EPA-600/3-77-037
April 1977
Ecological Research Series
             A  CONTROLLED  BIOASSAY SYSTEM  FOR
                     MEASURING  TOXICITY  OF  HEAVY
                                                  METALS
                                        Environmental Research Laboratory
                                       Office of Research and Development
                                      U.S. Environmental Protection Agency
                                             Duluth, Minnesota 55804

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                 RESEARCH REPORTING SERIES

Research reports of the Office of Research and Development, U.S. Environmental
Protection Agency, have been grouped into nine series. These nine broad cate-
gories were established to facilitate further development and application of en-
vironmental technology. Elimination  of  traditional grouping was consciously
planned to foster technology transfer and a maximum interface in related fields.
The nine series are:

      1.   Environmental Health Effects Research
      2.   Environmental Protection Technology
      3.   Ecological Research
      4.   Environmental Monitoring
      5.   Socioeconomic  Environmental Studies
      6.   Scientific and Technical  Assessment Reports (STAR)
      7.   Interagency Energy-Environment Research and Development
      8.   "Special" Reports
      9.   Miscellaneous Reports

This report has been assigned to the ECOLOGICAL RESEARCH series. This series
describes  research on the effects of pollution on  humans, plant and animal spe-
cies, and materials. Problems are assessed for their long- and short-term influ-
ences. Investigations include formation, transport, and pathway studies to deter-
mine the fate of pollutants and their effects. This work provides the technical basis
for setting  standards to minimize undesirable changes in living organisms in the
aquatic, terrestrial, and atmospheric environments.
This document is available to the public through the National Technical Informa-
tion Service, Springfield, Virginia 22161.

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                                                EPA-600/3-77-037
                                                April  1977
         A CONTROLLED BIOASSAY SYSTEM FOR

        MEASURING TOXICITY OF HEAVY METALS
                        By

           K. H. Mancy and H.  E. Allen
Department of Environmental and Industrial Health
             School of Public  Health
            The University of  Michigan
            Ann Arbor, Michigan,  48109
            Contract Number 14-12-591
                 Project Officer
                  Gary E. Glass
    Environmental Research Laboratory-Duluth
            Duluth, Minnesota  55804
    ENVIRONMENTAL RESEARCH LABORATORY-DULUTH
       OFFICE OF RESEARCH AND DEVELOPMENT
      U.S. ENVIRONMENTAL PROTECTION AGENCY
            DULUTH, MINNESOTA  55804

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                            DISCLAIMER
This report has been reviewed by the Environmental Research Laboratory-
Duluth, U.S. Environmental Protection Agency, and approved for publica-
tion.  Approval does not signify that the contents necessarily reflect
the views and policies of the U.S. Environmental Protection Agency, nor
does mention of trade names or commercial products constitute endorsement
or recommendation for use.
                                   ii

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                                  FOREWORD
     Our nation's freshwaters are vital for all animals and plants, yet our
diverse uses of water	for recreation, food, energy, transportation, and
industry	physically and chemically alter lakes, rivers, and streams.  Such
alterations threaten terrestrial organisms, as well as those living in water.
The Environmental Research Laboratory in Duluth, Minnesota develops methods,
conducts laboratory and field studies, and extrapolates research findings

     —to determine how physical and chemical pollution affects
       aquatic life

     —to assess the effects of ecosystems on pollutants

     —to predict effects of pollutants on large lakes through
       use of models

     —to measure bioaccumulation of pollutants in aquatic
       organisms that are consumed by other animals, including
       man

     This report defines the important parameters that must be measured or
controlled when heavy metals are introduced to the aquatic environment for
the purpose of determining their impact on that system.  The toxic response
of an organism is a function of the chemical forms of metals and their
availability to the organism.  System parameters which regulate the responses
must be studied for a complete understanding of the bioassay system.

                                      Donald I. Mount, Ph.D.
                                      Director
                                      Environmental Research Laboratory
                                      Duluth, Minnesota
                                     111

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                                ABSTRACT
Biological availability of metal micronutrients and metal toxicity are
believed to be dependent on metal oxidation state, complexation, and
solubility, as well as the physicochemical characteristics of the aqueous
phase.  Basic design criteria for fish bioassays which are capable of
elucidating the dependency of toxicity on the type and concentration of
various copper species were developed utilizing equilibrium chemical con-
cepts and appropriate analytical techniques.

In order to maintain a desired copper species in the bioassay medium,
synthetic waters were used under well defined physical and chemical conditions.
These solutions were synthesized in  accordance with equilibrium models, which
define the distribution of various copper species as a function of the solution's
physical and chemical characteristics.  An experimental system was developed
which permitted large volumes of the bioassay waters to be maintained at the
desired chemical equilibria for the duration of the experiment.

Monitoring of the bioassay system included measurements of (a) pH, (b) temper-
ature, (c) flow, (d) specific conductance, (e) calcium, (f) total alkalinity,
(g) dissolved oxygen, and (h) copper species.  Novel analytical procedures
were applied for the measurement and the differentiation of copper species.
This included the use of anodic stripping voltammetry and potentiometric mem-
bran electrodes.

The bioassay experimental system was tested  by performing toxicity tests at
pH 5.7, 7.0, and 8.1.  Results demonstrated the feasibility of conducting
bioassay experiments under conditions of chemical equilibria.

This report was submitted in fulfillment of Contract Number 14-12-591, by The
University of Michigan, under the sponsorship of the Environmental Protection
Agency.
                                     iv

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                                 CONTENTS
Foreword	ill
Abstract	   ±v
List of Figures	   vi
List of Tables	viii
Acknowledgments	   ix

     I  Conclusions	    1
    II  Recommendations	    2
   III  Introduction 	    3
    IV  Aqueous Chemistry of Copper	   14
     V  Analytical Chemistry of Aqueous Copper 	   32
    VI  Development of the Bioassay System	   58
   VII  Design of Bioassay Experiment	   69

References	   88
Appendices

     A  Copper Equilibria Computer Program Output	   96
     B  pH and Alkalinity Computer Program Output	103
                                    v

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                              FIGURES


No.                                                                Page

 1   Forms of Occurrence of Metal Species in Natural Waters          6

 2   Activity of Cupric Ion in Equilibrium with (a) CuO(s);
     (b) Cu2(OH)2C03  at Bicarbonate Ion Concentration of
     10~3 M; and (c) Cu3^0H)2(C03)2 at Bicarbonate Ion
     Concentration of 10~3 M                                        25

 3   Equilibrium Diagram for Copper                                 26

 4   Stability Field Diagram for Tenorite and Malachite as a
     Function of pH and Total Carbonate Concentration at
     Various Copper Concentrations                                  29

 5   Analytical Scheme for Anodic Stripping Voltammetric
     Determination of Metals in Natural Waters                      44

 6   Anodic Stripping Voltammetric Characterization of
     Metallic Species                                               46

 7   Cell and Electrode Arrangement for Anodic Stripping
     Voltammetry                                                    48

 8   The Effect of Calcium and pH on the Variation of the
     Peak Current During the Titration of 5 yg/ml Humic
     Acid with Copper                                               50

 9   The Effect of Calcium and pH on the Variation of the
     Peak Current During the Titration of 5 ug/ml Huraic
     Acid with Cadmium                                              51

                                   5 +
10   Graphical Representation of Cu^ -Carbonate Ion
     Titrations                                                     55

11   Physicochemical and Biological Interrelationships in
     Bioassay Experiments                                           59
                                  vi

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No.                                                               Page

12   Partial Pressure of Carbon Dioxide and Amount of
     Sodium Bicarbonate (Moles/Liter) Required to Maintain
     Total Carbonate = 10~2 Molar or Total Alkalinity =
     10 2 Equivalents/Liter                                        67

13   Partial Pressure of Carbon Dioxide and Amount of
     Sodium Bicarbonate (Moles/Liter) Required to Maintain
     Total Carbonate = 10~3 Molar or Total Alkalinity =
     10 3 Equivalents/Liter                                        68

14   Change of ASV Signal for Zn2+, Cu2+, and Pb2+ with Time       71

15   Schematic Diagram of Hatchery                                 73

16   Schematic Diagram of Fish-Conditioning Area                   74

17   Schematic Diagram of Bioassay Area                            80

18   Schematic Diagram of Bioassay System                          81

19   Bioassay Exposure Chamber                                     82
                                  vii

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                                 TABLES

No.                                                                 Page

 1   Distribution of Soluble Copper as a Function of
     pH and Total Carbonate                                          30

 2   Sensitivities of Instrumental Methods for Monitoring
     Metals in Water                                                 34

 3   Summary of Trace Metal-Humic Acid-Calcium Interactions
     as Observed During Titrations with Metal Ions                   52

 4   Equilibrium and Stability Constants                             57

 5   Random Sequence for Adding Fish to the Chambers                 77

 6   Ra'.idom Order for Placing Fish in Bioassay Chambers              78

 7   Summary of Bioassay Physicochemical Data                        83

 8   Summary of Fish Survival Data and Copper Measurements           84
                                  viii

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                             ACKNOWLEDGMENTS

The authors wish to acknowledge the valuable  contributions of the
students and staff of the Environmental Chemistry  Program, whose active
participation was essential for the completion of  this  study.   Special
thanks are due to J. Black, D.  Minicucci,  R.  Roberts, T.  O'Shea. D.
Johnson, B. Gulp, J. Twork, and R.  Lasheen.
In addition, we wish to acknowledge the contributions of  Dr. M. Bender,
Dr. W. Matson, and Dr. P. Colby during the initial phase  of  this study.
The authors also wish to acknowledge the continued interest  and helpful
suggestions of the Project Officer, Dr. Gary  E. Glass.
                                  ix

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                                SECTION I
                               CONCLUSIONS

It is imperative that rigorous physicochemical controls are applied in
heavy metal bioassay experiments, e.g., copper toxicity to fish,  in
order to elucidate the dependency of toxicity on the type and concen-
tration of given metal species.  Equilibrium chemical concepts can be
used to prescribe appropriate controls which must be verified by  appro-
priate analytical techniques.
Novel analytical procedures for the differentiation of copper species
were applied utilizing anodic stripping voltammetry and potentiometric
membrane electrodes.  These procedures are also applicable for trace
metal characterization in natural and waste waters.
Anodic stripping titration techniques were used to assess the competi-
tive interaction between trace metals (e.g., Cu, Pb, and Cd) and  alka-
line earth metals (e.g. , Ca and Mg) for organic complex formation.  Our
findings showed that copper exhibits higher tendency for organic  com-
plex formation in comparison to other trace metals.  Copper affinity
for organic matter is not affected by high levels of calcium in con-
trast to other trace metals, e.g., Cd.  Understanding this type of metal
interaction provides an insight into metal transformations in the
aquatic environment and their toxicity effects.

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                               SECTION II
                             RECOMMENDATIONS

Although water quality standards give tolerance limits in terms of total
metal concentrations, the toxicity of different metal species should be
ascertained.  This information is essential for the development of
toxicity models including physical, chemical, and biological interac-
tions.  These toxicity models may be able to predict the toxic effect
of certain metal species on a given aquatic organism under specified
environmental conditions.
Generally, current metal bioassay practices do not take into account
metal oxidation states, complexation, and solubility.  It is therefore
recommended that metal bioassay experiments be conducted under well-
defined physicochemical characteristics of the bioassay medium.
It is recommended that adequate analytical measurements be made so that
meaningful bioassay experiments can be conducted.  Such measurements
are required to insure that the added toxicant is present in the bio-
assay medium at the specified level,  and to define the physical and
chemical characteristics of the medium.
In view of the increasing awareness of the complexity of chemical trans-
formations in the aquatic environment, it is recommended that efforts
be made to develop reliable analytical systems capable of metal species
characterization.   Such analytical tools are essential for the under-
standing and prediction of toxicity to aquatic biota.

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                              SECTION III
                              INTRODUCTION

Fish bioassays are perhaps the oldest known procedures for determining
water "safety" and for detecting harmful toxicants.  The use of fish
to indicate the presence of polluting materials is similar to the 19th
century practice of using canaries to monitor the presence of toxic
gases in coal mines.  In Europe the toxicity of waste effluents is
commonly monitored by placing a known number of fish in a net or screen
box in the effluent stream or the receiving water and observing their
survival at daily intervals.  Consequently, fish bioassay tests can be
considered as nonspecific analytical procedures that use fish as in
situ biological sensors.
At the present time water quality criteria and standards are based
largely on data from bioassay and toxicity tests with fish and other
organisms.  In fact, acute and chronic toxicity experiments are being
increasingly used to establish guidelines for "safe" levels of toxicants
and other physical and chemical characteristics of the aquatic environ-
ment.  Because of the complexity of environmental factors of most nat-
ural aquatic systems, laboratory bioassay experiments, which permit the
maintenance of controlled conditions, are frequently preferred over
field experiments to study the effect of a given toxicant.  Neverthe-
less, the ultimate proof of results of laboratory bioassay experiments
must come from studies on natural aquatic communities.
In this report we describe procedures for conducting controlled labor-
atory bioassays.  The main thrust of these studies has been the provi-

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 sion  of rigorous physical and chemical controls of the bioassay experi-
 ment.  Only under  these conditions can the biological response be quan-
 tified and understood.  The unique aspect of the work was the attempt
 to  determine  the toxicity of different forms of the toxic element,
 rather than the effect of total concentration per se.  Thus, much
 effort was devoted to understanding the aqueous chemistry of copper and
 the development of analytical procedures for characterizing the various
 metal species.  We hope that the reported studies will offer a fresh
 outlook on the toxicity of heavy metals to fishes and will aid in
 improving the design of bioassay studies.
 The role of heavy metals in water quality and their pollutional effects
 on  lakes, estuaries, and marine environments have received national
 attention.  Metals such as copper, cadmium, zinc, and cobalt, frequently
 found in natural waters in the parts-per-billion range, may serve as
 essential micronutrients for enzymatic transformations in trace quanti-
 ties and may exert toxic effects on the aquatic biota at higher concen-
 trations.  Lead and mercury, in particular, are toxic at certain con-
 centrations and are considered to be direct signs of man-made pollution.
 The effect of trace metals on the quality of natural waters is dependent
 on  the distribution dynamics of these metals in the aquatic environment
 and the types of interactions with the aquatic habitat.   Meaningful
 investigations should be based on the characterization of the types and
 transport dynamics of metals in solution, whether they are adsorbed on
 suspended particulate matter,bound by or released from bottom sediments.
 The physicochemical characteristics of the aqueous phase and the avail-
 ability of organic ligands of natural or  pollutional origin greatly
 influence the distribution and effect of  trace metals on aquatic biota.
The establishment of tolerance limits and water quality standards has
been based on the assumption that the availability of metal micronutri-
 ents and the toxicity of metals to biological systems are solely depen-

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dent on metal concentration.  This assumption is not valid, since the
availability of metal micronutrients to biological systems and metal
toxicity effects are highly dependent on the degree and type of metal
complex formation.  The biological response to trace metals in aquatic
environments is greatly influenced by the presence of metal-binding
ligands of natural or pollutional origin and the form of the metal
species, that is, whether they are in solution or in the form of col-
loidal particulates.
Metals in aqueous environments are conceived to exist in a variety of
forms.  To differentiate simply between soluble and insoluble metals is
not easy.  Workers in the field frequently adopt an operational defini-
tion of "soluble" and "insoluble" metals by using 0.45y or O.ly membrane
filters.
Metal species in aqueous systems can be categorized as follows:
        I.  Soluble species
            1 - Free ions
            2 - Metal complexes with organic or inorganic ligands
       II.  Insoluble particulates
            1 - Colloidal particulates of metal complexes or
                aggregates of hydrous metal oxides
            2 - Metal complexes adsorbed on suspended particulates
An illustration of size distribution and the variety of species in the
soluble, colloidal, and particulate range is shown in Figure 1, reported
by Stumm and Bilinski    .  For example, soluble metals can exist as
 (a) inorganic complexes and ion  pairs, with the ligands such as
OH~, 02~, HC03~, COg"", Cl~, HPO^~, P0^~, S0^~, HS~, S2~, etc.; (b)
organic complexes with amino acids, polysaccharides, amino sugars,
fatty acids, organophosphorus compounds, porphyrin, etc.; and (c) poly-
nuclear complexes.

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< Membrane liltrable ^




l:rcc
metal
ions

Inorganic ion
pairs; inor-
ganic complexes

Diameter range: «
Organic com-
plexes ,
chelates

Metal species
complex bound
to high molec-
ular org. m£»t .
Metal species in
the form of
highly dispersed
colloids
Metal species
sorbed on
colloids



Examples :
Cu.aq
Fe.aq3*

Pb.aq3+




C»2(OH)2+
Pb(CO.)*'

CuCO-
AgSll
CdCl*
CaOH*
Zn(OH)3
Mc-SR
Mc-OOCR

cn,-c=o
M/ 2 \
,Cu
°\ /">
0— C — CH2
Me-lipids
Me— humic— acid
polymers

"lakes"
"Gelbstoffe"
Me polysac-
charides

PeOOU
Fc(OH)3

Mn(N) oxides
Mn 0 0
Na Mn 0
A- S
2
Mcx(OH)y,
MeCOj, MeS

etc . on clays ,
FeOOH or on
Mn(N) oxides


A«2S3fI2~
Figure 1.  Forms of occurrence of metal species in natural waters
                                                                 (1)

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Metal organic interactions have been mentioned repeatedly in recent
literature to explain anomalous metal distribution that cannot be ac-
counted for by known thermodynamic data   »»»»»»  _   Certain
organic constituents have been postulated to complex with certain
metals, and as a result the availability or toxicity of certain metals
to biological systems is altered.  Needless to say, some of these
deductions have been substantiated by the fact that a great many metal
species in natural waters are nonextractable with suitable reagents,
nonseparable by ion exchangers, and nondialysable.
The literature presents diverse opinions on the subject of aquatic metal
                                                                    (9)
distribution and organic-metal interactions.  Corcoran and Alexander
provided indirect evidence for the existence of trace metal organic
complexes in sea water by employing perchloric acid digestion.  More
direct evidence was provided by dialysis and neutron activation analy-
sis experiments performed by Slowey and Hood     and Slowey and
Jeffrey     .  More recently, Williams     has demonstrated the exis-
tence of organic copper complexes in sea water, and evidence for the
existence of organic complexes with other trace metals, particularly
                                            (13)
iron, has been reported by Koshy and Ganguly     and Koshy, Desai, and
Ganguly    .  Duursma and Sevenhuysen   , on the other hand, found that
trace metal-organic interactions are rather insignificant.  Recent
radio tracer studies indicate that certain organic compounds (leucine)
exhibit no chelation with trace metals at the levels normally present
in the sea    .  The existence of trace metal-organic complexes in
fresh water has been suggested by the anodic stripping studies of
Barsdate and Matson     , Matson and Roe     , and Allen, Matson, and
Mancy    .  Barsdate     provided further evidence through dialysis
studies.
The biological importance of trace metals chelated by organic substances
has also received attention.  DeKock     showed that iron became readily
available to plant cells and stimulated growth in the presence of humic

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 acids.  This  evidence  of  the  stimulatory  effect  of  chelated  iron-humic
                                                     (21)
 acid  substances was  later used by Prakash and Rashid      to  partially
 explain the increase in growth of marine  dinoflagellates  in  the pre-
                               (22)
 sence of humic acids.  Johnston   "   found that the  addition  of a mix-
 ture  of iron, manganese,  zinc, copper, and cobalt chelated with EDTA
 at pH 7-8  induced abundant growth of unialgal Skeletonema costaturo in
 most  sea waters enriched with phosphate,  nitrate, and silicate.  Jones
 (23)
      found that Escherichia coli does not grow in sea water  enriched
 with  glucose, ammonium, and phosphate ions unless the water  is treated
 with  chelating agents.  The inhibitory effect was attributed to the
 toxicity of heavy metal ions present in the water.  Enhancement of
 biological growth in the presence of chelating agents, such  as EDTA,
                                                    (24)
 has been further substantiated by Barber  and Ryther    , who found that
 the presence  of chelators increased phytoplankton growth  in  the Crom-
 well  Current.  Recently, the effect of nitrilotriacetic acid, a strong
 chelator, on  the growth of estuarine phytoplankton has been  reported by
                              (25)
 Erickson, Maloney, and Gentile
 Although the  literature suggests that some aquatic organisms can selec-
 tively concentrate certain metal ions by  secreting certain chelating
 agents, there is no  direct evidence in support of the predominance of
 such  trace organometallics in the aquatic environment.  It is not quite
 understood how organic matter in trace quantities can selectively
 interact with individual heavy metals, e.g.  Cu and Zn, in the presence
                                                          (26)
 of a  large excess of alkaline earth metals,  e.g. Ca and Mg    .  The
 question takes on added significance in view of  the toxicity studies
                     (27)       (28)                         (29)
 of Cairns and Scheier     , Wurtz     , Pickering  and Henderson    , and
Mount    .  These workers found that the  toxicity of zinc and copper
 to bluegills, freshwater mollusks, and minnows was less in hard water
 than  in soft water.   Although the mechanism of this inhibition is not
known, chemical interactions of the alkaline earths and trace organics
cannot be overlooked.

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In summary, considerable evidence indicates that trace metals in natu-
ral waters are, to a certain extent, present as complex species associ-
ated with trace organic materials.  Further evidence is available to
relate these complex organometallics to biological activity and the
toxic effects of trace metals in natural media.  However, little in-
formation is available on the dynamics of trace metals in natural
aquatic media and on the distribution and characterization of the
organometallics associated with these trace metals.  Furthermore, the
role of alkaline earth ions in the complexation of these trace metals
is presently only a matter of speculation.

TOXICITY OF METALS TO FISH
    (31)
Chen     discussed in his review how metals act in several ways as
poisons to fish.  The metals enter the organism in either of two ways
— through the gills while respiring or through the mouth with swallow-
ing,  The entrance of ionic poisons into the gills may be aided by the
excretion of external enzymes to facilitate the active transport of
ions against a concentration gradient.  Then, the gill epithelium is
destroyed by the action of certain heavy metals.  Therefore, the fish
dies of asphyxiation.
Toxic metals may travel to different parts of the body via the blood-
stream.  The poisons may react with cell protoplasm of various tissues
or organs.  Insofar as copper toxicity is concerned, a lack of agree-
ment exists concerning the actual "safe level" concentration in water
(32)
       Copper concentrations varying from 0.1 to 1.0 mg/1 have been
found by various investigators to be toxic.  On the other hand, con-
centrations of 0.015 to 3.0 mg/1 were reported as toxic, particularly
in soft water, to many kinds of fish,
EFFECTS OF CHELATORS ON TOXICITY
Various investigators seem to disagree on the influence of chelation
on metal toxicity to fish.  For example, a study of zinc toxicity to

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              (33)
 fish by  Lloyd      indicated  that  suspended  zinc  sulfate  complexes
 decreased  the  survival  time  of  rainbow  trout  in  concentrated  solutions.
           (34)
 Fitzgerald      showed that citric acid  chelation reduced  toxicity of
                                    (35)
 copper to minnows  500-fold.  Grande      in  his investigations with
 copper in several  Norwegian  lakes showed that one lake heavily colored
 with humic  colloids was less toxic to salmonid fishes than  the non-
 colored  lakes,  even though both lakes contained  the  same  copper concen-
 trations.   This phenomenon was  described as a masking effect by the
 organics on the metal.  Masking effect  refers to the organics' ability
 to reduce or "hide" the toxicity  of copper  to fish.  This apparently
 depends upon the type of metal  complex  present in the water system
 which in turn determines the metal's toxicity to organisms.
                         ( 36}
 Nielsen and Wium-Andersen     in  their  investigations of  copper toxi-
 city to plankton found that various organic substances also mask the
 effects of  metals  on algae.  The  organics appear  to  be polypeptides
                                                      (2)
 and other similar  substances.   Duursma  and Sevenhuysen    determined
 that other  organics (phenols, petroleum, and carbon  disulfide) present
 in the natural  system reduced the amount of free  copper in sea water.
 It is apparent  from these investigations that organic chelators defin-
 itely affect free  copper concentrations  in the aqueous system.  Whether
 the chelators enhance or detract  from metal toxicity is still uncertain.
SOURCES OF CHELATING AND SEQUESTERING AGENTS
The major chelating and sequestering agents found in the aquatic envi-
ronment include plant products, microbial metabolites, humic compounds,
lipid soluble fractions, detergents, and sewage effluent.  These sources
of complexing agents are discussed below.
                    (37)
In a review by Pauli     concern was given to the exogenic cycle of
matter and the manner of occurrence and distribution of minerals in the
outer surface of the earth.  The most effective method of transporting
                                  10

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elements from the lithosphere to the biosphere, including the fresh
water cycle, was reported to involve the biosynthesis of chelating and
sequestering agents.  These agents were classified as plant products,
microbial metabolites, or humic compounds,  Chelating agents produced
by organisms include lichen acids such as lakmus and orcein, which are
plant-derived polyphenols with two or more aromatic rings.
Humic compounds are derived from decomposition of organic residues
through biological activity and heterogenous catalysis.  The humic
compounds are collectors of copper, zinc, nickel, cadmium, lead, man-
ganese, and cobalt.  These chelates formed from humic substances and
trapped metals become available for use by various organisms in the
terrestrial and fresh water biocycle,
Slowey and Jeffrey     experimented primarily with copper complexes
found in sea water.  The largest amounts of extractable copper were
found at depths where sinking, decaying organisms were most likely to
release organic compounds into sea water.  Rupture of the phytoplankton
and zooplankton releases some of the organic complexing compounds to
the water.  Although the nature of these complexes is not resolved,
some evidence indicates that the copper complex is associated with the
phospholipid, amino lipid, or porphyrin fraction of these lipids.  Al-
though these studies concern sea water, fresh waters probably contain
similar complexing organics.
        (12)
Williams     demonstrated that organically associated copper may be
complexed with phospholipids, carotenoids, or other lipid-soluble frac-
tions.  However, it is not clear whether the copper is actually a mo-
lecular complex with an organic ligand or whether it is associated with
organic colloids by bonding within an organic matrix.
Detergents are obviously a significant component of domestic sewage.
                  (38)
According to Hynes    , raw sewage contains about 12 ppm of active
detergent.  A significant proportion of detergents remains undegraded
and eventually gets into water bodies.  Detergent builders are strong
complexing agents which must be considered in studies of ligands in
natural waters.
                                  11

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 EFFECT OF ALKALINE EARTH METALS ON METAL TOXICITY
 A first observation of  the metallic content of  natural water reveals  a
 predominance of  alkalies and alkaline earth metals.   Stumm and Morgan
 (26}
      were therefore prompted to pose the following questions:   (1) How
 can trace concentrations of  organic matter specifically interact  with
 individual metals? and  (2) Does not the presence of  Ca2  and Mg2  at
 concentrations many orders of magnitude larger than  potential organic
 complex-forming  species blur any complex-forming tendency of the  or-
 ganic functional groups? These pertinent questions  cannot be answered
 from  present available  information on trace metals in the aquatic
 environment.
 The effect of high concentrations  of  alkaline  earth  metals on transi-
 tion  or heavy metal-organic  complex  formation  can be illustrated  by
 considering the  effect  of Ca2  and Mg2   on EDTA  complex formation with
 Fe(III).   The formation of Fe(III)-EDTA complexes is given as
                 Fe3+ +  Y4~ = FeY~;    log K    =  25.1                (1)
The effect of Ca2  can be expressed as
                                               KFeY
            CaY2  + Fe3+ = FeY  + Ca2+;   log — - =14.4           (2)
where log K  Y = 10.7.
Based on the above relationships, FeY  cannot be formed to any signi-
ficant extent if the ratio  of  [Ca2 ] to  [Fe3 ] is sufficiently high.
This effect is of course dependent on the relative stability of the
formed complexes.  For example, in the case of L = citrate4" the
equilibrium
                      CaHl" + Fe3+ = FeHL -1- Ca2"1"                     (3)
is characterized by an equilibrium constant of log (1L., m /KC   ) = 7.4.
This indicates that the competitive effect of Ca2+ upon Fe3"1" complexing
by citrate is much smaller than that encountered with EDTA.
                                  12

-------
Numerous investigators have studied the effect of hardness by measuring
the toxicity of metal solutions with increasing hardness, and several
theories have been advanced to account for their results.  Early
workers attributed the decreased toxicity of solutions at high hard-
ness to complexation and precipitation of the metal ions from the
solution.  However, with the advent of continuous-flow bioassay tech-
niques, it has been found that some complexation may actually increase
the toxicity of metal solutions    .  Mount's study showed that at any
given hardness level, as pH increased, the solutions became more toxic.
He attributed this effect to the zinc complexes formed at higher pH
levels.
One of the major difficulties in interpreting the effect of hardness
on heavy metal toxicity stems from the inability to determine whether
the observed effects are a result of calcium-heavy metals competitive
interactions or are due to physiological effects that resulted from
                  0+      •) +
the presence of Ca   or Mg^ , or both.  For example, the effect of
hardness could be explained in terms of purely aqueous chemical inter-
actions.  Under a given set of conditions, certain heavy metals can
                                                                9 +
exist as soluble organic complex species.  At relatively high Ca^
concentrations, as a result of metal-ligand exchange reactions, these
heavy metals may be transformed into inorganic metal complexes that
may precipitate as hydroxy compounds at high pH.  This effect will be
more pronounced at higher levels of "hardness" and of course is largely
dependent on the pH.  Consequently, if the toxicity effects were pri-
marily due to the organic-metal complex, the presence of high levels
of calcium will result in a decrease of its toxicity.
On the other hand, the presence of high levels of Ca2  and Mg2  has
been thought to cause certain physiological changes in fishes that may
result in a decrease in the toxic effects of heavy metals.  This was
              (33)
shown by Lloyd     , who found that preconditioning of fish in hard
water will significantly decrease the animal's susceptibility to toxic
metals.
                                  13

-------
                               SECTION  IV
                      AQUEOUS  CHEMISTRY OF COPPER

The presence of  copper  in water  is ecologically significant because of
its extreme toxicity  to fish and other aquatic organisms.  Workers
have studied the toxicity of copper, but none have investigated the
relationship between  the speciation of copper and its toxicity.  A
comprehensive study of  the  toxicity of copper is required to provide
the information  necessary for  evaluating its environmental impact.

SOURCES
Most of the copper entering the aqueous environment is due to the
activities of man.  Natural weathering of minerals and the resulting
runoff, which contains  copper, contribute an almost insignificant
fraction of the  total copper input.  The copper washed out of the
atmosphere by precipitation is significant.  Lazrus, Lorange, and
     (39)
Lodge     found  an average of  21 Mg/1 Cu in precipitation in the
United States.   Kramer     found an average of 32 yg/1 total copper
and 22 yg/1 soluble copper in precipitation collected at 36 stations
in northern Ontario.  Although the copper reaching bodies of water by
precipitation is a natural occurrence, a large fraction of the copper
in the atmosphere is not of natural origin.
Large amounts of copper enter the natural water system from mining
operations and metallurgic processes.   The waste water from copper-
bearing acid mine drainage is usually stored in tailings ponds after
neutralizing the acidity.   However, the treated water in the tailings
                                                            (41)
pond may contain concentrations of copper as high as 50 ug/1    .   The
                                  14

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drainage from agina mines may have pH values as Low as 2,5.   The acid-
ity of the drainage water has been attributed to the metabolic activity
of certain bacteria, which enhance the oxidation rate of sulfide parti-
                                                          (42)
cles by a factor of about 2,000 at the optimum temperature    .   Pro-
      (43)
kovich     found concentrations of copper in water near old mines to be
as high as 1,000 mg/1.
Treatment of water with copper sulfate for control of algal growth has
                               (44 45 46)
been practiced since about 1900   '  '   .  In some cases lakes have
been treated with as much as 1.5 million pounds of copper sulfate over
                           (47)
a period of several decades
FORMATION OF COMPLEXES
The results from many studies indicate that copper in both fresh water
and sea water is held in solution by complexation with naturally
                                        (18}
occurring organic ligands.  Allen et al.     added 13 yg/1 of copper to
water from the Rouge River.  Results of analysis using anodic stripping
voltammetry indicated a logarithmic decrease in peak current with time.
The results were interpreted as a first-order complexation reaction of
copper and excess organic ligands in the river water.
Stability constants for copper hydroxide, oxide (tenorite), and basic
carbonates (azurite, Cu3(OH)2(C03)2; malachite, Cu2(OH)2C03) indicate
that the equilibrium concentration of copper in water with pH greater
than 7 would be very low.  In some waters, however, the amount of cop-
per apparently in solution exceeds theoretical limits by several orders
of magnitude.  This anomaly has also been explained as a result of
complexation of copper with organic substances called yellow acids,
humic acids, and fulvic acids.  Up to 50% of the copper in sea water
can be extracted with chloroform     indicating that much of the copper
                                                         (12)
in sea water is in some organic complex or sol.  Williams     found
that 5 to 28% of the copper in sea water was present as complexes with
stability constants greater than 1018.
                                   1.5

-------
 Studies concerned with the chemistry of copper in soil have also  indi-
 cated that copper forms complexes  with naturally occurring  ligands.
 If solutions of copper are stirred with ion exchange resins in  both
 the presence and absence of humic  and fulvic acid the resin will  take
 up less copper  when humic or fulvic acid is present

 PRECIPITATION
 In the absence  of organic ligands ,  copper  is very insoluble at  the pH
 values found in most natural bodies of water (pH > 7) .  When soluble
                                                                    •
 salts of copper (Cu(NC>3)2,  CuCl2 ,  CuSO^)  are mixed with natural water
 the cupric ion  precipitates as  copper hydroxide  (Cu(OH)2) and will
 evolve to tenorite (CuO)  or to  a basic carbonate,  either malachite
 (Cu2(OH)2C03) or azurite (Cu3(OH2) (C03)2) ,  depending on the carbonate
 and copper concentration.

 COPPER IN FRESH WATER  AND SEA WATER
Allen  et al.     , using anodic stripping voltammetry , found copper con-
centrations of  19 to 108 yg/1 in Rouge River water, 1.8 to 6.8 yg/1 in
Lake Erie, and  0.4  to  19 yg/1 in Lake Michigan.  Samples were acidified
before analysis  to  release copper from organic and inorganic complexes.
Analyses without prior acidification gave much lower results indicating
that most of the copper present was either in complexed or particulate
forms.
                                                                  (49)
The copper cycle in three Connecticut lakes was described by Riley
In all of the lakes studied both total and particulate (> 0.6 y) copper
concentrations reached a sharp maximum by November.  The minimum con-
centrations in all  defined forms of copper occurred from February to
September.
Copper in sea water exists primarily in nonionic forms, and its verti-
cal distribution is sharply defined    .  A peak concentration of over
20 yg/1 occurs near the base of the euphotic zone.  The concentration
decreases with depth into the aphotic zone to about 10 yg/1, after
which it stays nearly constant with increasing depth.
                                  16.

-------
SEDIMENTS
In many lakes the concentration of copper in sediments is quite high.
The sediments have served as a sink for the copper that has been added
to lakes for algal control.  Detailed profiles of copper in the sedi-
                                                     (47)
ment of several lakes are reported by Sanchez-Vazquez    .   In all
cases the copper concentration was at a maximum in the top layers of
sediment, indicating that most of the copper is from the activities of
man.

WATER QUALITY CRITERIA
The limit set for copper according to U.S. Public Health Service has
been changed twice.  The original standard, set in 1925, was 0.2 mg/1.
In 1942 the limit was raised to 3 mg/1, and in 1962 it was lowered to
the present standard of 1 mg/1.  The toxicity of copper to human beings
is low, but the present standard was set because of the poor taste
that copper imparts to water.  However, because copper is highly toxic
to many organisms the threshold concentration of copper is reported by
              (32)
McKee and Wolf     to be 0.1 mg/1 in water used for irrigation.  For
fish and other aquatic life the desirable concentration limit is 0.02
mg/1 in soft water and 0.05 mg/1 in sea water.

EQUILIBRIUM MODEL OF Cu2+-H20-C0? SYSTEMS
Of particular interest to the study of copper toxicity is the develop-
ment of an equilibrium model describing the homogeneous as well as the
heterogeneous equilibrium distribution of copper between the insoluble
and soluble hydroxy and carbonate complexes.  The equilibrium relation-
ships for prediction of the composition of an  aquatic system are most
easily solved by numerical digital computer techniques.
      2+
A   Cu  -H20-C02   equilibrium system has been studied in our laboratory
in an attempt to illustrate the chemical relationships that must be
considered in the interpretation of copper toxicity data.  However,
great care should be exercised in extending this equilibrium approach
                                  17

-------
to the prediction of metallic species present in natural waters.
Although it is possible to measure the concentrations of inorganic
complexing agents such as hydroxide, carbonate, bicarbonate, sulfate,
chloride, and ammonia and to incorporate these into equilibrium calcu-
lations, prediction of the effect of the organic constituents is not
possible at the present time,
                        9 +
In the ternary system Cu^ -H20-C02 we are concerned with
    (a) the solid phase
               CuO, N (tenorite)
                  (s)
               CuC03*Cu(OH)2, x  (malachite)
                            \s/
               (CuC03)2'Cu(OH)2, N (azurite)
                               \s)
and with (b) the aqueous phase
               H+, OH~,  C02(H2C03C) , HC03~,  C032~, Cu2+,  CuOH+,
               Cu2(OH)22 ,  Cu(OH)3~,  Cu(OK)h2~,  CuC03,  and
               Cu(C03)22~
                                  18

-------
The following equilibria relationships occur in the copper-carbonate

      (26)
system    ;
                                                        Log K



    CuO,, + 2 H+ = Cu2+ + H20                           7.65        (4)


  (tenorite)




    Cu2(OH)2C03(g) + 4 H+ = 2 Cu2+ + 3 H20 + C02  ,     14.16


     (malachite)
or
or
    Cu2(OH)2C03    + 3 H+ = 2 Cu2+ + 2 H20 + HC03~       6.34        (5)
               \s /




    Cu3(OH)2(C03)2, ,  + 6 H+ = 3 Cu2+ + 4 H20 + 2 C02, .21.24
                  \s^                                  \&)

       (azurite)
    Cu3(OH)2(C03)2, .  + 4 H+ = 3 Cu+ + 2 H20 + 2 HC03"   5.60        (6)
                  yS /




    Cu2+ + H20 = CuOH+ + H+                            - 8           (7)




    2 Cu2+ + 2 H20 = Cu2(OH)2+ + 2 H+                  -10,95        (8)





    Cu2+ + 2 C0g~ = Cu(C03)2~ + 2 H+                    10.01        (9)





    Cu2  + CO2" = CuC03                                  6.77       (10)





    Cu2+ + 3 H20 = Cu(OH)3~ + 3 H+                     -26.3        (11)





    Cu2+ + 4 H20 = Cu(OH)J~ + 4 H+                     -39.4        (12)




    H20 + C02 = H2C03                                   -1.5        (13)





    H2C03 = H+ + HC03"                                 - 6.37       (14)
                                  19

-------
    HC03~ = H+ + C0^~                                   -10.25       (15)



    C02 + H20 = HC03~ +  H+                              - 7.82       (16)



From relationships  13, 14,  and  15  it  is  possible to solve for [HC03~]

and [C03~] in terms of [H+]  and [H2C03]:
                          [HC03  ] =	                     (17)




                                   [H2C03]
                    [HC03~]
                      CT        [HT]'
                                      [H']2



The expression for total carbonate  is:


           [Total Carbonate] = CT =  [H2C03] +  [HC03~] +  [C03~]



or
                               [H2C03] ki      [H2C03]

              C  = [H2C03] +	 +	          (19)
               •L                 r TT~ 1             r TT ' T ?
This allows us to define  the a values as:
                    [H2C03]              [H+]2
                 	—	               (20)
                                  20

-------
                      [C032~]
                &2 = ~"~    Z	J       "~              (22)
                        CL       [H+]2
This permits the calculation of [H2C03] , [HC03 ], and [C032 ] if we know
the pH and the total carbonate concentration.
In the Cu-H20-C02 system there are two distinct  equilibrium situations
at any particular pH and carbonate concentration.  The first occurs when
the [Cu  ] is not large enough to cause precipitation, and the second
when the [Cu  ] is large enough to cause the precipitation of tenorite,
malachite, or azurite.  These two situations will be discussed separ-
ately.

Case I;  No Solid Species Formed
Expressions for each of the soluble species can  be obtained be rearrang-
ing equilibria relationships 7, 8, 9, 10, 11, and 12:
            [Cu2+](10~8)                              (l(T10-95)[Cu2+]2
 [CuOH ] =	,             [Cu2(OH)i  ] =	—	
                [H ]                                        [H]2

 [Cu(C03)2~] =  (1010'01)[C02~][Cu2+],   [CuC03] =  (106-77)[CO§"][Cu2+]
(10~26-3)[Cu2+]
      v
 [Cu(OH)3] = - v - ,        [Cu(OH)J~]  =
                    [H ]3

An expression for total copper concentration  is:

  [Total Copper] = [CuOH~l"] + [Cu(C03)2.~]+[CuC03] + [Cu(OH) 3] + [Cu(OH)2.~]+[Cu2+]
                                   21

-------
 This  can also be written as;
                   [Cu2
 [Total  Copper]  - -- v -- + (101 ° -01) [Cu^~] [Cu2+]
                         (10~26«3)[Cu2+]      (10"39-l*)[Cu2"f]
+(106-77)[C02i-][Cu2+] +	T-	+	r	  +[Cu2+]  (23)
                               [H+]3                [HY

                                              2+
These  expressions  neglect  the  dimeric  Cu2(OH)2  ,  which would be signi-
ficant only  at much  higher concentrations than  those present in natural
waters.

We can now define  6  as 3   =  [Cu2  ]/[Total Copper],  which  is equal  to:
                    o      o
                                                                     (24)
      ao~8)                                   do"26-3)
     — — +(io10-01)[co2i~]2+(io6'77)[co2i]+ — — --
        +                                        "1"
This is a significant equation because  it means  that  the  cupric  ion
                 2+
concentration, Cu  , can be calculated  by knowing  the pH, carbonate
concentration, and the total copper concentration  as  follows:
                        [Cu2+] = 3  [Total Copper]
where   6  = [Cu2 ]/ [Total Copper] as defined above.
         o

If we define Denom as,
Denom = (10~8)/[H+] + (1010« 01 ) [CO2,']2 +  (106 -77) [CO2;']
                                                           +
                            (10"26'3 )/[H]3 +  (10"39-1+)/[H]t* + 1   (25)
then similar equations can be constructed that can be solved for the
concentration of each of the copper species.
                                   22

-------
The resulting £> values are:





                B  = [Cu2 ]/[Total Copper] = l/(Denom)               (26)
                 o





           BI = [CuOH+]/[Total Copper] =  (10~8)/[H+](Denom)          (27)





          32 = [CuOH3"]/[Total Copper] =  (10~26'3)/[H+]3(Denom)      (28)





        B3 = [Cu(OH)5~]/[Total Copper] =  (ICf39^)/[E+]k(Denom)      (29)





          B4 = [CuC03]/[Total Copper] =  (106'77)[C0§~]/(Denom)       (30)





       B5 =  [Cu(C03)2]/[Total Copper] =  (1010-01) [CC)|~]2/(Denom)     (31)





From these equations:




                     [Cu2+] = B  [Total Copper]                       (32)
                               o




                     [CuOH+] = 3i[Total  Copper]                      (33)




                     [Cu(OH)3~] =  62[Total Copper]                   (34)




                     [Cu(OH)fp] =  63[Total Copper]                   (35)




                     [CuC03] = 6^[Total  Copper]                      (36)




                     [Cu(C03)2~]  = B5[Total  Copper]                  (37)






Case II;  Solid Species  Present



In the presence of  a solid  copper species, whether tenorite,  malachite,


or azurite,  the cupric ion  concentration at  equilibrium will  be depen-


dent on the  pH, total  carbonate,  and solid  species present.
                                   23

-------
Figure  2  shows  the  concentration  of  cupric  ion in  equilibrium with
tenorite, malachite, and azurite  in  a bicarbonate  concentration of  10~3
M.  At  any particular pH and bicarbonate  level,  the  precipitate that
dictates  the lowest cupric  ion  concentration will  be the  predominant
solid species.  Tenorite will be  the predominant form at  pH values
greater than 6  and  a bicarbonate  concentration of  10 3 M  (Figure 3).
Once the  solid  species has  been designated, the cupric ion concentration
will be dictated by that particular  solid species:

Tenorite:       [Cu2+] = (107-65)[H+]2                               (38)
or

                   +     (107.Q8)[H+]2
Malachite:      [Cu2 ] =	j-	                             (3^
or

                   +     (107-08)[H+]2
Azurite:       [Cu2 ] =	~r-	                             (40)
                          [P   ]2 3
                            C02

The concentrations of the soluble species are then easily calculated:
       ,      (10"8)[Cu2+]                          (10~26-3)[Cu2+]
  [CuOH+] = 	r	,          [Cu(OH)3"] »	
                [H']                                   [H']3

                                                    (10'10-95)[Cu2+]2
  [Cu(OH)2."] = 	+	>    [Cu2(OH)i ]
                      ~
  [CuC03] = (106'77)[Cu2+][COp,    [Cu(C03)i~] = (1010-01)[Cu2+][C02~]2
                                   24

-------
     0
OJ
     6
6
g



-   8
i
a
u f


o   10
rH




    12
      0
                                    o
                                    o
10     12     14
                                pH
    Figure 2.   Activity of cupric ion in equilibrium with

               (a)  CuO(s); (b)  Cu2(OH)zC03  at bicarbonate

               ion  concentration of  10~3 M; and  (c)  Cu,(OH)?(C03)2

               at bicarbonate ion concentration  of  lO""^ M.
                                25

-------
    7  U
LU

Q.

0.
CS
    o
    CJ
 9



10
   12



   13
      0
                                                                 10
12
            2463


                                          pH

Figure 3.  Equilibriun diagram for copper.  Total copper concentration equals 10~6 M


           and total carbonate concentration equals 10~2 M.

-------
Construction of the Equilibrium Model
A mathematical model can now be constructed that will yield the concen-
tration of each copper species when the total copper concentration,
total carbonate concentration, and pH are known.
This model is constructed by the following steps.
                                                            ? +
    1)  Assume no solid species are present and calculate Cu   as
        explained in case I,
    2)  Assume a solid species is present and calculate Cu2  as shown
        in case II.
    3)  If the value for Cu2  obtained in step  (1) is larger than Cu2
        calculated in step (.2) , then precipitation will occur and all
        calculations should proceed as described in case II.
If the value for Cu2  obtained in step 1 is smaller than Cu2  deter-
mined in step 2, then no precipitation occurs and the concentrations of
soluble species should be computed as described in case I.
This procedure may be repeated with different pH values and carbonate
concentrations to determine the variation in copper species under dif-
ferent chemical conditions.  This variation of parameters is a repeti-
tive process, which has been readily accomplished with the use of a
computer,  Sample computer program output is shown in Appendix A.
A particularly useful method of presenting these data is to plot the pH
vs. the log of the concentration of the different copper species, hold-
ing the total copper and carbonate concentrations constant.  An example
of this procedure is shown in Figure 3.  With a total copper concentra-
tion of 10 6 M and a total carbonate concentration of 10 2 M, cupric
ion predominates from pH 0 to 6, CuCOs from pH  6 to 8.3, and tenorite
from pH 8.3 to 12.5.
In the system described the fraction of the metal present as a given
species will be independent of the total metal concentration at any pH
and carbonate value as long as the total metal concentration does not
become so great as to permit dimeric metallic complexes and providing
                                  27

-------
 that no solid species are present.  If metal precipitates form, in-
 creasing  the total metal concentration will not change the concentra-
 tion of metal in solution or the species of the metals.  Rather, only
 the quantity of solid will  increase.  The pH region over which the
 solid  is  stable is dependent on the total amount of metal in the system
 (Figure 4).  The range of stability increases as the concentration in-
 creases.   Since the dominant form of the metal at pH values less than
 that at which tenorite forms is soluble copper carbonate, it should be
 expected  that the carbonate concentration in addition to the pH would
 influence  the lower limit of tenorite stability.  This is shown in
 Figure 4 by the curved portion of the lower stability boundary.  The
 effect of  carbonate is more pronounced as the total copper decreases.
 At high copper and carbonate concentrations malachite becomes a stable
 species, but it will transform to tenorite by decreasing either the
 total copper or carbonate concentration.  Again it should be mentioned
 that equilibrium calculations must be used with caution.   The equilib-
 rium constants used in the calculations are generally obtained at
 higher metal concentrations and ionic strength than are present in the
 natural environment.  Often, predicted stable solid species are not
 found under experimental conditions.  For example, although tenorite is
 the predicted stable solid, we did not find it to form within 1 week in
 laboratory systems.  Rather, solid copper hydroxide Cu(OH)2 was present.
 With the same model the predominance of any given copper  species as a
 function of pH and carbonate content can be calculated.   This is illus-
 trated in Table 1 for a total copper concentration of 1 x 10 6 M.
 Such equilibrium calculations are considered useful for toxicity studies.
 They can be used to predict the form of copper under defined experi-
 mental conditions.   This is particularly relevant since available ana-
 lytical techniques have not been refined enough to distinguish between
metal complexes in solution.
                                  28

-------
  2 —

  3
8
bo
o
   7
      T
                           TENORITE
         COPPER
         CONC,
         (MOLAR)
                                                     10
                                                       -5
                                                         10
                                                           -4
                             10
                                -6
                         10
                           -7
    0  5
7
 9
pH
11
13
     Figure 4.  Stability  field diagram for tenorite and
                malachite  as a  function of pH  and total
                carbonate  concentration at various copper
                concentrations.
                                29

-------
             Table 1.  DISTRIBUTION OF SOLUBLE COPPER AS A FUNCTION OF pH AND TOTAL CARBONATE
OJ
o
Moles/liter
PH
5.5
7.0
8.0
5.5
7.0
8.0
5.5
7.0
8.0
5.5
7.0
8.0
5.5
7.0
8.0

[co3lt
IxlO"5
IxlO"5
IxlO"5
5xlO~5
5xlO~5
5xlO~5
1x10'"
IxlO"1*
1x10""
5x10-"
SxlO"1*
SxlO'1*
1x1 0~3
lxlO~3
IxlO"3

Soluble Cu3
io"6
10-6.3
10-7.99
10~6
10-6.26
10-7.81
10~6
10"5-22
10-7.66
10~6
10"6
ID-MS
10~6
10~6
10~6.86

Cu2+
100
89
44
100
81
29
100
74
20
100
43
6
98
28
3

CuOH+

-------
                                    Table 1. (Continued)
Moles/liter Soluble copper (%)
PH
5.5
7.0
8.0
5.5
7.0
8.0
[co3]t
5xlO~3
5xlO~3
5xlO~3
IxlO"2
IxlO"2
IxlO"2
Soluble Cua Cu2+ CuOH+ Cu(OH)3 Ci
10~6 93 <1 <1
10~6 7 7 <1
10~6-17 <1 1 <1
10~6 89 <1 <1
10~6 4 <1 <1
10"6 <1 <1 <1
2 2 —
i(OHH CuCOa Cu(COs)2
cl 6 <1
cl 91 <1
cl 95 <1
cl 11 <1
cl 95 <1
a 91 8
a  Total copper = lO"6'00  M

-------
                               SECTION V
                         ANALYTICAL CHEMISTRY
                          OF AQUEOUS COPPER

Discussions in this section include a critical review of existing
methods for trace copper measurement, with emphasis on their applica-
bility and limitations.  More detailed discussions are presented on
electroanalytical techniques, specifically anodic stripping voltammetry
and potentiometric membrane electrodes.  These techniques have been
primarily used in this study in conjunction with the bioassay experi-
ments .

GENERAL REQUIREMENTS FOR TRACE METAL ANALYSIS
The principles and procedures of analysis for trace metals, including
copper, in the aqueous environment have been described by Hume    ,
Mancy    , Morrison    , and others.  The qualities most often noted
for an ideal method of trace metal analysis are detection limits, sen-
sitivity, selectivity, the ability to distinguish free from complexed
metal, and general applicability for use with natural samples.   The
analytical method should have low detection limits, sufficient  for
direct determination of the trace metal without pretreatment or precon-
centration steps.  Copper, for instance, occurs at levels of 10 7 to
10 8 M in natural waters, and an appropriate method should be able to
reach this level of detection with reliability.  Selectivity implies
the capability to distinguish a signal that is due strictly to  the
trace metal in question and not to the presence of any interfering ion
that may be present in the test solution.

                                  32

-------
Ideally speaking, the analytical procedure should also be capable of
defining the metal species under investigation.  Lack of information
on the speciation of trace metals has been a main obstacle to the
elucidation of their role in natural waters.  Information on the type
of species present (e.g., type of complexes and their stability and
rates of complex formation) is a prerequisite to a better understanding
of the distribution and function of trace metals in aquatic environ-
ments.
Copper does not exist in simple ionic form in natural waters.  The work
of Corcoran and Alexander   , Slowey and Hood    , and Foster and
      (54)
Morris     has led them to believe that as much as 50 percent of the
copper present in sea water is in the form of organic complexes.
                               / o / \             ^ o o \
Later work by Barber and Ryther     and Johnston     has shown that the
organic complexes of copper can be correlated to the biological activ-
ity in sea water.
The applicability of a method sums up a variety of factors of practical
consideration.  The reliability, reproducibility, and precision of the
method in question all fall under the heading of applicability.  Other
fundamental considerations are (1) the portability and ruggedness of
equipment for field operations, (2) the time required for a single
analysis, (3) the simplicity of the operation, and finally (4) economy,
space, and additional equipment requirements.

Available Analytical Methods for Trace Copper Analysis
The analyst can select among a number of analytical procedures with
different performance characteristics   ~    .  Various trace analysis
techniques and their general sensitivity ranges are shown in Table 2.
The following discussion will center on the applicability of some of
the more common techniques for copper and includes spectrophotometric,
radiochemical, and electroanalytical methods of trace copper determina-
tion.
                                  33

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       Table  2,   SENSITIVITIES  OF  INSTRUMENTAL METHODS

               FOR MONITORING METALS  IN WATER3
Method
Polarography
Molecular Absorption
Atomic Absorption (Flame)
Atomic Emission
Square Wave Polarography
Sweep Voltammetry
Emission Spectroscopy (Arc)
X-Ray Fluorescence
Pulse Polarography
Molecular Fluorescence
Spark Source Mass Spectrometry
Anodic Stripping
Neutron Activation
Atomic Absorption (Flameless)
Chemiluminescence
Emission Spectroscopy (Plasma)
Sensitivity
100
100
100
100
10
10
10
10
1
1
1
0.1
0.1
0.1
0.1
0.1
» PPb (yg/1)
- 1,000
- 1,000
- 1,000
- 1,000
100
100
100
- 100
10
10
10
10
10
10
10
10
For any method the sensitivity will vary according to the
particular metal.
                              34

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Optical Techniques — The optical techniques most commonly used in
trace copper determination include atomic absorption spectrophotometry
and colorimetry.  In recent years atomic absorption has become one of
the most popular methods of analysis for copper in natural and waste
waters because of its selectivity, speed, and relative simplicity of
analysis.  The technique requires as little as a 5- to 10-ml sample to
perform analysis, and sensitivities in the region of microgram of cop-
per per liter have been achieved with normal flame absorption techniques
       Non-flame techniques can improve on this sensitivity, but are
more costly and are severely lacking in precision.  The main limita-
tions of both flame and flameless atomic absorption techniques are:
(a) metal species cannot be determined; (b) the technique is strictly
a laboratory procedure and cannot be used in the field; and (c) cost
per analysis is high in comparison to electroanalytical procedures.
Colorimetry, in contrast to atomic absorption, is simple and inexpen-
sive.  The American Public Health Association et al.     prescribe
three colorimetric methods for use in copper determination, and these
methods are capable of achieving detection limits near 20 ppb depending
on the experimental conditions and the presence of interfering ions.
     1.  The cuprethol method achieves its maximum sensitivity with
         100-ml nessler tubes for visual comparison, but mercurous,
         nickel, silver, bismuth, and cobalt ions can  act as signifi-
         cant interferences.
     2.  The bathocuproine method is just as sensitive as the cupre-
         thol method, and no ions interfere significantly when in
         concentrations below the parts-per-million range.
     3.  The most specific reagent for copper, neocuproine, is most
         commonly used for analysis of aqueous copper.
                                 35

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 The main limitation with coloriraetric procedures is lack of selectivity
 and the effect of interfering ions.   In addition,  colorimetric  proce-
 dures require a number of preparatory steps as well as a separation or
 concentration step.  Colorimetric procedures are also incapable of
 metal speciation since complex forms  of copper may interact slowly  or
 not at all with the color-developing  compound.

 Radiochemical Techniques — Neutron activation analysis is  a very sen-
 sitive analytical procedure with detection limits  in the parts-per-
                     ( 58}
 trillion (ppt)  range    .   Hence, no  preconcentration step  is necessary.
 Nevertheless  the matrix effect can be quite significant and may result
 in supressing the signals.   Beyond matrix  effects,  the most serious
 limitations of  activation analysis are its inability to distinguish
 species of trace metals and the elaborate,  sophisticated, and expensive
 equipment  required  for  operation.

 Miscellaneous Techniques -— Several techniques that  are not easily
 categorized have been used  in recent  years  for trace analysis.  Mass
 spectroscopy  is  a very  sensitive  technique  for trace copper analysis,
 but extensive sample preparation  is required and the overall method is
                                                          (59)
 time consuming  and  expensive  and  requires  skilled analysts
 Electron spin resonance has also  been used  for the analysis of  copper
 (60)

 Electroanalytical Techniques  — Among the various electroanalytical
 techniques  particularly  applicable for  the  analysis  of  copper  in
 aqueous  samples  are  (a)  potentiometric membrane electrodes,  (b) conven-
 tional and  pulse  polarography,  and (c) anodic  stripping voltammetry.
These techniques vary in their  limit  of detection from  10~5 M for con-
ventional polarography  to 10  10 M for anodic stripping voltammetry.
Perhaps  the unique advantage of electroanalytical techniques  lies in
their ability to distinguish between  free and  complexed metallic species,
Additional advantages of these methods are  their capability of in situ
                                  3-6

-------
measurement, their high selectivity, and their easy adaptability for
field operations.  Furthermore, electroanalytical measurement of metals
in aqueous samples is considerably less expensive than atomic absorption
spectrophotometry.  The comparison is based on capital cost of equipment
and cost per analysis including manpower requirements.
The oldest of the above-mentioned techniques is conventional polaro-
graphy, with a detection limit for copper of about 10 5 M, a range of
practical operation of 10 2 to 10 5 M, and a precision of 1 - 3 percent.
If preconcentration steps are used, the sensitivity can be lowered to
about 10~7 M.
Cathode-ray polarography or oscillographic polarography has been used
for analysis of natural water and wastewaters with a detection limit of
10 7 M    '   .  This technique involves the use of a cathode-ray oscil-
loscope to measure the current-potential curves of applied (saw-tooth)
rapid potential sweeps during the lifetime of a single mercury drop.
Multiple  techniques are also applicable.  The peak current in the re-
sulting polarogram is related to the concentration of the electroactive
species for a reversible reaction.  Oscillographic polarography has  the
advantages of (1) relatively high sensitivity, (2) high resolution,  and
(3) rapidity of analysis.  Copper can be determined at the 50 ppb level
                                    ((."})
in natural waters by this technique
The basic polarographic techniques are  further modified in pulse and
differential pulse polarography, both of which may be used to increase
the sensitivity and minimize the effects of interferences     .  Pulse
polarography has  the advantage of extending the sensitivity  of deter-
mination  to approximately 10 8 M.  The  pulse technique is based on  the
application of short potential pulses of 50 msec  on either a constant
or a  gradually increasing background voltage.  Following  application of
the pulse,  current measurements are usually done  after the spike of
charging  current  has decayed.  The  limiting current in pulse polaro-
graphy  is larger  than  in classical  polarography.   In  derivative pulse
polarography, the voltage pulse is  superimposed upon  a slowly changing
                                   37

-------
 potential  (about  1 mV per sec) and  the difference  in current between
 successive drops  is recorded versus  the potential.
 Differential pulse polarographic techniques are based on measuring the
 current  twice during the lifetime of each mercury  drop and recording
 the difference    .  Current peak-shaped curves are obtained where the
 peak height is proportional to the metal concentration.  This technique
 is 100 - 1,000 times more sensitive  than classical polarography, and 10
 times more sensitive than nondifferential pulse techniques.
 Anodic stripping voltammetry (ASV),  sometimes referred to as inverse
 polarography, is a simple yet extremely useful electroanalytical tech-
 nique for trace metal analysis.  It  offers a detection limit of as low
 as 10 8 to 10 1° M for copper, which is well within the range of natur-
 ally occurring levels.  Moreover, ASV, along with other electrochemical
 methods, has the ability to distinguish free from complexed metal.  The
 method is highly specific for copper and, even at very low concentra-
 tions, the precision is near 5 percent.  The equipment required is
 relatively inexpensive, readily transported, and has been used in the
 field.
 The application of ASV to environmental studies has until recently
 received little mention in the literature.  Matson and coworkers have
 been the forerunners of bringing attention to this useful technique.
 Their work involves studies on the Great Lakes   '   , arctic lakes    ,
 and sewage effluents    .  Evidence for the existence of trace metal-
 organic complexes is clearly presented in these works; however,  the
 techniques utilized for the characterization of these complexes general-
 ly involved only acid digestion and  acid titration procedures.   The only
 other available study on the application of ASV to the characterization
                                                       r t o\
of trace metal complexation is that  of Zirino and Healy    ,   Their
study of the analysis of zinc in sea water by ASV included the observa-
 tion that the peak current  associated with zinc decreased as  the pH
increased.   This phenomenon was explained on the basis of the increased
formation of ZnCC>3 or Zn(OH>2 with increasing pH.
                                  38

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Other environmental studies with ASV have been primarily concerned with
the quantitative determination of trace metals in aquatic systems.
Peterson, Brant, and Mancy     recently used the composite mercury-
graphite electrode (CMGE) in the study of the -distribution of copper
and lead in a municipal water system.  This study included seasonal,
daily, and hourly variations in the concentrations of these metals at
various points along the distribution route of municipal water.  Varia-
tion in the concentrations of the metals was related to residence times
in the pipes as well as variations in the original source of the water
and in the rate of consumption.  Anodic stripping voltammetry was chosen
as the primary analytical technique for the characterization of copper
in the bioassay study.  In the following section a detailed discussion
of the experimental procedures and methods of metal speciation is
presented.

ANODIC STRIPPING VOLTAMMETRY
Introduction
Anodic stripping voltammetry (ASV) is a combination of  (a) a concentra-
tion step (pre-electrolysis, cathodic deposition, or plating)  in which
the metal ion  (or ions) of interest is reduced by controlled potential
electrolysis,  either by deposition as a metal onto a solid microelec-
trode or by formation of an amalgam with the mercury of a mercury drop
or mercury film electrode and  (b) a following step consisting  of an
anodic dissolution process (reverse electrolysis or stripping) in which
the metal is oxidized and returned to the solution.  As with other elec-
trochemical techniques ASV is a nondestructive method of analysis.
The concentration step is carried out for a definite time under repro-
ducible conditions such that the concentration onto the electrode is
quantitative (total plating) or such that a reproducible fraction of the
desired component is deposited from solution  (partial plating).  By
controlling the potential during this process, more easily electrolyzed
constituents may be separated.  The concentration step  is thus performed
                                   39

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by applying to the electrodes a potential which is cathodic of the
polarographic halfwave potential by some 0.3 to 0.4 volts (where the
current has its limiting value) and maintaining this potential for a
given period of time under reproducible conditions of stirring, type
and area of microelectrode, and composition of medium.
After a short "rest period" to allow the solution to become quiescent,
the stripping process is initiated.  The stripping step is performed by
a voltammetric scanning procedure, which produces a response proportion-
al to the amount of material deposited.  The resulting stripping "vol-
tammogram" produces peaks whose heights are generally proportional to
the concentrations of the corresponding electroactive metal ions and
whose potentials are a qualitative indication of the nature of the
species present in solution.  On this basis, the important character-
istics of the peak are its height (peak current, i , in microamperes),
its width at h±  (w, , in volts or millivolts),  and its potential (E ,
               p   -2                                               p
in volts).  These characteristics are affected  by the type of micro-
electrode used and by the rate of voltage change (sweep rate, v, in
mV/sec) used in the stripping process.
Anodic stripping voltammetry,  which is  also known as linear potential
sweep stripping chronoamperometry, stripping analysis, anodic amalgam
voltammetry, and inverse voltammetry, has been  successfully applied to
the analysis of Ag, Au, Ba, Bi, Cd, Ga, Ge, Hg, In,  K, Mn, Ni, Pb,  Pt,
Sb, Sn, Tl, and Zn    , in addition to  the analysis  of Cu.
                                  40

-------
Theory
In the plating step, metal deposition follows a first-order rate
expression

                            Q  = Q  (1 - e~kt)
                            vt   vo          '
where   Q  = number of coulombs equivalent to metal deposited at
             time t (coulombs)
        Q  = number of coulombs equivalent to total amount of metal
             in test solution (coulombs)
        t  = time of electrolysis (seconds)
        k  = rate constant defined by the expression (sec  )
                               k = DA/V6                            (41)
where   D = diffusion constant of the electroactive species in solution
            (cm /sec)
                                           2
        A = test electrode surface area (cm )
                                      3
        V = volume of the solution (cm )
        6 = effective thickness of the diffusion layer  (cm)
It is essential to conduct the plating step at conditions of constant
stirring, temperature, cell geometry, and supporting electrolyte.
The stripping step  is normally conducted in a quiescent solution.  The
voltammetric technique normally employed during the stripping phase is
potential sweep chronoamperometry (application of  an anodic linear
potential sweep); however, other voltammetric procedures may also be
applied.  Of those which have been attempted, square wave polarography
     , radio frequency techniques     , differential polarography     ,
                      (74)
and pulse polarography     have proven successful.
                                   41

-------
 The working electrode employed in this study was the thin-film mercury
 graphite electrode    .   The current-potential equations for the an
 stripping curves with mercury thin-film electrodes were derived and
 verified b
 equations:
graphite electrode    .   The current-potential equations for the anodic
                        srcur;
verified by Roe and Ton!    .   Their derivations result in the following
                             i  = nFA!C° v/e                        (42)
                          EP • Eo + ¥

 where   n   =   number  of  electrons in  the  electrode  reaction
         F   =   Faraday
         A   =  surface  area of  the electrode
         1   =   thickness  of the mercury film
         C   =   concentration of reduced metal  in  the electrode
         R
         v   =   rate  of potential  change
            =   nF/RT
         6   =   thickness  of the unstirred  layer of solution adjacent
               to  the  electrode
         Dn  =   diffusion  coefficient of  the oxidized  species in solution

 These equations predict  that  i  is  a  linear function of v, and E  is a
                               P                                 P
 linear function of  log v.   The equations  have been  experimentally veri-
 fied  and have  been  found  to be valid  at low sweep rates.  Large values
 of v  can cause departures from the  equations because of a breakdown in
 the assumptions that  (a)  5  is  a  constant  from one sweep rate to the next
 and (b)  the diffusion rate  of  deposited metal out of the mercury film is
 extremely rapid compared  to the  sweep rate.  The effect may be signifi-
 cant  for i  , but  is only  slight  for E  .
          P                           P
Regardless of which type  of electrode is used in ASV, the analytical
results obtained  from the stripping curve are always dependent in some
way on the sweep  rate and on some property of the geometry of the elec-
                                  42

-------
trode.  These factors profoundly influence the peak characteristics,
namely, the peak current, the peak potential, and the peak width.

I)iagnostic Criteria
Trace metals in the aquatic environment are frequently found in the
parts-per-billion range or lower (10~"8 - 10~10 M) .   With most analytical
procedures such low levels necessitate a preconcentration step before
the actual analysis.  Anodic stripping voltamraetry with the composite
mercury-graphite electrode (CMGE), however, has a great enough sensitiv-
ity to permit direct determinations of metal ions at these concentration
levels.  In addition, it is a nondestructive process which permits
repeated determinations on the same sample.  Therefore, ASV is a tech-
nique highly suited for environmental samples.
Additionally, ASV also offers the unique opportunity to study trace
metal complexation at the environmentally significant levels.  This is
based on the following diagnostic procedures for the differentiation of
free and complexed metals and for the characterization of metal com-
plexes: (a) acid digestions; (b) variations of ASV parameters, namely,
sweep rate, plating time, and plating potential; (c) titrations of
natural water samples with acid, metals which cannot be determined by
ASV, metals which can be determined by ASV, and titrations of the trace
metals in water samples with organic ligands.  Total metal content can
be determined by digesting the sample before ASV analysis, and filtra-
tion separates dissolved from particulate fractions on the basis of
particle size.  The analytical scheme is illustrated in Figure 5.
In a discussion of  the types of complex systems that might be encouraged
in bioassay waters, there are two broad classifications with several
subclassifications depending on the types of ligands present:
        Case I.   Excess trace metal  (M)
             a.    (M) and (MLL)
             b.    (M) and (MNL)
             c.    (M) and (MLL) and (MNL)
                                   43

-------
                            SAMPLE
                 digestion
         total
    digestion
                         filtration
                                         particulate
                             dissolved
             addition of materials
                                        variation of ASV parameters
         H
electroactive metals
non-electroactive metals
ligands
sweep rate
plating time
stripping medium
plating potential
mixing
      Figure 5.   Analytical scheme for anodic stripping voltammetric
                 determination of metals in natural waters,
                                  44

-------
        Case II.   Excess ligand,  (LL)  or (NL)
             a,   (LL) and (MLL)  and (MNL)
             b.   (LL) and (NL) and (MNL)  where the nonlabile
                  complex (MNL) is the stronger complex
             c.   (NL) and (MLL)  and (MNL)
             d.   (LL) and (NL) and (MNL)  where the labile
                  complex (MLL) is the stronger complex
In this classification M stands for the metal, LL is a labile ligand
which forms complexes whose rate of dissociation is faster than the rate
of plating, and NL is a nonlabile ligand which forms complexes whose
rate of dissociation is slower than the rate of plating.  Note that the
definition of labile and nonlabile ligands is an operational definition
based on the dissociation kinetics of their respective complexes.  This
should be differentiated from the definition of weak and strong com-
plexes which have therraodynamic significance based on stability con-
stants.  Note also that the above classification assumes a water sample
containing a mixture of available ligands.
Based on the above limitations of the terms labile and nonlabile, it
can be concluded that i  measurements can differentiate between free
                       P
metals and labile metal complexes on one hand and nonlabile metal com-
plexes on the other hand.  A further differentiation between labile and
nonlabile metal complexes is possible from data obtained during metal-
organic ligand titrations.  E  measurement can generally distinguish
between free metals and complexed metals, both labile and nonlabile.
The stability constants can easily by calculated from E  measurements.
Furthermore, the technique is suitable  for assessing rates of metal
complexation.  This is  illustrated  in Figure  6.
Results from the titrations of metal by the addition of organic ligands
provide evidence for metal-organic complexation; however, this technique,
in most cases, cannot be used  to characterize  the  type  of metal-organic
interaction.  On the  other hand, metal-organic titration  in which the
metal  is the titrant  is a useful diagnostic criterion for the character-
ization of both labile  and nonlabile ligand-metal  interactions
                                    45

-------
i  Measurements-
                             •FREE  METAL'
                           COMPLEXED METAL-
                        -LABILE METAL COMPLEXES'
                                                     •E  Measurements
                                                       P
E  and  i
 P      P

Measurements
                   L—NONLABILE  METAL COMPLEXES-
                                                          Metal Titrant
                                                          Titrations
  Figure 6.  Anodic stripping voltammetric characterization of metallic species.

-------
Experimental
Heath polarography system,  Model EUW-401,  (Heath Company,  Benton Har-
bor, Michigan) and a chopper stabilizer, Model EUA-19-4, were used in
this work.  A polarographic module interface unit, Model  1014, (Envi-
ronmental Sciences Associates, Inc., Newtonville, Massachusetts) in
conjunction with a Heath operational amplifier, Model EUW-19A, was used
to provide the necessary voltammetric deposition and stripping modes
for the simultaneous operation of four cells.  Voltammograms were re-
corded on a Honeywell Electronik 194 strip chart recorder  (Honeywell,
Inc., Fort Washington, Pennsylvania).
Measurements were performed in 100-ml fused quartz beakers, and the
electrodes were inserted into the cells through rubber stoppers.  The
cell and electrode arrangement is shown in Figure 7.
The reference and counter electrodes were  both separated  from the solu-
tion by 4-mm-diameter Vycor plugs.  These  plugs were attached to the
electrode proper (composed of 4-inch lengths of 4-mm-diameter Pyrex
tubing) by means of sleeves of Teflon tubing.
The counter electrode was filled with supporting electrolyte.  Electri-
cal contact was made by means of a platinum wire inserted in  the tube.
Contact to the solution in the cell was made through the porous Vycor
plug.  A saturated mercury-mercurous sulfate reference electrode was
                                             (78)
used and constructed as described by Lingane     .  The test electrode
was a composite mercury-graphite electrode  (CMGE).  The electrode sur-
                    2
face area was 2.2 cm  .  Detailed discussions on  the preparation and
properties of the CMGE are found elsewhere    '    .
A nitrogen-carbon dioxide gas mixture was used for deaeration of the
test solution.  The pH was controlled by using appropriate partial
pressures of  carbon dioxide.
                                   47

-------
                                  Test Electrode (CMGE)
Counter  Electrode  (Pt)



    Rubber
    Stopper
 Vycor Plugs
      and

 Mixture
\
V


















\
/
1






-
0
o
\
/
V-
n
'/
/
/
/
/
/
/
/
/
/
/
/











L









°0^
, < 	 Reference Electrode
/ (Hg/Hg2S04)
^ — Teflon Gas Purger









/
O GS
7


100 ml Quartz
Beaker





      Figure 7.  Cell and electrode arrangement  for anodic

                stripping voltammetry.
                              48

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STUDY OF COMPETITIVE METAL INTERACTIONS
Water samples containing 5 ppm humic acid were titrated with copper in
the presence and absence of 40 ppm calcium.   The titration curves shown
in Figure 8 illustrate the competitive interaction between copper and
calcium for humic acid ligands    .   The lack of peak current signal at
low Cu concentrations was believed to be a result of the formation of
strong nonlabile complexes.  At high pH values the capacity for non-
labile metal complex formation is greater than at low pH values.  After
the nonlabile ligand has been saturated, further addition of copper to
the test solution is accompanied by an increase in peak current signals.
The presence of calcium exhibits little or no effect on copper-humic
acid interactions.  Thus, copper competes favorably with calcium for
the formation of a nonlabile complex with humic acid.
For comparative purposes Figure 9 shows the same titration curves with
cadmium.  These titration curves are clearly linear, indicating predom-
inantly cadmium-humic acid labile interactions.  Furthermore, the pre-
sence of calcium causes a release of cadmium from its complexes and
allows for a more rapid interaction between the trace metal and the
test electrode.  This is evidenced by the increased peak currents ob-
tained in the presence of calcium.
Examples shown in Figures 8 and 9 represent two different heavy metal-
organic interactions.  Copper forms relatively stable, nonlabile com-
plexes with humic acid, but cadmium seems to form more labile com-
plexes.  Calcium competes favorably with cadmium, and not with copper
for complex formation with humic acid.  Additionally, humic acid
represents a mixture of labile and nonlabile ligand characteristics.
Because in the case of cadmium, the addition of calcium causes an in-
crease in peak currents, we believe that the cadmium-humic acid inter-
action is not exclusively labile.  A summary of metal-humic acid-calcium
interactions for a number of trace metals is given  in Table 3.
                                  49

-------
•H
            pH=8.5
     0
            pH-7.1
                                         10     12     14
                           Q
                 (Cu) x  10    (moles/liter)
Figure 8.   The effect of calcium and pH on the variation  of
           the peak current during  the titration of 5 wg/ml
           humic acid with copper.

                       O Calcium absent
                       O Calcium present
                              50

-------
    20
      0
    40
    20
    40

1

.H^ 20



      0
            pH=8.5
            pH=7.1
            pH=6.2
       0         2         4        6        8        10

                   (Cd)  x 10    (moles/liter)

   Figure 9.  The effect of calcium and pH on  the variation of
             the peak current during the titration of 5 pg/ml
             humic acid with cadmium.

                            O Calcium absent
                            D Calcium present
                              51

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                            Table 3.
     SUMMARY OF TRACE METAL-HUMIC ACID-CALCIUM INTERACTIONS
         AS OBSERVED DURING TITRATIONS WITH METAL IONS
Metal
Nature of Interactions
Bismuth
Cadmium
Copper
Indium
Lead
Thallium
Nonlabile interactions becoming
increasingly labile with increasing
pH; successful calcium competition
for labile ligands
Labile interactions; successful
calcium competition
Strong nonlabile interactions
increasing with increasing pH;
successful calcium competition
for labile ligands
Nonlabile interactions; successful
calcium competition for labile
ligands
Increasing lability with increasing
pH; successful calcium competition
increasing with increasing pH
Labile interactions; successful
calcium competition
                               52

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POTENTIOMETRIC MEMBRANE ELECTRODES
An Orion cupric ion selective electrode was used in this study to mea-
       2+
sure Cu   concentrations.  Absolute potentiometric measurements followed
a Nernstian relationship.  The detection limit for this electrode system
was 10  '  M.  Pertinent information on metal speciation can be achieved
from potentiometric titrations.
The formation of the soluble complex species CuC03 has been studied by
      /o/-)\        (81 }           ( 82^
Scaifev   , Silmanv   , and Stiff ^   .   This complex is found in
natural waters at pH 6.5 to 8.0; its concentration is governed by the
concentration of carbonate.
                                                     o+         -
The equilibrium constants for the reaction between Cu   and HC03  were
                                                           ( 82}
studied.  A modified procedure from that described by Stiff     was
used.  This included rigorous control of pH and total carbonate.  Based
                                     /go\
upon the techniques reported by Stiff    , but in which both copper
activity and pH were monitored , we have determined the equilibrium
constant for the reaction of cupric and bicarbonate ions to form CuC03.
Three concentrations of sodium bicarbonate (1.0 x 10 3, 2 x 10 3, and
5 x 10~3 M) and three concentrations of copper (1.58 x 10~6 , 7.9 x 10~6 ,
and 1.58 x 10~5 M) were used.
It was assumed that the following two equilibrium relationships govern
the copper-bicarbonate system:

         Cu2+ + HC03~ + CuHC03+, K  = [CuHC03+] /[Cu2+] [HC03~]        (44)
      Cu2+ + HC03~ t CuC03 + H+, K  =  [CuC03] [H+] /[Cu2+] [HC03~]      (45)
                                    53

-------
Under  these conditions,  the  total  copper  [Cu  ]  is  approximated by the
following equation:

[Cu  ]  =  [Cu2+] +  [CuC03] + [CuHC03+], or

                  [CuHC03+] + [CuC03] =  [Cut]  -  [Cu2+],               (46)

which  by substitution gives,

              ([Cutl - [Cu2+])/[Cu2+][HC03~] - K! + K2/[H+].          (47)

The values of ([Cu ] - [Cu2+])/[Cu2 ][HCO ~] when plotted against the
                            +
corresponding values of  I/[H  ] would be linear  if equations (45) and
(46) govern this system.  The slope of the line will be equal to K2,
and the intercept of the ([Cu^] -  [Cu2+])/[Cu2+][HC03~] axis will be
equal  to KJ.
The results obtained with different bicarbonate solutions are shown in
Figure 10.  The graphs have the following features:  (a) The intercepts
on the ([Cu ]  - [Cu2 ])/[Cu2  ][HC03~] axis are all very close to the
           t                                           +
origin showing that K} is close to zero and that CuHC03  is a virtually
                                                              	o
non-existent species.  (b) Below a pH value of 8.0 in 5.0 x 10   M
bicarbonate, pH 7.6 in 2.0 x 10 3 M bicarbonate, and pH 7.3 in 1.0 x
10~3 M bicarbonate equations  (44) and (45) do represent the system and
                                   2+
since KJ. is nearly zero  [Cut] - [Cu  ]  = [CuC03].   (c)  At pH values
greater than those indicated the plots are nonlinear, indicating the
formation of a complex or complexes in addition to CuC03.  The values
of K  were computed from the slopes of the linear  portions of the
graphs.  The values of K£ vary with varying total  ionic strength.  The
thermodynamic  equilibrium constant K2 '  was calculated from the equation:

                       (aCuC03)  (V)/(aCu2+)  (a-)'             (48)
                                  54

-------
    80
    70
    60
    50
    40
    30
      "A"

1 .0 x ID"3 M
   [HCO§]
o
    20
    10
     0
      0     40     80    120
           1/[H+] x IQ6
80

70

60

50

40

30

20

10
                                                                      5.0  x 10~3 M
                                                                        [HCO-]
                                                                               1	I
                            50   100  150  200  250
                              1/[H+] x 106
      20  40   60   80  100
        1/[H+]  x  106
     Figure 10.  Graphical  representation of Cu2+ - carbonate ion titrations.

-------
 The value  of  (a      )  is 1.0  since CuC03  is  an uncharged  species, and

                    3                                     (83)
 the other  activity  coefficients were  taken from Kielland      and Stiff

 (82)




 The stability constant  of  the complex species  CuC03  can be evaluated


 from  the following  equations:





                        Cu2+ + HC03~ £ CuC03  +  H+                    (49)
and        C032~ + H+ + HC03  ,  K3' =  [aHC03~]/[aC032~][aH+
(50)
      a

where   denotes activity.  In summing equations  (49) and  (50),
                         Cu2+ + C032~ J CuC03
                              -a
                               CuC03]
                           a  o+  a
                          [aCu2+][aC03
or                       pKit' = pK2' + pK3'.                        (52)





The equilibrium and stability constants for this reaction at different


bicarbonate ion concentrations are given in Table 4.



By using a value of -10.3 for pK3' as reported in the literature    ,


the pKit values determined from our experimental data were found to be "
-7.3.  We realize that pKV determined from our results is higher than

                       (82)              (81)
those reported by Stiff     and by Silman    .   Nevertheless, our re-


sults were quite reproducible as indicated in Table 4.
                                   56

-------
Table 4,  EQUILIBRIUM AND STABILITY  CONSTANTS



Bicarbonate
concentration        pK.2*              PK.1+'
    [M]	

  1.0 x 10~3         2.94             -7.3


  2.0 x 10~3         2.96             -7.3


  5.0 x 10~3         3.00             -7.3
                     57

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                               SECTION VI
                   DEVELOPMENT OF THE BIOASSAY SYSTEM

A vast amount of  literature on metal toxicity is based on bioassay
experiments.  Many of  the available data resulted from studies with
limited objectives, e.g., the determination of the effect of waste
discharge on a receiving water.  The remaining data are for the most
part ambiguous and difficult to interpret.  In fact, most bioassay
data, if not all, have resulted from studies based on limited or no
consideration of  the underlying chemistry.  Regrettably, these data
are the only information available on which water quality criteria
can be based.  At present, there is a pressing need for a better data
base — one which is founded upon scientifically conceived and rigor-
ously defined bioassay studies.
The chief task in this study was the design of a bioassay system in
which physicochemical and biological interrelationships were accounted
for and in which independent variables could be rigorously controlled.
The main interrelationships that need to be considered in the study of
metal toxicity are illustrated in Figure 11 in terms of (1)  physico-
chemical characteristics of bioassay water, (2) concentration of metal
species, (3) metal uptake by test organism, and (4)  biological response.
In this schematic representation the central factor is the physicochemi-
cal characteristics of the bioassay water.   These properties will dic-
tate the concentration of the toxic metal species in the bioassay water
(arrow 1-2)  and  influence metal uptake by the test organism (arrow 1-3)
and the biological response (arrow 1-4).   The concentration of toxic
                                  58

-------
1. Physicochemical character-
istics of Bioassay Water
>
2-1
\
1-2
\

/
2. Concentration of
Toxic Metal Species
2-3
\
3. Metal Up
Test Or
/
4-3
A
\
/
take by
ganism
3-4
\
s
4. Biological Response

< l~2
\
{ 1-4
^
Figure 11.  Physicocheraical and biological interrelationships
            in bioassay experiments .
                             59

-------
metal species will in turn influence the physicochemical characteristics
of the bioassay water (arrow 2-1) through adsorption, complexation, ion
exchange, and other reactions of the metal species.  Metal uptake by the
test organism is largely dependent on the concentration of the different
species of the toxic metal (arrow 2-3).  The biological response of the
test organism is determined by the rate and magnitude of the metal up-
take (arrow 3-4).  In turn the physiological status of the test organism
as indicated by the biological response will significantly influence
metal uptake (arrow 4-3).
Bioassays have been conducted in one of two ways:  (1) in static systems,
usually to establish the minimum dilution of an industrial waste in
which fish or other aquatic organisms survive, or (2) in dynamic systems,
either in the laboratory or in the field, to determine the toxicity of
a given material.  In neither approach, especially where metals are con-
cerned, has the investigator been able to effectively control the chemi-
cal and physical variables to permit his results to be applied to other
situations.  It is virtually impossible to control all the variables in
a laboratory bioassay.  Nevertheless, field experiments are not the com-
plete answer.  Although the field test indicates the toxicity of the
metal under consideration in the environment, the investigator cannot
use this method to understand the role of each of the important physico-
chemical variables in producing the observed effect.   The main physico-
chemical variables that need to be controlled are hydrodynamics, temper-
ature,  ionic strength, light, dissolved oxygen, pH, alkalinity, and the
chemical composition of the bioassay medium.  Some of these variables,
such as temperature and light, are usually considered by most investi-
gators, but others are rarely taken into account.
This section does not review the physical and chemical variables affect-
ing toxicity.   A critical review of some of these factors has been pub-
. .  .  ,   .    ,    (85)
lished  elsewhere
                                  60

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CHEMICAL SPECIES OF METAL TOXICANTS
Few bioassays have been conducted so that the results can be inter-
preted in terras of the toxicity of specific forms of the metal.   No
studies have been conducted that controlled pH,  alkalinity,  and  hardness
independently.  Mount     investigated the effect of pH and  total hard-
ness on the acute toxicity of zinc to fish.  Hardness was controlled at
values of 50, 100, and 200 mg/1 CaC03 by mixing  limestone spring water
and deionized water.  The pH was controlled by the addition  of strong
acid or base.  Addition of the strong acid or base will, besides con-
                                                    ( 7 f\~\
trolling the pH, change the alkalinity of the system    .  For example,
at a hardness of 100 mg/1 the alkalinities were  16, 56, and  102 mg/1
CaCOs at pH 6, 7, and 8, respectively.  The change in the alkalinity of
the system is due to the rapid establishment of  chemical equilibrium
between the solution and the atmosphere.
Most investigators consider alkalinity and hardness to be related since,
in nature, waters of high alkalinity tend also to be high in hardness.
These two properties are, however, basically not related since alkalin-
ity refers to the system's acid-neutralizing capacity and hardness re-
fers to its divalent metal concentration or its capacity to complex with
EDTA.  A solution of calcium chloride will have a high level of hardness
and a low level of alkalinity, whereas a sodium carbonate solution will
have a low hardness and a high alkalinity.
The concentrations of carbonate, bicarbonate, and carbon dioxide in an
aquatic system are of importance primarily because of their roles in pH
control, complexation of metals, and control of metal solubility.  Below
                                                            ( 86}
is presented  a simple method, developed by Roberts and Allen     , for
the control  of total carbonate or  total alkalinity and pH in the bioas-
say experiment.  This technique  is  applicable to natural as well as
synthetic waters.
The formation of significant quantities of metal carbonate or hydroxy
complexes will affect the chemical  relationships presented.  The amount
of base and  the partial pressure of carbon dioxide required to  establish
                                   61

-------
a desired set of conditions  can be predicted  if  the appropriate equilibr-
rium constants  in  the  equations presented  are included.

EQUILIBRIUM RELATIONSHIPS  IN THE CARBONIC  ACID SYSTEM
Equilibrium Relationships  of Total Alkalinity and pH
The total alkalinity of a  sample is defined as its acid-neutralizing
         / f\ /• \
capacity    .   This can be expressed as:
          Total Alkalinity = 2[C032"] +  [HC03~] + [OH~] -  [H+]       (53)
where the brackets indicate  concentrations, and  the alkalinity is ex-
pressed in equivalents per liter.  For a sample exposed to the atmos-
phere the concentrations of  carbonate and bicarbonate ions are related
to the partial  pressure of carbon dioxide  (P   ) by the following
                                            CO2
relationships:
                        [co32 ]
and
                                   Hf7-82p
                                           c°2
                        [HC03 ] = - - -                     (55)
                                       +
By substituting these relationships in equation (54) the following
equation results:
                      10-17. 85p        lO"7-82?
                               C02             C02
[Total Alkalinity] = - -r-; - + - T
                          [H }2            [H+]
                                                           +
                                                            1        (56)
                                  62

-------
In this expression for total alkalinity the pH and partial pressure are
the variables, but this expression will always equal zero unless a base
is added to the solution.   This can best be shown by an example.
In a solution of sodium bicarbonate, sodium carbonate, or sodium hydrox-
ide, the following anion-cation balance will hold:

                [Na+] + [H+] = 2[C032~] + [HC03~] + [OH~]           (57)

Rearrangement gives :

    [Total Alkalinity] = [Na+] = 2[C032~] + [HC03~] + [OH~] - [H+]  (58)

The total alkalinitv is equal to the amount of base in solution and
will equal zero if no base is present.
We can now substitute the sodium ion concentration for total alkalinity
in equation (57), which on rearrangement yields an expression for the
partial pressure of carbon dioxide as a function of the total alkalinity
(or sodium ion concentration) and the pH:
                           10-17.85 + 10-7.82[H+]

This equation is important because it shows that the [H ] can be con-
trolled by the partial pressure of carbon dioxide and the amount of base
added to the solution.  Both of these variables can be controlled by the
investigator.
A computer program for the computation of the P    required to establish
                                           (86)
equilibrium at any pH and total alkalinity     is presented. in
Appendix B.  Results for 10 l , 10 2 , and 10~3 eq/1 for the pH range
4.0  to  12.0 are presented in Appendix B.  The results computed and
the  titles given them in the computer output are total carbonate concen-
                                   63

-------
 tration (TC03),  the logarithm of the total carbonate concentration
 (LOG TC03),  the  logarithm of the partial pressure of carbon dioxide
 (LOG PC02),  and  the logarithms of the molar concentrations  of  C032~,
 HC03~,  and H2C03 (LOG C03,  LOG HC03, and LOG H2C03,  respectively).

 Equilibrium  Relationships of Total Carbonate and  pH  —  The  total  car-
 bonate,  [C032~]  + [HC03~] + [H2C03], is  also controlled by  the partial
 pressure of  carbon dioxide  and the pH according to the  following  rela-
 tionship:
                       10-18.15P         10"7-82P
                               CO2             CO2
  [Total  Carbonate]  =	+	+  lO"1
                          [H+]2             [H+j                co2    (60)

where                     [H2C03] =  lO"1-47?                          (61)
Solving equation  (60)  for  the partial pressure of carbon dioxide,

                             [Total Carbonate][H*]2
                                                                     (62)
an expression is obtained that can be used to calculate the necessary
partial pressure of carbon dioxide for any desired pH and total carbon-
ate concentration.  The concentration of sodium ion necessary to estab-
lish the desired conditions is given by equation  (60) , which can be
rewritten
                 (10-17.85+10-7.82[H+])p        [IlV-lO'1"-00
                                        co2
        [Na ] = - v --- T -     (63)
                           [H ]2                    [H+]

Addition of NaHC03, Na2C03 , or NaOH is necessary so that the total car-
bonate and pH may be independently varied.
                                  64

-------
Calculation Procedures for Synthetic Waters —
Case I:  Total Alkalinity and pH Control.
         The procedure for controlling total alkalinity and pH in solu-
         tions made from distilled or deionized water is very simple:
            1)  Decide what total alkalinity is desired and add that
                molar concentration of NaHC03, NaOH,  or half as much
                Na2C03 to the solution.
            2)  Mix carbon dioxide with air or a nitrogen-oxygen mixture
                and bubble the final mixture through  the solution,
                varying the amount of carbon dioxide  until the desired
                pH is obtained and maintained in equilibrium.
         The pH may be varied simply by adjustment of the amount of car-
         bon dioxide in the mixture, and the alkalinity will remain
         constant at any pH.
Case II: Total Carbonate and pH Control.
         The control of total carbonate and pH involves the following
         calculation:
            1)  Calculate the amount of NaHCOa, NaOH, or NaaCOs neces-
                sary for the desired total carbonate and pH  from the
                following equation which can be derived from equations
                62 and 63.


         [Total Carbonate] (10"17«85+10~7«82[H+])    [H+]2-10"llt •°°
             2)  After the addition of the sodium ion, the pH is adjusted
                with a mixture of carbon dioxide and air or a nitrogen-
                oxygen mixture as described in the total alkalinity
                case.  Once  the desired pH has been obtained, the total
                carbonate concentration should also be at the desired
                level.  These calculations and additions of base must be
                done for each pH and total carbonate concentration.
                                   65

-------
 Figures  12 and 13  show the partial pressures  of  carbon  dioxide  and  the
 amounts  of base necessary to  maintain total alkalinities  (eq/1)  or  total
                                         _ O        _ O
 carbonate concentrations  (moles/1)  or 10   and 10  .  There  is  a minimum
 pH that  can be maintained for any  designated  total  carbonate concentra-
 tion and a maximum pH  that can be  obtained  for any  desired alkalinity.
 The maximum pH decreases  by one unit  as  the alkalinity  decreases ten-
 fold.  In Figures  12 and  13 the maximum  pH  decreases  from 12 to  11  as
                                       — O      — O
 the total alkalinity decreases from 10   to 10    eq/1.  The  minimum pH
 that can be maintained  in the case  of total carbonate control is not
 directly proportional  to  the  total  carbonate  concentration.  As  the
                                 _2       —3
 total  carbonate decreases from 10   to 10  M the minimum pH increases
 from 4.0 to 4.7.
 Limitations — Although it is theoretically possible  to achieve  any pH
 and total alkalinity or total carbonate by  this procedure, practical
 constraints limit  its usable  range.   At  low pH's  and high total  alka-
 linity or total carbonate,  high concentrations of free carbon dioxide
 may be lethal  to fish.  At high pH's  the  high ionic strength present in
 the control of  total carbonate may prove  deleterious; also it may be
 difficult in practice to  control the  low  carbon dioxide pressure re-
 quired to maintain  a high pH.   These  problems should not develop in the
 control  of  bioassay chemical  conditions because extreme values of pH
 and total alkalinity or total carbonate would not be desired.
 Generally,  the  predominant  factor in  controlling  pH and alkalinity  is
 the carbonate  system.   However,  certain bioassay  conditions may  signi-
 ficantly  alter  the  total  alkalinity,  total  carbonate, and pH levels.
 For example, high hardness, alkalinity, and pH may cause carbonate pre-
 cipitation  and  reduce the  total  alkalinity.  High concentrations of
metal toxicants such as lead may also  result  in the formation of metal
hydroxides, which would lower  the alkalinity.   These conditions can be
 recognized  ff the pH obtained  is significantly lower than the predicted
value.
                                  66

-------
                               PH

Figure 12.  Partial pressure of carbon dioxide and amount
            of sodiun bicarbonate (moles/liter) required
            to maintain total carbonate = 10~2 molar (solid
            line) or total alkalinity = 10~2 equivalents/
            liter (dashed line).   Total carbonate cannot be
            below pH 4.0, and total alkalinity cannot be
            above pH 12.0.
                            67

-------
                                                       12
1/4
Figure 13.  Partial pressure of carbon dioxide and moles/liter
            sodium bicarbonate required to maintain total
            carbonate -10   molar (solid line) or total
            alkalinity = 10 3 equivalents/liter (dashed line).
            Total carbonate cannot be controlled below pH 4.7
            and total alkalinity cannot be controlled above
            pH 11.0.
                              68

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                            SECTION VII
                   DESIGN OF BIOASSAY EXPERIMENT

As stated earlier, it is essential that the chemical factors affecting
copper toxicity to fishes be controlled.  Although numerous reports
deal with the toxicity of copper, none includes an account of copper
species, e.g., free copper ions, inorganic and organic copper complexes,
and colloidal dispersions of copper species.
In our bioassay test facility the following variables were controlled:
(1) ionic strength; (2) inorganic ligands:  hydroxide and carbonate;
(3) calcium; and  (4) copper species.  The following variables were mea-
sured during the  testing of the bioassay facilities:  (1) toxicity as
measured by standard bioassay analysis techniques; (2) concentration of
copper species including free, labile, nonlabile, particulate, and
total; and  (3) characteristics of the bioassay water including pH, cal-
cium, carbonate,  dissolved oxygen, and turbidity.  The system design
insures that the  bioassay water is of the desired chemical composition
and that it does  not change during the course of the bioassay.  This
stable condition  is accomplished by using a synthetic bioassay water
that is allowed to reach chemical equilibrium before the beginning of
the bioassay, by  using continuous flow bioassays, and by measuring the
chemical composition of the bioassay water  throughout the test.  Special
precautions must  be taken to  insure that water samples do not change
between collection and analysis.
Since the attainment of chemical equilibrium is often a  slow process,
the bioassay water must be prepared and allowed to reach equilibrium
before exposing the test organism to it.  An example of  the  slow at-

                                   69

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 tainment of equilibrium is illustrated in Figure 14.   Lead,  copper,  and
 zinc nitrates were added at an initial concentration  of 2.5  x  10 6 M
 in an aqueous solution of 10 3 eq/1 total alkalinity  and pH  9.   Pulse
 polarographic measurement of copper,  lead,  and  zinc as a function of
                                                      / OJ \
 time indicates a slow process of metal transformation    .   Consider-
 able time is needed to assure the attainment  of equilibrium.   Conse-
 quently,  the common practice of adding the  toxic agent immediately
 before exposure of the test organism  should be  viewed with skepticism.
 Needless  to say,  analyses conducted to verify the existence  of a given
 metal species at  steady-state conditions  may  not be valid since  mea-
 surements are invariably made much later  than the moment of  exposure.
 This discrepancy  limits  our ability to extrapolate from data obtained
                   uch  ;
                   (89)
                                                            /• QO\
with "diluters," such as those described by Mount and Warner     and
Esve.lt  and Conners
Flow bioassays were deemed necessary  to insure  that chemical conditions
did not change during  the bioassay.   In addition  to the usually cited
deficiencies of static bioassays  (loss of toxicant through biological
uptake  or adsorption and the build-up of toxic  metabolic waste pro-
ducts), waste products released by the test organism may react with
copper  resulting in changing the  chemical species of the toxicant
present.

HATCHERY AND BIOASSAY SYSTEM
Fish Hatchery
A fish hatchery was established in which the necessary test organisms
(juvenile guppies,  Lebistes retieulatus) were raised and conditioned to
the required environmental conditions before their use in a bioassay.
Over 600 adult guppies were maintained in two 70-gal. aquaria.  Aquatic
plants covered the surface to provide cover for the young.  Illumina-
tion was maintained 18 hours a day, and newborn fish were removed daily
and transferred to 5-gal. aquaria.
                                  70

-------
   CK5
   c
 3  n
   ET
 IN3  (U
 + o
H-
rr  O
  00
  D
   r-o
   3
  Q-
                  100
                   50
                   20  —
                   10  —
                    5  —
                   2  —
                             10
                 pH = 9
                                                                      -3
                 alkalinity =10    eq/1—
20     30       40

   TIME  (Minutes)
50
60

-------
 The  juvenile fish  to  be  used  in  the  bioassay were approximately 10 days
 to 2 weeks  old.  Guppies in this age group  are no more susceptible to
 copper  poisoning and  stress than older  fish.  This factor was demon-
 strated during static bioassays in which  two groups of fish of differ-
 ent  ages were subjected  to  identical copper concentrations.  In all
 tests the older fish  died at  the same rate  as the younger fish.  Because
 it is easier to maintain a  large number of  younger fish, the 10-day to
 2-week-old  guppies were  used  as experimental organisms.
 Ann  Arbor tap water was  used  for the  hatchery water supply.  It was
 dechlorinated by passage through a charcoal filter, and the pH was main-
 tained  in the 7.1 - 7.4  range.  The  conditioned water was held in a
 250-gal. aluminum tank;  the water in  the  70-gal. aquaria was continu-
 ously exchanged.  The water supply system and hatchery are shown sche-
matically in  Figure 15.  Water level  in the 250-gal. reservoir was
maintained by a float valve.  The pH was measured, and the output from
 the  pH meter was used as the  input for a controller with a built-in
recorder, enabling the record to be checked for pH deviations outside
 the  acceptable range.  The controller was used to activate two peristal-
 tic  pumps to dispense acid and base.  Although this system is capable of
maintaining a narrower pH span, the limits were set to provide a pH
within a tolerable range without requiring the pumps to be activated
excessively.  At times only a single pump delivering acid was utilized;
the high pH  (greater than pH  10)  of the Ann Arbor water,  as compared to
the desired pH,  eliminated the need for addition of base.
Fish Conditioning
The fish-conditioning system,  shown in Figure 16, included a 250-gal.
equilibrium tank that contained water of the required chemical compo-
sition.   A heater and thermoregulator on a styrofoam float guided by
plastic rods maintained the conditioning water at 25° C.   Water from
the equilibrium tank flowed through a series of 5-gal. aquaria con-
taining the fish to be conditioned for a bioassay.  The aquaria were
equipped with heaters and constant-level siphons.
                                   72

-------
1
2
3
4
5
6
7
Charcoal Filter
Float Valve
pH Electrode
Heater-Thermostat
Acid and Base Lines
Air Stone
Water Outlet
Ann Arbor Tap
 Water Inlet
 8  Light Fixture
 9  Constant Head Siphon
10  Air Stone
11  Heater
12  Aquarium Filter
13  Rustrak RCpH Recorder-Contl.
14  Ministatic Acid and Base Pumps
      To Acid &
      Base Bottles
   pH Control Center


                                                                  11
                                                          10
                        Equilibrium Tank
                                                            70 Gallon Aquarium
                        Figure 15.  Schematic diagram of hatchery.

-------
          CONDITIONING WATER
          LINE
AQUARIUM WITH
HEATER AND
CONSTANT HEAD
SIPHON
                                         INDICATING LIGHT
                                         FLOAT WITH HEATER
                                         AND THERMOSTAT
                                5-GALLON AQUARIA
CONDITIONING EQUILIBRIUM TANK
                   Figure 16.  Schematic diagram of fish-conditioning area.

-------
The fish were acclimated for 10 days.   Two days before the beginning of
a bioassay, the conditioned fish were  transferred to their appropriate
bioassay chambers.   The conditioning water was then diverted to these
chambers.  This procedure eliminated stress and death of fish due to
damage or weakening by transfer from the conditioning aquaria to the
bioassay chambers.
Moassay Water
The test solutions  were prepared several days in advance of their use
to permit the solutions to reach chemical equilibrium and to allow time
for chemical analyses of the water before beginning a bioassay.  The
required chemicals  were added to charcoal-filtered, deionized water
contained in 250-gal. fiberglass-covered plywood tanks.  Sodium bicar-
bonate was added to the water to control the pH.  Then, carbon dioxide
and air were mixed  in a 3 x 1 x 1-foot baffled fiberglass-covered ply-
wood mixing box.  The gas mixture was  bubbled through the test water;
large aquarium stanes were used for gas dispersion.  The partial pres-
sure of carbon dioxide was adjusted until the desired pH of the water
was obtained.  The theoretical considerations of the control of pH in
this manner have been discussed previously.
To control hardness calcium sulfate was added to the equilibrium tanks.
Magnesium sulfate,  sodium chloride, and potassium sulfate, all neces-
sary ions for proper physiological function of the fish, were also
added to the water.  Sodium sulfate was added to obtain constant ionic
strength for the experiments.  The concentration of each of the chemi-
cals was:  [Ca2+] = 2 x lO"4 M; [C032~] = 1 x 10~3 M;  [Cl~] = 30 ppm;
  +                  2+
[K ] = 4 ppm; and [Mg  ] = 4 ppm.
Randomization and Exposure of Fish for Bioassays
In the bioassay, the fish must be assigned to the bioassay chambers in
a random manner since the  fish removed from the aquarium may be the
weakest.  Randomization prevents placing all  the weakest fish  in a sin-
gle chamber.  A computer program was written  that gives  a random
                                 75

-------
 sequence  for  assigning  fish  to bioassay  chambers.  Fish were placed in
 1-quart plastic  containers until each container had one fish, then a
 second fish was  added to  each chamber.   This process was continued
 until the required number of fish had been added to each container.
 The  fish  in each container were then transferred to the identically
 designated bioassay chamber.  Plastic containers were used because they
 could be  clustered in a small area around the aquarium.  This arrange-
 ment minimized the time required to accomplish the randomization and
 also minimized the time that a fish had  to be kept in the net.
 The computer  program (Table 5) provides  the random sequence for adding
 the  fish  to the  chambers.  The variables L and M in the program are,
 respectively, the number  of bioassay chambers to be used in the experi-
 ment and  the  total number of fish to be  placed in each chamber.  The
 computer  output  for 12 bioassay chambers and 20 fish per chamber has
 been included (Table 6).  This computer  program has as its essential
 part the  subroutine RANDU which generates the random numbers.  This
 subroutine produces the same results as  the subroutine of the same name
 available  in IBM's Scientific Subroutine Package/System 360.  The sub-
 routine has been rewritten for use on the IBM 1130 computer and was
 supplied by Professor L.S. Whitlock, Department of Biostatistics, School
 of Public Health, The University of Michigan.
 After all  the fish were randomized to their respective chambers, they
were allowed to acclimatize to a 30 ml/min flow of conditioning water
 for approximately 48 hours.   Then the actual test-water solutions con-
 taining copper flowed into the chambers at about 30 ml/min.  The bioas-
 say continued for 96 hours.   At 8-hour intervals the chambers were
checked  to measure various characteristics including pH, temperature,
resistivity,  flow rate,  and  dissolved oxygen.  Also, at these times
dead fish were removed from the chambers.  For experimental purposes
 lack of  pectoral fin movement indicated death.   The time and the day
of each fish death were recorded on data sheets throughout the 96-hour
experiment.
                                  76

-------
                              Table 5.

           RANDOM SEQUENCE FOR ADDING FISH TO THE CHAMBERS
     DIMENSION J(14)
     WRITE(1,200)
200  FORMAT('  GIVE ME 2 ODD RANDOM NUMBERS IN (14,IX,14)FORMAT.'/
    1'  BOTH NUMBERS SHOULD BE LARGER THAN 4000')
     READ(6,300)1X1,1X2
300  FORMAT(14,IX,14)
     1X1=1X1/2*2+1
     1X2=1X2/2*2+1
     WRITE(5,400)
400  FORMAT(1H1,8X,'RANDOM ORDER FOR PLACING FISH IN BIOASSAY CHAMBERS'
    HIT FISH NUMBER    1A  IB  2A  2B  3A  3B  4A  4B  5A  5B  6A  6B
    2 7A  7B'/1X,68('*'))
     L=12
     M=20
     DO 60 K=1,M
     DO 5 1=1,L
5    J(I)=-99
     NC=0
40   CALL RANDU(IX1,IX2,X)
     DO 20 1=1,L
     IF(X*L-I)10,20,20
10   IF(J(I))30,40,40
30   J(I)=NC+1
     NC=NC+1
     IF (NC-L)40,60,60
20   CONTINUE
60   WRITE(5,100)K,(J(I),I=1,L)
 100 FORMAT(1HO,6X,I3,3X,UI4)
     CALL EXIT
     END
                                   77

-------
                     Table 6.




RANDOM ORDER FOR PLACING FISH IN BIOASSAY CHAMBERS

FISH NUMBER 1A IB 2A 2B 3A 3B 4A 4B 5A 5B 6A 6B 7A 7B
********************************************************************
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
4
12
2
2
2
10
6
12
11
11
11
3
11
8
9
5
6
8
3
2
8
8
5
9
1
3
1
3
12
1
2
12
10
1
1
11
1
2
2
3
9
4
4
11
9
8
8
7
1
6
3
5
6
12
12
3
3
4
12
7
10
2
12
3
3
7
10
1
8
8
10
1
8
2
7
2
4
12
1
1
2
1
1
5
5
6
5
4
3
4
5
4
7
7
6
4
5
5
11
9
12
6
8
7
10
11
7
6
2
7
8
9
3
10
8
12
10
1
4
8
5
11
10
12
11
5
4
9
4
3
9
8
9
6
11
8
2
6
5
12
3
7
3
10
6
9
2
11
5
5
1
2
1
4
5
10
8
7
7
10
7
5
6
6
8
12
3
8
10
12
6
10
5
5
3
9
12
9
9
4
11
9
11
4
4
1
12
2
9
9
12
6
4
11
10
6
9
3
8
5
1
10
9
1
12
2
9
5
6
10
7
11
2
3
4
7
11
11
10
6
6
3
7
8
7
4
11
10
7
2
4
7
12
9
2
1
7
10
6
11
                         78

-------
Exposure Facility
The experimental facility is shown in Figures 17  and 18.   The test water
flowed from the equilibrium tank to the bioassay  chambers through 25-
foot coils of stainless steel tubing contained ir, three large water
baths connected by siphons and a pump.  Medical-grade silicone tubing
was used for connections.  The bioassay chambers  were contained in the
water bath to permit better temperature control and to ease the disposal
of effluent water.  Excess water in the bath was  removed by a standpipe.
The temperature of both the bath and the water contained in the exposure
chambers could be regulated with a half-degree precision.
The exposure chambers  (Figure 19) were fabricated from glass and sili-
cone sealant.  These chambers had a volume of approximately 1 liter, and
the flow of water through the chambers was baffled by the placement of a
plastic screen across  each end.  The flow pattern through the chamber,
as determined by dye tracing, was even.

TOXICITY EXPERIMENTS
Three bioassay experiments were conducted for the sole purpose of asses-
sing the validity of the newly developed concepts and techniques of
chemical control.  The time required to conduct one bioassay was between
2 and 3 weeks.  Physicochemical data for pH,  temperature, electrical
resistance, hardness,  alkalinity, and  total  carbonate are presented in
Table 7.  Alkalinity was determined by potentiometric titration, and
total carbon was determined by total carbon  analysis.
Data for bioassays at  pH 8.10, 7.00, and 5.67  (bioassays A, B, and C,
respectively)  are presented  in Table 8.  Percentage  survival was deter-
mined at 48 hours; in  most  cases  the mortality at  96 hours' exposure
was so  high  that  the data were not usable.   In bioassay  A all  fish died
at  the  highest copper  concentration.   In addition,  percentage  survival
was lower at  130  ppb total  copper than at 278 ppb.   In bioassays  B and
C  there were  no  anomalies  in the  data.
                                   79

-------
                                                                            DEMINERALIZED  WATER LINE
00
o
                   BIOASSAY

                EQUILIBRIUM TANKS


                             WALK V

                    PUMP OUTLET
                       WATER  BATH  TANKS
                                     Figure 17.   Schematic diagram of bioassay area.

-------
CONDITIONING
WATER LINE
        V
ON-OFF
 VALVE
                        EQUILIBRIUM TANK
                                                           WATER BATH
                                                              TANK
          Figure 18.  Schematic diagram of bioassay system.
                                  81

-------
MATERIALS;   DOUBLE STRENGTH GLASS
            PLASTIC SCREENING
            BONDED WITH SILICONS RUBBER SEALER
     ^
                                  4
                                                            2~

\


f


" 8 '""



3


16


— i
31
15 4 J
16
2
\
r5
G16

i
U. .„. r1
rL
h r^
5
3



|DIAX
	 — .
	 J
3
\
j





5
8
^


            Figure  19.   Bioassay  exposure  chamber.
                              82

-------
                              Table 7.   SUMMARY OF BIOASSAY PHYSICOCHEMICAL DATA
00
U)

Bioassay A Bioassay B
Variable
X n s X n s
pH 8.10 67 0.12 7.00 66 0.06
Temperature
(°C) 24.8 56 0.2 24.7 49 0.3
Electrical Resistance
(ohms) 3230 48 78 3200 18 51
Hardness
(meq/1) 0.404 6 0.010 0.284 6 0.032
Alkalinity
(meq/1) 0.975 6 0.012 0.789 6 0.013
Total Carbonate:
(mM) Measured 0.95 6 0.035 0.87 6 0.038
(mM) Calculated
from Alkalinity 0.98 	 	 0.97 	 	
Bioassay C
X3 n s
5.67 120 0.33
24.9 38 0.2
4007 84 129
0.412 6 0.012
0.130 6 0.002
0.95 6 0.036
0.75
       Arithmetic average

-------
     TABLE 8.  SUMMARY OF FISH SURVIVAL DATA AND COPPER MEASUREMENTS3
Total
Statistic Copper
Bioassay A X
(pH 8.10) N
s
X
N
s
X
N
s
X
N
s
X
N
s
X
N
s

<1
1
— — —
42
4
3
70
4
6
130
4
11
278
4
46
343
4
42

Soluble
Copper
— —
__ —
26
2
8
32
2
2
56
2
4
78
2
23
154
2
19

Partic-
ulate
Copper
— .__
— "— —
32
2
9
42
2
5
77
2
4
131
2
16
229
2
3

Free &
Labile Percentage
Copper Survival
	
—
6
2
2
10
2
1
22
2
1
39
2
1
51
2
1

100.0

92.5


42.5


7.5


15.5


0.0


. . . continued
Copper values are given in yg Cu/1.
                                  84

-------
                  Table  8.  (Continued)




  SUMMARY OF FISH SURVIVAL  DATA AND  COPPER MEASUREMENT5
Statistic
Bioassay B X
(pH 7.00) N
s
X
N
s
X
N
s
X
N
s
X
N
s
X~
N
s
Total
Copper
1
	
39
3
12
62
3
3
155
3
18
254
3
49
340
3
46
Soluble
Copper
__^
	
27
2
9
40
2
8
104
2
40
185
2
74
244
2
12
Partic-
ulate
Copper
	
—
16
2
7
26
2
2
54
2
10
68
2
34
95
2
18
Free &
Labile
Copper
	

10
2
4
15
2
0
71
2
38
96
2
56
109
2
58
Percentage
Survival
97.0

92.5


76.5


50.0


33.0


13.0

. . .continued
values are given in yg Cu/1.
                           85

-------
                          Table 8.  (Continued)




          SUMMARY OF FISH SURVIVAL  DATA AND COPPER MEASUREMENT3

Statistic
Bioassay C X
N
s
X
N
s
X
N
s
X
N
s
X
N
s
X
N
s
Total
Copper
<1
1
	
39
4
5
70
4
8
171
3
7
284
4
14
450
4
26
Parti- Free &
Soluble ulate Labile Percentage
Copper Copper Copper Survival
	 	 	 100.0
	 	 	
	 	 	
	 	 	 97.0
	 	 	
	 	 	
79.5
	 	 	
	 	 	
63.5
	 	 	
	 	 	
46.5
	 	 	
	 	 	
28.0
	 	 	
— — — 	
Copper values are given in Mg Cu/1,
                                  86

-------
In bioassays A and B total,  soluble,  particulate,  free,  and labile
copper were measured.  However, only  total copper  was determined in
bioassay C because at pH 5.67 total copper was equal to  soluble copper,
hence particulate copper was absent.   In addition, total copper at this
pH is equal to free and labile copper.
The performance characteristics of the bioassay experiments substantiate
the validity of the newly applied techniques of chemical controls.
Results of bioassays reported  in Table 8 need further validation due
to lack of replications and should not be used as fish toxicity data
per se.
                                   87

-------
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-------
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63.   Whitnack, G.C,   Applications of  Cathode-Ray Polarography  in the
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64.   Schmidt, H,, and M, von Stackelbert.  Modern Polarographic Methods.
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66,   Barsdate, R.J., and W.R. Matson.   Trace Metals in Arctic  and Sub-
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67.   Bender, M.E., W.R. Matson, and R.A. Jordan.  On the Significance
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68.  Zirino, A.R., and M.L. Healy.  Inorganic Zinc Complexes  in Sea-
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69.  Peterson, T.L., D.O. Brant, and K.H. Mancy.  Characterization  of
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70.  Barendicht,  E.  Stripping Voltammetry.   In:  Electroanalytical
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71.  Kodama, M.,  and T. Noda,  Application of a Mercury-Coated Platinum
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72.  Mirva,  T.,  S.  Oki, and A. Mizuike.   Anodic Stripping Voltammetry
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73.  Kemula,  W.   Application of  Stripping Processes  in  Voltammetry.
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74.  Christian,  G.D.   Anodic Stripping Pulse Voltammetry.   J. Electro-
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                                   93

-------
75.  Matson, W.R., D,K. Roe, and D.E. Carritt.  Composite Graphite-Mer-
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     1594-95, 1965.

76.  Roe, D,K,, and J. Toni.  An Equation for Anodic Stripping Curves of
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77.  O'Shea, T.A.  Anodic Stripping Voltammetric Study of the Competi-
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78.  Lingane, J,J.  Electroanalytical Chemistry.  2nd Ed.  New York,
     Interscience Publishers, Inc., 1958.  p. 316.

79.  Matson, W.R.  Trace Metals, Equilibria, and Kinetics of Trace
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80.  Scaife, J.F.  The Solubility of Malachite.  Can. J. Chem,  35:
     1332-40, 1957.

81.  Silman, J.   Reported in Stability Constants by L.G. Sillen and
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82.  Stiff, M.J.  Copper/Bicarbonate Equilibria in Solutions of Bicar-
     bonate Ion Concentrations Similar to Those Found in Natural
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83.  Kielland, J.  Individual Activity Coefficients of Ions in Aqueous
     Solution.  J. Amer.  Chem. Soc.  _59:1675-78, 1937.

84.  Sillen, L.G. r and A.E.  Martell.  Reported in Stability Constants
     by L.G. Sillen and A.E. Martell.  Special Publication No. 17,
     Chemical Society, London, 1964.

85.  Black, J.A., R.F. Roberts,  D.M. Johnson, D.D. Minicucci, K.H.
     Mancy, and  H.E.  Allen.   The Significance of Physical Variables
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     and Environmental Chemistry,  G.E. Glass (ed.),   Ann Arbor, Mich.,
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86.  Roberts, R,F., and H.E. Allen.  The Control of  pH and Total
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     Fish.  Soc.   101(4):752-56,  October 1972.
                                   94

-------
87.   Ernst,  R.,  H.E.  Allen,  and  K.H. Mancy.   Characterization  of Trace
     Metal Species and Measurement  of  Trace Metal  Stability  Constants
     by Electrochemical Techniques.  Water Res.  £:969-79, 1975.

88.   Mount,  D.I.,  and R.E. Warner.   A  Serial-Dilution Apparatus of
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     Washington, D.C., 1965.

89.   Esvelt, L.A., and J.D.  Conners.  Continuous-Flow Fish Bioassay
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     Engineering Research Laboratory,  SERL Report  No. 71-3,
     University  of California, Berkeley, 1971.
                                   95

-------
                    APPENDIX A







AQUEOUS COPPER EQUILIBRIA AT CONSTANT TOTAL CARBONATE




                AND VARIABLE pH VALUES
                        96

-------
0.010000
                            COPPIIU o.oooooi
PH
0.9
9.10
0.20
0.30
0.40
0.50
0.60
3.70
O.tO
O.OQ
I. CO
1.10
1.20
1.30
1.40
1.50
1.60
1.70
l.»3
1.90
2.00
2.10
2.20
2.30
2.*0
2.50
2.60
2.70
?.RO
2.90
3.00
3.10
3.20
3.31
3.'»0
3.50
3.60
3.70
3. "0
3.90
4.03
4.1 3
4.70
4.30
4.40
*.5T
*.'>0
4.70
«. °.o
4.90
CU
6.00
6.00
6.00
6. CO
6.00
6.00
6.00
6.00
6.00
6.00
6.03
6.00
••.03
6.00
6.30
fe.OO
6.00
6.00
6.0'J
6.00
6.00
6.00
<• .00
6.01)
6.10
6.00
6.00
6.00
6.00
6.00
A. 00
6.00
6.00
6. TO
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6. 00
6.00
6.00
6.00
6.00
6.00
CUOH
14.00
13.90
13.80
13.70
13.60
13.50
13.40
13.30
13.20
13.10
13.00
12.90
12. 33
12.70
12. AJ
12.50
12.40
:-'.30
12.23
12.10
12.03
u. 93
11.10
11.70
11.60
11.50
1 1.40
11.10
11.20
11.10
11 .00
10.90
10. "0
10.7'J
10.60
10. 10
10.40
10. 30
10.^3
in. 10
10.03
9.93
9.<*0
9.70
9.60
9.50
°.40
0.30
9.20
9.10
CUOH1
14.30
14.00
14.00
14. 30
14.00
14.30
14.00
14.00
14.30
14.00
- 14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14. JO
14.00
14.00
14.00
14.00
14.00
1 4. JO
14.00
14.00
14.30
14.00
14.00
14.00
14.00
1 4.00
14.00
14.00
14.00
14.00
I4.';o
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
CUOM4
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
'4.00
14.00
14.00
14.00
14.00
14.00
14.00
1't. 00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
|4. 00
14. 00
14.00
14. on
14.00
14.00
I'-.GO
14.00
1'. .00
14.00
14.00
14.00
14. 00
14.00
I4.no
14.00
14.00
ruco3
14.00
14.00
I'^.OO
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
i3.«3
13.63
13.41
13.23
13.03
12. <)3
12.63
1?.43
1?.23
1?.03
11. H3
11.63
11.43
11.23
11.03
10. »3
10.63
10.43
10.21
10.03
°. S3
9.63
9.43
9.23
9.04
B.F.4
S.64
8.44
«.25
6.05
C(J(C03)2
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.0'J
14.0')
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14. OO
14.00
14.00
!<•. 00
14.00
14.00
14.00
1 4.00
14.00
I '..00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
I4.no
14. 'JO
14.00
14.00
14.00
I4.no
14.00
14.00
14.00
14.00
14.00
14.00
14.00
13.63
CU2IOHI2
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
I '..00
14.00
14. OC
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14. PO
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
SOL. CU
6. 00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.30
6.00
6.00
6.00
6.00
6.00
6.00
6.00
ft. 00
6.00
6.00
6.00
6.00
6.00
t.OO
6.00
6.00
6.00
6.00
6 . JO
6.30
6.00
6.00
6.00
6.00
6.30
6.00
6.30
6.30
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
CU PUFC.
14.00
14.00
14.00
14.00
14.00
14.03
14.30
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.03
14.00
14.00
14.03
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.03
14.00
14.00
14.00
14.00
14.00
14.00
14.00
U.OO
14.00
SPECIES
NO PRFC.
NO PHEC.
NO PREC.
NO P3FC.
NO P9EC.
NO PHEC.
NO PREC.
NO "HEC.
NO PUPC.
NO PREC.
NO PREC.
NO P1EC.
NO °RFC.
NO P<*EC.
NO PREC.
NO PREC.
NO P1FC.
NO »REC.
NO PRFC.
NO P9FC.
NO PREC.
NO PREC.
NO PR?C.
NO OREC.
NO PRFC.
NO PREC.
NO PREC.
NO PR^C.
NO DRFC.
NO PREC.
NO PR^C.
NO PREC.
NO PREC.
NO PREt.
NO PRFC.
NO PREC.
NO PREC.
NO PRFC.
NO PRrC .
NO PREC.
NO PREC.
NO PRFC.
NO PREC.
NO PREC.
NO PR*C.
NO PREC.
NO PREC.
NO PR EC.
NO PREC.
NO PREC.

-------
en
PH
5.00
5.10
5.20
5.30
5.40
5.50
5.63
5.70
5.81
5. O1
6. no
6.10
"•.21
6.30
6.40
6.50
6.60
6.70
&.80
6.00
7.00
T.10
7.20
7.30
7.40
7.? 0
7.60
7.70
7.«0
7."0
1.00
?. 10
1.20
".30
1.40
8. 50
P. 60
J.70
1.10
1.00
9.00
9.10
9.20
9.30
9.41
9.50
9.60
9.70
9.90
9.90
CO
6.01
<-.01
6.02
6.02
6.94
6.05
6. J1
6.11
6.16
6. ?2
6. 10
6.39
6.4"
6.'0
6.71
6.13
6.~5
7.07
7.19
7.30
7.41
7.52
7.64
7.74
7.15
7 . •"">
6.17
1.17
" .21
1.19
1 .50
R.61
1 . 75
0.95
9.15
•3.35
0.55
9.75
9.T>
10.15
10.35
11.15
10.75
10.05
11.15
11 .35
11.55
11.75
11.95
12.15
CUOH
9.01
i.-n
8.82
1. 72
P.ft4
1. 5r
9.43
R.41
«. 3'.
R. 1?
(J. 30
•:, . 20
o.2V
0.10
P. 31
1.33
B. 3^
R. 37
8.31
R.40
8.41
1.42
R.44
R.44
1.45
1 . 't6
?.47
1.47
R.4H
R. VO
P. 50
R.51
». 5r)
P.")1!
1.75
8.15
fl.95
9.05
0.15
9.25
0 . 3 '•
0.45
c. 55
9.65
9.75
9.15
9. 95
10.35
10.15
10.25
CU1H3
14.00
14.00
14.00
1 4.00
1 * . 0 1
14.00
14.00
14.00
I'-.OO
14.00
14.00
14.00
14.00
1 4. 00
13.81
13.63
13.41:
13.27
13. OB
12. -50
12.71
12.52
12.54
12.14
11.95
11.76
11.57
11.37
11. in
10.99
1C. MO
10.61
10.45
10.35
10.25
10.15
10.05
9.95
9.15
9.75
9.65
'> . 5 5
".45
9.33
9.25
9.15
0.05
8.95
8.85
8.75
CUOH4
14.00
14.00
14.00
14.00
14. OO
14.00
1<». 00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14. JO
14.00
14.00
14.00
14.00
14.00
1 '«. 01
14. OC
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
I '..00
14.00
;4.oo
14.00
13. 55
13.75
13.^5
13.35
13. li
12.05
17.75
12.55
12.35
12.15
11.99
CDC 03
7.16
7.67
7.48
7.30
7.12
6.15
6.7Q
6.64
6. 51
6.40
6.31
6.23
ft. 17
ft. 13
6. 10
6.07
6. Of
6.04
6.03
6.03
6.02
6.02
6.02
ft. 02
6.02
6.0?
6.02
6.02
6.03
6.03
6.04
6.05
6.09
ft. 19
6.21
'..39
6.49
6.59
6.69
6.PO
6. "0
7.01
7.11
7.22
7.33
7.44
7.56
7.68
7. BO
7.93
C U I CO i 1 2
13. ?4
1 2 . f 6
12.47
1?. 10
11.73
11 .37
11.03
10. 70
10.39
10. 11
9.84
9. f. 1
9.39
Q. 19
9.01
p.es
6.69
1. 55
l.'l
R. 2°
R. 16
P. 04
7.93
7.f'2
7.71
7. f I
7. SO
7.40
7.30
7.21
7. 11
7.02
6.96
6.96
6.96
6.96
6.V6
6.^-7
6.07
6. '"8
6.")
7. no
7.01
7.03
7.05
7.07
7.10
7.14
7.18
7.23
Ci)2inH|2
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
1 4.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
12.05
12.25
12.45
12.65
12. «5
13.05
13.25
13.45
13.65
13.15
14.00
14.00
14.00
14. OO
14.00
14.00
14.00
14.00
SOL. CU
6.00
6.00
6.00
ft. 00
6.00
6.00
6.00
ft. 00
6.00
fr.UO
6.00
6.00
6.00
6.00
6.00
6.00
6.10
6*00
6.00
6.00
6.00
6.00
6.00
6.10
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.03
6.12
6.20
6.2fl
6. 36
6.44
6.51
6.58
6.64
6.7(1
6.76
6.81
6.16
6.91
6.97
7.02
7.08
7.14
CU PRFC.
14.03
14.00
14.00
14.00
14.01
14.00
14.10
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.10
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
7.15
6.62
6.43
6.32
6.25
6.?0
6.16
6.1 3
6.11
6. 10
6.01
6.07
6.06
6.06
6.05
6.04
6.04
6.03
SPF.C IFS
NO PREC.
NO PREC.
NO °RFC.
NO PRFC.
NO PREC.
NO PREC.
NO PREC.
NO PREC.
NO PRFC.
NO PRCC.
NO PREC.
NO PREC.
NO PRFC.
NO PREC.
NO PRFC.
NO PRFC.
NO PP?C.
NO PRFC.
NO PREC.
NO PREC.
NO PRFC.
NO PRFC.
NO PRFC.
NO PREC.
NO PREC.
NO PREC.
NO PRFC.
NO PR=C.
NO PRFC.
NO PREC.
NO PREC.
NO PRFC.
TE^ORITE
T^'JORl TF
TFNORITE
TENOR1TE
TENORITE
TFNORITE
TENORITE
TENORITE
TFNORITE
7FNORITF
TENOR1TE
TENORITE
TENORITF
TENORITE
TENORITE
TFNORITF
TENORITE
TENO«m

-------
PH
11.00
10. 11
10.20
10.13
10.40
10. SO
10.60
10.70
10. PO
10.90
11.03
11.10
11.20
11.30
11.40
11.50
11. 40
11.70
11.80
11.93
12.00
12.10
12.20
12.10
12.40
12.53
12.60
12.70
12. «1
12.90
13.00
11.10
13.20
13. 3J
11.40
1.1. 50
13.60
13.70
13.AO
13.90
14.00
CU
12.35
12.55
12. 75
12.95
13. 15
13.35
13.55
13.75
13.95
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.10
14.00
14.00
14.00
14.00
14.11
14.00
14.00
14.30
14.00
14.00
14.00
CUOH
10. J5
10.45
10. 55
10.65
10.75
10.85
10.95
11.05
11.15
11.25
11.35
11.45
11.55
11.65
11.75
11.85
11.95
12.05
12.15
12.27
12.35
12.45
12.55
12.65
12.75
12. 85
13.02
13.25
1 3.48
13. Tl
13.95
14.00
14. 90
14.00
14.00
14.00
14.00
14.30
14.00
14.00
14.00
CUOH3
R. 65
fi.55
P . '*K
E.35
B.75
".15
P. 05
7.95
7.35
7.75
7.65
7.55
7.45
7.35
7.25
7.15
7.05
6.05
6. 85
6. 75
6.65
6.55
6.45
6. 35
6.25
6.15
6. 12
6.15
6. 11
6.21
6.25
6.30
*>. 35
6.41
6.48
6.55
6.67
6.70
6.78
6.86
6.95
CUOH4
11.75
11.55
11.35
11.15
10.95
10.75
10.55
10.35
10.15
9.95
o. 75
9.55
0.35
9. 15
fl.95
6. 75
P. 55
8.35
8.15
7.S5
7.75
7.55
7.35
7.15
6.05
6.75
6.62
6.55
f .4"
6.41
6.35
6.30
6.25
6.21
6.18
6.15
6.12
6.10
6.08
6.06
6.09
CI/C03
8.06
8.19
».33
8.48
".63
8.79
9.<36
9.13
9.30
9.48
0.^6
9. =4
10.03
10. 72
10.41
10.61
10.80
11.00
11.19
11.39
11. '-9
11.79
11.09
12.18
1?.3S
1 ? . 58
1?. 15
13.18
11.51
1 3 . B4
14.00
14.00
14.no
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
CU(C03)2
7.21
7.37
7.45
7.54
7.65
7.77
7.P9
?.C3
8.11
8.34
8.50
8.67
8.84
9.02
".21
9.39
0.58
9.78
c.97
10. 16
10.36
10.56
10. 75
10.05
11.15
11.35
11.61
11.94
12.27
12.61
12.95
13.29
13.65
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
CU2(nHI2
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
SOL. CU
7.21
7.28
7.36
7.44
7.52
7.59
7.64
7.67
7.67
7.64
7.50
7.51
7.43
7.13
7.24
7.14
7.04
6.93
6.83
6.72
6.62
6.51
6.40
6.29
6.17
6.05
6.00
6.00
6.00
6.30
6.00
6 . ,10
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
CU PRE
*• 03
6.02
6.02
6.02
6.01
6.01
6.01
6.01
6.01
6.01
6.01
6.01
6.02
6.02
6.03
6.03
6.04
6.05
6.07
6.09
6.12
6.16
6.22
6.32
6.49
6.94
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14. OO
14.30
14.00
14.00
14.00
14.00
 SPEC IPS
 TFNORITE
 TENORITE
 TENORITE
 TFNORITF
 TFNORIT?
 TENORITE
 TENORITE
 TENORITE
 TENORITE
 TENQRITF
 TENOR ITE
 TENORITE
TENORITE
TFNORITF
TFNnaiT*
TENORITF
TENORITE
TENORITE
TENORITE
TENORITE
TENOTITE
NO "RFC.
NO P»FC.
NO PREC.
NO PRSC.
NO PRFC.
NO "REC.
NO P*EC.
NO PREC.
NO PRFC.
NO PRFC.
NO PREC.
NO PREC.
NO PR«=C.
NO PREC.
NO PREC.

-------
                     o.oonoo
                                                 COPPER= 0.000001
o
o
PH
0.0
9.10
9.20
0.30
0.43
0.10
0.60
0.73
0.*}
0.90
.00
.10
.21
.31
.*0
.50
l.AO
1.70
I. HI
I. 93
3.00
2.10
2.2 1
z.-n
2.40
2.50
2. AT
2. TO
2.»0
Z.93
3.00
3.10
3.70
3.30
3.40
*.50
3.*0
3.70
3. fa
3.90
4.00
4.10
4.20
4.30
4.40
4.50
4.60
4.70
4.80
4.10
cu
6.00
6.00
6.00
IS. 00
IS. 00
6.00
ft. 00
•>. oo
s.oo
ft. 00
ft. 00
ft. 00
ft .00
ft. 03
*. 30
6.00
ft.OJ
ft . m
ft. 00
ft. CO
ft. 00
ft. 00
ft. 10
f..?1
ft. 00
ft. 00
6.00
ft. 00
A. 00
ft. 00
ft. 00
6.00
ft. 00
ft .00
6.0J
ft. 00
6.00
6.00
ft .01
ft. 00
6.00
ft. 0-1
ft. 00
ft. 03
6.00
ft. 00
6.00
6.00
6.00
6.00
CD'IH
14.03
13.90
13.33
13.7Q
13.60
13.50
1 3.40
13. 3D
13.20
13. 10
13. QT
12.90
U.SO
12.70
12.63
12. SO
I2.'«0
1 '. > 3
1 7 . 1 0
12. 10
12.00
1 1 .90
1 l.flO
11.70
11 .SO
1 1 .-JO
11 .'.0
11.. 40
11. '0
11.10
11.00
1C. 90
10.30
10.70
10.60
13. SO
10.40
10.30
10.20
10.10
10.00
9.9Q
9.40
9.70
9.60
9.50
9.40
9.30
9.20
9,10
CUOH3
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.T.O
14.00
14.00
I'-.on
1 4.00
14.00
14.00
14.00
I 4.00
14.00
1 '..00
I '..00
14.00
I '..00
14.00
1 4.00
14.00
14.00
1 4.00
14.00
14.00
14.00
14.00
1 4.00
14.00
14.00
14.00
14.00
14.00
1 4.00
14.00
14.00
14.00
14.00
I'.OO
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14,00
CHr


-------
PM
5.00
5.10
5.23
5.30
5.43
?.53
5.60
5.70
5.«0
5.93
6.00
6.10
5.?0
6.30
"..*3
6.53
A. 60
6.73
A. 80
S.90
7.00
7.10
7.20
7.30
7.40
7. 53
7.61
7.73
7.13
7.10
9.00
8.10
«.?0
1.33
8.40
8.50
K.60
<».70
«.«3
1.90
9.03
9.n
9.20
0.30
9.43
9.53
«.60
9.70
9. A3
9.90
CU
6.03
5.00
6.03
C.OO
6. no
6.01
6.01
6.01
6.02
6.03
6.04
6.06
6.39
6.1?
r>.!6
6.21
6.26
6.32
6.39
'..47
6. 55
6.ft4
6.75
ft. 9S
7.15
7.35
7.55
7.75
7.?5
8.16
«. 15
«. 55
•?. 71
M.9S
". 15
".35
'•(.•i-i
9. 75
T.ns
10.15
10.35
I').S5
10. T>
10.95
11.15
11. 35
11.55
11.75
11.95
12.15
CUOH
9.0'3
8.?0
8.80
8.70
8.63
8.51
fl.41
0.31
8.7?
ff.13
r.T.
7.<>6
7.3T
7.92
7.76
7.71
7.66
7.62
7.59
7.57
7. 55
7.54
7.55
7.55
7.75
7.95
7.95
8.05
*. IS
1.25
P. 35
r,.4S
a.ss
H.65
W.75
fi.85
".95
c'.05
". 15
9.25
9.35
".AS
''.Si
9.65
".75
9.85
'».95
10.05
10.15
10.25
CUOH3
I*. 00
14.00
14.00
14.00
I4.no
14.00
14.00
14.00
14.00
14.00
14.03
14.10
M.79
13.52
13.26
13.01
I?. 76
12.52
12.29
12.07
11.85
11.04
11.45
11. J5
11.25
11.15
11. OS
10.95
10.85
10. 7S
10.65
10.55
10.45
10.35
10.25
10. 15
10.05
Q.95
P. 85
9.75
^.65
9.SS
9. '.5
9.35
0.25
". 15
o.05
fl.95
fl.85
8.75
CUOH4
14.00
14.30
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
I4.no
14.00
14.00
14.00
I4.no
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.30
14.00
14.00
14.00
14.00
14.00
14.00
14.00
13. "5
13.75
13.r.5
13.35
13.15
12.95
12.75
12.55
12.35
12.15
11.95
cur. 03
».R5
8.66
B.47
P. 27
8.09
7.90
7.72
7.54
7.37
7.21
7.05
6.°1
6.77
6.65
6. 54
6.45
6.37
6.30
6.24
'..20
*. 16
6.13
6.13
6.22
6.31
6.41
6.50
6.60
t: . t 9
6.79
6. P9
6.99
7.09
7.19
7.29
7.39
7.49
7.59
7.69
7.80
7.90
«. PI
8.11
8.22
fl.33
8. 44
8. 56
8.6*
8.80
8.93
CUICr)3)2
14.00
14.00
14.00
14. CO
13.70
13.33
12. "6
12.60
12.25
11.91
11. 5}
11.28
10.99
10.71
10.46
10.22
10. 01
0.81
°.62
o.46
9.30
9.16
f>.05
9.03
9.01
".00
8.99
R.98
8.97'
8.97
8. 96
P. 96
P. 96
fi.96
8.96
8.96
8.96
fl.97
8.°7
«.98
8.99
9.00
9.01
9.03
9.05
9.07
9.10
9.14
9.18
9.23
Cll2IPH)2
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
1'..00
14.00
14.00
14.00
10.05
10. ?5
10.45
10.65
1U.05
11.05
11.25
11.45
11.65
11. «5
12.05
1?.25
12.45
12.65
12.85
13.05
13. ?5
13.45
13.65
13.85
14.00
14. "0
14.00
14.00
14.00
14.00
14.00
14.00
SOL. CU
6.00
6.00
6.00
6.10
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.02
6.13
6.24
6.35
6.45
6.55
f .66
6.76
6.86
6.16
7.06
7.16
7.26
7.36
7.46
7.56
7.65
7.75
7.85
7.04
8.03
8.12
8.20
8.28
8.34
8.39
8.43
ft. 44
CU PREC.
14.00
14.30
14.00
14.00
14.00
14.03
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
7.26
6.58
6.37
6.26
6.19
6.14
6.11
6.03
6.06
6.05
6.04
6.03
6.02
6.02
6.92
6.01
6.01
6.01
6.01
6.01
6.00
6.00
6.30
6.00
6.00
6.00
6.00
6.00
SPECIES
NO p«*r..
NO PREC.
NO PREC.
NO PREC.
NO PRFC.
NO PR=C.
NO PREC.
NO PRCC.
NO PREC.
NO PRFC.
NO PREC.
NO PRFC.
NO PREC.
NO PREC.
NO PREC.
NO PREC.
NO PREC.
NO PREC.
NO PRFC.
NO PREC.
NO PREC.
NO PRFf.
TFNflRITF
TENORITE
TENQRITE
TEWJRITE
TC^IRITF
TENORITE
TENORITF
TENflRITE
TENTITE
T?NORITF
TFNORITE
TFNORITF
TENORITE
TENORITF
TENORITF
TENORITE
TENHR1TE
TFNPRITF
TF'JORITF
TfN'IRtTF
TENORITF
TEN1RITF
TENORITF
TE>gO
-------
PH
13.01
10.10
in. ?o
10.30
13.40
10.51
lO.ftO
11.70
10. JO
10.00
11 .00
11.10
11.20
11.30
11. *0
11.50
11.61
11 .70
ll.'O
11. Oil
12.00
12.11
12.20
12.30
12.40
12 .50
12.60
12.71
12. "1
1 ?.91
13.00
13.10
13.20
13.30
13.40
13.50
13.61
13. TO
13.80
13. "0
14.00
CU
17. 15
1'.55
12,75
1 7 .qc>
13.15
13.35
1? . 51;
13.75
13. "5
I'-.OO
14.00
14.11
1 4 . 00
14,00
1-..00
14.10
14.01
14.01
1 '•• . 00
1 'i . 0 0
14.00
14.00
14.00
14.00
14.00
14,00
14.10
14.00
14,03
14,10
14.00
14,00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
CUD'<
10.35
11. '-5
10.55
11.55
10.75
10. 95
10.^5
11.0*
11.15
11.75
11. 35
11.45
1 1. 55
1 1 . 65
11.75
11.85
11.15
1?. 15
12.15
17.25
12.35
12.45
12.55
12.65
12.75
12.85
13.02
13.25
1 1.48
1 3.71
13.95
14. CO
14.00
1 4. 10
14.00
14.00
14.00
14.00
14.00
14.00
14.00
CUO'13
8.65
8.^5
F, . 4 5
°. ?"•
Q.?5
°. 15
1.05
7. 5 5
7.85
7.75
7.65
7.55
7.45
7.35
7.25
7.15
7.05
ft. 95
6.85
6. 75
6. 65
6. "5
6.45
6.35
6.25
6.15
6.12
ft. 15
6.18
6.21
6.25
6.30
6.35
6.41
6.48
6.5S
6.62
6.70
6.78
6.86
6.95
CIH)H4
11.75
11 .55
11.35
11.15
1(1. ^
10.75
10.55
10.35
10. 15
9. Q5
9. 75
0.^5
9.35
9. 15
K.05
8.75
0.55
8.35
(!. 15
7.T-,
7.75
7. 55
7.35
7. 15
6.95
ft. 75
ft. 62
6.55
ft. 48
ft. 41
ft. 35
6.30
ft. 25
6.21
6. 18
ft. '15
ft. 12
ft. 10
6.08
6.06
6.05
euros
9.06
•3.10
9.33
9.48
9.63
9.79
9.96
10. 13
10.30
10.^8
10.66
13.^4
11.03
11.22
11.41
11.61
11. °0
12.^0
12.10
12.39
12.59
12.79
12.99
13.18
13.38
13.58
13.35
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
CUICH3)2
9.29
9.37
1.45
9. 54
9.65
9.77
9 . °9
1 0 . P 3
10.13
10. 34
10. 50
1 0 . ft 7
10. B4
11.02
11.21
11. 39
11.5ft
11 .78
11. °7
12.16
12.36
12.56
I?. 75
12. 35
13. 15
13.35
13.61
13.94
1 4 . (; 0
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
CM2 (riH»Z
14.00
14.00
14.00
14.00
14.00
14.00
1 4.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14. CO
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
SOL. C'J
P. 44
8.41
f .36
8.29
8.21
P. 13
8.04
7.94
7.84
7.75
7.65
7.55
7.44
7.34
7.24
7.14
7.04
6.93
6.83
6.72
ft. 62
6.51
6.40
6.29
ft. 17
ft. 05
6.00
ft. 00
6.00
ft. 00
ft. 00
6.00
6.00
6.00
ft. 00
6.00
6.00
6.00
6.00
6.00
6.00
CU I'PFC.
6.00
6.00
6.00
6.00
6.00
6.00
6.00
6.00
ft. 01
6.01
6.01
6.01
6.02
6.02
6.03
6.03
6.04
6.05
6.07
6.03
6.12
6. 1 «>
6.22
6.32
6.49
6.94
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14.00
14. OC
14.00
14.00
14.00
14.00
SPFCIFS
TENORITF
TFNORITE
TENORHF
TFNORITE
TENORITE
TFNORITE
TFNORITE
TENHRITE
TFNORITE
TCNORTTE
TENORITE
TFNORITF
TENORITE
TFNORITF
TFNORITE
TFNORITE
TFNORITE
TFNHRITE
TENOR ITE
TF'JP'MTP
TENORITE
TFNORITF
TENORITE
TFNORITE
TENHRIT?
TENORITF
NO PRCC.
NO PRFC.
NO PRFC.
NO PRFC.
NO PREC.
NO PREC.
NO PRFC.
NO PREC.
NO PREC.
NO PRCC.
NO PREC.
NO PREC.
NO PREC.
NO PREC.
NO PREC.

-------
                    APPENDIX B







COMPUTATION OF EQUILIBRIUM pH AND TOTAL ALKALINITY




 AT DIFFERENT PARTIAL PRESSURES OF CARBON DIOXIDE
                        103

-------
PH» 4.00

EQUILIBRIUM
PIC02>
ATM
***INITIAL***
NA2CC3
MG/L
EOUIV
CAC03
********* **********FINAL*************** *******
H2CC3 HC03 COS
************* PCLAR *************
ALKAL INITV
EU/L
MAXIMUM
CALCIUM
MCLflR
FINAL
IONIC
STRENGTH
MG/L

0.2222E
0.3703E
0.5184E
0.6665E
0.8146E
0.9628E
0.1110E
0.1259E
0.1407E
0.1555E
0.1703E
0.1851E
0.1999E
0.?l*7E
0.2295E
0.2443E
0.2592E
0.2740E
0.28R8E
0.3036E
0.3184E
0.3332E
0.3460E
0.3628E
0.3776E
0.3925E
0.4073E
_JU«221JL
0.4369E
0.4517E
01
01
01
01
01
01
02
02
02
02
02
02
02
02
Of
02
02
02
02
02
02
02
02
02
02
02
02
02
02
02
12.387
24.775
37.162
49.550
61.938
74.325
P6.713
99.100
111.488
123.876
136.263
UB.651
161.039
173.426
185.814
198.201
210.589
222.977
235.364
247.752
260.139
272.527
284.915
297.302
309.690
322.078
334.465
346.853
359.240
371.626
10
20
30
40
50
60
70
80
90
100
110
120
130
140
150
160
170
180
190
200
210
220
230
240
250
260
270
280
290
300
0.7530E-
0.1254E
0.1756E
0.2258E
0.2760E
0.3262E
0.3764E
0.4^66E
0.4768E
0.5269E
0.5771E
0.6273E
0.6775fc
0.7277E
0.7779E
0.8281E
0.8783E
0.9284E
0.9786E
0. 1028E
0.1079E
0.1129E
0.1179E
0.1229E
0. 1279E
0.1329E
0.1380E
0.1430E
0.1480E
0.1530E
01
00
00
00
00
00
00
00
00
00
00
00
00
00
00
00
00
00
00
01
01
01
01
01
01
01
01
01
01
01
0.2997E-03
0.4995E-03
0.6993E-03
0.8991E-03
0. 1098fc-02
0. 1298E-02
0.14986-02
0.1698E-C2
0. 1998E-02
0.2097E-02
0.2297E-C2
0.2497E-02
0.2697E-C2
0.2897E-C2
0.3096E-02
0.3296E-02
0.3496E-02
0.3696E-02
0.3896E-02
0.4095E-02
0.4295E-02
0.4495E-02
0.4695E-02
0.4895E-02
0.5094E-02
0.5294E-C2
0.5494E-02
0.5694E-02
0.5894E-02
0.6093E-02
0.1502E-09
0.2503E-09
0.3505E-09
0.4506E-09
0.550et-09
0.6509E-09
0.7510E-09
0.85l2b-09
0.9513E-09
0. 1C51E-08
0. 1151E-08
0. 1251E-08
0. 135U-08
0. 1452E-08
0. 1552C-08
0. 1652E-08
0. 175^E-08
0.1R52E-08
0. 1952E-08
0.2052E-08
0.2153E-08
0.225JE-08
0.2353b-08
0.2453E-08
0.2553E-08
0.2653E-08
0.2753E-08,
0.2853E-08 '
0.2954E-08
0.3054E-08
0.1908E-03
0.3996E-03
0.5994E-C3
0.7992E-03
0.9990E-03
0.1198E-02
0. 1398E-02
0.15'^8E-02
0.1798E-02
0. 199RE-02
0.2197E-02
0.2397E-02
0.2Sr;7fc-02
0.27')7E-02
0.2997E-02
0.3196E-02
0.3396E-02
0.35T6E-02
0.3796E-02
0. 3996E-02
0.4105E-02
0.4395E-02
0.45g5E-02
0.4795E-02
0.4995E-02
0.5194E-02
0.5394E-02
0.5594E-02
0.579AE-02
0.5994E-02
0.3335E
0.2C01E
0.1429E
0.1112E
0.9099E
0.7699E
0.6672E
0.58676
0.5268E
0.4766E
0.4351E
0.4003E
0.37C7E
0.3451E
0.3228E
0.3033E
0.2H59E
0.2705E
0.2566E
0.2441E
0.2327E
0.2224E
0.2129E
0.2042E
0.1962E
0.1888E
0.1819E
0.1756E
0.1696E
0.1640E
02
02
02
02
01
01
Cl
01
01
01
01
01
01
01
01
01
01
01
01
01
01
01
01
01
01
01
01
01
01
01
0.2990E-03
0. 49965-03
0.6994E-03
0.8992E-03
0. 1099E-02
0. 1298E-02
0. 1498E-02
0.1MPE-02
0.1898E-02
0.2098E-02
0.2'297E-02
O.P497E-02
0.2697E-02
0.2897E-02
0.3097E-02
0.3296E-02
0.3496E-02
0.3696E-02
0. 3896E-02
0.4096E-02
0.42T5E-02
0.4495E-02
0.4695E-02
0.4895E-02
0.5095E-02
0.5294E-02
0.5494E-02
0.5694E-02
0.5894E-02
0.6094E-02

-------
PH= 5.00

EQUILIBRIUM
PJC02)
ATM
***INITIAL***
NA2C03
MG/L
EBUIV
CAC03
************* ******FINA|_**********************
H2C03 HC03
************* KCLAR
CC3
*************
ALKALINITY
EC/L
MAXIMUM
CALCIUM
MOLAR
FINAL
IONIC
STRENGTH
MG/L

0.1555E 00
0.3036E 00
0.4517E 00
0.5998E 00
0.7*796 00
0.8960E 00
0.1044E 01
0.1192E 01
0.134CE 01
0.1488E 01
0.1636E 01
0.1784E 01
0.1932E 01
0.2080E 01
0.2229E 01
0.23776 01
0.2525fc 01
0.2673E 01
0.2821E 01
0.29C.9E 01
0.31l7fc 01
0.3265E 01
0.3413k 01
0.3562E 01
0.3710E 01
0.3B58E 01
0.4006E 01
0.4154E 01
0.4302E 01
0.4450E 01
12.387
24.775
37.162
49.550
61.938
74.325
66.713
99.100
111.488
123.676
136.263
14H.651
161.039
173.426
Ifl5.814
198.201
210.589
222.977
235.364
247.752
260.139
272.527
284.915
297.302
309.690
322.078
334.465
346.853
359.240
371.628
10
20
30
40
50
60
70
80
90
100
110
120
130
140
150
160
170
IPO
190
200
210
220
230
240
250
260
270
280
290
300
0.5269E-02
0.1028E-01
0.1530E-01
0.2032E-01
0.2534E-01
0.3036E-01
0.3538E-01
0.4040E-01
0.4541E-01
0.5043E-01
0.5545E-01
0.6047E-01
0.6549E-01
0.7051E-01
0.7553E-01
0.8055E-01
0.8556E-01
0.905flE-01
0.9560E-01
0.1C06E 00
0.1056E 00
0. 1106E 00
0.1i:>6F 00
0.1206E 00
0.1257E 00
0.1307E 00
0.1357E 00
0.1407E 00
0.1457E 00
0.1508E 00
0.2097E-03
0.4095E-C3
0.6093E-C3
O.F091E-03
0.1008E-02
0.1208E-02
0.140BE-C2
0.1608E-C2
0.1B08E-02
0.2007E-C2
0.2207E-C2
0.2407E-02
0.2607E-C2
0.2R07E-02
0.3006E-02
0.3206t-02
0.3406E-02
0.3606E-02
0.3806E-C2
0.4005E-02
0.42C5E-02
0.44C5E-C2
0.4605t-02
0.4805E-C2
0.5004E-02
0.5204E-02
0.5404E-02
0.5604E-02
0.5804E-02
0.6003E-02
0.1051E-08
0.2052E-08
0.3C54E-08
0.405bE-08
0.5056h-08
0.6058E-08
0.7C59E-08
0.8061E-08
0.9062E-08
0.1C06E-07
0.1106E-07
0.1206E-07
0.1306C-07
0. 1406E-07
0.1507E-07
0.1607E-07
0.1707E-07
0. 1807E-07
0.1907E-07
0.2007E-07
0.2107E-07
0.2208E-07
0.2308E-07
0.2408E-07
0.2508E-07
0.2608E-07
0.2708E-07
0.2808E-07
0.2908E-07
0.3009E-07
0.1998E-03
0.3996E-03
o.sgg'.E-os
0.7992E-03
0.99^0E-03
O.H9RE-02
0. 139RE-02
0.1598E-02
0.179RE-02
0.1998E-02
0.2197E-02
0.2397E-02
C.2597t-02
0.2797E-02
0.2997t-02
0.3196E-02
0.3396E-02
0.3596E-02
0.3796E-02
0.3996E-02
0.4195E-02
0.4395E-02
0.4595E-02
0.4795E-02
0.4995E-02
0.5194E-02
0.5394E-02
0.55S4E-02
0.5794E-02
0.5994E-02
0.4766E 01
0.2441E 01
0.1640E 01
0.1235E 01
0.991CE 00
0.8272E 00
0.7C99E 00
0.6217E 00
0.553CE OC
0.49POE 00
0.4529E 00
0.4153E 00
0.3H35E 00
0.3562E 00
0.3325E 00
0.3118E 00
0.2935E 00
0.2772E 00
0.2627E 00
0.24-36E 00
0.2377E OC
0.2269E 00
0.2171E 00
0.2081E 00
0.1998E 00
0.1921E 00
0.1850E 00
0.1784E 00
0.1722E 00
0.1665E 00
0.2098E-03
0.4096E-03
0.6094E-03
0.8092E-03
O.l009fc-02
0.120PE-02
0.1408b-02
0.1608E-02
0.1806E-02
0.200Rt-02
0.2207E-02
0.2407E-02
0.2607E-02
0.2807E-02
0.3Q07E-02
0.3206E-02
0.3406E-02
0.3606E-02
0.3806E-02
0.4006E-02
0.4205E-02
0.4405E-02
0.4605E-02
0.4805E-02
0.5005E-02
0.5204E-02
0.5404t-02
0.5604E-02
0.5804E-02
0.6004fc-02

-------
PH= 6.00

EQUILIBRIUM
PCCC2)
ATM

***INITIAL***
NA2CC3
MG/L

EQUIV
CAC03
MG/L
*******************FINAL*************** *******
H2C03 HC03 C03
************* PCLAR *************



ALKALINITY
EG/L

MAXIMUM
CALCIUM
MOLAR

FINAL
IONIC
STRENGTH


0.1488E-01
0.2969b-01
0.4450E-01
0.5931E-01
0.7412E-01
O.B893E-01
0.1037E 00
0.1185E 00
0.1333E 00
0.1481E 00
0.16Z9E 00
0.1777E 00
0.1926E 00
0.2074E 00
0.2222E 00
0.2370E 00
0.2518E 00
0.2666E 00
0.2814E 00
0.2962E 00
0.3110E 00
0.3258E 00
0.3407E 00
0.3555E 00
0.3703E 00
0.385U 00
0.3999E 00
0.4147E 00
0.4295E 00
0.44436 00
12.387
24.775
37.162
49.550
61.938
74.325
86.713
99. 100
111.486
123.876
136.263
148.651
161.039
173.426
185.814
198.201
210.589
222.977
235.364
247.752
260.139
272.527
284.915
297.302
309.690
322.078
334.465
346.853
359.240
371.628
10
20
30
40
50
60
70
80
90
100
110
120
130
140
150
160
170
IflO
190
200
210
220
230
240
250
260
270
280
290
300
0.5043E-03
0.1006E-02
0.1507E-02
0.2009E-02
0.2511E-02
0.3013E-02
0.3515E-02
0.4017E-02
0.4518E-02
0.5020E-02
0.5522E-02
0.6024E-02
0.6526E-02
0.7028E-02
0.7529E-02
0.8031E-02
0.853?E-02
0.9035E-02
0.9537E-02
0.1003E-01
0. 1054E-01
O.H04E-01
0. 1154E-01
0.1204E-01
0.1254E-01
0. 1304E-01
0. 1355E-01
0.1405E-01
0.1455E-01
0.1505E-01
0.2007E-03
0.4005E-03
0.6003E-03
0.6001E-03
0.9998E-03
0.1199E-02
0. 1399E-02
0. 1599E-C2
0. 1799E-02
0. 1998E-02
0.2198E-C2
0.2398E-02
0.2598E-02
0.2797E-02
0.2997E-02
0.3197E-02
0. 33976-02
0.3597E-02
0.3796E-02
0.3996E-02
0.4196C-02
0.4396E-02
0.4595E-02
0.4795E-02
0.4995E-02
0.5195E-02
0.5395E-C2
0.i594E-02
0.5794E-02
0.5994E-02
0.1C06E-07
0.2C07E-07
0.3C08E-07
0.4C10E-07
0.5Cllfc-07
0.6C12E-07
0.7013F-07
0.8015E-07
0.9016E-07
0. 1C01E-06
0.1101E-06
0. 1202E-06
0. 1302E-06
0. 1402E-06
0. 1502E-06
0. 1602E-06
0. 170^6-06
0. lPO?b-06
0. 190
-------
PM» 7.00

EQUILIBRIUM
PCCC2)
ATM
***INITIAL***
NA2C03
HG/L
E8UIV
CAC03
************ *******FINA L******* ***************
H2C03 HC03
************* KCLAR
CC3
*************
ALKALINITY
EQ/L
MAXIMUM
CALCIUM
MOLAR
FINAL
IONIC
STRENGTH
MG/L

O.U79E-02
0.2959E-02
0.4438E-02
0.59186-02
0.7398E-02
0.86776-02
0.1035E-01
O.U83E-01
0.13316-01
0.1479E-01
0.16276-01
0.1775E-01
0. 19236-01
0.2071E-01
0.2219E-01
0.2367E-01
0.25156-01
0.26636-01
0.2811E-01
0.2959E-01
0.3107E-01
0.3255E-01
0.3403E-01
0.3551E-01
0.3699E-01
0.3847E-01
0.3995E-01
O.M43E-01
0.4290E-01
0.4O8E-01
12.387
24.775
37.162
49.550
61.938
74.325
86.713
99.100
111.48R
123.876
136.263
148.651
161.039
173.426
1R5.814
198.201
210.589
222.977
235.364
247.752
260.139
272.527
284.915
297.302
309.690
322.078
334.465
346.853
359.2*0
371.628
10
20
30
40
50
60
70
80
90
100
110
120
130
140
150
160
170
1RO
190
200
210
220
230
240
250
260
270
280
290
300
0.50136-04
0.1002E-03
0.1504E-03
0.2005E-03
0.25066-03
0.3008E-03
0.3509E-03
0.4010E-03
0.4512E-03
0.5013E-03
0.5515E-03
0.6016E-03
0.6517E-03
0.7019E-03
0.7520E-03
0.8021E-03
0.85236-03
0.90246-03
0.95266-03
0. 1002E-02
0. 10526-02
0.1103E-02
0.1153E-02
0.12036-02
0.1253E-02
0.1303E-02
0.1353E-02
0.1403E-02
0.1453E-02
0.1504E-02
0.1996E-03
0.3992E-03
0.5988E-C3
0.7984E-03
0.9980E-03
0.1197E-02
0.1397E-02
0.15966-02
0. 1796E-02
0.19966-02
0.2195E-02
0.2395E-02
0.2594E-02
0.27946-02
0.29946-02
0.3193E-02
0.33936-02
0. 35926-02
0.3792E-02
0.3992E-C2
0.4191E-02
0.4391E-02
0.4590E-C2
0.4790E-02
0.4990t-02
0.5189E-02
0.53B9E-02
0.5588E-02
0.5788E-02
0.5988E-02
0. 1000E-06
0.2000E-06
0.3C01E-06
0.4C01E-06
0.5C01E-06
0.600?E-06
0.7002t-06
0.8002E-06
0.9C03E-06
O.lCOOE-05
0.1100E-05
0. 12006-05
0.1300E-05
0.14006-05
0. 15006-05
0.166E-C2
0.5010E-02
0.4554E-02
0.4175E-02
0.3853E-02
0.3b78E-02
0.3340E-02
0.3131E-02
0.29476-02
0.2783E-02
0.2636E-02
0.2505E-02
0.2385E-02
0.2277E-C2
0.2178E-02
0.20B7E-02
0.2004E-02
0.1926E-02
0.18556-02
0.17896-02
0.1727E-02
0.1670E-02
0.
-------
                                                                      PH- B.QO
O
CO

EQUILIBRIUM
PICG2)
ATM

***INITIAL***
NA2C03
MG/L

EBUIV
CAC03
MG/L
***************** **FINAL**********************
H2CC3 HC03 C03
************* KCLAR *************



ALKALINITY
EC/L

MAXIMUM
CALCIUM
POLAR

FINAL
IONIC
STRENGTH


0. 14596-03
0.2925E-03
0.43926-03
0.5858E-03
0.73246-03
O.B791E-03
0.1025E-02
0.1172E-02
0.1319E-02
0.1465E-02
0.1612E-02
0. 175Hfc-02
0.19056-02
0.2052E-02
0.2198E-02
0.2345b-02
0.2492E-02
0.263BE-02
0.278SE-02
0.2932E-02
0.3078E-02
0.3225E-02
0.3372E-0?
0.351BE-02
0.3665E-02
0.3812E-02
0.3958b-02
0.4105E-02
0.4251E-02
0.4398E-02
12.387
24.775
37.162
49.550
61.938
74.325
86.713
99.100
111. 488
123.876
136.263
146.651
161.030
173.426
105.814
196.201
210.589
222.977
235.364
247.752
260.139
272.527
284.915
297.302
309.690
322.078
334.465
346.853
359.240
371.628
10
20
30
40
50
60
70
80
90
100
110
120
130
140
150
160
170
mo
190
200
210
220
230
240
250
260
270
280
290
300
0.4944E-05
0.9913E-05
0. 1488E-04
0. 19B5E-04
0.2482E-04
0.2978E-04
0. 3475E-04
0. 3972E-04
0.4469E-04
0.4966E-04
0.5463E-04
0.5960E-04
0.6457E-04
0.6954E-04
0.7450E-04
0.7947E-04
0.8444E-04
0.8941E-04
0.9438E-04
0.9935E-04
0. 1043F-03
0.1092E-03
0. 1142E-03
0. 1192E-03
0.1241E-03
0.1291E-03
0.1341E-03
0.1391E-03
0.1440E-03
0.1490E-03
0. 1968E-03
0.3946E-03
0.5924E-C3
0.7902E-03
0.9881E-03
0.1185E-02
0. 13fi3t-C2
0. 1581E-C2
0.1779E-02
0.1977E-02
0.2175E-02
0.2372E-02
0.2570C-C2
0.2769E-02
O.P966E-02
0.3164F-02
0.3361E-02
0.3559E-02
0.3757F-02
0. 3955E-02
0.4153fc-02
0.4350C-C2
0.4548E-C2
0.4746fc-02
0.4944E-C2
0.5142E-02
0.5340E-02
0.5537E-C2
0.5735E-02
0.5933E-02
0.9865E-06
0.1977E-05
0.2969E-05
0.3960E-05
0.4952E-05
0.5943E-05
0.6935E-05
0.7926E-05
0.8916E-05
0.9909fc-05
0. 1090E-04
0.1189E-04
0.128«E-04
0. 1387E-04
0. 1486E-04
0.158^-04
0.1684E-04
0. 1784E-C4
0.1883E-04
0.19R^E-04
0.2C81E-04
0.218CE-04
0.2279E-04
0. 23786-04
0.2478E-04
0.2577E-04
0.2676E-04
0.2775E-04
0. 28746-04
0.29736-04
0.1998E-03
0.3996E-03
0.5994E-03
C.7992E-03
0.9990E-03
0.1198E-02
0.1398E-02
0.1598E-02
0. 179PE-02
0. 1998E-02
0.2197E-02
0.2397E-02
0.2597E-02
0.2797F-02
0.2997E-02
0.3196E-02
0. 33966-02
0.35966-02
0.3796fc-02
0. 3996E-02
0.4195t-02
0.4395E-02
0.4595E-02
0.4795E-02
0.4995E-02
0.5194E-02
0.5394E-02
0.5594E-02
0.5794E-02
0.5994E-02
0.5080E-02
0.2533E-02
0.1687E-02
0.1265E-02
0.1012E-02
0.8432E-03
0.7226E-03
0.6322E-03
0.5619E-03
0.5C57E-03
0.4597E-03
0.4214E-C3
0.3t"50E-03
0.3612E-03
0.3371E-03
0.3160E-03
0.2974E-03
0.2P09E-03
0.2661E-03
0.2528E-03
0.2407E-03
0.2298E-03
0.2198E-03
0.21C6E-03
0.2022E-03
0.1944E-03
0.1872E-03
0.1805E-03
0. 17436-03
0.1685E-03
0.2007E-03
0.4015E-03
0.6023E-03
O.fiOUE-03
0.1003E-02
0. 1204E-02
0.1405E-02
0. 1606E-02
0.1807fc-02
0.2007E-02
0.2208E-02
0.?409E-02
0.2610E-02
0.2811E-02
0.3011E-02
0.3212E-02
0. 34136-02
0.3614E-02
0.3815E-02
0.40156-02
0.42166-02
0.4417E-02
0.4618E-02
0.4819E-02
0.5019E-02
0.52206-02
0.5421E-02
0.56226-02
0.58226-02
0.60236-02

-------
PH» 9.00

EQUILIBRIUM
PIC02)
ATM
***INITIAL***
NA2C03
MG/L
EQUIV
CAC03
******* ************FINAL**********************
H2C03 HC03
************* PCLAR
CC3 ALKALINITY
************* EC/L
MAXIMUM
CALCIUM
POLAR
FINAI.
IONIC
STRENGTH
KG/L

0.1278E-04
0.2625E-04
0.3971E-04
0.5317E-04
0.6663E-04
0.80096-0'.
0.9356fc-04
0.1070E-03
0.1204E-03
0.1339E-03
0.1474E-03
0.1608E-03
0.1743E-03
O.I877E-03
0.2012E-03
0.2147fc-03
0.2281E-03
0.2416E-03
0.2551E-03
P.26B5E-03
0.2820E-03
0.2954E-03
0.3089E-03
0.3224E-03
0.3358E-03
0.3493E-03
0.3627E-03
0.3762E-03
0.3897E-03
0.*031E-03
12.387
24.775
37.162
49.550
61.938
74.325
86.713
99.100
111.488
123.876
136.263
148.651
161.039
173.426
185.814
198.201
210.589
222.977
235.364
247.752
260.139
272.527
284.915
297.302
309.690
322.078
334.465
346.853
359.240
371.628
10
20
30
40
50
60
70
80
90
100
110
120
130
140
150
160
170
180
190
200
210
220
230
240
250
260
270
280
290
300
0.4333E-06
0.8894E-06
0.1345E-05
0.1801E-05
0.2257E-05
0.2714E-05
0.3170E-03
0.3626E-05
0.4082E-05
0.4538E-05
0.4994E-05
0.5450E-05
0.5907E-05
0.6363E-05
0.6819E-05
0.7275E-05
0.7731E-05
0.81B7E-05
0.8644E-05
0.9100E-05
0.9b56t-05
0. 1C01E-04
0. 1046E-04
0.1092E-04
0.1138E-04
0.1183E-04
0.1229E-04
0.1274E-04
0.1320E-04
0.1366E-04
0.1725E-03
0.3S4lb-03
0.5357E-03
0.7173E-03
0.8988E-03
0.1080E-02
0. 12626-02
0.1443fc-C2
0.1625E-C2
0.1806E-02
0.1988E-02
0.2170E-02
0.2351E-02
0.2533E-02
0.2714E-02
0.2896E-C2
0.307PE-02
0.3259E-02
0.344U-02
0. 362?E-02
0.3804E-02
0.3986E-C2
0.4167E-02
0.4349E-02
0.4530E-02
0.4712E-02
0.4894C-02
0.5075E-02
0.5257E-02
0.5438E-02
0.8645E-05
0. 1774E-04
0.2684E-04
0.3595E-04
0.4505E-C4
0.5415E-04
0.6325E-04
0.7235E-04
0.8145E-04
0.9C55E-04
0.9966E-04
0.1C87E-03
0.1178E-03
0.1269E-03
0.136UE-03
0.1451E-03
0.1542E-03
0.1fc33E-03
0. 17?4fc-03
0.1815E-03
0.1906E-03
0. 1997E-03
0.2C88E-03
0.2179E-03
0.227CE-03
0.2361E-03
0.2452E-03
0.2543E-03
0.2634E-03
0.2725E-03
0. 1998E-03
0.39T6E-03
0.5994E-03
0.799PE-03
0.9990E-03
C.H98E-02
0.1398E-02
0.1598E-02
0.1798E-02
0.1998E-02
0.2197E-02
0.2397E-02
0.2597E-02
0.27T7E-02
O.P997E-02
0.3196E-C2
0.3396E-02
0.3596E-02
0.3796E-02
0.3996E-02
0.4195E-02
0.4395E-02
0.4595E-02
0.4795E-02
0.4995E-02
0.5194E-02
0.5394E-C2
0.5594E-02
0.5794E-02
0.5994E-02
0.5796E-03
0.2624E-03
0.1866E-03
0.1394E-03
0.1H2E-03
0.9255E-04
0.7923E-04
0.6926E-04
0.6152E-C4
0.5534E-04
0.5028E-04
0.4608E-04
0.4252E-C4
0.3947E-04
0.3683E-04
0.3452E-04
0.3248E-04
0.3067E-04
0.2905E-04
0.2760E-04
0.262RE-04
0.250PE-04
0.2399E-04
0.2299E-04
0.2207E-04
0.2122E-04
0.2043E-04
0.1970E-04
0.19C2E-04
0.1838E-04
0.2084E-03
0.4173E-03
0.6262E-03
0.8351E-03
0.1044E-02
0.1252E-02
0. I'f616-02
0.1670E-02
0.1879E-02
0.2088E-02
0.2297E-02
0.2506E-02
0.2715E-02
0.2924E-02
0.3133E-02
0.3341E-02
0.3550E-02
0.3759E-02
0.3068E-02
O.M77E-02
0.43R6E-02
0.4595E-02
0.4804E-02
0.5013E-02
0.5222E-02
0.5430E-02
0.5639E-02
0.5848E-02
0.6057E-02
0.6266E-02

-------
PH'10.00

EQUIL IBRIUH ***INITIAL*** *************«,*****FINAL**********************
PICC2)
ATM
NA2C03
MG/L
ECUIV
CAC03
H2C03 HC03
************* NCLAR **
C03
***********
ALKALINITY
EQ/L
MAXIMUM
CALCIUM
KOLAR
FINAL
IONIC
STRENGTH
MG/L

0.3694fc-06
0.1109E-05
0.1848E-05
0.258fiE-05
0.3328E-05
0.4067E-05
0.4807E-05
0.5547E-05
0.6286E-0!>
0.7026E-Ot>
0.7766fc-05
0.8506t-05
0.9245E-05
0.9985E-05
0.1072E-04
0.1146E-04
0.122CE-04
0.1294E-04
0.1368E-04
0. 1442E-04
0.1516E-04
0.1590E-04
0.1664E-04
0.1738E-04
0.1812E-04
0.1886E-04
0.1960E-04
0.2034E-04
0.2108E-04
0.2182E-04
12.387
24.775
37.162
49.550
61.938
74.325
86.713
99.100
111.488
123.876
136.263
148.651
161.039
173.426
185.814
198.201
210.589
222.977
235.364
247.752
260.139
272.527
284.915
297.302
309.690
322.078
334.465
346.853
359.2*0
371.628
10
20
30
40
50
60
70
80
90
100
110
120
130
140
150
160
170
180
190
200
210
220
230
240
250
260
270
280
290
300
0. 1251E-07
0.3758E-D7
0.6264E-07
0.8771E-07
0. 1127E-06
0.1378E-06
0. 1629E-06
0.1879E-06
0.2130E-06
0.2380E-06
0.2631E-06
0.2882E-06
0.3132E-06
0.3383E-06
0.3634E-06
0.3884E-06
0.41 35E-06
0.4386E-06
0.4636E-06
0.4887C-06
0.5137E-06
0.5388E-06
0.5639E-06
0.5889E-06
0.6140E-06
0.6391E-06
0.6641E-06
0.6892E-06
0.7U3E-06
0.7393E-06
0.4984E-04
0.1496E-03
0.2494E-03
0.3491E-03
0.4489E-03
0.5487E-03
0.6485E-03
0.7483t-C3
0.8480E-03
0.9478E-03
0.1047E-C2
0.1147E-02
0.1247E-02
0.1347E-C2
0.1446E-02
0.1546E-02
0.1646E-02
0. 1746E-02
0.1845E-02
O.C1945E-02
0.2045E-02
0.2145E-02
0.2245fc-02
0.2344E-02
0.2444E-02
0.2544E-02
0.2644E-02
0.2743E-02
0.2843E-02
0.2943E-02
0.2497E-04
0.7498E-04
0.1249E-03
0.175CE-03
0.225CE-03
0.2750E-03
0.325CE-03
0. 3750E-03
0.425CE-03
0.4750E-03
0. 525CE-03
0.5750E-03
0.t250E-03
0.6751E-03
0.7251E-03
0.7751E-03
0.0251E-03
O.P751E-03
0.9251E-03
0.9751E-03
0. 1C25E-02
0.1C75E-02
0.1125E-02
0.1175E-02
0.12256-02
0.1275E-02
0.1325E-02
0.1375E-02
0.1425E-02
0.1475E-02
0.1998E-03
0. 3996E-03
0.5994E-03
0.7992E-03
0.9990E-03
0.1 198E-02
0.1398E-C2
0. 15986-02
0.1798E-02
0. 1998E-02
0.2197E-02
0.2397E-02
0.2597E-02
0.2797E-C2
0.2997E-02
0. 3106E-02
0.3396E-02
0.3516E-02
0.3796E-02
0.3996E-02
0.4195E-C2
0.4395E-02
0.4595E-02
0.4795E-02
0.4995E-02
0.5194E-02
0.5394E-02
0.5594E-02
0.5794E-02
0.5994E-02
0.2006E-03
0.6A83E-04
0.4009E-04
0.2863E-04
0.2227E-04
0. 1FS22E-04
0.1541E-04
0. 1336E-04
0.1179E-04
0.1C54E-04
0.9545E-05
0.8715E-05
0.8017E-05
0.7423E-05
0.6911E-C5
0. 64656-05
0.6074E-05
0.5726E-05
0.5417E-05
0.5139E-05
0.48886-05
0.4661E-05
0.4454E-05
0.4264E-05
0.4090E-05
0.3930E-05
0.3781E-05
0.3644E-05
0.3516E-05
0.3397E-05
0.2247E-03
0.4745E-03
0.7243E-03
0.9742E-03
0.1224E-02
0.1473E-02
0.1723E-02
0.1973E-02
0.2223E-02
0.2473E-02
0.2722E-02
0.2972E-02
0.3222E-02
0.3472E-02
0.372?E-02
0.3971E-02
6.4221E-02
0.4471E-02
0.'. 721E-02
0.4971E-02
0.5220E-02
0.5470E-02
0.5720E-02
0.5970E-02
0.6220E-02
0.6470E-02
0.6719E-02
0.6969E-02
0.72196-02
0. 74696-02

-------
                                  TECHNICAL REPORT DATA
                           (Please read Instructions on the reverse before completing)
 REPORT NO.
 EPA-600/3-77-037
                            2.
                                                          3. RECIPIENT'S ACCESSION NO.
 TITLE AND SUBTITLE
  A CONTROLLED BIOASSAY SYSTEM FOR MEASURING TOXICITY
  OF HEAVY METALS
            5. REPORT DATE
              April  1977 issuing date
            6. PERFORMING ORGANIZATION CODE
. AUTHOR(S)
  K. H. Mancy and H. E. Allen
                                                          a. PERFORMING ORGANIZATION REPORT NO.
. PERFORMING ORGANIZATION NAME AND ADDRESS
  Department of Environmental & Industrial Health
  School of Public Health
  The University of Michigan
  Ann Arbor, Michigan  48109
             10. PROGRAM ELEMENT NO.

               1BA608
             11. CONTRACT/GRANT NO.
               Contract No.  14-12-591
12. SPONSORING AGENCY NAME AND ADDRESS
  Environmental Research  Laboratory-Duluth
  Office of Research  and  Development
  U.S. Environmental  Protection Agency
  Duluth, Minnesota   55804
             13. TYPE OF REPORT AND PERIOD COVERED
              Final  Project Report	
             14. SPONSORING AGENCY CODE
                      EPA/600/03
15. SUPPLEMENTARY NOTES
16. ABSTRACT
  Biological availability of metal micronutrients  and metal toxicity are believed  to
  be dependent on metal  oxidation state, complexation, and solubility as well  as the
  physicochemical characteristics of the aqueous phase.   Basic design criteria for
  fish bioassays which are capable of elucidating  the dependency of toxicity on the
  type and concentration of various copper species were developed utilizing equili-
  brium chemical concepts and appropriate analytical  techniques.
  In order to maintain a desired copper species in the bioassay medium, synthetic
  waters were used  under well-defined physical and chemical conditions.  These solu-
  tions were synthesized in accordance with equilibrium models, which define the
  distribution of various copper species as a function of the solution physical and
  chemical characteristics.  An experimental system was developed which permitted
  large volumes of  the bioassay waters to be maintained at the desired chemical
  equilibria for the  duration of the experiment.
  Monitoring of the bioassay system included measurements of (a) pH, (b) temperature,
  (c) flow,  (d) specific conductance, (e) calcium, (f) total alkalinity, (g) dissolved
  oxygen, and (h) copper species.  Novel analytical procedures were applied for the
  measurement and the differentiation of copper species.  This included the use of
  anodic stripping  voltammetry and potentiometric  membrane electrodes.
17.
                               KEY WORDS AND DOCUMENT ANALYSIS
                  DESCRIPTORS
b.lDENTIFIERS/OPEN ENDED TERMS
                                                                           COSATI Field/Group
  Experimental  Design
  Bioassay
  Toxicity
  Fishes
  Chemical  Equilibrium
  Metal Complexes
                              06/F
                              06/T
18. DISTRIBUTION STATEMENT

  RELEASE TO  PUBLIC
19. SECURITY CLASS (This Report)
 UNCLASSIFIED
21. NO. OF PAGES
    121
                                              20. SECURITY CLASS (This page)
                                               UNCLASSIFIED
                           22. PRICE
EPA Form 2220-1 (9-73)
                                            111
                        U S GOVFDNMENI PRINTING Of fKt. 1977- "> 5 7 - 0 56 ; 5t>00

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