Ecological Research Series
              AVAILABILITY, UPTAKE  AND
TRANSLOCATION OF  PLUTONIUM  WITHIN
                    BIOLOGICAL SYSTEMS:
A Review of the Significant Literature
                  Environmental Monitoring and Support Laboratory
                         Office of Research and Development
                        U.S. Environmental Protection Agency
                              Las Vegas, Nevada 89114

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                  RESEARCH REPORTING SERIES

 Research reports of the Office of Research and Development, U.S. Environmental
 Protection Agency,  have been grouped into  five series. These five  broad
 categories were established to facilitate further development and application of
 environmental technology. Elimination of traditional grouping was consciously
 planned to foster technology transfer and a maximum interface in related fields.
 The five series are:

      1.    Environmental Health Effects Research
      2.    Environmental Protection Technology
      3.    Ecological  Research
      4.    Environmental Monitoring
      5.    Socioeconomic  Environmental Studies

 This report has been assigned to the ECOLOGICAL RESEARCH series. This series
 describes research  on the effects of pollution on humans, plant and  animal
 species, and materials. Problems are assessed for their long- and short-term
 influences. Investigations include formation, transport, and pathway studies to
 determine the fate of pollutants and their effects. This work provides the technical
 basis for setting standards to minimize undesirable changes in living organisms
 in the aquatic, terrestrial, and atmospheric environments.
This document is available to the public through the National Technical Informa-
tion Service, Springfield, Virginia 22161.

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                                                EPA-600/3-76-043
                                                April 1976
AVAILABILITY, UPTAKE AND TRANSLOCATION OF PLUTONIUM WITHIN
BIOLOGICAL SYSTEMS:  A Review of the Significant Literature
                            by
           Anita A. Mullen and Robert E. Mosley
   Monitoring Systems Research and Development Division
      Environmental Monitoring and Support Laboratory
                  Las Vegas, Nevada 89114
           U.S. ENVIRONMENTAL PROTECTION AGENCY
            OFFICE OF RESEARCH AND DEVELOPMENT
      ENVIRONMENTAL MONITORING AND SUPPORT LABORATORY
                  LAS VEGAS, NEVADA 89114

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                                DISCLAIMER

     This report has been reviewed by the Environmental Monitoring and Support
Laboratory-Las Vegas, U.S. Environmental Protection Agency, and approved for
publication.  Mention of trade names or commercial products does not constitute
endorsement or recommendation for use.
                                  NOTICE

     Effective June 29, 1975, the National Environmental Research Center-Las
Vegas (NERC-LV) was designated the Environmental Monitoring and Support
Laboratory-Las Vegas (EMSL-LV).  This Laboratory is one of three Environmental
Monitoring and Support Laboratories of the Office of Monitoring and Technical
Support in the U.S. Environmental Protection Agency's Office of Research and
Development.

     Effective January 19, 1975, the U.S. Atomic Energy Commission (AEG) was
reorganized into two separate agencies, the Nuclear Regulatory Commission and
the U.S. Energy Research and Development Administration (ERDA).
                                     ii

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                                 ABSTRACT

     This report is a selective review of the literature on the availability
of plutonium in the environment and its cycling throughout representative
biological systems ranging from large biomes covering hundreds of miles to
the molecular transformations within individual cells.  No attempt was made
to develop a comprehensive bibliography.  Rather, references were selected
for inclusion as representative documentation for the vast spectrum of
material that is available on the subject.

     Important general references are listed separately.  Thereafter the
literature is described in essay form on a subject basis.  References
cited by number in the text are listed in complete bibliographic form at
the end of the report together with an author index.  The majority of the
material reviewed is limited to relatively recent publications.
                                   111

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                                CONTENTS
                                                                  Page
Abstract                                                           1:L1
Acknowledgments                                                     v^
Conclusions                                                          1
Introduction                                                         *•
                                                                     r\
  Objective
  Approach                                                           3
  Organization            .                                           3
Bibliographies                                                       3
Potential Sources of Plutonium                                       5
  General                                                            5
  Weapons Development                                                "
  Weapons in Transport                                               8
  Fuel Processing and Reprocessing                                   8
  Fires                                                              9
  Waste Disposal from Plants                                         9
  Shipment                                                          10
  Nuclear Reactors and Radioisotopic Generators                     10
Transport of Plutonium from Sources to Biosphere                    13
  Introduction                                                      13
  Chemical and Physical States of Environmental Plutonium           14
    Soil                                                            14
    Air                                                             16
    Water                                                           17
  Biological Aspects of the Physicochemical Complexities            20
  of Plutonium
    Mammals                                                          21
    Plants                                                          24
                                     IV

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                                                                 Page
Biological Deposition and Effects                                 24
  Deposition and Effects of Pu in Aquatic Environments            24
  Deposition, Effects and Countermeasures of Pu in Soil,          27
  Plant Environments
    Soil                                                          27
    Plants                                                        29
  Absorption of Plutonium by Animals                              31
  Biological Behavior of Inhaled Plutonium in Animals             37
  Bone Deposition                                                 39
  Biological Effects of Plutonium in Animals                      40
    Blood                                                         41
    Bone                                                          41
    Liver                                                         42
    Lung                                                          43
    Lymph                                                         4 3
    Other Effects                                                 44
  Countermeasures for Plutonium in Animals                        45
  Deposition of Plutonium in Man                                  46
  Effects of Plutonium in Humans                                  49
  Countermeasures for Humans Exposed to Plutonium                 49
  Contamination
Present Analytical Methods                                        49
  General Methods                                                 50
  Autoradiography                                                 52
  Instrumental Analysis                                           52
  Bioassay for Plutonium                                          53
  In Vivo Measurement of Plutonium                                54
  Monitoring Instrumentation                                      55
References                                                        57
Author Index                                                      85

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                             ACKNOWLEDGMENTS

     The computer search of the Nuclear Science Abstracts was provided by
the Nevada Applied Ecology Group, Oak Ridge National Laboratory.  Special
thanks to all who aided in the search for material .from 1970 to 1975
inclusive.

     Much information was derived from the extensive indexed bibliography
of plutonium documents prior to 1970 that was compiled by Dr. M. G. White,
formerly of EPA and now with ERDA, Nevada Applied Ecology Group, Nevada
Operations Office and R. 0. Houston of EPA, EMSL-LV, who helped research
and organize the documents, and D. Wickman, EPA, EMSL-LV, who ordered many
of the documents referenced.

     Special appreciation is due to J. M. Burford, D. D. Wicker and the
many others who sorted and indexed the abstracts and documents.

     The efficient typing and preparation of the manuscript by P. A. McGill
and S.  F. Forshee are deeply appreciated.

     Special thanks to Omer Mullen for assisting with the sorting, editing
and proofing of the material.

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                               CONCLUSIONS

     This review has briefly covered many topics but has addressed only a
small portion of the articles published over the past 25 years.

     As additional energy requirements increase the number of nuclear power
generators, processing and reprocessing plants throughout the world, the
potential increases for the distribution of plutonium within a variety of
biomes previously only touched by worldwide fallout.  At this time it appears
that more is known of the interactions of plutonium within a desert ecosystem
than in any other system.  The largest single group studying plutonium in the
environment is the Nevada Applied Ecology Group (NAEG) which is supported by
the Energy Research and Development Administration.  Its main concern has
been the area of the Nevada Test Site and its immediate environs.

     Additional studies are needed to determine the physicochemical inter-
actions of plutonium in areas receiving moderate to heavy rainfall, which
results in lush vegetation with its attendant litter and greater quantity
of decomposition products affecting soil chemistry and fertility.  Tempera-
ture extremes causing seasonal variations in plant constituents, endemic
and transient wildlife populations, invertebrate community size and average
age of individuals within these ecosystems all must be taken into considera-
tion.

     Differing soil types are known to affect the translocation and avail-
ability of plutonium.  What is not clearly defined is the effect of adding
amendments to the soil in order to increase fertility.  Nitrogen and trace
minerals may be added to soil in many chemical forms, each of which may
change the availability of plutonium for absorption by flora or fauna.

     The selectivity of organisms for iron and plutonium present in the
same biomass is not thoroughly understood.  This may be affected by
seasons, nutritional needs, altitude and age of the receptor.

     Investigations underway by NAEG indicate that the availability of
plutonium may be increased each time it is ingested by an animal, passes
through the digestive processes and is excreted.  This may concentrate
plutonium in vegetation growing on plots of ground previously fertilized
by the application of contaminated manure.  More work must be done before
these complex interactions can be understood.

     Some work has been completed on the behavior of plutonium in aquatic
systems.  The reactions of plutonium particulates in fresh and saline water
are just beginning to be understood.  Incorporation of particulates within
bottom sediments may isolate plutonium and prevent dissolution for long
periods of time but violent agitation, e.g., storm wave action, mechanical
mixing or erosion and flooding of large land areas and deposition of the

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 sediments  on farm lands may  cause  a  release of this stored plutonium and
 increased  uptake  by  biological  systems.  This mechanism should be better
 understood since  most  nuclear power  plants are located near large sources of
 water  increasing  the potential  for low level chronic release in effluent
 streams.

     The role, microorganisms play  in affecting availability of plutonium
 in soil-plant relationships  is  being studied on a  limited basis.  Little
 is known of plutonium  interaction  with microorganisms present in the
 gastrointestinal  tract.  Work has  started on the absorption of plutonium
 by algae.   Ingestion of this algae by aquatic organisms may lead to further
 concentration and transfer up the  food chain.

     There are many  studies  underway by  the NAEG as summarized by Dunaway and
 White2"* in 1974.   Many of these studies  seek to further information on the
 behavior of plutonium  in fauna  found within a desert environment.  These studies
 encompass  tissue  concentration  of  plutonium in cattle grazing in contaminated
 areas, rodents and lizards inhabiting the same area, transfer of various forms
 of plutonium to milk by dairy cows,  and  concentration of plutonium in tissues
 and eggs of chickens ingesting  differing forms of  plutonium.

     Many  authors have devoted  much  time to extrapolating the results
 obtained with laboratory animals to  man.  This comparison though perhaps
 valid  in many instances, may prove to be of limited use when the compli-
 cating factors and discriminating  mechanisms inherent in man's physiology
 are taken  under consideration.  The  effects of chronic plutonium contami-
 nation in  humans  necessary for  a true evaluation of its hazards will continue
 to be  a source of conjecture for many years in the future.  Man can only hope
 to limit the concentration of this potentially hazardous radionuclide in the
 environment until additional knowledge is forthcoming.
                               INTRODUCTION

 OBJECTIVE

      The objective  of  this  review is  not  to  develop  a  comprehensive  biblio-
 graphy  on plutonium, but  rather  to identify  and  comment  upon  the  significant
 and  relevant  literature representing  the  many  detailed studies  of the inter-
 action  of plutonium in biological systems.   Material included in  this review
 has  been selected to provide  an  overview  of  the  subject  and to  indicate
 where deficiencies  may exist  in  biological data.

     Plutonium, produced  for  the first  time  in 1941, has been released and
 distributed throughout the  biosphere  during  the  last 25  years.  The  physico-
 chemical  changes induced  by natural processes  may  enhance the biological
 availability  and increase the concentration  of plutonium in specific biomes
 thereby  causing greater biological effects and risk  to mankind.

     The  concern then  is  to limit the production and control  the  use of
plutonium until such time as  the very complex  interactions with biological
systems have  been investigated and a  better  understanding of  potential

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hazards has been achieved.  The need for energy makes It imperative that
mankind make most efficient use of available sources and channel the
potential of plutonium-derived energy to the benefit of all, while mini-
mizing the deleterious effects on man and his environment.  Only by an
overall understanding of what has been found in the past can guidelines be
set for the future.

APPROACH

     Literature included in this report was selected on the basis of its
significance and relevance from a variety of bibliographies, general
references and abstracts.  Where the original reference  was not available
to judge the value of the reference, the abstract was consulted to determine
its applicability.

ORGANIZATION

     This report lists annotated bibliographies, important general references
and abstracts.  Thereafter, the literature is reviewed in essay form on a
subject basis.  The subject headings are covered in individual sections
and under each of these, several subsections review literature on a given
topic.  The 419 references reviewed are cited in the appropriate subject
divisions of this report and in complete bibliographic form at the end
of the report in the order mentioned in the text.  Finally, an author
index  of the references cited is included to facilitate the location of
particular material.
                              BIBLIOGRAPHIES

     The  following bibliographies  are closely related to the subject.

     (1)  Environmental Aspects of Plutonium - A Selected Annotated
Bibliography,  September,  1972, Environmental Plutonium Data Base Group,
Oak Ridge National Laboratory, Oak Ridge, TN, ORNL-EIS-72-21, 387 p.

     (2)  Ibid (Suppl. 1) August,  1973, ORNL-EIS-73-21, 479 p.

     (3)  Ibid (Suppl. 2) February, 1974, ORNL-EIS-74-21, 272 p.

     These  three volumes  contain over 2,000 references  on the environmental
aspects of  plutonium and  uranium at the Nevada Test Site (NTS).  The litera-
ture was  selected to meet the needs of the Nevada Applied Ecology Group  (NAEG)
of the Nevada  Operations  Office, Las Vegas, Nevada.

     (4)  Environmental Aspects of the Transuranics - Compiled and edited by
Martin, F.  M. ,  Sanders, C. T., and Talmage, S, $. , Oak Ridge National
Laboratory, TN, ORNL-EIS-74-2 (Suppl. 3) December 1974, 226 p.

     This bibliography contains 528 references on the environmental aspects
of uranium  and the transuranic elements, with a preponderance of material
about plutonium.  Indexes are given for authors, subjects, categories,

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 keywords, geographic locations,  permuted titles,  taxons,  and publication
 descriptions.

      (5)  Biological Effects of  Plutonium.   A Bibliography  - Compiled
 by Suratno,  P., CE-BNL-Bib-6, 60 p.

      This bibliography was compiled  mainly  from Nuclear  Sciences  Abstracts
 for the period 1969-1973.   It was produced  in support  of  the continuing
 work at Berkeley Nuclear Laboratories on the health physics problems
 associated with the transuranic  elements.   Its arrangement  is  in  three
 sections:  A.   Biological Effects (47 refs); B.   Physiological Aspects
 (110 refs);  and C.   Techniques of Measurement (60 refs).

 General Reference Reviews and Summaries

      The following reports or references give reviews  and summaries of
 available information related to the subject.

      (1)  Uranium-Plutonium-Transplutonic Elements, Hodge,  H,  C,, Stannard,
 J. N., and Hursh, J. B., Eds., Springer-Verlag, New York, 995  p., 1973.

      This book is separated into three broad categories and 21 chapters
 which deal separately with uranium,  plutonium, and the transplutonic
 elements.  In general the sections discuss  history; chemical and  physical
 properties;  distribution, excretion, and effects; body burdens; bioassay;
 health physics aspects; and environmental considerations.

      (2)  Radiobiology of Plutonium, Stover, B. J. and Jee, W, S. S.,  Eds.
 The J. W. Press, University of Utah, Salt Lake City, Utah,  552 p., 1972.

      This book is separated into 25  chapters dealing with general topics
 of historical background,  metabolism and effects  of plutonium  as  pertains
 to beagle studies,  metabolism of 239Pu in rodents, use of chelating agents*
 and plutonium in man.

      (3)  The Metabolism of Compounds of Plutonium and Other Actinides,
 ICRP Pub. 19,  Pergamon Press, New York, 59  p«, May, 1972.

      This report reviews literature  up to 1972 relating  to  the chemistry  of
 plutonium as related to biological behavior, e*g., entry  of plutonium
 by inhalation,  absorption  and retention of  deposited actinides.   Three
 tables  are given which summarize principal  inhalation  studies  with plutonium
 compounds in experimental  animals, absorption of  actinides  from  the gastro-
 intestinal tract  and distribution of actinide compounds  in  experimental
 animals and  man.

      (4)  "A Review  of Transuranic Elements in Soil, Plants, and  Animals  »
J. Environ.  Quality  2;1» pp,  62-65,  1973.

     Published  information concerning the distribution and  fate  of neptunium,
plutonium, americium and curium  in terrestrial ecosystems is reviewed in this
article.

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     (5)  "Physiopathology of Plutonium Contamination:  Fundamental
Concepts," Saenz, M« D. L. and Ramos, E,, translated by Ralph McElroy Co.
for Los Alamos Scientific Laboratory, Los Alamos, LA-TR-74-16, 40 p.,
1973, translated 1974.

     Published information concerning the physiopathology of plutonium
was reviewed including internal contamination, effects, dose and therapy.

     (6)  "Consideration of Reactor Accident Exposure Guides for Plutonium,"
Bair, W. J., BNWL-SA-4968, 23 p., 1974.

     This report reviews some of the biological aspects needed to be
considered in establishing an emergency reference dose for plutonium.  The
discussion is limited to plutonium entering the body through the respiratory
tract.

     (7)  "Evaluation of Plutonium-Contamination in Radioactive Waste
Disposal Areas with Respect to Their Potential Hazard and Eventual
Disposition," McCurdy, D. E., M. L. Wheeler, M. D. McKay, and R. K.
Lohrding, Quarterly Report Transuranic Solid Waste Management Research
Programs October-December 1973, Los Alamos Scientific Laboratory, p. 17-55.

     This report evaluates the plutonium contaminated waste disposal areas
of the Los Alamos Scientific Laboratory  (LASL) and analyzes the risk of
natural and man-made disasters that could be causal events for releasing
stored waste to the environment, the resulting abiotic transfer processes for
movement of the released  plutonium-contaminated waste, biological trans-
port mechanisms, radiological assessment of the intake of transuranic
elements and the radionuclide inventory of waste disposal areas at LASL
are also discussed.

     (8)  The Dynamics of Plutonium in Desert Environments, Dunaway, P. B.
and M. G. White, Eds., NVO-142, 369 p., July, 1974.

     This is a progress report of projects of the Nevada Applied Ecology
Group, U.S. ERDA, Nevada Operations Office and contains summaries of
work presently underway as well as preliminary reports of completed
projects.  The several topics covered are soils, statistics, vegetation,
animals, resuspension, distribution, and inventory modeling and infor-
mation as applies to plutonium on the Nevada Test Site and surrounding
environs.
                  POTENTIAL SOURCES OF PLUTONIUM

GENERAL

     Plutonium is a radioactive element of increasing importance as the
need for energy sources becomes more acute.  It is one of the most toxic
radionuclides because of its biological behavior as well as its physical
characteristics.

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      As summarized by Saenz and Ramos1 in 1973,  the Importance of plutonium
 has been apparent since it was produced for the  first time by Seaborg and
 others as reported by Weast2 in 1966.   Although  the existence of plutonium
 in nature has been proven, its natural quantities are so low (Levine and
 Seaborg3 in 1951 and Hoffman1* et al.  in 1971)  that the risk comes from
 its artificial production.

      Of the radioactive plutonium isotopes, 239Pu is the one most used in
 the nuclear industry where it is produced by irradiation of 238U in nuclear
 reactors.  Its high specific activity and long half-life determine its
 high radiological toxicity (Saenz and Ramos1).

      Increasing use of plutonium in nuclear fuel, industrial, biomedical and
 space applications, as well as weapons development, increases the possibility
 of accidental releases contaminating the biosphere.

      Since 1952, weapons tests and nuclear accidents in space have released
 plutonium into the biosphere with resultant fallout over the entire earth
 (according to the estimates of Cherdyntsev5»®  et al. in 1968 and 1970)
 approximating a concentration of 239Pu of 10 12% when mixed in the first
 few centimeters of soil.  The values of liberated plutonium given by the
 Scientific Committee of the United Nations and reported by Norwood7 in 1971,
 assumes an activity of 0.5 mCi of 239Pu, equal to eight tons.

      Global air activity values for 239Pu were shown to be 10 15 Ci/m3 of
 239Pu in 1963 as reported by DeBortoli8 et al. in 1966 and 10~17 to 10~16 Ci/m3
 in the USA in 1965 as calculated by Drobinski9 et al. in 1966.  These values
 appear to coincide in order of magnitude with  later maximums arrived at
 by Volchok and Kleinman10 in 1971.

      Although the risk from weapons testing would appear to be low, the
 increasing use of plutonium in industry, medicine and aeronautics has caused
 Seaborg11 in 1970 to forecast that  these applications will use 60 to 80 tons
 of 239Pu in the next few decades and up to 6 tons of 238Pu by the end of the
 century.

      Environmental contamination as the result of accidents was reviewed by
 Eisenbud12  in 1973.   The events discussed include:  fallout from the thermo-
 nuclear  detonation of March 1,  1954;  the accident to the Windscale Reactor
 number one  of October  1957;  the Oak  Ridge plutonium release of November
 1959; the Army Stationary Low-Power Reactor (SL-1);  the Houston incident
 of March  1957;  the  Fermi fuel  meltdown; the abortive reentry of the
 SNAP 9-A; and  the  plutonium fire at Rocky Flats.

WEAPONS DEVELOPMENT

     The first  formal  environmental contamination experiments with plutonium
were carried out under the auspices of the U.S.  Atomic Energy Commission
 (AEG) at the Nevada  Test  Site.   In  1956 a test (Project '56) was carried
out to study the behavior of  released  plutonium  in air, soil, and the desert
environment.  No biological work was  included  in this experiment.  As        •'
reported by Stannard13  in 1973,  "Operation Plumbbob" conducted in 1957  a

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full scale biomedical program was undertaken by a group from the University
of Rochester (Wilson   et al.) along with more elaborate studies of aerosols,
cloud physics, decontamination and area monitoring studies by a group from
Sandia Corp. (Cowan15) and the United States Air Force.  The levels of
ground contamination were on the order of 2.6, 40 and 560 micrograms per
square meter of desert soil.

     "Operation Roller Coaster" was a. joint United States-United Kingdom
venture and was carried out in a remote area of north central Nevada.
There were four test firings and the emphasis was placed on inhalation
of plutonium from cloud passage according to Wilson and Terry16'17 in
1965 and 1968.

     An environmental inventory of the Los Alamos area was reported for the
period July 1, 1972 to March 31, 1973, by Hakonson18 et al. in 1973.  Included
in this survey was the Trinity area, scene of the world's first nuclear deto-
nation (July  1945).  A similar study of the 2,200 hectares surrounding Los Alamos
was reported by Johnson19 in 1972.  These environmental assessments included
careful search of the administrative records to determine the extent to
which the land might have been used or involved in the Laboratory's activities.
Extensive measurements of the radiation levels in the field, and radiochemical
analysis of numerous soil and vegetation samples were completed.  Results
showed that all measured values were comparable to reported worldwide levels,
and no radiation or radioactive contamination observations were encountered
that are of radiological health or environmental concern.  In a similar
                              n ft
report by Hakonson and Johnson   in 1973, a comparison was made between the
data collected 20 years earlier and that collected in 1972.  The major change
observed was an increased migration of plutonium downward through the soil
profile.  Concentrations of plutonium in vegetation and rodents were too low
to make valid comparisons.
                                                  o i
     The comprehensive review prepared by Eisenbud   in 1973 of fallout
distributed from nuclear explosion includes the amounts and distribution
of radioactive debris produced in weapons tests; methods of estimating
how radioactive debris is partitioned among the three components of fallout
(the fallout in the immediate vicinity of the explosion, the debris injected
into the troposphere, and the debris injected into the stratosphere); and
the behavior of plutonium released in weapons tests.

     In the quarterly reports of the New York Operations Office Health and
Safety Laboratory, Hardy  '   tabulates the data from the fallout monitoring
program.  The plutonium 'content of atmospheric samples obtained through
Project Air Stream during 1970 and 1971 and the plutonium content of
stratospheric samples obtained through the High-Altitude Balloon Sampling
Program during 1969 and 1970 are included in the report for March-June 1971.
The June-September 1972 report shows plutonium deposition at worldwide land
sites and those collected at Atlantic Ocean weather stations.

     Environmental surveys of the main areas of plutonium deposition at the
Nevada Test Site are included in the Nevada Applied Ecology Group Progress
Report edited by Dunaway and White   in 1974.

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 WEAPONS IN TRANSPORT

      American bombers carrying nuclear weapons have crashed  in Spain  in  1966
 and Greenland in 1968.  Problems connected with the detection and decontami-
 nation of these areas were discussed by Hvinden25  in 1973.   The area  contami-
 nated in Spain was cleared sufficiently after 2 or 3 months  to allow  agricul-
 tural use.  In Greenland, measurements of fish, birds,  foxes and seals showed
 no contamination above allowable limits.

      In a follow-up study, Hanson26 in 1972 was able to detect small  amounts
 of 239Pu and 21*°Pu in lichens collected near the Greenland site during
 July-August of 1968.  This activity was associated with -particles estimated
 to be 0.5 to 1.0 micrometers in diameter containing slightly  greater than
 background levels of plutonium radioactivity.

 FUEL PROCESSING AND REPROCESSING

      Increasing utilization of nuclear fuels will  result in  wide-scale
 plutonium recovery processing, reconstitution of fuel,  transportation
 and extensive handling of this material.   According to  Paxton27 in 1972,
 there have been only six supercritical accidents spread over 20 years of
 processing fissile material.  None of these was associated with mechanical
 processing, storage or transportation.  All occurred in recovery plants  and
 all occurred with aqueous solutions; four involved highly enriched uranium,
 and two involved plutonium.  One of those involving plutonium occurred at
 the Los Alamos Scientific Laboratory - December 30, 1958, and the other at
 the Recuplex Plant  at Hanford, Washington, April  7, 1962.   The results  of
 these 6 accidents have been 2 deaths, 19 significant overexposures to
 radiation, no equipment damage, and negligible loss of  fissile materials.
 In no case was there any danger to the general public.

      Many safeguards are built into facilities used for processing of nuclear
 fuels so that confinement of radionuclide release  to the environment  is kept
 to a minimum.   It is a well recognized problem that plants handling long-
 lived radionuclides may release extremely low levels of radionuclides daily
 through filters or other pathways.  Although these releases  may not be
 generally measured in terms of air concentration,  after many years of opera-
 tion  small quantities of the material can be detected outside of the  facility.
 This  has happened at Rocky Flats and other areas according  to Biles28 et al.
 in 1971.

     A  study was  made of the possibility of hazardous environmental plutonium
 releases  caused by uncontrolled oxidation of plutonium within the enclosure
 at Rocky  Flats  (Hunt  ).   The report reviews the existent observed data on
 restricted plutonium release and then constructs a release  model relating
 the free-release  source strength (i.e., the source strength outside the
 enclosure) and  the parameters describing the release.  Based on both the
 observation data  and the model calculations, the conclusion follows that
 restricted release is  unlikely to lead to dangerous free-releases of a
 plutonium aerosol.   The model calculations specifically show that the maxi-
mum credible accident  assumed,  would be safe by about 2 to  5 orders of
magnitude.  However,  the calculations indicate that maximizing either the

                                       8

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Plutonium release rate or the asymptotic temperature reached in an enclosure
can cause a free-release exceeding the safe rate.

FIRES

     The pyrophoric nature of plutonium metal has led to several plutonium
fires.  The major fires occurred at Rocky Flats in 1957 and 1969 and resulted
in airborne plutonium being distributed over a wide area surrounding the
facility and resulted in contamination of the soil as reported by Hardy and
Krey30 in 1970.

     An analysis of 46 incidents involving plutonium metal or compound under
thermal stress has been reported by Mishima and Schwendiman31 in 1971.  These
incidents, occurring in the nuclear industry during the period from 1952 to
1967, are cataloged as to the nature of the material involved, the initiating
event and the consequences.

WASTE DISPOSAL FROM PLANTS

     For several decades, transuranic-containing solid waste has been placed
in shallow burial grounds at several of the AEC contractor facilities.  The
hazards of the release of this radioactivity from the waste burial environ-
ment at the Los Alamos facility was discussed by McCurdy32 et al. in 1974.
The possible catastrophic events leading to the release of these materials
are addressed and a model is proposed for the movement of these radioactive
materials throughout the environment.

     A review of processing of plutonium-contaminated liquid wastes at
Los Alamos was reported by Emelity and Christenson33 in 1971.  This report
lists the quantities of liquid wastes processed during the years 1966
through 1970.

     Over the past 20 years, the National Reactor Testing Station (NETS) in
Idaho has accepted radioactive waste for burial.  Wastes received included
materials contaminated with transuranic elements, primarily from AEC's Rocky
Flats operation, as well as items contaminated with fission products and neutron
activated materials from various onsite operations.  Early disposal methods
varied from stacking of drummed and boxed waste into pits to random dumping
of various categories of waste into common trenches or pits.  Reassessment of
past disposal practices and concern for long-term isolation of transuranic
wastes led to the establishment of above-ground retrievable storage in
1970.  However, about 54,000 cubic meters of alpha waste remains buried at
NRTS.  The plan for retrieval of this waste for treatment and storage in
a Federal depository was reported by Hickman3"* in 1974.

     Liquid wastes from the plutonium finishing plant at Richland, Washington,
have been discharged to subsurface disposal facilities  (enclosed trenches)
since the startup of the facility approximately 22 years ago.  The removal
and disposal of this contaminated soil is discussed in AEC Environmental
Statements35*36 of January and April of 1972.

     New guidelines for the interim storage of solid transuranic wastes were
proposed by the H-Division staff of Los Alamos37 in 1974.  A summary  of

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 current practices in solid waste disposal  is  included.

      A review of radioactive waste  disposal and  its  eventual pollution  of
 groundwater from this waste was  completed  in  1974 by Todd  and McNulty

 SHIPMENT
                                                                A Q
      It has been estimated by the U.S.  Atomic Energy Commission   in 1972
 that by the year 2000 the total  nuclear power generating capacity  in the
 United States will be approximately 1200 gigawatts.   Over  the next 25 years,
 the expansion will increase the  need for shipping 239Pu (produced  in light
 water reactors and liquid metal  fast-breeder  reactors)  from fuel reprocessing
 plants to fuel fabrication plants where these fissile isotopes  can be recon-
 stituted into reactor fuel elements.

      If aqueous solvent extraction  processes  are employed  for separating
 plutonium in the spent fuel, the plutonium will  emerge  in  the reprocessing
 plant product streams as nitrates in aqueous  solutions.  The problems and
 hazards inherent in shipping these  solutions  were discussed by  Ulrica1*0 in
 1974.  In a conference on transportation of nuclear  material, Wischow1*1 in
 1970 reported case histories of  misrouted  and misplaced shipments.  Corrective
 action and regulations pertaining to physical prohibition  of fissile materials
 were briefly discussed.  Hazards associated with transportation accidents  in
 which plutonium was subjected to gasoline  fires  were reported by Mishima and
 Schwendiman42 in 1973.

      Becker"*3 in 1971 reported on steps that  LRL was taking to  improve  controls
 over packing of plutonium shipments.  Two  case histories were cited. The
 importance of quality control in packaging Pu(N03)if  solution and PuOa powder
 was discussed in an article by Brown and Heaberlin'*'* in 1974.   An  industry-
 wide survey was conducted of 775 shipments involving 6,200 packages over a
 period of 3 to 4 years.  Package closure error could result in  breaking of
 containment during shipment or a series of errors might release the contents
 during normal shipment.  A listing  of closure faults is provided with the
 number of times the faults have  been observed.

      This is not a complete record  of these faults as records on errors were
 not  kept  consistently as required to be kept. Although increased  quality
 control procedures implemented since 1972-1973 have  decreased  the  frequency
 of these  errors,  they have not been entirely  eliminated.

     The  problems  of calculating risk of criticality in transporting fissile
materials were  addressed by Thomas1*5 in 1971. Proper packaging and container
sizes are discussed.

NUCLEAR REACTORS AND RADIOISOTOPIC  GENERATORS

     Plutonium  is  produced by irradiating  uranium with  neutrons in a nuclear
reactor.  Its isotopic  composition  depends on the time  in  the  reactor and
the type of reactor.   This plutonium may be recycled in light-water reactors.
Utilization in  this  way will require reprocessing and fuel refabrication.
                                     10

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     The introduction of commercial fast-breeder reactors is not expected to
occur until the mid-1980's according to Pigford and Mann1*6 in 1973.  The
fast-breeder reactor has excellent fuel economy.  It is fueled with plutonium
and "breeds" plutonium from 238U, creating a surplus for the first fuel charge
of other fast-breeder reactors.  The danger from this type of reactor, accord-
ing to Hamilton   in 1971 is that the chain reaction can easily run away,
creating a supercritical mass which would eventually be blown apart.  The
total radioactivity in the yearly amount of fuel discharge, at the time of
discharge, is greater for the fast-breeder reactor than for the light-water
plant because of the need for more frequent replacement of core fuel in the
breeder.  The yearly amount of plutonium in the fast-breeder fuel cycle is
about eight-fold greater than in the light-water nuclear plant.  An experi-
mental 1000 Mw(e) fast-breeder reactor will contain about 1.8 to 2.8 tons
of plutonium corresponding to 2.2 to 3.5 million curies of plutonium
according to the U.S. Atomic Energy Commission48 report of 1972.  A flow
sheet listing the material and environmental release for a breeder reactor
nuclear power plant was presented by Pigford and Mann1*6 in 1973.

     A historical resume of the breeder reactor program and its impact on the
environment was given in a report by Daub49 in 1972.  A program plan listing
fuel materials and accident conditions for a liquid metal fast-breeder reactor
(LMFBR) was'presented by the Argonne National Laboratory50 in 1972.

     Safety aspects of nuclear fuel cycles were reviewed by Wymer51 in 1971.
Future developments and requirements for the nuclear fuel cycle are also
discussed.  Plutonium utilization in boiling water reactors is summarized
in a report by General Electric Company,52 released in 1971.  Activities
associated with t;he AEC-sponsored Integrated Safeguards Experiment and
Plant Instrumentation Program are reported.  A review of plutonium utili-
zation in thermal reactors was presented by Schmid53 in 1973.  Recycling,
fueling, decontamination and safety are discussed.  The drawbacks of nuclear
power were discussed by Engstroem54 in 1972.  Siting of power plants, radia-
tion hazards and environmental impact are included.  .The dangers associated
with nuclear power reactors were described by Lapp55 in 1975.  The possibil-
ities of accidents occurring are presented in an analytical diagram called
an "accident tree."  Of all the possibilities, the probability of a few
hundred of the most important was determined and of these 15 were judged
to be the major sources of risk.  An assessment of accident risks in U.S.
commercial nuclear power plants was reported by Rasmussen56 in 1974.

     The health physics aspects of fast-breeder reactors were discussed by
Heine and Hart57 in 1971.  The specific aspects are grouped according to
segments of reactor operations.  Under certain hypothetical accident
conditions where fuel vaporization could occur, plutonium could be the
dominant hazard.

     The results of a radiological environmental survey in the vicinity of
EBR-II were given by Oltman58 et al. in 1973.  Measurements were made at
radial distances up to 5 miles from the effluent stack.  Soil samples were
obtained and analyzed for plutonium content.  No plutonium deposition pattern
could be constructed.
                                      11

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       A series of studies to evaluate the  fractional  airborne release of
 Plutonium under various postulated reactor accident conditions was completed
 by Mishima and Schwendiman59 in 1972.   Data generated in earlier laboratory
 scale studies were also reviewed.   Topics  dealing with the acceptability  of
 nuclear fission for energy sources were discussed by Gofman60 in 1972.   The
 biological problems associated with nuclear fission,  the fission product
 problem, the plutonium problem and the major accident problems are included.
 The University of Washington nuclear reactor plutonium contamination incident
 of June 13, 1972, was discussed by Hilborn6l et al. in 1972.

      Both major and minor accidents involving the release of aerosols of
 sodium are possible during the generation  and maintenance of a fast reactor.
 These aerosols could be radioactive, involving not only the radioactive
 sodium, but also fission products, uranium and plutonium.  This problem
 was reviewed by Kotrappa62 in 1973.  The use of nuclear power generators,
 batteries and heat sources for use on land, in space  vehicles, ocean diving
 equipment and for powering artificial pacemakers, organs, and other medicinal
 uses increase  the possibility of environmental contamination from these
 sources.  A detailed nuclear safety evaluation of the SNAP-19 C generator
 was conducted by Conway and Dobry63 in 1973.  In 1972 Weiner6"* reported on
 the safety testing of a 238Pu fueled Radioisotope Thermoelectric Generator
 (RTG) for the Navy Transit Satellite.   The containment of the 238Pu fuel
 was checked for all normal reentry environments and for all accident
 environments during prelaunch, launch pad, early launch abort and pre-
 orbital insertion abort.

      The operating experience and advances in the safety of space nuclear
 power systems fueled with 238Pu were reviewed by Dix65 in 1972.  The opera-
 tional safety experience with eight NASA launches of  spacecraft powered
 with nuclear energy since 1968 including two Nimbus weather satellites,
 the Apollo 12, 13, 14, 15 and 16 lunar landing missions, and the Pioneer  10
 Jupiter mission was discussed.  The concept of isotopic power generators  is
 explained in a 1971 article by Penn66.  The heart and pacemaker batteries
 and the UKAEA RIPPLE (radioisotope-powered prolonged  life equipment) generator
 is described.   In 1971 Williams57 developed models for estimating the extent
 to which radioisotopic fuel forms will vaporize if released to the fires
 resulting from the catastrophic aborts of  rocket booster vehicles.  Pad
 aborts of both liquid- and solid-propellant systems are considered and  aborts
 after lift-off are also treated.  Estimates are given for the size and
 activity spectra of the radioactive particles produced when the vapor
 recondenses.   Numerical results are specifically calculated for certain
 238Pu(>2  fuel  forms (microspheres,  solid-solution cermet, and plutonia-
 molybdenum cermet).   Tables  for the ingrowth of daughter products  from  the
 decay  of  plutonium isotopes  used in heat sources are  given  in an  article
 by Haas and Campbell68 in 1972.   The typical composition of the plutonium
 metal  used in  the fabrication of the sources is also  listed.

     The  general  considerations of radioisotopic power and  its international
 aspects were discussed by Cellini and Stadie69 in 1972. The work of  the
OECD Nuclear Energy  Agency since 1967  in studying radioisotope batteries,
particularly for  cardiac pacemakers is discussed.  This work includes heat
source development,  capsule  evaluation, safety assessments of radioisotope
heat sources under normal and accident conditions, radiation hazards

                                        12

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evaluations, and human  implantation  of  238Pu  fueled pacemakers which began
in April 1970.

     Safety tests  of  plutonium fueled devices  for medical applications were
presented by Warner70 in  1972.  Results  demonstrate that plutonium devices
are  suitable for widespread  implantable  medical applications.  The current
status of radioisotope-powered artificial organs was given in 1972 by
Toyoshima and  Hart71.   The basic problems of  energy sources for implant-
type organs and the status of  high reliability long-life (10 years) 238Pu
power sources  were discussed.

     The use of radioactive  material as  a power source implanted in the
human body such as 238Pu  batteries to drive cardiac pacemakers, becomes
more important.  In most  cases the amount of  radioactive material implanted
in the body is considerable  and the  nuclide hazardous  (Hunzinger72).
Bearers of isotopic batteries  are not necessarily old people bound to
hospital beds.  They  include all ages and persons in their full professional
activity.  A bearer of  an isotopic battery can hardly be confined to a
controlled zone, the  establishment of which is a well known principle of
control in radiation  protection, nor can the  person be treated like a
consignment of radioactive material  and  be subject to the well established
test procedures for these packages.  The safety principles of isotopic
sources implanted  in  a  freely  moving human are outlined in this paper.
The  focus is on the exposure of other people  to penetrating radiation,
on partial loss of the  radioactive material,  i.e., exposure to contamina-
tion and the total loss of control over  the source.  Safety in industrial
fabrication of implantable devices powered by  radioisotopes was discussed
by Matheson73  in 1971.
            TRANSPORT  OF  PLUTONIUM FROM  SOURCES TO BIOSPHERE

 INTRODUCTION

      The presence  of  free  plutonium in  the atmosphere, ground and water can
 lead  to direct  contamination  by  inhalation or through ecological cycles.
 Understanding the  transport mechanisms  for the various physical and chemical
 forms of plutonium from  the contaminating source to the biosphere is compli-
 cated by the nature of plutonium chemistry.  The fate of the released pluto-
 nium  is uncertain  and complex.

      Studies have  been conducted by several investigators to determine the
 problems associated with plutonium cycling in the environment and reported
 in 1973 by  Carfagno and  Westendorf7\   These authors discuss the research
 being done  to determine  the mobility of radionuclides in major segments of
 terrestrial  environments.  Reciprocal transfers of environmental radio-
 activity are investigated  for three major components—soil, plant, and
 animal.

      The environmental levels of plutonium at U.S. Atomic Energy Commission
 installations were  summarized in a report by the U.S. Environmental Protection
Agency75 in  1973.   The sources and activities of plutonium found in environ-
mental samples are  given.
                                      13

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      The behavior of plutonium in ecosystems  and  its biological effects on
 natural fauna and flora as applied to desert  ecosystems at the Nevada Test
 Site were presented by Romney and Davis76  in  a report  published in  1971,

      The environmental monitoring program  for a nuclear fuel  reprocessing
 plant (located in the South Carolina coastal  plain), as based  on a man-
 environment ecosystem concept, was described by Platt   et al. in 1973.  The
 principal pathways to man—atmospheric,  terrestrial, and aquatic—are each
 subdivided into natural, recreational and  domestic  components.  The use of
 ecologically defined samples provides rates of movement and bioaccumulation
 that lead to ecological models that are transferable  to other nuclear
 industries.

      Research to provide data on the mobility and transport of plutonium
 in environments associated with nuclear  power production facilities of
 the humid eastern United States was outlined by Reichle78 et al. in  1974.

      A model for predicting the redistribution of particulate contaminants
 from soil surfaces was proposed by Travis79 in 1975.   The usefulness of
 this predictive tool is demonstrated by  calculations involving mixture
 of particulate 238Pu02 in highly erodible  soils under  dust storm conditions.
 Time dependent surface concentration and breathing  zone exposure isopleths,
 evolving from a small contaminated area, show the potential hazard  from wind
 eroding toxic materials.

      Wind distribution of plutonium released  by the fires at  Rocky  Flats
 was discussed by Poet and Martell80 in 1974.

 CHEMICAL AND PHYSICAL STATES OF ENVIRONMENTAL PLUTONIUM

      Since the chemical and physical states of plutonium determine  its
 transport throughout ecosystems, these states will  be  discussed separately.

 Soil

      Soil chemistry mechanisms partially control  the biotic availability  of
 plutonium over long time periods.   The chemical form of radionuclides  in
 soil in the vicinity of Oak Ridge National Laboratory  and the character of
 secondary reaction products and the availability  of nuclides  to plants
 after weathering under natural environments was included in the report by
 Carfagno  and Westendorf7>t in 1973.

      Miner81'82  et al.  in 1972 and 1973  give  the  results of studies on the
 chemical  behavior of plutonium in soil-water  environments.  Plutonium
 dioxide and  soluble plutonium were shaken  with a  soil-water mixture and
 the plutonium in  the various phases was  measured.

      Isotopic  ratios and chemical constituents of plutonium in  soils and
sediments of  the  Argonne National Laboratory  were listed in 1974  by Sedlet83
et al.

     The chemical  nature of  waste solutions discharged to the soil  of  the
Hanford facility  and a  study of their present form  were reported  in 1973

                                       14

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by Swanson81*.  The leach behavior of plutonium in these soils was also
investigated.

     As stated by Krey85 in 1974 and Poet and Kartell86 et al., the knowledge
of the chemistry of fallout plutonium in soil is not very complete.  They
tabulate the mobility of plutonium in soil from Rocky Flats,  Rhodes87'88
in two articles published in 1957, pointed out the polymerization known to
occur in soil.  The slow rate of diffusion and leaching of plutonium by
groundwater at the Hanford facility was reported by Hajek89 in 1966.  Self-
diffusion of plutonium  in soils will only occur due to migration of molecular
size particles through  soil solutions.  He concluded that plutonium could be
expected to move about  85 mm in 2.4 X 105 years by this process.  This was
predicted by using soils moistened with 20% water content by volume.  This
is a high water content when considering desert environments.

     The rate of transport of plutonium out of a waste burial area by
migrating soil moisture is dependent on many factors.  The chemical forms
of the isotopes and their respective solubility in water must be determined.
The amount of dissolution of plutonium from waste is strongly affected by
the chemical form of plutonium present, and by the chemistry of the leaching
solution.  In general,  soluble salts increase the solution rates, as do
organic complexing agents  [Los Alamos Scientific Laboratory (LASL)90 in 1974].
In 1974, LASL9"and Patterson91 et al. commented that plutonium leaching
from soil appears to be more affected by solubility than by exchange
reactions.

     Plutonium absorption by soils is equally complex, being strongly
affected by pH, as well as the concentrations of other metallic ions.
Plutonium appears to be present as a polymer in most solutions and is
highly sorbed by natural materials.  In general, it behaves as a cation
at pH values up to about 12, at which point it exhibits strong anionic
properties.

     Dissolution and absorption of plutonium can be quantitatively described
in terms of a distribution coefficient  (ratio of material concentration on
solid phase to concentration in liquid phase, Kd), as in Christenson92 et al.
in 1958.  A table showing solution percentage and distribution coefficients
for plutonium in waste  and soil was given in the LASL90 report.

     The distribution and characterization of plutonium in soils from the
Nevada Test Site were reported by Reichle78 et al. in 1974.  The leach-
ability of plutonium from soils containing different particle sizes was
also given.

     The physical aspects of plutonium in soils affect its solubility and
transport by resuspension.  Plutonium resuspension from soil is the subject
of reports by Michels93, Sehmel and Lloyd9"*'95.  These reports cover resuspen-
sion of plutonium from  the vicinity of Rocky Flats  and investigate effects
of particle size and wind speed.

     The evaluation of  the resuspendable amounts of plutonium-238  in soil
from the vicinity of the Mound Laboratory at Miamisburg, Ohio, was  given
in summary reports by Carfagno and Westendorf96,97,98 in 1972 and  1973.

                                      15

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 Results of analysis of  silt  samples  from the  Great Miami River were also
 given.

 Air

      The importance of  knowing the chemical and  physical forms and the uncer-
 tainties involved in estimating inhalation exposures  from  the 239Pu contami-
 nation at Rocky Flats were reviewed by  Boss"  in  1972.

      In 1972 Volchok100 et al. investigated the  plutonium  in the neighborhood
 of Rocky Flats as pertains to airborne respirable particles.  Experiments
 were carried out to determine the respirable  fraction of the resuspended
 239Pu.  The results indicate that in this area,  with  concentration in excess
 of 1 fCi/m3, the respirable fraction is about 0.2 to  0.4.

      In 1972 Fish101 et al.  discussed  experimental data and analytical models
 to be considered in assessing the transport of plutonium aerosols following a
 hypothetical reactor accident.  Behavior of released  airborne materials
 within the reactor containment system, as well as in  the atmosphere near
 the reactor site boundaries, were semiquantitatively  predicted from experi-
 mental data and analytical models.

      The results of a simulated airborne release of uranium, representing
 plutonium, during the burning of contaminated waste were discussed by
 Mishima and Schwendiman102 in 1973.  A source term containing both a
 fractional release and aerodynamic size distribution  is required to evaluate
 the potential downwind hazard from a postulated  inadvertent release.  The
 approach taken in this  study of a potential accident  situation in a radio-
 chemical processing plant was to measure the  fractional airborne release
 and aerodynamic size distribution released using a simple  but realistic
 situation.  The release of uranium  (as a stand-in for plutonium) was mea-
 sured during the burning of flammable, solid  materials containing either
 uranium dioxide powder or uranium nitrate solution.   The flammable materials
 used were representative of the typical types of solid waste generated by
 such a process and were packaged in  a  typical manner.

      In 1972 Mishima and Schwendiman103 characterized radioactive particles
 in a plutonium processing plant exhaust system.  Filter and cascade impactor
 samples were taken of the stack gases  and various exhaust  streams.  The  aero-
 dynamic characteristics, the amount  and distribution  of particles and  their
 associated radioactivity were measured.  The general conclusion that was
 reached was that  the plutonium present appeared  to be attached to large,
 nonactive  particles.

     A  series  of  studies to  determine  the fraction of plutonium made  airborne
 and characteristics  of  aerosols produced by overheating plutonium metal  and
 several plutonium compounds  was undertaken by Schwendiman!°"* et al. in 1958.
 Plutonium  in various  phases  of processing and use commonly appears  in  three
 forms—plutonium  metal,  plutonium compounds as powders, and liquids contain-
 ing soluble plutonium compounds.  All  three forms can be found in various
atmospheres and configurations.   Of  the materials studied, plutonium
compounds  in the  form of powders  released the largest amounts of plutonium
aerosol.   The  chemical  compound,  temperature  and airstream velocity obviously

                                        16

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had a great effect on the percentage released.  The highest rate of release
was from partially oxidized plutonium oxalate, 0.82 wt % per hour at 1000  C
in 100 cfm per second air.  Plutonium fluoride release rates were the lowest
for the compounds considered, 0.007 to 0.05 wt % per hour.

     Overheating plutonium metal created less airborne material than its
compounds.  The size distribution of the oxide produced can vary with the
oxidation conditions and, since the oxide is friable, the mechanical
processes to which it is subjected.  The median mass diameter of the
particles airborne during one of the experiments was 4.2 micrometers.

     Heating liquids containing soluble plutonium compounds released the
smallest amounts of material.  Using slow heating rates, the maximum amount
airborne from a concentrated plutonium nitrate solution over a 2-hour period
was 0.03 wt %.  When a  greater heating rate and greater volume were used,
the maximum amount airborne was 0.18 wt % during a 63-minute sampling period
at a full rolling boil.  Tables are presented in the report giving the results
of the study in greater detail.

     In 1973 Clarke105  devised a model for determining dispersion of effluent
from stacks of nuclear  installations and evaluated the inhalation doses.

     Plutonium isotopes 239Pu, 2I>0Pu and 238Pu have been injected into the
stratosphere as a result of atmospheric nuclear weapons tests and have
reached the ground as particulate fallout.  A satellite bearing a Systems
for Nuclear Auxilliary  Power generator (SNAP-9A), containing 17 kCi or 1 kg
of.238Pu reentered the  atmosphere in the southern hemisphere in 1969.  The
resulting burn-up during reentry turned the 238Pu into small particles at
an altitude of about 50 km.  The global inventory and distribution of fall-
out plutonium from these and other sources were compiled by Hardy106 et al.
in 1973.  The surface air concentrations and characteristics of the SNAP-9A
burnup in the vicinity  of Richland, Washington, were also reported in 1973
by Thomas107.

     A mathematical model describing the behavior of particulate Plutonium
in the lower atmosphere was proposed by Travis108 in 1973.  The model accounts
for particles settling  or impinging on the earth's surface as well as particle
re-entrainment or resuspension.

     The radioactive fallout rates and mechanisms describing the behavior
and characteristics of  radioactive debris from Chinese nuclear weapons
tests were reported in  1973 by Thomas109 et al.

     The nature and distribution of radiation to be expected in the strato-
sphere, including that  resulting from nuclear tests, contributions from
plutonium released on burnings of SNAP generators, galactic rays and solar
proton flares were included in a report by Webb110.

Water

     The dissolution of 238Pu in environmental systems was the subject  of  a
report by Patterson91 et al.  He points out that some experimenters have

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 erroneously assumed that  the  problem of plutonium transport is connected
 with the "solubility"  of  PuOa, whereas the real problem is the rate of
 dissolution.   Plutonium dioxide  reacts chemically with aqueous solution to
 release plutonium ions.   These ions, in turn, will react quickly with
 other agents in the solution  to  form soluble complexes or will react with
 the water itself to form  insoluble  suspensions.  The term "solubility"
 does not apply in the  thermodynamic sense to the plutonium dioxide water
 system.  Because the dioxide  cannot be formed in the presence of water, no
 equilibrium can be achieved between the dioxide  and the plutonium ions in
 solution.  The dissolution reaction for Pu02 is irreversible, the dioxide
 continuously goes into solution.  In a static system a state of pseudo-
 equilibrium may be reached, with the plutonium in solution remaining
 constant.  The concentration  of  ionic species of plutonium is controlled
 by the solubility product of  the hydrous oxides that are precipitated from
 solution according to  Polzer111  in  1971.  The rate of dissolution of the
 dioxide is independent of the concentration of the plutonium ions in
 solution.

      As stated in 1974 by Patterson91 et al., the definition of plutonium in
 solution is not well defined.  The  "solution" may contain fine particles of
 undissolved oxide, precipitated  hydrous oxide, polymers in colloidal form,
 and various complex ions  of plutonium.

      As shown by Polzer111 and Silver112 in 1971, the rate of dissolution
 of PuC>2 is dependent on many  factors, including pH, temperature, oxidizing,
 reducing, or complexing agents and  surface areas of the oxide (particle
 size and shape).

      The history of the sample is also important , including  the  duration  and
 temperature of heat treatment, time, temperature, and radiation history
 since treatment.  According to Kapshukov113 et al. in 1971, the high specific
 alpha activity affects several of these factors, 238Pu(>2 might be expected to
 dissolve more rapidly  than 239Pu02. The self-irradiation of the 238Pu02
 causes damage to the crystal  lattice, but this damage may be annealed out
 by heat treatment.   It was found that the lower the temperature at storage,
 the greater the retention of  damage.

      In 1974  Patterson91  et al.  concluded that Pu02 initially rapidly dissolves
 or reacts when first placed in contact with a neutral aqueous medium.  The rate
 may vary from sample to sample under nominally the same conditions but is
 generally greater than 100 ng/m2s for 238Pu and on the order of magnitude
 of 1 ng/m s for    Pu.  After  a few  hours the rate decreases  to a value that
 generally remains constant, until the concentration of alpha activity becomes
 high enough to  initiate competing reactions, such as coagulation of  suspended
material  or precipitation from a colloid.  The change in rate of dissolution
after a  few hours is unknown  but in 1969 Kubose111* et al. speculated that  a
hydrous  coating  may form,  protecting the material from further rapid dissolu-
tion.

     In an electrochromatographic experiment, Lingren115 discovered  in 1968
that the  dissolution of high-fired  Pu02 in seawater yielded negatively and
positively charged  carbonate  complex ions as well as neutral colloids.  The

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latter may be in the form of very small hydrous oxides as stated by Lloyd and
Haire116 in 1973.

     The dissolution of airborne plutonium in water has been measured in
a scrubber in parallel with an air filter by Hayden117 in 1972.

     Some investigations have been done on the dissolution of 238Pu02 micro-
spheres in water (Hudson118 in 1968; Adams and Fowler119 in 1974).  In 1973
Raabe120 et al. reported on in vitro solubility of respirable particles of
238Pu and 239Pu.

     A report by Curtis and Bentz121 in 1972 discusses the chemistry of
plutonium in natural waters.  The assumption that the form of plutonium in
natural water would be that of the tetravalent state is questioned.  Acci-
dentally released "soluble" plutonium is alleged to form plutonium-polymers,
or becomes solubilized ionic trivalent plutonium.

     Nature provides an abundance of organic materials which may act as
reducing agents toward an oxidizing material such as the Pu + 4 ion, so that
were the Pu(IV) polymer to depolymerize, it might yield trivalent plutonium
ions.  On the other hand, plutonium dioxide falling into ocean waters may
spend years dissolving.  As radioactive decay and natural causes slowly
degrade the oxide, plutonium may be released to the ocean water very slowly
and at such low concentrations that polymer formation may not be possible.
Several things may occur to the released soluble plutonium:  it may become
attached to a bit of sediment or organic material and remain so attached in
the tetravalent state; it may be reduced to Pu(III); or it may be oxidized
to a higher form.  A computer program for predicting the final form of
"soluble" plutonium in given conditions of acidity, potential, and complexa-
tion factors was proposed by Silver122 in 1971-

     In 1972 Dix and Dobry123 summarized some of the environmental phenomena
that modify plutonium reactions in water.  They listed the results of Metz
who placed plutonium sulfate solution in synthetic seawater and found that
it "disappeared" from solution with a half-life of about 40 hours.  Of the
plutonium recovered, 50% was found in a tan flocculent deposit on the bottom
of the tank, 20% was found on the walls of the tank below the surface and
about 30% was found on the walls above the water line which was exposed to
the spray from air bubbles resulting from circulation and aeration.  Metz's
experiments with a fresh specimen of 238Pu02 molybdenum cermet yielded surface
dissolution rates of 0.01 yCi/day per square millimeter of exposed surface
area (0.05 yg/day per g of Pu02) and plate out of an insoluble plutonium
product was observed on .,the,container surface.

     In 1972 Langham124 studied the fate of Pu02 following the Thule accident.
He found that particles agglomerated on inactive debris and only about 1% was
suspended as very fine particulates in water derived from melting ice.  In
1966 Lai and Goya125 found that plutonium metal chips placed in seawater
reacted completely in about 10 hours, accompanied by the evolution of hydrogen
which served to continuously expose fresh metal surfaces until the reaction
went to completion.  The products of the reaction were small particles of black
material (Pu02) that settled to the bottom and a white or pale green gelatinous
product [Pu(OH)iJ of low density which was suspended in the water.   The

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 solubility of the above products appeared quite  low.  As stated by Taube126
 in 1964 the equilibrium constant for hydrated  plutonium oxide in distilled
 water is 7 X 1(T56.

      In 1971 Aarkrog127 re-examined the area of  the accidental  release of
 plutonium at Thule,  Greenland.   The investigation  included  samples of  sea-
 water, bottom sediments, bottom animals,  zooplankton, fish,  seabirds,  seals,
 seaplants, and lichen.   The results showed that  the levels  had  decreased
 but plutonium was still present in concentrations  significantly above  the
 fallout background.   Samples of bottom sediments showed a vertical movement
 of plutonium in the sediments down to a depth  of at least 10 cm.  Bivales
 as far as 30 kilometers from the point of impact contained  plutonium from
 the accident.

      In 1968 Kubose128  et al. found that when  238Pu02 microspheres,  100 ym
 in diameter, were implanted in situ in the Pacific Ocean on the surface of
 bottom sedimets they became encrusted as a result  of biochemical action.
 The microspheres were clearly visible in each  aggregate and appeared to be
 little altered by submergence after 5 months in  the sea.  The aggregates
 were composed of sand grains and organic detritus, which was lightly cemented
 around the microsphere  by a dense matrix of mucilage and small  diatoms.   This
 phenomena lowered the rate of dissolution of 238PuOa markedly.  Observations
 were also made on microspheres buried within bottom sediments.  The plutonium
 adsorbed to the sediments and remained localized at the site of the  micro-
 sphere.

      The particulate size and physicochemical  state of plutonium in  the
 Bikini Atoll Lagoon is  reported by Nevissi and Schell129 in 1975.  They
 found that 16 years following the last nuclear test on the  Atoll, the
 plutonium was neither buried totally in the lagoon  sediments nor has  it been
 discharged completely to the ocean.  Small particles and/or  ions are  released
 at the sediment-water interface and are transported by the  currents.   These
 particles and ions agglomerate during transport  away from the sediment surface.

      In 1972 an evaluation of the potential for  concentration and transfer of
 plutonium in Lake Michigan was done by Wahgren and Nelson130.   The present
 levels,  distribution and biogeochemical behavior of fallout plutonium  are
 presented.

 BIOLOGICAL ASPECTS OF THE PHYSICOCHEMICAL COMPLEXITIES OF PLUTONIUM

     A detailed summary of the chemical and physical properties of plutonium
was given  by Taylor131  in 1973.   Saenz and Ramos1  also summarize  the physical
and chemical aspects of plutonium as applied to  physiological interactions
among biological  systems.   A brief summary of  the  most important  biological
aspects of  the  chemistry of plutonium in this  review concerned  its  states of
oxidation,  its  tendency to hydrolyze, and its  capacity to  form  complexes.  The
polyvalence  shown  by plutonium from Pu(III) to Pu(VII), and the proximity of
its oxidation potentials permit  equilibrium states in solution  among its  ions
of different valences.   This coexistence  brings  about a very complex chemistry
of this nuclide in its  biological behavior.
                                        20

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Mammals

     If plutonium in ionic form enters the mammalian body, three main types
of reaction may be expected to occur.

Hydrolysis to yield polymeric or colloidal species - Plutonium can hydrolyze
to insoluble hydroxide, passing through intermediate states of polymeriza-
tion with formation of colloidal aggregates.  This phenomena can occur at
physiological pH, qualitatively and quantitatively influencing its distri-
bution and deposit in the various  systems, organs, and tissues.

Complexing by proteins or other biological macromolecules - Plutonium deposited
in a localized area will complex with structural proteins.  Particulate forms
may be removed by phagocytic action.

Complexing or chelation with small molecular weight components of animal cells
and tissues - Plutonium complexes with anions of mineral acids (nitrates,
carbonates, etc.) and of organic acids (citrates, proteinates).  Stability
depends on the complexing bonders.  Bonders of great chemical stability can
form complexes which dissociate at physiological pH.  Other nonbiodegradable
bonders form biologically stable complexes.  Thus, the interaction of plutonium
with a broad variety of biological bonders produces simultaneous mechanisms of
dissociation and formation of new  complexes which influence retention, excre-
tion and elimination therapy.

     Plutonium compounds occurring as airborne contaminants are either very
insoluble in body fluids, e.g., PuOa, or are relatively soluble and trans-
portable in the body, e.g., plutonium nitrate.

     Insoluble plutonium translocates from the respiratory tract very slowly,
over many years, and this  imparts most of the emitted radiation energy to
lungs and associated tissues such  as thoracic lymph nodes.  Insoluble pluto-
nium inhaled as particles normally remains in the particulate form in the
body for a relatively long time.   The size of the particles determines the
deposition and mobilization in the respiratory tract.  The transportable
compounds inhaled are diffused in  the blood and organic fluids, and are
transported to be deposited in organic tissues or excreted in the urine.

     Of the nontransportable compounds, a fraction of that deposited in
the upper respiratory channels is  exhaled and another part is moved by
ciliary action toward the esophagus and gastrointestinal tract.  The remain-
ing fraction which is deposited in the lower portion of the lungs is either
localized or is slowly moved either toward the gastrointestinal tract and
eliminated in the feces, or toward the blood and lymph by solubilization of
the particles or by transport through the pulmonary epithelium.  The
mechanics of elimination of radioactive particles from the respiratory
tract are basically ciliary action, phagocytoses and transport to the
circulatory system (blood and lymph).

     Saenz and Ramos1 presented the kinetics of movement of plutonium parti-
cles in the respiratory tract and Bair132 et al. in 1973 summarized the depo-
sition arid translocation of inhaled plutonium in great detail.


                                      21

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      Diffusion of plutonium in the blood and interaction with proteins
 discussed by Saenz and Ramos1 and Taylor131.   Plutonium reaching  the blood
 may be absorbed and excreted in the urine without  change,  if the  compound
 is stable; disappear in a few minutes to be  mobilized by the liver, if
 strongly hydrolyzed; or form complexes with  proteins  if the compound is
 unstable.  Approximately 90% of serum protein is protein bound, mainlv^to
 the beta globulin (transferrin) to form a very stable complex  (Palmer    in
 1956 and Belliayer131* in 1959).  The interactions  of  plutonium with protein
 are influenced by the physicochemical characteristics  of the plutonium com-
 pound that reaches the blood.

      Many authors have studied the interactions of Pu(IV)  and serum proteins
 to establish that transferrin is the major plutonium  binding protein in  man
 [(Stover135 et al. in 1968; Bruenger135 et al.  in  1970); dog  (Stevens137
 et al. in 1968; Muntz and Barren138 in 1947); horse (Turner and Taylor139  in
 1968); rabbit (Taylor1"*0 in 1969); and rat  (Boocock and Popplewell1 "*l in
 1965 and Turner and Taylor1"*2)].

      As summarized by Taylor131, transferrin is a  glycoprotein with a molec-
 ular weight of about 90,000 in which aspartic acid, glutamic acid, lysine,
 leucine and alanine are the most abundant amino acids.  The transferrins of
 all the species studied bind 2-gram atoms of iron  to  form  a complex which
 dissociates in acid solution but is stable up to pH 9 to 10 as reported
 by Feenez and Komatsu1"13 in 1966.  The effect of pH on  the binding of
 Pu(IV) to transferrin was studied by Chipperfield  and Taylor1"*1* in 1970,
 who showed that maximum binding occurred at  pH 7.   The  mechanisms of Pu(IV)
 binding by transferrin are not fully understood.   In  1968  Stover135  et  al.
 and Turner and Taylor139, on studying dog and horse sera,  concluded  the
 bicarbonate ions were required for the binding of  Pu(IV) to transferrin  in
 a similar manner to their requirement in the binding  of iron  (Feenez  and
 Komatsu143).

      Chipperfield and Taylor11*1* question the requirement for bicarbonate
 in the binding of Pu(IV) to transferrin. Plutonium can be displaced  from
 its combination with transferrin by the addition of iron;  little  plutonium
 is found bound to iron-saturated transferrin (Popplewell and  Boocock1"*5  in
 1968;  Stevens137 et al.; Turner and Taylor139; Massey and  Lafuma1"*6  in  1968).
 Under  their experimental conditions, Chipperfield  and Taylor11*1*  found that
 the binding of iron and plutonium to transferrin was  similar  in  that  both
 increased up  to  pH 7;  above pH 7 the Pu(IV)  transferrin complex was  less
 stable  than the  Fe(III) complex, but this may be a reflection  of  the greater
 tendency  of plutonium to hydrolyze.  The results of these  studies indicate
 that Pu(IV) can  bind at the iron binding sites in  transferrin  but can also
 bind at other  sites  in the molecule.

     The  stability of  the Pu(IV) transferrin complex  has not  been determined.
 Stover135 et al.  concluded it had a high stability but  less  so than Fe(III)
 transferrin.   Popplewell and Boocock11*5 showed that Pu(IV) can be displaced
by  citrate  or  DTPA.

     A small proportion of plutonium may be  associated  with  albumin and with
the gamma and  other  globulins (Turner and Taylor139;  Stevens137  et al.;  and
Popplewell  and Boocock! "*5) .

                                       22

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     As summarized by Saenz and Ramos1 the distribution from the blood and
deposit in the various organs and tiss,ues are determined by the stability
of the plutonium compounds which have passed to the blood or which have
been formed by hydrolysis at physiological pH.  In the case of heavily
complexed compounds, deposit takes place preferentially in the bone at a
proportion of about 80% to that in liver.  If the compound has been hydro-
lyzed, the deposit takes place preferentially in the liver, at a proportion
of 7Q to 80% to that in bone.

     Deposit of plutonium takes place on the surface of the bone as found by
Stover and Eyring11*7 in 1970, mainly in the cells of the endostium, and to a
lesser degree on the periostium.  The selectivity for the cells of the endo-
stium is associated with the presence in those cells of a .glyeoprotein,
sialoprotein, to whose carboxylic groups plutonium is joined to form a very
stable complex  (Chipperfield and Taylor11*1* and Herring11*8 et al. in 1962).
This incorporation includes mechanisms for the dissociation of the Pu-
transferrin complex at the level of the bone surface and diffusion of the
plutonium from  the blood, with the changes of pH derived from the presence
of specific organic acids of the bone (lactic, citric) mainly influencing
this process.

     Experiments with different species of animals and in different parts of
the skeleton in 1972 showed the nonhomogeneity in the distribution in the bone
in medullary spaces, and later, according to Jeell>9, at the base of the cranium.

     Plutonium  can be localized in the red bone marrow before being deposited
in the bone medullary spaces, and later, according to Jee11*9 and Erleksova150
in 1960, the plutonium mobilized from the bone deposits can be found either
recirculating in the blood, or in spaces in the bone marrow, or retained in
cells of the reticuloendothelial system.

     The accumulation and retention in the medullary macrophages seem  to be
associated with processes of phagocytosis and existence of the iron storage
protein, hemosiderin (Vaughn151 et al.).  Their conclusion is that plutonium
reaching the blood by any route of entry, if in monomeric form, is deposited
initially in the skeleton rather than in the liver; if in the polymeric form,
it is likely to be deposited in the liver rather than the skeleton though it
may subsequently translocate to the skeleton.  During their research of the
subject they concluded that little is known at present about the mechanisms
and the form in which plutonium translocates from the site of entry to the
blood stream, though experimental results suggest it is initially translocated
in the form of soluble complexes and may later take a more colloidal form.
Initially it can be expected to be deposited primarily in the skeleton rather
than in the liver.  Plutonium, from occupational and fallout exposure, reach-
ing the circulation in man after some years, has been assessed as redepositing
roughly half in bone and half in liver (Taylor152).

     The fractions of available plutonium which are deposited in the liver
and the characteristics of their distribution are a function of the physico-
chemical state of the compound and of the route of penetration  (Saenz and
Ramos1).  In general, low relations of liver/bone deposits indicate the
entrance in blood of complexed transportable compounds.  High values suggest
the existence of hydrolized plutonium.

                                        23

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      By intravenous injection of complexed  compounds in dogs and rabbits,
 Cochran153 et al.  observed in 1962  a diffuse  initial distribution in all
 liver structures and,  later,  an irregular redistribution and accumulation
 in cells of the endothelial network according to  the degree of polymeriza-
 tion reached in their  interaction with organic fluids.

      With easily hydrolyzable compounds, the  initial distribution is not
 homogeneous, and the plutonium aggregates are localized  in macrophages mainly
 in peripheral zones of the hepatic  lobes  (Lindenbaum151f  et al.  in 1957).

      Mechanisms of interchange between plutonium  and proteins  seem   to be
 involved in incorporation into the  liver, especially processes  of phago-
 cytosis in redistribution.

      In this respect,  histological  studies  of dog livers in vivo (Stover and
 Eyring147 in 1970) and in vitro (Bruenger136) show the existence of plutonium
 associated with ferritin in the liver. This  leads to the  supposition  that
 the incorporation takes place according to  the following reaction:

      Pu IV-transferrin + ferritin = transferrin + Pu IV-ferritin.

      The processes of  phagocytosis, on the  other  hand, facilitate the
 mobilization and interchange  of plutonium between the hepatic compartment
 and the Kupffer cells.

 Plants

      In the area of plant uptake of plutonium, the determining  factors are
 plant species and vigor, soil type  and chemical form or  solubility of  the
 deposited material (Romney and Davis155 in  1972 and Menzel156  in 1965).

      Plutonium is tenaciously held  by the soil, preventing plant incorpora-
 tion, particularly if  salts,  acids, detergents and organic compounds are
 absent (LASL37 in 1974).  Soil characteristics, such as  soil structure,
 organic content, pH and the amount  of clay  present, influence  the availabil-
 ity of plutonium to some extent but probably  no more than  a factor of  ten
 (Romney and Davis155).
                    BIOLOGICAL DEPOSITION AND EFFECTS

DEPOSITION AND  EFFECTS  OF  PU IN AQUATIC ENVIRONMENTS

     When plutonium is  released to  surface  or groundwaters it may be concen-
trated by and in plants and  in silt sediments and suspended materials in the
water.  The concentration  of plutonium in sediments or in aquatic organisms
frequently exceeds  their concentration in the surrounding water, often by
several orders  of magnitude  as reported by  Sayre157 et al. in 1963.  The
plutonium, thus concentrated,  may be released suddenly at some future time
in quantities far exceeding  their maximum*permissible concentrations.

     The available  data concerning  the dissemination of plutonium in the
aquatic environment were summarized by Noshkin158 in 1971.  The most studied

                                      24

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isotope has been 239Pu derived from worldwide fallout.  Essentially all the
published work has been concerned with levels in the marine environment where
Plutonium is found widespread among planktonic, pelagic and benthic organisms.
The concentrations are higher in organisms feeding on sediment or on surfaces
than in those drawing on the water itself.  There is some evidence that
Plutonium concentrations are increased in organisms of high trophic levels.
Bone and liver are major repositories for plutonium in marine vertebrates
while muscle tissue of both marine vertebrates and invertebrates contain
relatively lower concentrations.  In marine sediments, as in soils, plutonium
is more mobile than was originally expected.  Fallout 239Pu appears to contri-
bute more than fallout 90Sr or 137Cs to the artificial radiation exposure of
many marine species.

     The results of studies of the concentration of plutonium in marine
invertebrates of the North Atlantic Ocean and their ecological relationships
are presented by Noshkin159 et al.  Organisms from the near-shore environment
were selected to show effects  on plutonium uptake  of variations in feeding
habits, association with sediment, or with absorptive surfaces and of trophic
levels.  Of the invertebrates analyzed, the mussel clam, oyster and scallops
are filter feeders, subsisting on tiny organisms and organic detritus removed
from suspension.  Plutonium-239 concentrations in the mussel body average
51 dpm/100 kg, a level 300 times that in seawater.  The shell of the mussel
was found to have, on the average, 64% more plutonium than the same weight of
soft tissue.  Fallout plutonium concentration in the mussel has increased by
a factor of 2 in 6 years, as compared to results of Pillar160 et al,,
reported in 1964.  Deposition as recorded in Tokyo by Miyake161 et al. in
1970 has increased only by 25%.

     The 239Pu concentration in starfish collected as they were feeding on
the mussel beds, although different in each sample, was about four times
that found in the mussel upon which each was feeding  (Noshkin159).  The
marine worm, Nereis, contained the highest concentration found in marine
invertebrates.  This worm is a nonselective deposit feeder and ingests
quantities of surface sediment.  Concentrations in other invertebrates are
also given in the report by Noshkin159 et al.  Significant concentrations
of 2  Pu were also found in Sargasso weed.  This weed may cycle considerable
quantities of 239Pu to specific near-shore regions.

     In 1971 Wong162 et al. reported on the plutonium concentrations in
organisms of the Atlantic Ocean.  The results showed that 239Pu was concen-
trated in some tissues of each organism so far examined.  In fish, 239Pu
concentrations range from 0.2 to 140 dpm per 100 kg fresh weight; in benthic
invertebrates from 23 to 140 dpm per 100 kg; in plankton, from 130 to 340 dpm
per kg and in Sargasso weed, up to 1280 dpm per 100 kg.

     In 1971 Patin163 et al. reported on a series of experiments performed
to study the accumulation of 23 Pu by live and dead Misgurnus Fossilis
spawn.  The intensity of the accumulation was found to be related to the
phase of embryogenesis because of change in the membranes of the spawn.
The bulk of the plutonium was adsorbed on the membrane of live spawn.   In
dead spawn, the adsorption of plutonium was virtually irreversible.
                                     25

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      Wong16"*  et  al.  in  1970  reviewed data  from a number of studies of 239Pu
 uptake and retention by marine animals and sediments and the content of 239Pu
 in seawater from various  locations.  It was concluded that most of the plutonium
 released to the  near-shore marine environment concentrates in sand and silts of
 the sea floor and beaches.

      Hodge165 et al. in 1973 found  that repeated measurements of  23aPu in
 several organs of albacore tuna  suggest that the upper layers of  the North
 Pacific Ocean can retain  large fractions of this nuclide for periods of a decade
 or more.  During the period  from 1965 to 1971 the reported rate of input from
 fallout decreased by about one-half in 1 year.  During this same  period, Z39Pu
 concentrations in albacore liver decreased to one-half in about 3.5 years.

      In 1974 Adams and  Fowler166 placed goldfish in aquaria containing 238Pu02
 microspheres.  At intervals  up to  302 days the fish were analyzed and tables
 were presented to show  alpha activity in the gills, intestine, muscles, bones,
 and total body.   Results  are also  presented for radioactivity in  the aquaria
 water and for radioactivity  of the  shell and tissues of inhabitants of the
 aquaria after 185 days.

      Examples of variations  in bioconcentration and concentration factors for
 plutonium in aquatic ecosystems  are given  by Kneip and Lauer167 in 1973.  Pro-
 blems associated with making general statements regarding concentration factors
 are discussed.

      Two reports covering the years 1971-72 and 1973 present the  data of studies
 on the plutonium concentration along freshwater food chains of the Great Lakes
 (Ouchi168 et al. and Bowen and Noshkin169).  Results indicate that bottom
 feeding fish, e.g., shad, sucker,  carp, bullhead, and rum, had higher 239Pu
 concentrations than predators such  as perch or bass.

      Results of studies to determine the 239Pu levels in water, sediment, plank-
 ton and fish samples collected from Lake Michigan were reported by Marshall170
 et al. in 1972.

      Gromov and Spitsyn171 in 1974  studied the change in the physicochemical
 state of 239Pu due to its assimulation by  phytoplankton.  The stabilization
 of the physicochemical  form  of 239Pu alters the state of plutonium  in  the
 marine environment,  which, in turn, leads  to variation in the entrapment of
 this  element  by  bottom  sediment  and by suspended matter.

      Gromov and  Spitsyn172 also  studied the absorption of 239Pu by  a  culture  of
 single-cell green algae and  by a natural phytoplankton community.  It  was  found
 that  the natural community concentrated the plutonium to a  greater  extent  than
 the pure culture did.   At 15 to  17  days, the isotopes are released  into  the
 seawater in a physicochemical form  in equilibrium with the  oceanic  environment.

     Folsom173 et al. presented  data in 1972 on the  concentration of  238Pu,
 239Pu, 2"*°Pu  in  IAEA seaweed samples.

     In 1971  Wong171* et al.  found that seaweeds concentrate plutonium and that
seaweeds may  be  a sensitive  indicator for  the detection of  variations  of pluto-
nium concentration in the marine environment.

                                    26

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     Zlobin and Perlyuk175'176 in 1973, studied the photosynthesis and mecha-
nism of the action of cyanide on cell respiration and 239Pu accumulation by
marine algae.  The introduction of this respiratory inhibitor  into the
energy system of algae caused a response reaction in the form of a decrease
in plutonium accumulation by plant cells.  It is postulated that assimilation
of colloidal particles of plutonium occurs as a result of ultraphagocytosis.

     A study of plutonium and polonium inside giant brown algae is reported
by Wong177 et al. in 1972.  It was found that large variations occurred
between specific tissue types.  Sampling thin parts of brown algae or thin
outer layers should provide great sensitivity for detecting early changes
in the extremely small traces of plutonium that are now anticipated in the
sea from reactor effluents, nuclear fuel processing or fallout.

DEPOSITION, EFFECTS AND COUNTEKMEASURES OF PU IN SOIL, PLANT ENVIRONMENTS

Soil

     A great deal of study  has been done concerning the deposition, concen-
tration and mobility of plutonium in the soil of the Nevada Test Site (NTS).
Langham178 et al. in 1966 and Langham179 in 1968 assessed the problem of
plutonium contamination at  NTS and elsewhere and suggested that the hazard
to man was minimal as long  as particles deposited on vegetation and soil
remain deposited.  In 1970  Romney180 et al.  observed a slight downward move-
ment into undisturbed soil  profile at NTS during the 10 years following
deposition.  Although the solubility of plutonium fallout particles in the
presence of soil is poorly  understood, plutonium infiltration to a depth of
12 cm was attributed to the downward movement of high density particles and
not solubilization.  In 1970 Mork181 noted very similar downward movement in
the same sampling area 2 years post-detonation where plutonium was found to
be predominantly associated with >44 y particles.  Price182 in 1973 review-
ed the distribution and fate of plutonium in terrestrial ecosystems and iden-
tified areas needing further study.

     In 1973 Tamura183 et al., discussed studies characterizing plutonium-soil
interactions at NTS.  Samples were taken to provide adequate activity levels
to permit determination of  plutonium on different size and mineral fractions.
The soils were then subjected to extraction of the plutonium content by nitric
and hydrofluoric acid.

     The ecological aspects of plutonium dissemination in terrestrial environ-
ments at the NTS were covered in a report by Romney and Davis155 in 1972.
Emphasis was placed upon standardization of analytical methods, delineation of
contaminated areas, problems of resuspension and redistribution, food chain
transport and ecological effects.

     Anspaugh181* et al. , Eberhardt185 and Gilbert186 gave summaries of test
results and statistical analysis of soil plutonium studies at NTS.  Inventory
of 239Pu and 21*°Pu at various depths in surface soil samples was determined
and results of resuspension and redistribution studies were given.

     Soils of five areas located on the Test Range Complex at NTS  were charac-
terized and soil profiles prepared, identified, and classified by  Leavitt187
in 1974.
                                     27

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      The plutonium content  of  soil  at  Nagasaki, Japan, 24 years after detona-
 tion of a nuclear device, was  reported by  Sakanoue and Tsuji188 in 1971-  The
 results indicate that  plutonium contaminants  in soil  are not easily removed
 by natural forces.

      The literature regarding  effects  of the  liquid waste disposal operations
 at Hanford on the soil environment  was summarized by  Rautson18  in 1973.  The
 summary is divided into studies pertaining to:  the vadose  zone below a depth
 of 6.1 meters (lower vadose zone);  the saturated zone; and  the vadose  zone
 above 6.1 meters (upper vadose zone).  Migration of wastes  discharged  to  the
 lower vadose zone over the  past 20  years by the mechanisms  of  diffusion,
 leaching and particulate transport  is  discussed.

      A program to examine soil-actinide relationships in sediments from a
 disposal facility and characterization of  actinide-bearing  soils was covered
 in a report by Ames190 in 1974.  At least  two types of plutonium were  found
 by autoradiographic and microprobe  examination of core samples.  Plutonium
 particles (up to 10 urn in diameter  and 60  wt  % Pu02)  were the most conspicuous
 form.  The second form of plutonium occurred  in lesser concentration  (<0.4 wt %
 Pu02) but was found associated with silicate  hydrolysis.

      The isotopic analysis  of  soils in the vicinity of the  Lawrence Livermore
 Laboratory as measured in 1973 was  given by Silver191 et al. in 1974.

      Mass isotopic analysis of fallout in  the soils of north central and
 southeastern sections of Utah  collected in 1971 indicates that the NTS was
 the probable secondary source  according to a report by Hardy192 et al.  in
 1972.  Isotopic ratios and  composition of  the fallout in soil were given.

      Environmental monitoring  reports  by Reynolds Electrical and Engineering
 Company, Inc.,193 in 1969 and the National  Environmental Research Center19"*  in
 1973 gave the soil fallout  composition from testing activity as determined  in
 areas surrounding the NTS.   Plutonium-239  soil measurements, depth profiles
 and physical state were given  for the  Rocky Flats vicinity  by Poet and
 Martell86 in 1972.

      In 1974 Healy195  proposed an interim  standard for the  upper limit of
 concentration of plutonium  in  soils in inhabited areas.  Available information
 on possible  sources of exposure of  people  living in an area where  soils  are
 contaminated with plutonium was analyzed to derive estimates of intake.   An
 evaluation of protective guidelines was reported by Anspaugh184 et al.

      Bliss and Dunn196 reported in  1971 on the results of studies  underway to
 determine plutonium in soil from areas outside the NTS.  Plutonium was detected
 in  four locations and showed concentrations in the top 3 cm  of  soil that  were
 10  to 100 times  greater than the concentration of soils in  other areas.

     The  role  of  soil  microorganisms in the movement  of plutonium was  noted
by Au     in 1974.   The study was designed  to  determine the  ability  of micro-
organisms to absorb  plutonium,  to quantify the uptake and to determine the
microbial population of soils  of the NTS.
                                      28

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     The effects of soil heating by  239Pu on  field flora were studied by
Krivolutskii and Fedorova198  in 1973.  The results indicate a sharp decrease
in the number of soil organisms as a result of heating the medium with239Pu.

     The feasibility and environmental impact of cleaning up plutonium contam-
inated areas in Nevada were discussed by Wallace and Romney199 in 1974,  It
was felt correction of the damage done to the fragile environment during the
course of  decontamination may require greater efforts than the decontamination.
Alternate  procedures were discussed.

Plants

     In 1973 Francis200 found that a survey of literature associated with the
movement of plutonium in soil and uptake in plants reveals that a major portion
of the investigations pertains to soils developed under arid or semiarid
climates.

     Short-term studies summarized by Price182 in 1973 indicate less than 10%
plutonium  uptake by plants.   Fallout plutonium uptake by plants was studied
by Nishita2  l et al. in 1965  who reported that ladino clover uptake was very
low.  Another study by Selders202 et al. in 1955 found that only trace amounts
of alpha activity were detectable in tomato plants grown in fallout-contaminated
soil, and  no uptake was recorded for bean, barley or tumbleweed.  In 1948
Jacobson and Overstreet203 reported  that plutonium uptake by barley from
clay suspensions was greatest for tetravalent plutonium.  That the sorption
of tetravalent plutonium to root surfaces from culture solutions was linear
with respect to concentrations, whereas, leaf concentrations appeared to be
curvilinear, was also noted in 1955  by Rediske2"*0 et al.  The uptake into
shoot tissue by tumbleweed from solution cultures was slightly less than for
bean, barley or tomato.

     In 1971 Cummings and Bankert205 used pot culture to compare the uptake
by plants  of 238Pu from nine  different soils.  Plutonium uptake was found to
be 7 X 10~6% to 280 X 10~6% of plutonium applied.

     In 1963 Wilson and Cline206 compared the uptake of plutonium from soil
by barley  with that of tungsten and  lead.  Using a modified Neubauer technique,
it was found that uptake of Pu(IV) from an acid soil was greater than uptake
from alkaline or calcareous soils, and plutonium uptake was less than tungsten
or lead.   Cline207 in 1967 reported  the uptake of americium and plutonium from
two of the same soils and indicated  concentration factor (CF) values [CF =
yCi/g acceptor (e.g., plant)/yCi/g precursor  (e.g., soil)] of 0.003 for ameri-
cium uptake from either soil  and 0.0002 and 0.0001 for plutonium uptake from
acid and alkaline soil, respectively.

     Neubold and Mercer208 and Neubold209 reported that although plutonium
uptake by  perennial ryegrass  was low, it increased over the 2-year period
of study.  It'was detectable  in the  third harvest of the first growing season
for one of the three soils studied (an acid soil) and the fourth harvest
showed barely detectable uptake from all three soils.  Results from the next
growing season indicated that  uptake from the acid soil was nearly four times
greater than the previous year's maximum.  In 1970 Romney    et al. studied


                                      29

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 the uptake  of  fallout plutonlum by  clover over a long period under greenhouse
 conditions.  A substantially  greater  foliage uptake of plutonium, a seven-
 fold increase,  from a highly  contaminated soil occurred after 5 years of
 cropping as compared to the first year.  It was postulated  that  the increase
 in uptake was  due to either more perennial roots coming into contact with
 Plutonium particles as  the plants aged or that natural organic materials
 resulting from root tissue decay complexed with the plutonium and resulted
 in increased uptake.

      As chelates enhance plant uptake of heavy metals, the  influence of
 chelating agents on the uptake of plutonium was studied by  Hale  and Wallace
 in 1970. They compared the effects of chelating agents DTPA, EDDHA and Fe-
 EDDHA on the uptake of  heavy  metals from soil by bush beans.

      Romney212 et al.  reported results from studies of perennial vegetation
 growing in  Area 13 of  NTS.  Relatively uniform distributions of  239~21*0pu anci
 2ttlAm were  found among individual samples of the same plant species.  However,
 there were  considerable variations  in the contamination levels between differ-
 ent species, presumably from  superficial entrapment of resuspended particulate
 material.  Concentrations in  Eurotia  lanata were three to five times higher
 than in other  species  sampled from  the same study  site.  Additional studies
 by Romney213 et al. were reported in  1975.

      Price2lk  studied  tumbleweed and  cheatgrass uptake of plutonium and demon-
 strated that shoot uptake is  clearly  influenced by the chemical  form of the
 transuranic.  Test results indicate that organic acid complexes  of plutonium
 such as oxalate or citrate can increase plant uptake when added  to soil as
 compared to uptake from dilute nitric acid solutions.

      In 1972 Smith215  et al.  studied  the radionuclide content and botanical
 composition of the diet of beef animals grazing on the Area 18 range of the
 Nevada Test Site by analyzing rumen samples collected from  fistulated  steers.
 The radionuclide concentrations were  generally low with periodic increases
 in individual  isotope  levels  which  could be traced to a specific contaminating
 event.

      Klepper and Craig216 in  1973 report studies of the interactions of pluto-
 nium aerosols  with plant foliage.   The studies include exposure  of 2-week-old
 bean seedlings to 239Pu nitrate aerosols of 1 to 3 U size at wind  speeds  of
 16.4 m/min.

      When samples of foliose  and fruticose lichens were collected  from 27
 sampling sites  in the Thule,  Greenland, region following the crash of  a bomber
 carrying nuclear weapons aboard, Hanson26 found slightly higher  than back-
 ground amounts  of 239Pu and 21ft)Pu associated with  particles of estimated  0.5 to
 1.0  ym diameter.

      In  1969 Alvarez-Ranies and Santos217 studied  the contamination  of terres-
 trial gastropoda feeding on plants  exposed to the  plutonium aerosol  from the
 collision of planes  carrying  atomic weapons over Palomares, Spain.   Plutonium
values in soil,  plants  and the gastropoda flesh and shell were  tested and the
 interacting relationships noted.

                                      30

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ABSORPTION OF PLUTONIUM BY ANIMALS

     Several possible routes of entry exist by which terrestrial animals can
incorporate plutonium into their bodies.  Herbivores and small animals may
ingest radioactivity by foraging on  contaminated vegetation, drinking contami-
nated water or consuming radioactive soil while eating or grooming the fur.
Inhalation may also represent  an important avenue of intake for burrowing
animals and for animals living in an environment where considerable resuspen-
sion is evident.  Predators, however, would probably derive most of their
intake from the consumption of smaller prey animals which contain plutonium
in their internal organs or on their pelts.  Man enters into this terrestrial
food web by consuming the meat from  certain sport game and domestic grazing
animals and the milk from dairy herds  (LASL37).  Ingestion has been considered
to be the primary route of entry involved in the food chain transport of resid-
ual plutonium in the environment  (Olafsen and Larson218).

     Ershov219'220 et al. in 1970 and  1971 studied the penetration of 239Pu
through the epidermis during superficial contamination using piglets.  The
results, shown graphically, indicate that at the beginning, activity penetrat-
ing the skin at any fixed depths very  rapidly increased.  After 12 hours of
contact the amount taken up per unit time decreased and became balanced, i.e.,
the amount of the absorbed and expelled radioactive preparation was equal.
The kinetics of microdistribution in the skin and the dose distributions in
structural layers of skin are  also given.

     An early work by Finkle221 in 1945 considered the distribution of pluto-
nium in a dog 16 days after administration of a lethal dose of plutonium
nitrate.  The skeleton  contained 25% of the injected dose, liver 31%, muscle
8% and spleen 3.5%; 10% was excreted.  The plasma fraction of the blood con-
tained 80% of the plutonium found therein.  Studies with intravenous injec-
tion of plutonyl nitrate into  mice found concentration of 27% of the dose
after 64 days.  When plutonyl  citrate  was injected the liver content fell
from 35% to 14% on the  31st day and  to 7% on the 64th day showing a half-time
value of 20 days.  Mice receiving plutonyl nitrate intramuscularly had liver
retention that showed the same 20-day  half-time.  Plutonyl citrate injected
into the peritoneal cavity of  mice was slowly absorbed with a concentration
occurring in the femur and liver; plutonyl nitrate, intraperitoneally, behaved
similarly but was somewhat more slowly absorbed and therefore, was retained
longer by the liver.

     In 1948 Hamilton222 noted that  plutonium is not absorbed to any signifi-
cant degree by way of the digestive  tract.  The metabolism of plutonium follow-
ing intramuscular injection is essentially the same after administration  as
Pu  , Pu+tf and Pu+6.  He suggests that plutonium is converted by the body into
one valence state regardless of the  valence administered.

     Results of the biomedical program in conjunction with the  1957  "Operation
Plumbbob," were reported by Stannard13  in 1973.  The primary purpose  of  the
program was the direct determination of the uptake and retention of  plutonium
by a variety of animals as a function  of time of exposure and the relation
of the amounts found to air and ground levels.  There were two major phases
and groups of exposed animals—those exposed during cloud passage and  those


                                      31

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 exposed during long-term post-event periods.   Rats and dogs were used in the
 former; dogs, sheep and burros for the latter.   For the cloud passage phase,
 animals were placed from 164 to 656 meters downwind from Ground Zero.  For the
 resuspension phase the animals were placed in areas with levels of ground
 contamination of 2.6, 40 and 560 micrograms per square meter of desert floor
 soil.  The animals remained in place for periods ranging from 4 to 160 days.
 The plan was to measure the accumulation of plutonium in the lung and other
 tissues as a function of time and various aerosol parameters.

      The dogs exposed to the cloud passage at the time of detonation showed
 generally higher burdens than the animals exposed to resuspended plutonium,
 even though the former were not at the range of maximum airborne concentra-
 tions at ground level.  There was no appreciable increase in lung burden
 with time and very little difference in burden in animals placed at several
 different surface contamination areas.  This may have been difficult to
 ascertain as radiochemical analysis at that time lacked sensitivity for very
 low levels .

      Results reported from Operation Roller Coaster have not all been
 declassified at this time but what is available is included in papers by
 Wilson and Terry16'17 and Stewart223 et al.  The initial lung concentrations
 for dogs, sheep and burros were very similar despite wide differences in lung
 size, etc.  There was, however, a marked difference in clearance pattern among
 the three species.  The lung clearance for the dog closely resembles that
 found in laboratory studies with inhaled
      A review of the comparative metabolism of radionuclides in mammals was
 published by Stara22"* et al.  in 1971.   They note that the rate of absorption
 through the gastrointestinal  tract was low when 239Pu was given to rats.
 Katz and Weeks225 reported absorption  was 0.001-0.004%.   Plutonium-239 (N03K
 orally administered to rats was found  to be absorbed between 0.002 and 0.003%,
 with a maximum of 0.009%.   Carritt226  et al.,  demonstrated in 1947, that the
 amount absorbed is related to the dose;  with a lower dose, the absorption was
 0.30%, whereas with a dose approximately tenfold higher,  the absorption was
 decreased to 0.01%.  Absorption of 239Pu (N03)i, given orally to swine was
 reported as 0.002% by Weeks227 et al.  in 1956.  Inhaled  239Pu02 in dogs was
 absorbed in a range from 0.1  to 16.9%  with a mean value  of 3.7% as calculated
 by Morrow228 et al. in 1967.

      In 1972 Kashima229 et al.  conducted studies to clarify the relation between
 the  physicochemical form of plutonium  and its distribution in all organs and
 tissues after  subcutaneous injection in  mice.   The forms used were monomeric
 plutonium with pH adjusted to 1,  4.5,  7.2, and 13.0 immediately before injec-
 tion; and polymeric plutonium solution with its pH adjusted to 9.0.  After
 subcutaneous injection of  the monomeric  form,-239Pu remained mostly at the
 injected site  for the  56 days studied.   Concentration of plutonium in bone
 increased with  time, the level of plutonium in liver, especially around the
 central vein, was relatively  high in the initial 3 days, but was lower at 21
or 56 days.  The  distribution pattern  was only slightly  affected by changing
pH or time prior  to  injection after preparation.   With polymeric plutonium,
however, relatively  low plutonium concentrations were observed only in the'
bone and liver.
                                      32

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     Ballou    et al, in 1972 found that about 0,08% of an oral dose of 239Pu
citrate was retained in dogs after 3 days, distributed mainly between skeleton
(50%) and liver (25%).  After intravenous administration the liver retained
32 to 45%3
-------
 at  generally  comparable,  decreasing rates.  Polymeric plutonium was as much as
 10  to 30 times  more  concentrated in predominantly red (erythropoietic) marrow
 than in fatty marrow.  Monomeric plutonium was 1.3 to 2,8 times more concen-
 trated in the red marrow.  Mice injected with the same plutonium solution as
 the rabbits showed,  in comparison, a higher percent of the injected monomeric
 Plutonium in  the  femurs and a lower percent in the liver; they showed a higher
 uptake of polymeric  plutonium in the marrow and a higher uptake of both forms
 in  the spleen and lungs than did the rabbits.
             \                                                           9 3 Q
      In 1974  Zapol'skaya235 et al. intravenously injected male rats with    Pu
 chloride salt solution and determined the biological half-life in various
 tissues, organs and  albumin proteins.  The results indicated that the exchange
 of  plutonium is determined by the exchange of the albumins with which they
 form bonds.

      Durbin236  et al.  constructed a kinetic model to describe the transport
 and deposition  of intravenously injected Pu(IV) citrate in the rat.

      The implications  of  the solution of the model led to the following working
 hypothesis:  (a)  Unbound  plutonium reacts with protein, presumably transferrin,
 in  extracellular  fluid compartments as well as in plasma.  The Pu-transferrin
 complex is the  most  likely form in which orally administered plutonium is
 carried once  it reaches the plasma.   (b) Little, if any, protein bound plutonium
 is  excreted or  deposited  in the liver.  Formation of the Pu-transferrin complex
 is  probably not necessary preliminary to liver deposition of diffusible pluto-
 nium.   (c) Both unbound plutonium and bound plutonium are sources of plutonium
 deposited in  bone.   The surface of the reticulocyte (where iron is released
 from the Fe-transferrin complex) is considered the most likely site of dissoci-
 ation of the  Pu-transferrin complex.
                                                    j-.
      A good review of  the metabolism of monomeric and polymeric plutonium in
 small animals and the  radiotoxic implications for man was prepared in 1971
 by  Lindenbaum2 3 7.

      Matsuoka 38  et  al. compared autoradiograms obtained from animals admini-
 stered monomeric  or  polymeric plutonium.  Results were compared with those
 obtained from animals  administered metabolically stable reference particles
 of  known particle size.   It was found that the behavior of plutonium soon after
 injection is  largely influenced by its particulate character rather than by
 its  chemical  nature.   Results following inhalation of a plutonium nitrate
 aerosol  showed  no obvious  translocation to the liver, despite the water solubil-
 bility of the nitrate  form.  The initial uniform distribution of inhaled pluto-
 nium changed  to a nonuniform distribution after 3 months.

     Baxter and Sullivan239 studied the gastrointestinal absorption and reten-
 tion of plutonium chelates administered to rats.  They found the intestinal
absorption of plutonium nitrate is increased about 700-fold when chelated with
DTPA.  Within 2 days,  virtually all of the absorbed plutonium-DTPA complex  is
excreted  in urine.   Thus,  plutonium retention in the liver and skeleton  is
quite  low, but  more  than  twice as high as when DTPA is not present.   Citrate
is less effective than DTPA in increasing absorption, but more effective  in
increasing retention.
                                       34

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     In 1972 Mays^and Dougherty2"*° reported results of beagle studies,  The
initial distribution of plutonium was uniform throughout beagle liver follow-
ing intravenous; injection of monomeric plutonium in citrate solution.  Most
of the initial burden appeared in the hepatic cells.

     Subsequently, much of the activity shifted into the liver reticulo-
endothelial cells lining the sinusoids; the rate of translocation was great-
est at the higher dose levels, presumably  due to release of the nuclide from
cells killed by the irradiation.  At very  high doses, the skeletal burden may
actually increase due to the amount released from the liver.  At long times
after injection the nuclide distribution is very nonuniform within the liver,
being highest in the portal region and lowest in regenerative nodules that
grow after the time of injection.  In the  lower-level 239Pu beagles, the net
liver burden remains roughly constant at about 30% of the injected activity
for about the first 1,000 days after injection.  The loss of liver 239Pu during
this period is at least, in part, compensated by the influx of 239Pu released
from the skeleton.  After 1,000  days, the  liver burden at the lower levels
decreases with a half-time of about 8 years.

     Placental transfer of 239Pu in rats and mice has been reported to be
inversely related to the size of dose given to the dam,  (Finkel21*1 in 1947,
Wilkinson and Hoecker21*2 in 1953) with maximum transfer  of 3 to 8% of the
dose, when the dose was given about 1 week after conception.  Detectable
amounts of    Pu were found in milk of mice, rats and cats after administer-
                                                                      O it 0
ing plutonium to the lactating dam.  A more recent study by Ovcharenko    in
1971 showed that when pregnant rats were administered Pu-citrate intravenously
on the day of birth and then allowed to give birth and nurse their young, the
lactation did not significantly  reduce the content of plutonium in the organs
of the lactating mothers.  Sacrifice of the mother rats  after 1 and 10 days
indicated 84.4% and 96.4% of the administered activity was retained in the
body.  The content of 239Pu in the entire  litter of baby rats fed by treated
mothers from 1 to 10 days of age was 4.24  to 7.36% of the amount introduced.
After 20 to 30 days 1.44 to 2.32% remained in the bodies of the baby rats.

     In 1972 Sikov and Mahlum21*11 reviewed  the deposition and retention of pluto-
nium in immature, developing animals.  The absorption of 239Pu from the gastro-
intestinal tract of juvenile rats and dogs relative to absorption in adults was
shown graphically.  As would be  expected,  the graph shows enhanced uptake of
plutonium from the gut of the neonatal rat and dog is about 100-fold greater than
in the adult of these species, with a gradual decrease to adult values to about
the time of weaning.

     Ballou21*5 reported in 1958  that age appears to have a direct effect on
absorption in rats.  He noted 0.25% absorption of  239Pu  nitrate at 1 day of
age, decreasing to 0.10% at 7 to 13 days of age, 0.02% at weaning and 0.003%
in adult animals.

     Finkel21*6 in 1947 indicated that the  absorption of  plutonium bound to
protein, as in milk, may be as much as 20-fold greater than in the uncombined
form.  This might increase the hazard to juveniles whose diet is primarily
milk.  On the other hand, McClellan2"7 et  al. in 1962 reported there  are
discrimination processes, which  result in  the concentration of plutonium  in

                                      35

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 breast milk being substantially  lower than that in the diet or the plamsa of
 the mother.

      Sikov and Mahlum21"*  indicated that there is a possibility that there is
 greater hazard to the gastrointestinal tract from ingestion of insoluble
 Plutonium in the infant  than  in  the adult.  It has been shown that the gastro-
 intestinal passage time  of  particles in juvenile rats does not differ from
 that in adults, although the  average rate of passage is substantially reduced
 and there is evidence of prolonged retention in the stomach and ileum.  This
 results in an increase in the average time that the plutonium is in contact
 with any given segment of the tract, and a corresponding increase in the
 radiation dose in the young,  which is further increased because of the smaller
 organ diameters.

      Sikov and Mahlum2"*1* also found that at 1 day after injection into
 adult rats, a much greater  fraction of polymeric than of monomeric plutonium
 was localized in the liver.   In  the femur, the initial deposition of mono-
 meric plutonium was much higher  than that of polymeric.  In the newborn
 there was an equal fraction of the administered dose of monomer and polymer
 in the femur at 1 day after injection and the concentration of the two forms
 in the liver were more similar than in the adult.  With the passage of time,
 the monomer was removed  from  the liver of both age groups and the polymer was
 retained.   This retention was more marked in the adults than in newborns,
 while the initial redistribution of plutonium from liver to bone was more
 marked in the juvenile rat  than  in the adult.  As a result, at 30 days and
 again at 90 days, the patterns of partition of the two forms of plutonium
 were not markedly dissimilar  in  either the animals injected at birth or as
 adults.  Buldakov 21f 8 et  al. in 1970 found similar patterns in studies on
 lambs.

      The interpretation  of  data  on immature animals is complicated by the
 fact that the size of individual organs or organ systems relative to that of
 the total body changes during development.  Sikov and Mahlum21*1* summarized
 the changes in the size  of  various organs with age.  They also found that
 although the adult and newborn rat had a similar fraction of administered
 polymer in the liver at  1-day after ingestion, the corresponding concentra-
 tion in the liver of the  newborn was 20-fold that in the adult.  In the adult,
 the concentration of polymeric plutonium in the liver decreased by about half
 in the  first 90 days  after  injection; in the newborn, the dilution produced
 by growth  resulted in an  80-fold decrease in concentration.  At both ages
 there was  a net increase  in the  fraction of the dose in the femur.  This
 resulted in a slight  increase in concentration in the adult, while there
was  a 20-fold decrease in the newborn.  It is therefore evident that age
at  the  time  of injection  influences the total radiation dose as well as
the  temporal  distribution of  the dose.

     Incorporation  of  239Pu into the hair of rabbits was reported by
           2^9
Jaworowski     et  al.  in 1971.  They found that when doses of 0.1, 1.0 and
10.0 yCi of     Pu chloride, dissolved in 0.5% phenol, pH 5.5, were injected
subcutaneously into red-haired New Zealand rabbits, the mean incorporation
of  239Pu in the anagenetic hair  of the first fleece (grown 22 days after  dosing)
ranged  from 1.22  X  10" to 2.73 X 10"3% of the dose per gram of hair.  There
appeared to be  an inverse relationship of percent incorporation to  dose
amounts.  Twice as much was incorporated at the 0.1 yCi per animal,  as

                                      36

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at 1.0 and 10.0 yCi per animal.  In the growth of the second fleece, the
rate of incorporation was 10 times lower at 10,0 yCi than at the lower
doses.  At the dose levels studied, the amount in hair was similar to that
in muscle but much lower than bone or  liver.

     A recent report of the biological effects of plutonium in experimental
animals was presented by Bair    in 1975.


BIOLOGICAL BEHAVIOR OF INHALED PLUTONIUM IN ANIMALS

     Airborne plutonium particles are  similar to most other particles when
they are inhaled in that deposition in the respiratory tract is primarily
dependent upon the physical properties of the particles and the respiratory
characteristics of the individual inhaling the particles.  After deposition
in the respiratory tract, plutonium may be retained in the lung for a long
time, be translocated to other tissues in the body, accumulated in the lymph
nodes associated with the respiratory  tract or excreted in urine and feces.
The actual deposition 'of inhaled plutonium depends largely upon the physical
and chemical characteristics of the inhaled material (Bair251 and Lafuma252).

     Bair251 reviewed the literature in 1974 and presented figures and tables
showing lung clearance and translocation values of several investigators.  He
notes that within the first week after exposure, a fraction of the deposited
plutonium is cleared from the respiratory tract and excreted.  In the case of
PuOz inhaled by beagle dogs, all plutonium cleared from the respiratory
tract is excreted in feces except for  a small fraction that may be solubilized,
absorbed into the circulating blood, and either cleared by the kidneys or
deposited (by the blood) in another tissue.  The magnitude of the fraction
cleared within the first week depends  upon the fraction of readily soluble
material present and also upon the distribution of the deposited plutonium
within the respiratory tract.  Plutonium particles deposited on the ciliated
epithelium of the upper respiratory tract are trapped in mucus, propelled to
the esophagus and swallowed.  Plutonium deposited in the lower regions of
the lung, in the alveoli, is not available for clearance and may be incorpor-
ated into the cellular structure of the lung and retained for a long time.
The clearance of plutonium deposited in the lower lung is generally assumed
to be exponential over a reasonably long period of time.

     Bair251 gives retention half-times of several plutonium compounds.  The
retention half-times for organic complexes of plutonium, plutonium nitrate
and fluoride range from less than 100  days to about 300 days in rats and dogs.
The retention half-times for Pu02 are  substanially longer, ranging from  200
to 500 days in rats, and from 300 to 1000 days in dogs.  The wide range of
values observed in dogs is largely due to extensive experimentation with a
variety of plutonium oxides with different particle size characteristics.
Studies with 238Pu02 in dogs indicate  a much shorter lung retention time than
is observed for 239Pu02.  This appears to be due to instability of 238Pu02
particles, possibly caused by radiolysis in tissue fluids.

     In 1970 Bair253 presented figures showing the relationship of particle
size and isotope on retention of plutonium in lung as well as for oxides
prepared under different conditions.   Oxide prepared by calcining the  oxalate

                                     37

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 at 1000°  C was retained with  a half-time of 650 to 950 days compared with 300
 to 400 days for the oxalate calcined at 350° C.  Oxides prepared from metal
 powder at temperatures  of 123° C  to 450° C were retained in lung longer than
 the low-fired oxalate.   Small particle size high-fired oxide was retained with
 a half-time of 400 to 500 days, compared with half-times up to 900 days for
 the larger particle size  high-fired oxide.  Retention of the small particle
 size 238Pu02 was less than for the comparable 239Pu02.

      Plutonium may be removed from lung via bronchial and tracheal mucociliary
 processes and excreted  but this accounts for only part of the plutonium cleared
 from the  respiratory tract.   Another fraction is accumulated by the lymphatics
 and deposited in lymph  nodes.  Data from an 11-year study with beagle dogs were
 presented by Park25" et al. in 1972.  After 5 years both lung and thoracic lymph
 nodes each contained 30%  of the plutonium initially deposited in the lower
 respiratory tract.  After 11  years the lung burden had decreased to about 9%
 and the thoracic lymph  nodes  had  increased to 40%.  Translocation of plutonium
 from lung resulted in levels  in liver of about 10% of the alveolar-deposited
 plutonium, in bone of about 5% and in the abdominal lymph nodes of about 7%.

      The  relative distribution among body tissues of plutonium translocated
 from lung is essentially  the  same for all plutonium compounds.  In beagle
 dogs, within several months after  inhalation of plutonium nitrate, the fraction
 remaining in lung decreased to 40% or less of the amount deposited in the
 lower respiratory tract according to Ballou    et al. in 1972.

      Translocation from the lung  resulted in bone accumulating about 30% and
 liver about 10% of the  alveolar-deposited plutonium.  A small percentage was
 found in  spleen, lymph  nodes, and the soft tissues and the remainder was
 excreted  in urine and feces.

      Data presented by  Park255 et al. in 1973 showed that 238Pu02 may be
 cleared from lung more  rapidly than 239Pu02-  It has also been found that
 translocation of 239Pu  from lung  to other tissues in the body may be greater
 than for  238Pu.

      Distributions of plutonium in tissues of beagle dogs 5 years after inhala-
 tion of 238Pu02  and 239Pu02 are also compared by Park25  et al.  Both oxides
 were calcined at 350° C.   The particle size of 238Pu02 was 0.1 pm  (CMD) and of
 239Pu02 was 0.1  to 0.5  ym (CMD).  The ultrafilterability was 1 to 2% for the
 238Pu02 and <1%  for the 239Pu02.  After 5 years, only 10% of the body burden
 of the  238Pu was in lung  compared with 46% for   39Pu.  Accumulation in thoracic
 lymph nodes was  3 times greater for 239Pu than for  238Pu, however,  the bone
 burden  of 238Pu  was 12  times  that of 239Pu.  This illustrates  that  the behavior
 of  38Pu02  in the body  may differ significantly  from that of  239PU02.

     In 1972  Craig256 et  al.  exposed beagle dogs to  239Pu02 aerosols as part
 of a low-level effects  study  of the oxides of both  238Pu and  239Pu.  The alveo-
 lar deposition of 239Pu02 aerosols was measured  as a function of the aerodynamic
 particle  size distribution of the aerosol and the tidal volume of the dog during
 exposure.

     Matsuoka257 et  al.  studied the distribution of plutonium nitrate aerosol
by rats.  Rats sacrificed immediately after exposure showed deposition in the

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tracheo-bronchial and pulmonary region.  One day after exposure, plutonium
was mainly in the lung with very  small amounts in the intestine.  One month
after inhalation, the plutonium deposited in the lungs still remained, with
very small amounts translocated to  the trabecular bone of the vertebra and
to the ileum.  Isolated lung  specimens showed no deposition in the tracheo-
bronchial region and very  little  deposition in the pulmonary region.

     In 1972 Suzuki258 et  al.  also  exposed rats to submicron aerosols of
plutonium nitrate and determined  the  retention and excretion.  Excretion of
plutonium was mainly observed in  the  feces on the first  day  following inhala-
tion and the daily fecal excretion  decreased to 1/40 to  1/100 of the total
intake of plutonium on the third  day  and thereafter.  The urinary excretion
was less than 1/10 of that of the feces.

     Ballou259 et al. studied the deposition of inhaled  239Pu citrate in dogs
and found that 239Pu reached  a peak concentration in blood after 6 hours, then
decreased with clearance kinetics similar to intravenous administration.  One
day after exposure the lung,  skeleton, liver, blood and  intestines retained
most of the  inhaled  239Pu. After 100 days 36% of the initial deposit was
retained; 33% in the lung, 44% in the skeleton and 17% in the liver.  Although
the concentration in specific lymph nodes  (hepatic, splenic, and tracheo-
bronchial) was among the highest  of any  tissues analyzed, the total amount
deposited in a selection of 12 lymph  nodes  (~10 g tissue) was always less
than 0.6% of the body burden.

     Wandall260 reviewed the  mechanisms  involved in the  lung clearance of
inhaled 238Pu02 and  239Pu02 aerosols  in  dogs and rats and the subsequent
translocation of the solubilized  fraction throughout the body.

     Dionne  and Sanders261 found  that 239Pu02 particles  inhaled by rats were
deposited in alveoli and rapidly  phagocytized and concentrated within pulmonary
macrophages.  A hybrid computer alveolar model was developed in order to demon-
strate the distribution of alpha  energy  in the alveoli and the modifying
influence of pulmonary macrophages  on the alveolar alpha-energy from phagocy-
tized particles.

BONE DEPOSITION

     Atherton262 et  al. administered  239Pu(IV) intravenously to beagles in:
(1) the Pu-transferrin complex, (2) the 0.08 M  citrate buffer and (3)as a
suspension of near colloid size particles at pH ~6.  The skeletal retention
of plutonium, when injected in a  monomeric form, is 50%  of the  injected dose.
The retention of polymeric plutonium  in  the skeleton was less than  1/20 that
seen in monomeric or transferrin  complexed plutonium at  comparable  times.
The uniformity of skeletal distribution  of polymeric plutonium  from bone  to
bone is markedly less than that seen  with the two monomeric  forms.  Reduced
skeletal retention in animals injected with polymeric plutonium indicates  a
different skeletal retention  mechanism in animals injected with polymeric
plutonium.   The authors postulated  that  skeletal retention of the polymer is
not associated with bone per  se but likely increases with the amount  of red
marrow a bone contains.
                                       39

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      Stover263  et al.  reported the  retentions  of plutonium in the humerus and
 third lumbar vertebra, of dogs  injected with  0.016, 0,048, and 0.095 yCi/kg
 239Pu.   The dogs survived from 1600 to 5200  days after injection,  The data
 show that the rate of decrease of retention  is greater during the first  2
 years than at later times.   There appear  to  be two phases in the time depend-
 ence  of skeletal retention.   Two things  are happening during the first  years
 after injection.  Both the rates of the remodeling processes which remove
 plutonium from the bone surfaces and the  fraction of skeletal plutonium  that
 remains on the surfaces are decreasing.   Both  of these factors contribute to
 a decreasing loss of plutonium from the skeleton.  The remaining plutonium
 which has been buried during the remodeling  of the bone is much less access-
 ible and is lost from the skeleton  only at a very slow rate, if at all.

      Another 1972 report by Stover 26l( et  al. compared the skeletal and hepatic
 dose rates from intravenously  injected dogs.   Young adult dogs were administer-
 ed 0.0059 yCi 239Pu(IV)/kg in  0.08  M citrate buffer, pH 3.5.  The dogs were
 sacrificed at 1648 and 2546 days.   The cumulative rad  doses to the skeleton
 and liver were compared and the results show that the average dose to the liver
 exceeds that to the skeleton,  except at the  2.8 UCi/kg dose level, after 1300
 days.
                 1 J. Q
      In 1972 Jee    found that the  effectiveness of the various routes of
 administration in delivering plutonium to bone, in decreasing order is:
 intravenous>intraperitoneal>subcutaneousIntramuscular>inhalation>oral>direct
 application upon skin.   Reticuloendothelial  cells in the marrow compete  with
 bone for polymeric plutonium and thus decreases the plutonium available  for
 bone surfaces.   The uptake in  young and adult  bones differed by a factor of
 two.  The fate of the plutonium surface deposits is modified by bone growth,
 modeling and remodeling in growing  animals and remodeling in adults.  These
 age-related processes remove the plutonium from bone surfaces and/or bury
 the surface deposits with new  bone.

      A  very complete summary of the metabolism of plutonium after administration
 in different forms and by different routes up  to 1972, is given  in  tabular  form
 in the  International Commission on  Radiological Protection  (ICRP) Publication  19265.

 BIOLOGICAL EFFECTS OF PLUTONIUM IN  ANIMALS

      In animal  experimentation, the acute forms of toxicity are manifested
by  an irradiation syndrome  comparable to  that  produced by a total irradiation
with x-rays-  Forms of  a chronic nature can  produce broncho-alveolar  tumors
in  the  lung,  bone  cancers,  and bone fractures  in the skeleton, and, with less
frequency,  liver tumors.   Saenz and Ramos1 present lethal and tumor-causing
doses of plutonium in dogs,  rabbits and mice.

     It has been shown  that  the distribution of plutonium and the radiation
dose among  the  tissues  in the  body  will vary depending upon the  physical and
chemical characteristics  of  the plutonium inhaled.  The biological  effects
that occur will  depend  upon  the radiation exposure and the relative radia-
tion sensitivity of  each  tissue into which plutonium is transported and
deposited.  These  are primarily blood, bone, liver, lung and lymphatic  system.


                                      4Q

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Blood

     Plutonium is cleared from the circulating blood within a few hours after
absorption.  Therefore, the effects seen in blood cells are probably due to
irradiation of hematopoietic tissue in which plutonium is deposited or to
irradiation of blood circulating through plutonium-containing tissues.

     In 1972 Dougherty266 reported blood cell alterations in dogs receiving
a single intravenous injection of a wide range of dose of 239Pu (0.016 to
2.8 yCi/kg).  Dogs receiving the four highest dose levels showed changes that
were dose-dependent and were most marked in granular leukocytes which were
maximally depressed at 2 to 3 weeks post injection.  A sustained lymphopenia
occurred at the two highest dose levels.  A transient early thrombocytopenia
and anemia were also found in higher level dogs.  A moderate anemia reappeared
as the animals became terminal.

     In 1972 Nabors267 noted that beagles bearing 239Pu burdens show the most
marked alterations in serum alkaline phosphatase and serum glutamic oxaloacetic
transaminase and serum glutamic pyruvic transaminas.e.  Animals with low dose
levels appear to have a much longer latent period prior to the appearance, of
altered serum biochemistry than do animals with higher dose levels.

Bone

     The most sensitive effect of plutonium deposited in bone is radiation-
induced osteogenic sarcoma.  Because plutonium deposits on bone surfaces,
a large fraction of the alpha energy is absorbed in sensitive cells.  Although
osteogenic sarcoma has been reported in rats that accumulate doses of less
than 10 rads, statistically significant increases in tumor incidence have not
occurred at doses less than 30 to 50 rads (Bair250).  The available data suggest
that the dose-effect curve for dogs is different from that for rodents.  Mays
and Lloyd258 conclude that the incidence of plutonium-induced osteogenic sarcoma
is 0.38%/rad for beagle dogs, 0.10%/rad for mice and 0.06%/rad for rats.
                                                           n q n
     No bone sarcomas have been reported in dogs following    PuOa inhalation
but they are the important late biological effects in dogs which have survived
intravenous injection of plutonium citrate for more than 3 or 4 years (Mays
et al.).

     Osteogenic sarcoma has been observed in mice, rats, rabbits, and dogs
after intravenous injection of several 239Pu compounds including organic
complexes and plutonium nitrates.

     Osteogenic sarcoma has also been observed in rats after inhalation of
238Pu02, but has not been reported in any animal species after inhalation of
239Pu02.

     In 1973 Moskalev and Strelt'sova270 found that optimun osteosarcomogenic
doses to rat skeleton were in the range of 0.7 to 1.8 krads for  239Pu.  The
osteosarcoma incidence was identical for both male and female rats but  signifi-
cantly age dependent for each sex.
                                      41

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      Pesternikov and Bukhtoyarova27*  discussed the quantitative assessment of
 osteosarcoma growth rate in rats  following a single intraperitoneal administra-
 tion of 2.5 mCi/kg of 239Pu citrate.  Osteosarcomas are shown not to predomi-
 nantly originate at the sites  of  the  highest isotope concentration.  Of some
 tumors, primary multiple growth was characteristic.  Possible extrapolation for
 determining the osteosarcomas  development time was noted.

      James272 reported a study in which 6 to 8 week-old rats were given a
 single injection of 4.5 yCi/kg soluble Pu(N03K, a dose level expected to give
 an approximate 80% yield of osteosarcomas.  Localized dose rates were measured
 at fixed reference distances between  5 to 20 ym from bone surfaces by counting
 gamma-flux entering small cylindrical targets located in a thin nuclear emul-
 sion in contact with a bone section.  Dose rates 1 day after the injection
 ranged from 22 to 57, 12 to 37, and 7 to 26 rads/day at distances of 5, 12.5,
 and 20 ym, respectively, from  a number of different endosteal surfaces.  These
 local dose rates changed with  time by factors of the order of 2, in either
 direction, depending on the prevailing conditions of remodeling.  A cumulative
 alpha dose of about 2000 rads  (delivered to primitive cells at trabecular
 surfaces over a period of 36 weeks) may be associated with a 10% probability
 of developing a tumor in an individual femoral epiphysis.  Taylor  73 et al.
 studied the general syndrome induced  by 239Pu(IV) injected intravenously in
 beagle dogs.  The most critical factors were the induction of bone cancer,
 hematopoietic changes and liver lesions.  Some of the less serious end points
 were pathological fractures, dental changes and atrophy of the turbinates.
 These latter conditions produced  functional impairment in only part of the
 dogs and principally at the higher dose levels.

      Taylor and Mays271* gave young adult St. Bernard dogs a single intravenous
 injection of    Pu,  because of their  high spontaneous incidence of bone cancer.
 The appearance time of radiation  induced osteosarcomas in this giant breed
 was approximately one-half of  that observed in beagles when both were injected
 with 0.3 yCi 239Pu/kg of body  weight.

      In 1972 Moskalev275 reviewed the results of studies by Soviet scientists
 on problems of the biological  action  of 239Pu.  Main attention was given  to  the
 analysis  of late effects (tumor and nontumor effects) developing  in  the body
 as a result of injury by 239Pu, dose-effect curves for bone and luiig  tumors,
 estimation of minimum carcinogenic dose levels, and determination  of  doses
 not  affecting the natural life span.

 Liver

     Liver accumulates  levels  of  plutonium similar to bone.  However,  liver
 tumors have not  been  a  common  finding in experimental animal studies.   Bile
 duct tumors  have been observed to occur in beagle dogs at doses as low as
 60 rads.   However, not  only was the incidence very low, but bile  duct tumors
also occurred in control dogs  (Taylor276 et al.).

     Kashima277  et al.  demonstrated in 1973 that liver function at 21 days
post exposure was markedly reduced in rats injected with  239Pu at birth or
at 7 days  of  age.  The  differing  effects produced by monomeric and polymeric
plutonium  were explained by differences in radiation dose to the  liver.

                                      42

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Lung

     Inhalation of relatively soluble plutonium compounds, such as organic
complexes, plutonium nitrate and plutonium oxide, has resulted in primary
pulmonary neoplasia in rodents, rabbits and dogs (Bair25i).

     Pulmonary neoplasia has also been observed in beagle dogs, baboons
(Metivier278 et al.) and rodents after inhalation of 239PuO .   Tumors were
observed in animals that had lung doses less than 10 rads.  Statistically
significant increases in pulmonary neoplasia occurred at doses of about
30 rads and above.  In several experiments, increased incidences of lung
tumors occurred at doses between 10 and 100 rads.  In an 11-year study of
239Pu02 in beagle dogs, nearly all animals which had depositions of between
about 3 nCi and 50 nCi per gram of lung had lung tumors.  Dogs which had a
deposit of more than 50 nCi per gram of lung died early due to radiation
pneumonitis and fibrosis  (Park251* et al.).  The mean dose to the lungs of
the dogs that developed neoplasia ranged from 1200 to 4000 rads.  As in the
case of osteogenic  sarcoma, the dose-effect curve for pulmonary neoplasia
in dogs appears to differ from the rodent dose-effect curve.

     Howard279  described the results of a long-term study to evaluate the
pathologic effects in the lungs of beagle dogs following 239PuO  particle
inhalation by 40 dogs exposed to 239Pu02 particles having a CMD of 0.1 to
0.5 ym; 22 died with primary lung neoplasia 38 to 110 months post exposure;
8 died of pulmonary fibrosis; 5 were sacrificed for radionuclide distribution
measurements; and 5 were still alive over 9 years post exposure.  Initial
alveolar deposition ranged from 0.5 to 3.5 yCi and accumulated radiation
dose was 2500 to 12,000 rads.  Most lung tumors were found to be bronchial-
alveolar carcinomas of peripheral origin, with two peripheral squamous cell
carcinomas and three epidermoid carcinomas.  Also in this group of dogs
there were three thoracic sarcomas and two with malignant lymphomas-

     Sanders and Park280 and Sanders281'282  reported on the carcinogenicity
of inhaled 238Pu and 239Pu in soluble and insoluble forms by rats and dogs.
It was concluded that exposure of rats to even small amounts of 238Pu resulted
in significant incidence of tumors in lungs.  The type of metaplasia and neo-
plasia in rats and dogs indicates the epithelial, either alveolar or bronchi-
olar, origin of most of the observed tumors.

     Anderson283 et al. reported on lung_irradiation of Syrian  (golden) ham-
sters with static plutonium microspheres.  Exposure was accomplished by inject-
ing 10 ym microspheres into the jugular vein.  The microspheres were trapped in
the capillary bed of the lung.  Activity range was 0.01 to 59 pCi/sphere.  Of
1,142 hamsters exposed to 5,700,000 microspheres, only two lung tumors were
observed.   Although the experiment was not complete at the time of the report
the authors felt that the volume of tissue irradiated was an important factor
and that locally concentrated alpha irradiation is less damaging in the hamster
lung than the. same amount of energy delivered over larger volumes.

Lymph

     Plutonium accumulates in lymph nodes following deposition  of plutonium in
the respiratory tract.  Months or years after the contaminating event,  lymph

                                      43

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 nodes may attain concentrations of plutonium many times the average concentra-
 tions remaining at  the  site of deposition and consequently the accumulated
 radiation dose to lymph nodes may be greater than to any other tissue.

      Although dogs  have been studied for 11 years after inhaling a variety
 of plutonium compounds, primary neoplasia of lymphatic tissue has not been a
 common observation  (Bair  53).  Metastases of tumors from lung to lymph nodes
 have occurred, but  none has  been the primary cause of death.  It can be
 concluded from the  many rodent and dog life-span experiments with  38Pu and
 239Pu, in which many lymph nodes have been exposed to doses ranging from near
 background to thousands of rads delivered at a wide range of dose rates, that
 the lymph nodes are not especially susceptible to the carcinogenic action of
 alpha radiation from plutonium.

      Dagle284 reported  on lymph node clearance of 239Pu02 administered as
 insoluble particles from  subcutaneous implants in adult beagles to simulate
 accidental contamination  of hand wounds.  The popliteal lymph nodes were
 monitored following implantation ranging from 9.2 to 39.4 uCi 239Pu02 into
 the hind paw.  Groups of  dogs were sacrificed 4, 8, 16, and 32 weeks later
 and it was found that the popliteal lymph node contained from 1 to 10% of
 of the implant dose. Histopathologic changes in the popliteal lymph nodes
 with plutonium particles  were characterized primarily by reticular cell hyper-
 plasia, increased numbers of macrophages, necrosis and fibroplasia.  Eventu-
 ally, the plutonium particles became sequestered by scar tissue that often
 replaced the entire architecture of the lymph node.  There was slight clearance
 of plutonium from the popliteal lymph nodes of dogs monitored for 32 weeks.

 Other Effects
            f\ Q |-
      Stover    et al. found correlation between dose level and life-shortening
 in beagle dogs.   Sikov  and Mahlum21*1* found that 12 yCi/fcg of 239Pu, which is
 well below the acute lethal range for the adult rat, produces extensive
 prenatal death within 3 to 5 days when administered to the pregnant rat at
 9  days>of gestation.  There was evidence that-these deaths-occur by alterations
 of the nutritive function of the fetal membranes.  In contrast, no prenatal
 mortality is  produced at  doses beyond the lethal level for the mother,
 150 nCi/kg,  if administered at 15 or 19 days of gestation.  In 1972 Ovcharenko286
 found similar results when studying the effects of plutonium on reproductive
 ability of rats.  He found that the character and depth of radiation-induced
 changes depended upon the amount of activity injected and the time elapsed
 since injection.

      In the course  of investigating the postulated development of the offspring
 of  experimental  animals,  a number of peculiarities were observed:  decrease in
 viability, delayed  physical development, variations in weight, disturbance  of
 blood  formation,  change in radiosensitivity and depression of sex  function.
                                                   /
      Sanders287'288   and Sanders and Jackson289 studied the synergistic  effects
 of plutonium  and  asbestos administered to rats.  It was  found that coadmini-
stration  by  inhalation  of 239Pu02 with asbestos resulted in less excretion of
239Pu  than was the  case with 239Pu02 only.  Focal granulomatous  lesions were
common  around bronchiolar areas of the lung in animals receiving asbestos.
Plutonium oxide particles tended to concentrate in these asbestos-induced

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lesions forming hot spots of alpha activity.  When the same combination was
intra-abdominally injected into  rats, asbestos acted in an additive manner with
   Pu02 in inducing mesotheliomas, the two agents combined having an effect
equal to the sum of their effects when administered separately.  A similar
additive response was found when benzpyrene and 239Pu02 were injected with
resulting formation of abdominal sarcomas.

COUNTEKMEASURES FOR PLUTONIUM IN ANIMALS

239  A number of chelating agents have been used in experiments to remove
   Pu by increasing the excretion rate.   Smith290, as reported in 1958,
testing 10 of the amino-acetic  acid-type  chelators administered intraperi-
toneally in a dose of 1.5 mM/kg in an equivalent amount of calcium gluconate
1 hour after intravenous injection of 239Pu to rats.  The most effective agent,
diethylene-triamino-pentaacetic acid (DTPA), decreased deposition of plutonium
in the liver and bone by about  99%; the next most effective, diamino-diethylether-
tetraacetic acid  (DDETA), reduced deposition by about 95%.  Even when DTPA
calcium gluconate plus 25,000 IU of Vitamin A was administered 39 days after a
dose of   39Pu, it reduced retention in bone by 25% and in liver by 80%.

     Zirconium citrate and certain phosphate compounds, especially hexameta-
phosphate (HMP) reduced retained 239Pu by 50 to 70% if administered immediately
after a dose of 239Pu according to Wagner and Temple201 and Rosenthal292.
However, all of the above mentioned treatments have certain drawbacks:  DTPA
                          O O O
and its analogs increased    Pu concentration in the kidney, while zirconium
and phosphates increased concentration in the liver.  Desferriox-amine (DFOA)
appears to be as effective or nearly as effective as DTPA if given within a
few hours after 239Pu exposure; it did not show significant 239Pu renal concen-
                           n n o
tration according to Taylor

     Nenot294 et al. found that the therapeutic effectiveness of DTPA on bone
deposits of plutonium can be appreciable.  DTPA treatment begun 3 weeks follow-
ing intramuscular injection of  239Pu decreased bone burdens by a factor of one-
half to one-third in 3 months.

     Pretreatment of rats with  tetracycline was found to decrease retention
of 239Pu in bone to about 60% of control  values according to Taylor295 et al.
Tetracycline increased the initial rate of clearance of 239Pu from plasma.
Treatment with tetracycline was found to  have no effect on the distribution
of intramuscularly-injected 239Pu, even when treatment was started immediately
after contamination.

     Morin231 et al. studied the effect of treatment by DTPA following intra-
muscular and inhaled 238Pu nitrate by rats.  Treatment was initiated 24 hours
after exposure and repeated twice a week.  Protection of or removal from bone
by DTPA was clearly demonstrated at the 30th day when the respective bone
burdens of control and treated  animals were in a ratio of 8:1, a decrease from
45 to 17% of bone activity.  Postponing treatment until the 21st day showed
the importance of early treatment.

     Sanders296 and Sanders and Meier297  studied the influence of fasting on
the efficiency of DTPA in enhancing the excretion of intraperitoneally injected
238Pu in female rats.  Complete fasting for 10 days had no significant effect

                                       45

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 on the cumulative  excretion of injected 238Pu.  In animals given DTPA within
 the first hour after  238Pu there was a 2.2-fold increased excretion, and a
 1.4-fold increased excretion when DTPA was administered 14 days after 238Pu,
 irrespective of fasting.  They concluded that distribution of 238Pu bound to
 liquids in extracellular  fluids and bone was not significantly altered by
 fasting so as to influence the chelation of 238Pu or 239Pu by DTPA.

      Studies of the translocation of plutonium from wound sites and effects
 of treatments were reported by Bestline298 et al., Lebel and Watters299,
 Watters and Lebel300,  Watters and Johnson301, and Gomez302 et al.  The investi-
 gators found that  the  effect of DTPA treatment was greater for Pu(N03),t than
 for Pu02 implants. A time study of the buildup of plutonium in the lymph
 node draining the  wound site disclosed that the lymphatic system provides
 rapid translocation from  the wound.  Rapid surgical excision, to be highly
 effective, should  be  performed as soon as possible.  If accumulation of
 plutonium is detected  in  lymph nodes, lymphadenectomy should be considered.

      Rosenthal303  et  al. , Rosenthal301* et al. and Baxter305 et al. studied
 the effectiveness  of  DTPA and glucan, a polysaccharide derived from yeast
 cell walls, as agents  for  plutonium decorporation.  DTPA therapy removed over
 96% of intravenously  injected monomeric 239Pu from beagle dogs.  Removal from
 spleen, lungs, kidneys, testes, muscle and lymph nodes ranged between 50 to
 90%.  The total skeletal  content was also reduced by 50%.  Adjunct therapy
 with glucan did not result in significant additional removal of plutonium.
 Use of the same treatment following polymeric 239Pu injection of mice showed
 that at 47 days after  administration, the net removal was 8.5% of the injected
 dose by glucan, above  11.5% by twice weekly DTPA treatments and 19.5% by
 combined therapy.

      In 1972 Ballou and Hess306 found that about 50% of the plutonium excreted
 into the perfused  intestine of the rat during the first hour after plutonium
 injection arrived  by way  of the bile.  After DTPA treatment, the proportion
 excreted in bile was  increased to 75% and the bile plutonium concentration
 increased 15 to 20-fold.

      Rahman307 et  al.  in  1973, successfully encapsulated EDTA and DTPA within
 lipid spherules (liposomes).  Encapsulated EDTA given intravenously to mice
 was retained longer in tissues than nonencapsulated EDTA.  Encapsulated DTPA,
 given to mice 3 days after 239Pu injection, removed an additional  fraction of
 plutonium in the liver, presumably intracellular, not available to nonencapsu-
 lated DTPA.   It also increased urinary excretion of plutonium.

      Inhaled insoluble plutonium is not effectively mobilized by a wide variety
 of  expectorants, mucolytes agents, bronchodilators, surface-active  agents,
 phagocytic  stimulating procedures, chelating and other physiologically  active
materials  (Smith308).  Recent experiments have shown that pulmonary  lavage
with physiological  saline may remove 50% or more of Pu02  from the  lung  of  rats
[(Sanders309); dog  and  rat (McDonald310); monkeys  (Kunzle-Lutz311 et  al. ,
Nolibe312);  and baboons (Kunzle-Lutz313 et al,)],

DEPOSITION OF  PLUTONIUM IN MAN

     Plutonium has  found its way to man, in readily measurable  quantities,  only
through occupational exposure, where the route is usually direct—by ingestion,

                                     46

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inhalation, or by way of a Plutonium-contaminated wound (Bair and Thompson31").

     In 1959 Langham315 found that alpha radiation from plutonium on the skin
surface does not penetrate to the sensitive basal layer of the epithelium.
Absorption of plutonium through the skin occurs only to a very slight degree,
and probably only when the skin is damaged.  Lister316 studied fecal and
urinary plutonium excretion from two subjects who sustained high levels of
contamination on the uncut skin of the hand from accidental contact with
acid plutonium solutions.  In one case, where the contaminant was mixed
plutonium isotopes in aqua regia and nitric acid, the measurements covered
150 days.  In the other case, contamination with a solution of plutonium in
dilute hydrochloric acid containing EDTA and a detergent  was followed for
110 days.  The excretion patterns showed marked differences from those found
by Langham315, particularly in the high and variable amount of plutonium
excreted in feces relative to urine.

     Laylee317 also studied excretion of plutonium following a lacerated wound
contaminated with 100 nyCi of Pu02.  No translocation values were given.

     In October 1970, a 59-year-old employee of Dow Chemical Company died
from complications following open heart surgery.  This employee had an
estimated systemic burden of 0.22 yCi of plutonium or 541% of the maximum
permissible system burden, at the time of his death.  The employee had been
involved in several contamination incidents and had two contaminated puncture
wounds prior to November 1965.   He had excreted measurable amounts of pluto-
nium in his urine since 1960 and had an estimated 21% of the maximum permis-
sible systemic burden.  In July 1965, he received another puncture wound
which left 18.5 yg of plutonium.  A surgical incision removed 5.9 yg in
1967, a second incision removed all but about 0.3 yg.  Autopsy and tissue
analysis showed plutonium concentration>skeleton>liver>lung>ttacheobronchial
lymph nodes>spleen>kidneys.  The total extrapolated system burden from
autopsy samples was 0.07 yCi or about 177% of the maximum permissible amount
(Lagerquist318 et al.).

     In 1973, Popplewell3l9 reviewed the chemistry of plutonium incorporated
in the body.  Subjects covered are transport within the body in relation to
use of chelates, especially Ca-DTPA; studies of transferrin complexes in
body fluids; intracellular uptake in liver and deposits in bone.

     Keely and Wenstrand32° calculated the urine to fecal ratios for two
workers following inhalation of soluble plutonium nitrate.  Twenty days post
exposure the residual body burden was primarily in the skeleton.  After 80
days and 250 days post-exposure <10 nCi was in the skeleton and <2 nCi total
plutonium in liver or lung as determined by in vivo counts.  Urinary excre-
tion was erratic after 20 days post exposure but indicated a 1.8-day half-
life during the initial time of translocation.

     Lagerquist321 et al. studied 19 cases where tissues were obtained  from
autopsy and analyzed for plutonium.  Twelve of the cases had no known record
of plutonium exposure.  All of these had some detectable plutonium in their
organs but generally an amount that is representative of less than 1% of  the
Maximum Permissible Systemic Burden  (MPSB),  In the seven  others,  the  distribu-
tion varied from case to case, depending on the mode  of entry,  the chemical form
and the length of time since exposure.  A  typical  distribution  was not  found, but

                                     47

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 on the average,  the  lung and tracheo-bronchial lymph nodes had the highest con
 centration of material  followed by the liver, the bones, and other tissues.

      Przyborowski322  and Vaane323 et al. studied the deposition of plutonium
 particles in the human  respiratory tract after inhalation.  A combined experi-
 mental mathematical  procedure was used,

      Mclnroy321*  et al.  studied autopsy samples from 70 former employees of
 Los Alamos Scientific Laboratory who had exposures ranging from 4 to 30 years.
 Exposure in most cases  was  to inhaled Pu02 aerosols.  Thirty-three of the
 measured cases had plutonium deposition in th tracheobronchial lymph nodes.
 No abnormalities of  the lymph nodes were found.

      Richmond325 presented  a summary of information on human experience with
 plutonium.  Groups studied  ranged from exposed GI's in 1944 to 1945 weapons
 development, to  occupationally exposed workers at Los Alamos and Rocky Flats,
 to inhalation intake and burden in man of fallout 239Pu in New York.  Data
 are presented in tabular and graphical forms with tissue distribution included.

      The retention half-time for Pu02 inhalation in humans was reported as
 250 to 300 days. The relatively low values for human beings, compared with
 dogs, suggest either that man clears plutonium particles from his lungs more
 rapidly than dogs or that the materials inhaled in human accident cases were
 more soluble than plutonium dioxide  (Bair326).
      Durbin    reviewed old  data and applied more recent knowledge  derived
 from animal experiments.  Much  of  the older information was derived from intra-
 venous administration  of tracer amounts of 239Pu citrate to men suffering incur-
 able diseases in 1945  and 1946.  A few days after injection, human  soft  tissues
 (other than blood and  liver)  contained as much as 20% of the plutonium dose.
 Five to 15 months after injection  the average liver plutonium  content  was 31%
 of the dose for three  cases  with presumably normal liver function.   Four to
 457 days after injection, mean total skeletal plutonium was 49% for  the seven
 cases judged to have most nearly normal liver and skeletons.

      Plutonium is transported in blood combined with transferrin, the  iron  trans-
 port protein,  and is stored  in  the liver in association with stored iron.  After
 being bound to transferrin,  plutonium partially traces the behavior of the  carrier
 protein.   The early phases of plutonium transport which are apparently associated
 with extracellular fluid mixing, were prolonged in individuals with impaired  cir-
 culation.

      Maximum urinary plutonium  excretion occurred before the bulk of plutonium
was  protein-bound.  Minimum  urinary excretion coincided with the time  of maximum
Pu-transferrin binding.   These  observations were taken to mean that some pluto-
nium is  filtered by the  kidney  in  the form of a low-molecular  weight chelate.
Urinary plutonium  excretion was reduced by one-half in those persons who were
anemic, presumably because of their more efficient Pu-transferrin binding.

     Fecal excretion of  plutonium  apparently represents secretion  in bile and
other digestive juices.   Fecal  excretion was reduced by one-half or more in
those persons whose gastrointestinal tracts were judged not to be normally
stimulated.

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     Average soft tissue release half-time was estimated to be.not less than
480 days and bone surface turnover for the whole adult human skeleton was
estimated to be about 5% per year.  For an individual on a diet adequate in
iron and with normal iron stores, the model predicts that bone and liver will
contain equal amounts of plutonium 15 years after exposure.

EFFECTS OF PLUTONIUM IN HUMANS

                328
     Hemplemann    et al., reporting a 27-year study of Los Alamos workers,
found that to date, none of the medical findings in the group can be attri-
buted definitely to internally deposited plutonium.  The bronchial cells of
several of the subjects showed moderate to marked metaplastic change, but the
significance of these changes is not clear.  Diseases and physical changes
characteristic of a male population entering its sixth decade were observed.
Because of the small body burdens, on the order of the maximum permissible
level, in these men so heavily exposed to plutonium compounds, it was concluded
that the body has protective mechanisms that are effective in discriminating
against these materials following some types of occupational exposures.  This
is presumably explained by the insolubility of many of its compounds.  Pluto-
nium is more toxic than radium if deposited in certain body tissues, especially
bone; however, from the practical point of view, plutonium seems to be less
hazardous to handle.
                                3 O Q
     In 1974 Tamplin and Cochran    reported two cases of cancer presumably
caused by plutonium contaminated puncture wounds of the hand.  At this time
these were the only reported cases of cancers in people working with plutonium.

COUNTERMEASURES FOR HUMANS EXPOSED TO PLUTONIUM CONTAMINATION

     The most successful treatment of inhaled plutonium particles appears
to be pulmonary lavage, followed by chelation therapy (Nolibe330, Smith331,
McClellan332 et al.).  Lavage appears to remove about half the inhaled parti-
culates.  Chelation by means of administration of DTPA either by aerosol or
intramuscular injection has been successful in removing soluble translocated
plutonium (Nenot    et al. and Schofield and Lynn334).

     Treatments of contaminated wounds include immediate use of a venous
tourniquet; flushing and decontaminating the wound site; excising tissue;
and administering chelating agents (intravenously, orally, and by aerosol
inhalation) (Jolly335 et al., Hesp and Ledgerwood336 and Larson337 et al.).


                        PRESENT ANALYTICAL METHODS

     A thorough evaluation of present analytical methods for the transuranic
elements including plutonium in environmental samples has been prepared by
Bernhardt338 and is in press at this time.

     It has been noted by numerous authors that analytical results are only
as good as the methods by which the samples are collected.  Among the  environ-
mental samples posing the greatest sampling problems are air and soil.   In
1972 Hull3   reported tentative methods for the sampling and analysis  of
radioactive substances in air as formulated by the Radioactive Substances


                                     49

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 Subcommittee  of  the  Intersociety Committee on Methods for Air Sampling and
 Analysis.

      In 1973  Eberhardt  and Gilbert3"*0 discussed the problems associated with
 sampling of soils  for environmental levels of plutonium.  The general statisti-
 cal considerations of sampling, problems associated with compositing samples
 and consistent  use of random  sampling as a basic technique were examined.

      The literature  contains  many methods for determination of plutonium in
 biological and  environmental  samples.  Most of these methods are variations
 of basic procedures  which utilize the precipitation, ion exchange and complex
 forming properties of plutonium followed by electroplating and counting.

 GENERAL METHODS

      In 1971  Wong341 reported on the radiochemical determination of plutonium
 in seawater sediments,  and marine organisms.  Seawater samples of 50 to 60
 liters were collected at various depths; 50 to 100 g of dried shallow water
 sediments or  a  larger aliquot from  core samples was used.  The author provides
 a table of average 239Pu concentrations in marine organisms to allow selection
 of proper sample size for optimal sample counting rate; the activity should be
 about 0.5 to  5  dpm.  An iron(II) hydroxide coprecipitation method was used for
 the concentration of plutonium in seawater.  A nitric-hydrochloric acid leach-
 ing method was  adapted  for the treatment of sediments and ashed organisms.
 The sensitivity for  this method is  0.004 dpm per 100 liters of seawater  (using
 50-liter sample),  0.02  dpm per kg of sediments  (100-g sample) and 0.002 dpm per
 kg of marine  organisms  (1-kg  sample).  Factors influencing the uncovering,
 contamination and blank activity were discussed.

      Darrall3"*2  et al.  described a  method for the simultaneous determination
 of 21|1Pu and  plutonium  alpha  activity in effluent samples from nuclear
 installations.   The  plutonium was isolated by coprecipitation on barium sulfate
 followed by extraction  into  di-(2-ethylhexyl)-phosphoric acid, which was incor-
 porated in a  liquid  scintillator for counting in a liquid scintillation spectro-
 meter.   Interferences from alpha- and beta-emitting radionuclides were studied
 together with interferences  from nonradioactive elements.  The lower limit of
 detection is  in  the  region of 1 pCi.  This is a modification of the method
 described by  Sill31*3 and by  Sill and Williams3"1* in 1969 who reported that
 plutonium will  coprecipitate  on barium sulfate provided it was in the tervalent
 or quadrivalent  state and the coprecipitation was carried out in the presence
 of potassium  ions.   By  coprecipitation after selective oxidation the elements
 can be  separated.  The  barium sulfate precipitate may be alpha counted directly
 or the  plutonium can be electrodeposited for alpha spectrometry.  Talvitie3"*5'31*6
 described  methods  for radiochemical determination of plutonium in environmental
 and biological samples  by ion exchange and an electrodeposition method for
 alpha spectrometric  determination.  Sample preparation procedures were given
 for urine,  animal  tissue, bone, saline and nonsaline water, siliceous and
 limestone  soil,  and  glass fiber air filters.  Samples were prepared  as azeo-
 tropic  6M  hydrochloric  acid solutions of Pu(IV), stabilized with hydrogen
 peroxide,  and absorbed  on anionic resin from 9M hydrochloric acid solution.
 Coadsorbed iron  was  removed from the resin with 7.2M nitric acid.  Plutonium
was selectively  eluted  with 1.2M hydrochloric acid - 0.6% hydrogen peroxide
 and electrodeposited from 1M  ammonium sulfate at pH 2 for alpha spectrometric
 determination.   Minimum detectable  activity for 1000-min counts was  0.02  pCi
                                      50

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 239Pu.  Puphal  and  Olson31*7 describe a method for electrodepositing various
'alpha-emitting  nuclides singly or in combination onto  a stainless  steel  cathode
 from a mixed ammonium oxalate-ammonium chloride electrolyte.   The  nuclides were
 deposited  as hydroxides to better than 98% in 50 minutes.   Interferences are
 also discussed.

      Gillette3lt8 et al. investigated the determination of  plutonium in soil.
 The method involves a complete dissolution of the soil by  a fusion technique
 followed by chemical separation of the plutonium by barium sulfate, extraction,
 electrodeposition,  and alpha pulse height analysis.

      Hampson and Tennant31*9 reported the simultaneous  determination of acti-
 nide nuclides in environmental materials contaminated  by controlled discharges
 of liquid  wastes.   Multielement actinide analysis is achieved by extracting
 the whole  group, or part of it, in the tri-n-octylphosphine oxide-n-heptone-
 nitric  acid-sodium nitrate system, stripping into ammonium carbonate  solution
 and electrodeposition, followed by solid-state alpha spectrometry.  Up to 2 kg
 of edible  seaweed or 10 kg of fish flesh can be handled, with detection  limits
 (in terms  of activity to double background) of 2 X 10  6 and 4 X 10 7  pCi/g,
 respectively, for a 1-week counting time.  Sensitivities for precision with
 4 percent  standard deviation are 4 X 10 "* and 8 X 10 5 pCi/g, respectively,
 which corresponds to levels associated with fallout.

      Piltingsrud and Stencel350 report a direct method for evaluating 239Pu
 content in large soil samples (up to 1 kg) by means of a phoswich  detector.
 Sample  239Pu + 21tlAm specific activity must be >20 pCi/g.   When high  levels
 of other  gamma emitters, a ratio >1000:1, are present  causing analysis
 interference, specific radiochemical analysis of the sample is indicated.

      In 1973 Thomas351 presented a method for measuring 238Pu and  239Pu  in
 air samples.  The membrane filter was prepared by fusion and the plutonium
 was retained on an ion exchange column and eluted with HC1 containing HI.
 After electroplating, the plutonium isotopes were alpha counted.

      Novoselova352 et al. described the optimum conditions of concentration,
 separation and measurement of 239Pu after the latter had been added to ashed
 bone tissue.  Bone tissue was ignited in a muffle furnace  at 600°  C,  and the
 inorganic  residue was dissolved in HN03.  The plutonium was extracted with
 TBP, reextracted with aqueous hydroxylamine HC1, and precipitated  from solu-
 tion into  bismuth phosphate.  The latter was washed, mixed with a  phosphor,
 and the plutonium was determined in a scintillation counter.

      Ershova353 et al. described a method for determination of 239Pu  based
 on liquid-liquid extraction by monoisoactyl methyl phosphonate and mixed
 isoamyl phosphate at a phase volume ratio of at least  100.  This method  is
 used for the estimation of the 239Pu content in lung tissue, bone  tissue,
 surface waters,  and other environmental objects.

      Ghysels351* reported the extraction of plutonium was effected  with liquid
 ion exchangers  (e.g., tertiary amines such as triisooctylamine).   The pluto-
 nium was put back into organic solution by extraction  with DEHPA.   After mixing
 with a liquid scintillator, the counting yield was determined (close  to  100%).
 This method was  intended for determination of plutonium in biological substances.
 It  was applied  to  the analysis of radioactive dusts of different  origin.

                                     51

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      Schieferdecker355  presented  a method  for separation of plutonium in pCi
 amounts from biological material  by extraction with bis  (2-ethyl-hexyl) phosphate
 (DEHPA).   It was possible  to  identify individual transuranic elements after joint
 carrier-free preparation by electrodeposition and subsequent alpha spectrometry.
 Amounts below 0.2 pCi can  be  determined.

      In 1973 Budnitz356 provided  an overview of the techniques for measuring
 plutonium in various media.   The  emphasis  was on measurements for surveillance
 and protection in environmental and occupational situations.  Overviews were
 first provided for the characteristics of, sources of, and typical levels of
 plutonium concentrations.  The various measurement techniques were discussed.

 AUTORADIOGRAPHY

      Fission track autoradiography has been used by several investigators to
 study the distribution of  plutonium in bone and other biological samples
 (Hamilton357, Becker358 et al., Jee359'360 et al. and Schlenker and Oltman361).
 The autoradiographs are produced  by placing thin bone sections in contact with
 plastic films which register  fission fragment tracks.  The bone and film are
 inserted into a nuclear reactor where neutrons induce fissioning of plutonium.

      Some plastics show not only  the fission tracks but an image of the bone
 itself.  The image shows no detail but does reveal where the boundaries between
 the bone and marrow spaces lie so that positioning of the fission tracks rela-
 tive to the boundaries may be easily determined.  Lexan is the plastic which
 has been most widely used  to  produce images of irradiated materials  (Schlenker
 and Oltman361).
                            Q C O
      Lindenbaum and Russell    found good  agreement between liver burdens
 determined autoradiographically or radiochemically.  Using livers from mice
 injected intravenously wiht polymeric 239Pu, at dose levels ranging from
 6.6 to 93 yCi/kg, the quantitative technique for assay of plutonium deposited
 in animal tissue was tested for linearity  in the relationship between photo-
 graphic exposure time and  number  of alpha  tracks formed.

 INSTRUMENTAL ANALYSIS

      In 1973 Winkle and Hoetzi363 reported a sensitivity of 0.04 to 0.01 pCi/m3
 for long-lived alpha-emitters in  air.  The sensitivity was obtained by  alpha
 spectrometry of electrostatically-precipitated aerosols  in a large Frisch  grid
 ionization chamber after decay of the natural activity for 2 to 4 hours  and
 100 minutes  counting.   This sensitivity corresponds to about 0.1 to 0.01 of
 the  maximum  permissible concentrations (168/h/wk) for 239Pu.

     Liquid  scintillation  counting of plutonium in biological  samples  is  the
 subject of a  report by  Joshima and Matsuoka36\  The liquid scintillation
method without  ashing was  established for  plutonium in animal  tissues  by use
of a commercially  available tissue solubilizer.  Samples up to  250 mg  wet
weight could  be handled with  this procedure.  The samples were  then  counted
in a liquid  scintillation  spectrometer set for the optimal gain setting for
plutonium.
                                       52

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     Matsuoka and Joshima365 developed a method for determining particle size
of plutonium in solution by a diffusion chamber.  The diffusion chamber was
prepared from a plastic ringe (20 mm  x 5 mm), both sides of which were
covered with millipore membrane of adequate pore size of Visking membrane.
The diffusion chamber containing 0.1 ml of test solution was placed in a plastic
container with 5 ml to 10 ml saline solution and incubated 1 hour with shaking
in a 37° C water bath.  The "diffusibility" of plutonium particles from the
diffusion chamber was estimated from the sample measurement of extra fluid at
the final point by liquid scintillation counting and for external counting at
17 keV L-x-ray of the diffusion chamber before and after the incubation.  The
particle size of "monomeric" and polymeric plutonium was estimated by this
method.  This technique may be useful in future studies of the effect of
chelating agents on plutonium in vitro.

BIOASSAY FOR PLUTONIUM

     A review of current methods for bioassay of plutonium is given by Low-
Beer366.  Collection and initial handling of samples is included.

     The history of the plutonium bioassay program at Los Alamos Scientific
Laboratory from 1944 to 1972 was reviewed by Campbell367 et al.  Methods of
urine sample collection, radiochemical separation, and counting are described.

     A number of investigators have published methods for determination of
plutonium in urine samples.  In Japan, Kara    et al. used a method of ion
exchange, and counted with a 27T gas flow counter.  When plutonium is ingested
or inhaled, fecal samples are collected for 3 to 5 days after intake.  In France,
Ventadour    et al. have a rapid method using a large surface area xenon-methane
proportional counter having low background noise.

     In India, Iyer and Kamath370 have developed a rapid determination of
uranium, thorium, plutonium, and americium sequentially in a single urine sample.
The actinides are precipitated with BiPOij and the hydrolysis method used for
separation.

     In Russia, Mikhailova371 et al. developed a method in which Pu(IV) as the
cupferronate is separated from macro impurities by chloroform extraction from
a nitric acid medium using zirconium carrier or no carrier.  About 3 percent
of the 239Pu remains in the aqueous phase.  After removal of the chloroform from
the organic phase, the 239Pu is precipitated with ammonia, using a lanthanum
carrier and is counted after further processing.   The plutonium can also be
electro-precipitated.  Golutvina372 et al., in determining 239Pu in the presence
of enriched uranium, used a method based on boiling urine with nitric acid and
hydrogen peroxide, concentration of isotopes by precipitation and subsequent
extraction with precipitates of bismuth phosphate (239Pu) and lantan fluoride
(enriched uranium).  The precipitates are mixed with a fluorescent compound
and counted in a layer of hard scintillator.  The efficacy of registration
of alpha-particles comprises 90 to 95%.

     In Germany, Schieferdecker373 detected plutonium in urine and feces based
on DEHPA extraction, and subsequent counting by alpha spectrometry.  Activity
amounts of 0.1 pCi can be determined.


                                      53

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      In England,  Popplewell and  Stradling374  developed  a  2  to  3 hour  deter-
 mination for plutonium in urine.   The  procedure  utilizes  the fact  that plutonium
 binds to transferrin  in solutions of  pH >6.   This  serum  protein,  M.W. 88,000,
 was filtered from aqueous solutions with appropriate membranes.  Another  rapid
 method was developed by Bates375  et al.  for determination of plutonium in urine
 within 4 hours of receipt, excluding counting time.  The  method consists  of
 evaporating a 250-ml sample to dryness with nitric  acid,  baking at 550° C to
 remove organic matter, dissolution of  the residue and removal  of interfering
 condensed phosphates by boiling with a catalyst.  The plutonium was isolated
 by an anion-exchange column and  finally  counting the plutonium on  a tray  in a
 low background counter.

      In Italy, Camera376 et al.  described a method  for  the  determination  of
 alpha activity in urine by means  of extraction,  in  a beaker, of radionuclides
 in a solution of triactylphosphine oxide (TOPO)  on  a base of Mitene 350/80
 (polyethylene).  Testa377 et al.  reported using  wet mineralization, plutonium
 coprecipitation,  a passage in an  anion-exchange  resin column,  an electroplating
 procedure and alpha counting on  an automatic  solid  state  detector.  They  report-
 ed recovery of 78 ± 22%, with a  sensitivity limit of 0.04 pCi/1 or urine,

      Campbell and Mclnroy378 discussed the various  aspects  of  handling human
 autopsy tissues and methods used  for the determination  of plutonium.  It  was
 pointed out that  5 mrem/y has been proposed as the  maximum  permissible level of
 plutonium for the general human population, which is equal  to  approximately
 three 1 ym particles of plutonium deposited per  person  per  year and the analytical
 chemist is confronted with what appears  to be an almost insurmountable task to
 detect this level of plutonium in man.   The interlaboratory calibration of count-
 ing methods, using standards of  239Pu, 238Pu,  and 236Pu plated on  stainless steel
 or samples of ashed beef bones,  liver  and lung spiked with  239Pu,  was discussed.
 Methods used for  the chemical preparation of  the ashed  tissue  samples for radio-
 metric analysis were included.

      Once analyzed the process of determining body  content  from urine data is
 just begun.   Evaluation of excretion data by  computer programs, models, and
 equations was reported by many investigators  (Henle and Bramson379, Heid380'381
 et al.,  Osanov382'383 et al.,  Snyder38",  Nelson385  and  Beach386).

 IN VIVO MEASUREMENT OF PLUTONIUM

      The detection and measurement of  plutonium  in  wounds was  the  subject of
 an article by Sharma and Somasundaram387 in 1971.   A thin Nal(Tl)  scintillation
 detector was optimized for the detection of soft x-rays for use as a  239Pu wound
 monitor.   The  resolution obtained for  17-keV  x-rays was between 60  and 65%. Data
 on monitor sensitivity,  background,  minimum detectable  239Pu activity and varia-
 tion of  the  count  rate in the  plutonium  channel  with depth  of  plutonium source
 in a simulated  human wound were presented.

     Ohlenschlaeger  and  Fromhein388  reported  on  the operational and technical
 features of  a measuring  system for the determination of radionuclides,  particu-
 larly 239Pu, in contaminated wounds.   Special consideration was given to  the
 integration  of  different  detectors and devices into a mobile and portable measur-
 ing unit with central  control.  Experimental  results gave proof of the  system
performance.
                                        54

-------
     As in vivo determination of plutonium is only as good as the standardi-
zation of the systems used for its detection; a number of authors have reported
on calibration of these systems.

     An IAEA plutonium interlaboratory calibration experiment was described
by Button    in 1973.  Calibration factors for the in vivo measurement of 238Pu
and    Pu in human lungs as a function of body size were obtained using inhaled
  Cr  and    Pd particles as simulants.  Optimum subject-detector geometry for
the in vivo measurement of low-energy photon emitters in human lungs was deter-
mined.  A mathematical model for americium and plutonium distribution in human
lungs was developed.  The performance of scintillation detectors inside a
shielded whole-body counting room was evaluated.

              390
     Tomlinson    et al. reported in 1973 on chest wall tissue measurements for
lung counting applications.  Because the half-layer for 17-keV x-rays in tissue
is only 6 to 7 mm, the effective thickness of tissue overlying the lungs was
determined by ultrasonic measurements over the second, third and fourth rib.

     In 1973, Swinth and Dean391 discussed an intercalibration program consist-
ing of tabulation of photon intensities for actinides, an intercomparison of
counting systems using a standard consisting of either a source or a phantom,
and counting of a subject or subjects who had been accidentally exposed to
plutonium.

     Tomlinson392 et al. reviewed the basic, whole body counting program for
the Mound Laboratory since December 1969,  The phoswich detection system is
described.  Detection limits were given as a function of the subject's tissue
thickness between the lungs and detectors.  For a typical subject with an
effective tissue thickness of 2.3 cm over the lungs, the system has a detection
limit of 4 nCi.

     Detection systems with various modifications of Nal(Tl) crystals and propor-
tional counters have been reported by many investigators.  The problems of
detecting low-energy photons from plutonium  deposited in the human body were
discussed by these investigators and detectors modified to attempt to obtain
direct plutonium measurement for assessing body burdens  (Yaniv393, Newton391*
et al., Ishihara395 et al.,.Tomitani and Tanaka396, Sharma397 et al., Dolgirev39i
et al., Clemente399, Boulay*00, Morsy1*01, Loessner"02, and Newton1*03 et al.).

     Swinth lf°" et al., Swinth and Ewins405,  Swinthlf06, and Moldofsky"0 7 discuss
in vivo plutonium measurements in tracheobronchial lymph nodes by means of intra-
esophogeal probes designed to minimize attenuation of the x-rays associated with
plutonium.

MONITORING INSTRUMENTATION

     As the primary hazard of plutonium to humans is inhalation of respirable
aerosols and particulates, improvements and modifications in air filters and
monitoring instrumentation have been continually under investigation.

     Davis1*08, Angel and Anderson1*09, and Ettinger"10 et al. discussed  the use
of high-efficiency particulate air  (HEPA) filters for plutonium particle  and
aerosol entrapment.

                                      53

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     Plutonium monitoring instruments for laboratory use were reported by
 Binard1*11, Hardy1*12 et al., Nichols"13, Singh and Chugh1*", and Kikkawa1*15
 et  al.

     The  instrument described by Nichols413 was found to eliminate greater
 than 90%  of  the  2,250 dpm alpha natural background activity collected in
 order  to  detect  48 RCGS of soluble 239Pu within 1-1/2 hours.  The instrument
 utilized  two ZnS scintillation detectors.  It was believed that by using
 discriminators,  lower concentrations of 239Pu could be detected.

     Singh and Chugh1*11* reported using an alpha spectrometer technique using
 a silicon surface barrier detector.  The system is capable of indicating
 alarm  at  a level of 8 MPC hours in the presence of natural activity.

     Portable plutonium detectors for use in the field were discussed by
 Burton and Anastasi1*16 who used a probe with a CaF(Eu) scintillator with a
 pulse  height analytical capability to detect fissile material in the presence
 of  background.   The absolute sensitivity is ~24 counts/min/yg/m2 of 239Pu
 on  a uniformly contaminated surface with no overburden.

     Cohen and Gundersen1*17 discussed field experience with the mobile AEC
 measurement  van.

     The  necessity of surveying large areas of land has required the develop-
 ment of instruments for aerial surveys.  Stuart**18 reported on such a system,
 set to sense the 60-keV x-ray from 21tlAm, a decay product of 21tlPu.  The
 detector  was mounted inside an Air Force helicopter and flown over known
 concentrations at the Nevada Test Site.

     Boyns and Anderson1*19 discussed an airborne spectrometer designed to
 search for ruptured plutonium capsules in event of an accident involving a
 space mission utilizing 238Pu thermoelectric generators.  A spectrometer
 capable of_detecting 2 8Pu in air below the maximum permissible concentra-
 tion of 10    yCi/cm3 without interference from naturally occurring airborne
 alpha emitters was mounted in an Airborne Radiological Monitoring System
 (ARMS).   Information was included on the aircraft positioning system, air
 sampling, and the procedures to be followed when actually using the spectrome-
eter.
                                    56

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281.   Sanders,  C.  L.,  "Carcinogenicity of Inhaled Plutonium-238 in the Rat,"
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295.  Taylor, D. M.,  Chipperfield,  A. R.,  and  James,  A.  C.,  "The Effects  of
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 296.   Sanders,  C.  L.,  "Effect  of  Fasting  on Removal of  Injected  239Pu02  by
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 298.   Bestline,  R.  W.,  Watters, R. L.,  and Lebel,  J. Z. ,  "Study  of  Translocation
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310.  McDonald, K. E., "Removal  of  Inhaled Z39Pu02  From Rat  Lung by Pulmonary
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317.  Laylee, A. M. ,  Fraser,  D.  C., and Johns,  T. J.,  "Plutonium-Contaminated
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 328.   Hempelmann,  L.  H.,  Richmond,  C.  R., and  Voely, G.  L., "Twenty-seven
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 329.   Tamplin, A.  R.  and  Cochran, T.  B.,  A Report on the Adequacy of Existing
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 335-   Jolly,  L. J., McClearen,  H. A.,  Poda,  G. A., and Walke,  W. P.,  "Treatment
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339.  Hull, A. P.,  "Sampling and Analysis  of Radioactive Substances in the
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342.  Darrall, K. G., Hammond,  G. C.,  and  Tyler,  J.  F.,  "The Determination of
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348.  Gillette, J.  H. , Curtis,  M. L., Nunn, E.  B., Frye,  J.  0., and Bishop,
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 354.   Ghysels, J. P., "Determination of Pu in the Environment by Extraction
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 368.  Kara,  T.,  Motoyama,  S.,  Tomita,  K.,  and Suzuki, N., "Bioassay Aspect for
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 369.  Ventadour,  J.,  Philippan,  A.  M., Boulay, P., and Cheviot, J.  J., "Rapid
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 376.  Camera, V., Barassi,  G.,  and Legras,  J.  J.,  "Rapid and  Simple Method  for
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 380.  Heid, K. R. ,  Jeck, J.  J.,  and Anderson,  B.  V.,  "Interpretation of Data on
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 384.   Snyder,  W.  S.,  "Method of  Interpreting  Excretion Data Which Allows
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       of Body Content of Plutonium from Urine Data," Health Phys. 24, pp.  9-
       16,  January,  1973.

 387.   Sharma,  R.  C.  and Somasundaram, C., "Detection and  Measurement  of 239Pu
       in Wounds," BARC-559,  23 p.,  1971.

 388.   Ohlenschlaeger,  L. and Fromhein, 0., "Structure and Function of a
       Medical Measuring System for Wounds," Strahlentherapie (STRAA)  146:4,
       pp.  422-432,  October,  1973 (in German).

 389.   Button,  J.,  "IAEA Plutonium Interlaboratory  Calibration Experiment,"
       COO-3382-13,  pp. 6-7,  1973.

 390.   Tomlinson,  F.  K., Brown, R.,  Anderson,  H., and Robinson,  B., "Chestwall
       Tissue Measurements for Lung Counting Applications," MLM-2047  (OP)
       6  p.,  1973.

 391.   Swinth,  K.  L.  and Dean, P.  N., "Intercalibration for Low-Energy Photon
       Measurements,"  Health  Phys. ^5:6, pp. 599-603, December,  1973.

 392.   Tomlinson,  F.  K., Brown, R.,  Anderson,  H., and Robinson,  B., "Applica-
       tion of Phoswich Detectors for Lung Counting Plutonium-238," MLM-2048
       (OP),  6 p.,  1973.

 393.   Yaniv,  S.  S.,  "Plutonium and Americium  Measurement  in Humans by X-
       and  Gamma-Ray  Spectral Analysis," WASH-1241, 94 p., April, 1973.

 394.   Newton,  D.  J.,  Rundo,  J. ,  and Taylor, B,  T., "Progress in Instrumentation
       and  Calibration Techniques for the Assessment of Lung Burdens  of  239Pu,"
       EUR-4612, pp. 469-482,  1971.

 395.   Ishihara, T., Nahara,  N.,  linama, T.?Tonaka, E., and Yashiro,  S., "Tl
       Crystal  of Large Area," NIRS-Pu-7, pp.  31-34, 1972.

396.  Tomitani, T. and Tanaka, E.,  "Large Area  Proportional Counter  for
      Assessment of Plutonium Lung  Burden," NIRS-Pu-7, pp. 34-37, 1972.

397.  Sharma, R." C., Nilsson, I.,  and Lindren,  L., "Twin  Large  Area  Propor-
      tional Flow Counter for the Assay of Plutonium in Human Lungs," AE-463,
      49 p.,  December, 1972.

398.  Dolgirev, E. I., Kaidanovsky, G., and Shamar, V., "In Vivo Counting of

                                     82

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      Transuranium Isotopes  in the Human Body - 239Pu and 2lflAm," CONF-
      720503, pp.  497-502, May,  1972.

399.  Clemente, G.  F. ,  "In Vivo  Measurements of 239Pu in Man," CONF-720503
      pp. 503-507,  May,  1972.

400.  Boulay, P.,  "Direct Measurement  of Pulmonary Plutonium Contamination,"
      Advances in  Physical and Biological Radiation Detectors,  Vienna,  IAEA,
      pp. 287-297,  1971.

401.  Morsy, S. M.,  "Direct  Methods for the Assessment of 239Pu and  235U
      Body Burdens," CONF-711104,  pp.  115-127,  1972.

402.  Loessner, V.,  "Assessment  of Low Energy Photon Emitters  in Man:
      Technical Aspects of Detection," CONF-720503,  pp.  459-463, May, 1972.

403.  Newton, D.  J., Fry,  F. A., Taylor, B, T., and Eagle, M.  C.,  "Factors
      Affecting the Assessment of  239Pu jn Vivo by External  Counting Methods,"
      CONF-711104,  pp.  83-96,  November,  1971.

404.  Swinth, K. L., Park, J.  F. ,  and  Moldofsky,  P.  J.,  "Counting Plutonium
      in the Tracheo-Bronchial Lymph Nodes," Health Phys. 22, pp. 899-904,
      June, 1972.

405.  Swinth, K. L.  and Ewins, J.  H. ,  "Biomedical Probe  Using a Fiber Optic
      Coupled Scintillator," BNWL-SA-4761,  17 p.,  1973.

406.  Swinth, K. L. , "Radiation  Instrumentation:   In Vivo Plutonium Measure-
      ments," BNWL-1751 (Pt.  2),  pp.  89-92,  April,  1973.

407.  Moldofsky, P.  J.,  "Avalanche Detector Arrays for In Vivo Measurement
      of Plutonium and  Other Low-Activity,  Low-Energy Emitters," CONF-711111,
      pp. 55-63, 1971.

408.  Davis, W., Jr.,  "High  Efficiency Particulate Air Filters:  State of the
      Art Summary  Pertaining to  Plutonia Aerosols,"  ORNL-TM-4463, 10 p.,
      April, 1974.

409.  Angel, K. C.  and  Anderson, H.  F.,  "Different Type  Glove Box Filters Used
      for 238Pu Work,"  CONF-690103-P2, pp.  981-993,  1972.

410.  Ettinger, H.,  Elder, J., and Gonzales, M.,  "Performance of Multiple
      HEPA Filters Against Plutonium Aerosols Progress Report, July  1 -
      December  31,  1972," LA-5170, 27  p., January, 1973.

411   Binard  L.,  "Air  Monitoring  in Plutonium  Laboratories,"  Safety Measure-
      ments in Nuclear  Research. Freium, G., Ed.,  Mol, Belgium, Centre  d'Elude
      de 1'Energie  Nuclaire,  pp. B.7.1-B.7.3, 1972.

412.  Hardy, R. W.,  Knowlen,  R.  B. ,  Sandifer, C.  W.,  and Plake, W. C.,
      "Personnel Plutonium Monitor," U.S. Patent  3,670,164,  June 13, 1972,
      Filed August  18,  1970.

413.  Nichols, C.  E. , "Plutonium Sensitive  Alpha  Air Monitor,"  Health Physics
                                        83

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      Operational Monitoring, _2, Willis, C. A., Ed., New York, Gordon
      and Breach, Science Pub., Inc., pp. 1117-1125, 1972.

414.  Singh, A. N. and Chugh, P. K., "Air Monitor for Continuous Measurement
      of Airborne 239Pu," CONF-701138, pp. 585-593, November, 1970.

415.  Kikkawa, M., Yashikawa, K., and Sumita, H., "Study on Routine Operation
      for Plutonium Air Monitor," NIRS-Pu-7, pp. 73-74, 1972.

416.  Burton, B. S.  and Anastasi, M. N., "Portable Plutonium Detector,"
      Anal. Instrum. 10, pp. A7-A16, 1972.

417.  Cohen, I. and Gundersen, G., "Field Experience With the AEC Measure-
      ment Van," CONF-710617-3, 23 p., 1971.

418.  Stuart, T. P., "Use of Aerial Surveys for Determining Plutonium Concen-
      tration," EGG-1183-1517, 10 p., April, 1971.

419.  Boyns, P. K. and Anderson, C. N., "Airborne Alpha Spectrometer:  Systems
      and Procedures in Support of the Apollo Program," ARMS-69.6.11, 20 p.,
      November, 1969.
                                     84

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                               AUTHOR INDEX
    Author/Reference

         A

 Aarkrog, A./127
 Adams, W. H./119.166
 Alvarez-Raines, C./217
 Ames, L. L./190
 Anderson, H. F./283
 Angel, K. C./409
 Anspaugh, L. R./184
 Argonne National Laboratory/
   50
 Atherton, D. R./262
 Atomic Energy Commission,
   WASH/35,36,39,48
 Au,  F. H. F./197
         B
 Bair,  W.  J./132,250,251,
   253,314,326
 Ballou,  J.  E./230,245,253,
   259,306
 Bates, T.  H./375
 Baxter,  D./232,239,305
 Beach, S.  A./386
 Becker,  R.  C./43,358
 Belliayer,  Y.  A./134
 Bernhardt,  D./338
 Bestline,  R.  W./298
 Biles, M.  B./28
 Binard,  L./411
 Bliss, W./196
 Boyns, P.  K./419
 Brown, C.  L./44
 Bruenger,  F.  W./136
 Bowen, V.  T./169
 Boss, M. R./99
 Boocock, G./141
 Boulay, P./400
 Budnitz, R. J./356
Buldakov, L. A./248
Burton, B. S./416
Bustad, L. K./224
Button, J./389
                                      85
     Author/Reference

          C

 Camera, V. G./376
 Campbell, E. E./367,378
 Carfagno, D. G./74,96,97,98
 Carritt,  J./226
 Cellini,  R. F./69
 Cherdyntsev, V. V./5,6
 Chipperfield, A. R./144
 Christenson, C. W./92
 Clarke, R. H./105
 Clemente, G. F./399
 Cline, J. F./207
 Cochran,  T.  H./153
 Cohen, I./417
 Conway, T. W./63
 Cowan, M./15
 Craig, D. K./256
 Cummings, S. L./205
 Curtis, M. L./121
         D
Dagle, R. E./284
Darrall, K. G./342
Daub, W. 0./49
Davis, W., Jr./408
DeBortoli, M./8
Dionne, P. J./261
Dix, G. P^/65,123
Dolgirev, E./398
Dougherty, J. H./266
Drobinski, J. G./9
Dunaway, P. B./24
Durbin, P. W./236,327
         E
Eberhardt, L. L./185,340
Eisenbud, M./12.21
Emelity, L. A./33
Engstroem, S./54
Erleksova, E. V./150

-------
 Ershov, E. B./219,220
 Ershova, Z. V./353
 Ettinger, H./410
 Feenez, R. E./143
 Finkel, M. P./241.246
 Finkle, R. D./221
 Fish, B. R./101
 Folsom, T. R./173
 Francis, C./200
Howard, E. B./279
Hudson, J./118
Hull, A. P./339
Hunt, D. C./29
Hunzinger, W./72
Hvinden, T./25
ICRP Publication/265
Ishihara, T./395
Iyer, R. S./370
 General Electric Company/52
 Ghysels, J. P./354
 Gilbert, R. 0./186
 Gillette, J. H./348
 Gofman, J. W./60
 Golutvina, M. M./372
 Gomez, L. S./302
 Gromov, V. V./171,172
Jacobson, L./203
James, A. C./272
Jaworowski, Z./249
Jee, W. S. S./149,264,359,360
Johnson, L. J./19
Jolly, L. J./335
Joshima, H./364
         H
 Haas, F. X./68
 Hajek, B. F./89
 Hakonson, T.  E./18,20
 Hale, V. Q./211
 Hamilton, D./47
 Hamilton, E.  I./357
 Hamilton, J.  G./222
 Hampson, B.  L./349
 Hanson,  W.  C./26
 Kara, T. S./368
 Hardy, E. P./22,23,30,192,106
 Hardy, R. W./412
 Hayden,  J. A./117
 Healy, J. W./195
 Held,  K.  R.7380,381
 Heine, W.  F./57
 Hemplemann, L.  H./328
 Henle, R.  C./379
Herring,  G. M./148
Hesp,  R./336
Hickman,  W.~ W./34
Hilborn,  J. W./61
Hodge, V. F./165
Hoffman,  D. C./4
                                                        K
Kapshukov, I. I./113
Kashima, M./229,277
Katz, J./225
Keely, R. B./320
Kikkawa, M./415
Klepper, B./216
Kneip, T. J./167
Kotrappa, P./62
Krey, P. W./85
Krivolutskii, D. S./198
Kubose, D. A./114,128
Kunzle-Lutz, M./311,313
Lai, M. G./125
Lafuma, J./252
Lagerquist, C. R./318.321
Langham, W. H./124,178,179,315
Lapp, R. E./55
Larson, H. V./337
Laylee, A. M./317
Leavitt, V. D./187
Lebel, J. Z./299'
                                     86

-------
 Levine, C. A./3
 Lindenbaum, A./154,237,362
 Lingren, W. E./115
 Lister, B. A. J./316
 Lloyd,. M.  H./116
 Loessner,  V./402
 Los Alamos Scientific Laboratory/
   37,90
 Low-Beer,  A./366
         Me

 McClellan, R.  0./247.332
 McCurdy, D.  E./32
 McDonald, K.  E./310
 Mclnroy, J.  F./324
 Nichols, C. E./413
 Nishita, H. E./201
 Nolibe, D./312,330
 Norwood, W. D./7
 Noshkin, V. E./158,159
 Novoselova, G. P./352
          0
 Ohlenschlaeger,  L./388
 Olafson,  J.  H./218
 Oltman,  B.  G./58
 Osanov,  D.  P=/382,  383
 Ouchi, S./168
 Ovcharenko,  E. P./243,286
         M
 Marshall,  J./170
 Massey,  P./146
 Matheson,  W.  E./73
 Matsuoka,  M./238,257,365
 Mays,  C. W./240,268,269
 Menzel,  R.  G./156
 Metivier,  H./278
 Michels, D./93
 Mikhailova,  0. A./371
 Miner, F.  J./81.82
 Mishima, J.  M. /31,42,59,102
   103
 Miyake,  Y./161
 Moldofsky,  P.  J./407
 Morsy, S.  M./401
 Morin, M./231,296
 Mork, H. M./181
 Morrow,  P.  E./228
 Moskalev,  Y./270,275
 Muntz, J.  A./138
        N
Nabors, C.  J./267
National Environmental  Research
  Center-Las Vegas/194
Nelson, I.  C./385
Nenot, J. C./294,333
Neubold, P./208,209
Nevissi, A./129
Newton, D.  J./394,403
 Palmer,  R. F./133
 Park,  J. F./254.255
 Patin, S. A./163
 Patterson, J. H./91
 Paxton,  H. C./27
 Penn,  A. W./66
.Pesternikov, V. M./271
 Pigford, T. H./46
 Pillar,  K. C./160
 Piltingsrud, H./350
 Platt, R. B./77
 Poet,  S. E./80,86
 Polzer,  W. L./lll
 Popplewell, D. S./145,319,374
 Price, K. R./182,214
 Przyborowski, S./322
 Puphal,  K. W./347
         R
Raabe, 0". G./120
Rahman, Y./307
Rasmussen, N. C./56
Rediske, J. H./204
Reichle, D. E./78
Reynolds Electrical and
  Engineering Company/193
Rhodes, D. W./87,88
Richmond, C. R./325
Romney, E. M./76,155,180
  210,212,213
Rosenthal, M. W./234,292,304
                                    87

-------
 Routson, R. C./189
 Saenz, M. D. L./l
 Sakanoue, M./188
 Sanders, C. L./280,281,282
   287,288,289,296,297,309
 Sayre, W. W./157
 Schieferdecker, H.7355,373
 Schlenker, R. A./361
 Schmid, L. C./53
 Schofield, G./334
 Schwendlman, L. C./104
 Seaborg, G. T./ll
 Sedlet, J./83
 Sehmel, G. A./94,95
 Selders, A. A./202
 Slkov, M. R./244
 Sharma, R. C./387,397
 Sill, C. W./343,344
 Silver, G. L./112.122
 Silver, W. J./191
 Singh, A. N./414
 Smith, D. D./215
 Smith, V. A./331
 Smith, V. H.7290,308
 Snyder, W. S./384
 Stara, J. P./224
 Stannard, J. N./13
 Stevens, W./137
 Stewart, K./223
 Stover, B. J./135,147,263
   264,285
 Stuart, T. P./418
 Suzuki, M./258
 Swanson, J. L./84
 Swinth, K. L./391,404,405
   406
Tomlinson, F. K./390,392
Toyoshima, E./71
Travis, J. R.779,108
Turner, G. A./139.142


         U

Ulrich, W. C./40
U.S. Environmental Protection
  Agency775


         V

Vaane, J. P./323
Vaughn, J./151
Ventadour, J./369
Volchok, H. L.710,100


         W
Wagner, R. W./291
Wahgren, M. A./130
Wallace, A./199
Wandell, J./260
Warner, E. E./70
Watters, R. L.7300,301
Weast, R. C./2
Webb, G./110
Weeks, M. H./227
Weiner, R./64
Williams, D. C./67
Wilkinson, P. N./242
Wilson, D. 0./206
Wilson, R. H.714,16,17
Winkle, R./363
Wischow, R. P./41
Wong, K. M./162,164,174,177,341
Wymer, R. G./51
Talvitie,  N.  A./345,346
Tamplin, A. R./329
Tamura, T./183
Taube, M./126
Taylor, D. M./131,140,293,295
Taylor, G. N.7152,233,273,274
  276
Testa, C./377
Thomas, C. W. 7107,109,351
Thomas, J. T./45
Todd, D. K./38
Tomitani, T./396
                                        88
                                               Yaniv, S. S./393
Zapol'skaya, N. A./235
Zlobin, V./175,176
                                                                     GPO 690— SOI/Z51

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                                    TECHNICAL REPORT DATA
                             (Please read Instructions on the reverse before completing)
 1. REPORT NO.
  EPA-600/3-76-043
             3. RECIPIENT'S ACCESSION-NO.
 4. TITLE AND SUBTITLE
  AVAILABILITY, UPTAKE AND TRANSLOCATION OF PLUTONIUM
  WITHIN BIOLOGICAL  SYSTEMS:  A Review  of the Significant
  Literature
             5. REPORT DATE
               April 1976
             6. PERFORMING ORGANIZATION CODE
 '. AUTHOR(S)
  Anita A. Mullen  and Robert E. Mosley
             8. PERFORMING ORGANIZATION REPORT NO
 9. PERFORMING ORG ANIZATION NAME AND ADDRESS
  Environmental Monitoring and Support Laboratory-LV
  U.S.  Environmental  Protection Agency
  P.  0. Box 15027
  Las Vegas, NV  89114
             10. PROGRAM ELEMENT NO.

               1FA083
             11. CONTRACT/GRANT NO.
 12. SPONSORING AGENCY NAME AND ADDRESS

  Same as above
                                                             n/a
             13. TYPE OF REPORT AND PERIOD COVERED
              Final  FY75	
                                                            14. SPONSORING AGENCY CODE
                                                              EPA-ORD
                                                             Office  of Health and Ecologi-
 15. SUPPLEMENTARY NOTES
 16. ABSTRACT
  This report is a  selective review of  the  literature on the  availability  of
  plutonium in the  environment and its  cycling throughout representative biological
  systems ranging from large biomes covering  hundreds of miles to the molecular
  transformations within individual cells.  No attempt was made to develop  a
  comprehensive bibliography.  Rather,  references were selected for inclusion
  as  representative documentation for the vast spectrum of material that is
  available on the  subject.
  Important general references are listed separately.  Thereafter the literature
  is  described in essay form on a subject basis.   References cited by number in
  the text are listed  in complete bibliographic form at the end of the report
  together with an  author index.  The majority of the material reviewed is  limited
  to  relatively recent publications.
                                 KEY WORDS AND DOCUMENT ANALYSIS
                  DESCRIPTORS
                                               b.lDENTIFIERS/OPEN ENDED TERMS
                           c.  COSATI Field/Group
 Chemical Analysis
 Ecology
 PhysicoChemical Properties
 Plutonium Isotopes
 Radiation Effects
 Radiation Hazards
  biological accumulation
  environmental
    contaminants
06F
06R
06T
07D
18B
 3. DISTRIBUTION STATEMENT

 RELEASE TO PUBLIC
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