra, P., 25
Morel, F.M.M., 68
Mostaghimi, S., 6
Moyle, P., 35
Mulla, D., 26
Myers, W.L., 70
Nagarkatti, P.S., 6
Napier, T.L., 8
Nassauer, J., 36
Neale, C.M.U., 84
Newbold, J.D., 38
Norton, B.C., 2
Norton, S., 11
Novotny, V.,  19
Olson, T.M., 60
Orlob, G., 35
Orth, D.J., 6
Pace,  M.L., 75, 83
Papellis,  L., 31
Parker, G., 69
Patil, G.P., 70
Patten, B.C., 2
Paul, M.J., 66
Perry, J.A., 26
Petersen, G.W., 12
Peterson, B., 72
Pickett, S.T.A., 20
Polasky,  S., 36
Porter, K.G., 2
Prasad, R., 84
Prins, T., 51
Puente, C.E., 74
Quinn, J.,  35
Rambo, A.T., 29
Rasmussen, T.C., 2
Rassenti, S., 79
Rastetter, E., 10
Reice, S.R., 32
Revesz, K., 90
Rhoads, B.L., 33
Richerson, P., 35
Richey, I.E., 34
Roberts, P.V., 56
Rustad, L., 11
Sabatier, P., 35
Saiers, J.E., 62
Sankowski, E.T.,  17
Santelmann, M., 36
Schauman, S., 28
Schindler,  D.E., 75
Schlager, E., 76
Schmandt, J., 77
Schneider, D., 25
Schubauer-Berigan, J., 51
Seyfried, M.S.,  84
Shabman, L.A., 6
Shaw, D., 31
Shepherd, A., 2
Sipes, J., 17
Skelly, D.,  3
Slaughter,  C.W., 84
Smith, C., 16
Smith, V., 79
Smith, R.L., 90
Sobsey, M.D., 80
Sparks, R.E., 25
Spencer, W.E.,  89
Standley, L., 38
Steenhuis, T.S., 14
Stephenson,  K., 6
Stevenson, R.J., 81
Straffin, E.G., 82
Strayer, D.L., 83
Suchanek, T., 35
Sun, C., 84
Sweeney, B., 38
Tarboton, D.G., 84
Taulbee, W.K., 66
Teeri, J.A.,  87
Turco, R.P., 40
Tyler, S., 31
Vallino, J., 10
Valocchi, A.J., 85
Vieux, B., 17
Villa, F.,49
Voinov, H.,  49
Voinov, A.,  49
Vorosmarty, C., 72
Wainger, L., 49
Walter, L.M., 87
Ward, A.D., 8, 57
Wardrop, D.H., 47
Webler, T.,  4
Weinberg, M., 35
White, D., 36
White, D.S., 89
White, D.C., 25
White, D., 57
Wiley, M., 81
Willett, K.,  17
Wilson, D.,  33
Woodin, S.A., 65
Xia, R.,25
Yoshinari, T., 90
Zak, D.R., 87
                                             92

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