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                AIR POLLUTION ASPECTS

                         OF

               AEROALLERGENS (POLLENS)
                 Prepared for the
   National Air Pollution Control Administration
Consumer Protection & Environmental Health Services
   Department of Health, Education, and Welfare
            (Contract No. PH-22-68-25)
          Compiled by Harold Pinkelstein, Ph.D,
                Litton Systems, Inc.
           Environmental Systems Division
                 7300 Pearl Street
              Bethesda, Maryland 20014

                   September 1969

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                          FOREWORD


       As the concern for air quality grows, so does the con-

cern over the less ubiquitous but potentially harmful contami-

nants that are in our atmosphere.  Thirty such pollutants have

been identified, and. available information has been summarized

in a series of reports describing their sources, distribution,

effects, and control technology for their abatement.

       A total of 27 reports have been prepared covering the

30 pollutants.  These reports were developed under contract

for the National Air Pollution Control Administration  (NAPCA) by

Litton Systems, Inc.  The complete listing is as follows:


    Aeroallergens (pollens)       Ethylene
    Aldehydes (includes acrolein  Hydrochloric Acid
      and formaldehyde)           Hydrogen Sulfide
    Ammonia                       Iron and Its Compounds
    Arsenic and Its Compounds     Manganese and Its Compounds
    Asbestos                      Mercury and Its Compounds
    Barium and Its Compounds      Nickel and Its Compounds
    Beryllium and Its Compounds   Odorous Compounds
    Biological Aerosols           Organic Carcinogens
      (microorganisms)            Pesticides
    Boron and Its Compounds       Phosphorus and Its Compounds
    Cadmium and Its Compounds     Radioactive Substances
    Chlorine Gas                  Selenium and Its Compounds
    Chromium and Its Compounds    Vanadium and Its Compounds
      (includes chromic acid)     Zinc and Its Compounds
                          I

       These reports represent current state-of-the-art

literature reviews supplemented by discussions with selected

knowledgeable individuals both within and outside the Federal

Government.  They do not however presume to be a synthesis of

available information but rather a summary without an attempt

to interpret or reconcile conflicting data.  The reports are

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necessarily limited in their discussion of health effects for

some pollutants to descriptions of occupational health expo-

sures and animal laboratory studies since only a few epidemio-

logic studies were available.

       Initially these reports were generally intended as

internal documents within NAPCA to provide a basis for sound

decision-making on program guidance for future research

activities and to allow ranking of future activities relating

to the development of criteria and control technology docu-

ments.  However, it is apparent that these reports may also

be of significant value to many others in air pollution control,

such as State or local air pollution control officials, as a

library of information on which to base informed decisions on

pollutants to be controlled in their geographic areas.  Addi-

tionally, these reports may stimulate scientific investigators

to pursue research in needed areas.  They also provide for the

interested citizen readily available information about a given

pollutant.  Therefore, they are being given wide distribution

with the assumption that they will be used with full knowledge

of their value and limitations.

       This series of reports was compiled and prepared by the

Litton personnel listed below:

       Ralph J. Sullivan
       Quade R. Stahl, Ph.D.
       Norman L. Durocher
       Yanis C. Athanassiadis
       Sydney Miner
       Harold Finkelstein, Ph.D.
       Douglas A. Olsen, Ph0D.
       James L. Haynes

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       The NAPCA project officer for the contract was Ronald C.



Campbell, assisted by Dr. Emanuel Landau and Gerald Chapman.



       Appreciation is expressed to the many individuals both



outside and within NAPCA who provided information and reviewed



draft copies of these reports.  Appreciation is also expressed



to the NAPCA Office of Technical Information and Publications



for their support in providing a significant portion of the



technical literature.

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                          ABSTRACT






        Aeroallergens  (pollens) are airborne materials



which elicit a hypersensitivity response in susceptible in-



dividuals.  The two major responses exhibited are allergic



Rhinitis and bronchial asthma.  The importance of aero-



allergens as air pollutants is shown by the statistic that



an estimated 10 to 15 million people suffer from seasonal



allergic rhinitis (hay fever) in the United States.  The



pollens of wind-pollinated plants are the most important of



the aeroallergens, and ragweed pollen is commonly found in



this group.  Ragweed pollen is the cause of more than 90



percent of pollinosis in this country.  Other aeroallergens



include molds, house dust, danders, and a miscellaneous



group of insecticides, cosmetics, paints, and vegetable



fibers.  There is evidence to indicate that the aeroallergens



and other air pollutants can act synergistically in affect-



ing human health.



        Most of the aeroallergen investigations have been



concerned with ragweed.  The plant is found primarily in the



North Central and Northeastern parts of the United States/



but it has spread to some degree to the remaining portions



of the country.  Ragweed grows best in soil which has been



disturbed, and therefore is found in abundance both in farm-



lands and in urban areas.  Pollen counts are taken daily in



many local areas throughout the country.  These counts are

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used as guidelines for anticipating and understanding  the



incidence of pollinosis in a given area rather than as



standards.



        Local programs of ragweed eradication generally



have met with little success in controlling pollen concen-



trations.  The pollen can be windborne for many miles, and



therefore pollen entering a city from the outside usually



is sufficient to cause pollinosis in the susceptible popula-



tion.  An adequate program for control would perhaps require



an approach on a regional rather than a local basis.  There



are no adequate estimates of cost values for illnesses



caused by aeroallergens, nor are there estimates for the



cost of abatement on the scale that would be required  for



adequate control.



        The gravity slide method has been accepted as  the



standard procedure for pollen sampling by the Pollen Survey



Committee of the American Academy of Allergy.  However,



because of inherent limitations in the procedure, other



methods have been devised and are used for special sampling



situations.

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                          CONTENTS



FOREWARD

ABSTRACT

1.  INTRODUCTION  	   1

2.  EFFECTS	   3

    2.1  Effects on Humans	   3
         2.1.1  Natural Effects  	   3
         2.1.2  Dose-Effect Relationship 	   6
         2.1.3  Synergistic Responses and Effects
                  of Unknown Substances  	   9
         2.1.4  Effects of Vegetable Dusts	13
         2.1.5  Effects of Molds	16
    2.2  Effects on Animals	18
    2.3  Effects on Plants   	19
    2.4  Effects on Materials	19
    2.5  Environmental Air Standards 	  19

3.  SOURCES	22

    3.1  Natural Occurrence  	  22
    3.2  Production Sources  	  26
    3.3  Product Sources	27
    3.4  Environmental Air Concentrations  . . -	27

4.  ABATEMENT	34

5.  ECONOMICS	40

6.  METHODS OF ANALYSIS	43

    6.1  Qualitative Methods	43
    6.2  Quantitative Methods	45

7.  SUMMARY AND CONCLUSIONS	51

REFERENCES

APPENDIX

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                        LIST OP FIGURES
 1.  Seasonal Variations in Respiratory Illness  	  67

 2.  Seasonal Fluctuations in Respiratory Illness
     for Several Years   	67

 3.  Ragweed Pollen Concentrations During the 1958
     Ragweed Season at and near Ann Arbor, Mich	68

 4.  Variations in Ragweed Pollen Concentrations Close
     to the Pollen Source	69

 5.  Ragweed Pollen Concentration Patterns (grains/m3) ...  70

 6.  Seasonal Average Concentration of Ragweed Pollen
     (grains/m3)   	70

 7-  Diurnal Ragweed Pollen Concentration Patterns at
     a Location Distant from a Local Source  	.71

 8.  Diurnal Pollen Emission Patterns from Fields of
     Ragweed	71

 9.  Diurnal Pollen Emission Patterns from Fields of
     Timothy, Corn and Castor Bean Plant   	72

10.  Weekly Average Pollen Counts, El Paso, Tex	73

11.  Pollen Count for New York City, 1946-1954   	74

12.  Ragweed Pollen Refuges in the United States   	75

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                   LIST OF TABLES
1.
2.
3.
4.
5.

6.

7.

8.

9.
10.
11.
12.

13.

14.
15.
16.

17.
18.
Common Aeroallergens 	

Pollen Seasons Throughout the United States 	
Most Common Aeroallergenic Fungi 	
Summary of Differences Between Hospital Admission
Rates for Days of High and Days of Low Air Pollution .
Percentage Distribution of Emergency Visits by
Month, 1960 . 	 	
Average Daily Number of Emergency Clinic Visits
for Asthma, 1962, 1961, 1957 	
Average Daily Number of Emergency Clinic Visits
for Asthma, September, 1964, 1965 	
Annual Admissions, Brisbane Children's Hospital . . .
Comparison of Mold and Pollen Counts (Israel) ....
Maximum Desitometric Readings for Compound A ....
Dermal and Bronchial Reactivity to Candida
Albicans (81 Patients) 	
Abundance of Ragweeds According to Land use
Categories 	


Pollen Counts, St. Louis Site, 1963-1964,


Recommended Conditions for Use of Common Germicidal
76
77
79
87

88

89

90

91
92
93
94

95

96
97
98

100
101

Substances (At Room Temperature, 25°C) for Fungi .  .  .  102

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                  LIST OF TABLES (Continued)
19.  Resource Costs of Diseases Associated with
     Air Pollution	   103

20.  Asthma-Hay Fever Purchased Acquisition  of
     Prescribed Medicine,  July 1964-June 1954	   104

21.  Six Most Frequent Causes of Non-major Activity
     Limitation, July 1963-June 1965	   105

22.  Average Number of Persons Reported as Limited in
     Activity Due to Selected Chronic Conditions, July
     1961-June 1963   	   106

23.  Death Rate (1950 to 1966) and Deaths (1965  and
     1966) from Selected Causes	   107

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INTRODUCTION



        Aeroallergens (pollens) are airborne materials that



elicit a hypersensitivity or allergic response in susceptible



individuals.  The most common aeroallergens are the pollens



of wind-pollinated plants—especially ragweed pollen,  which



is the main cause of hay fever.  Not all individuals are



allergic or susceptible, but those who are become sensitized



by initial exposure (s) to the allergen (e.g., pollen) and



respond with acute allergic symptoms on subsequent challeng-



ing exposures.  In addition to the pollens, aeroallergens



include molds, danders, house dust, cosmetics, and others.



These will be discussed briefly in this report; the major



emphasis will be on the pollens, and especially ragweed



pollen.



     ,'  There is no question of the adverse effects of aero-



allergens on human health.  It has been estimated that



between 10 and 15 million people in the United States are



affected by seasonal allergic rhinitis (hay fever).  In



addition, many individuals exhibit the more severe syndrome



of bronchial asthma, and it is believed that 5 to 10 percent



of untreated hay fever patients develop the latter illness.



        Ragweed pollen, the most common and the most import-



ant aeroallergen in North America, is the cause of more than



90 percent of all pollinosis.   Although most common in the



Northeastern and North Central part of the United States,



the plant is found in other regions of the country as well.

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Ragweed grows best in soils which have been disturbed by



plowing or other means and, therefore is found in abundance



along railroad tracks, highways, in new urban subdivisions,



and in farmlands.  It has been estimated that one-third of



the 60 million acres of wheat stubble fields are infested



with ragweed.  Since more and more soil has been disturbed



over the years, both ragweed growth and the number of hay



fever sufferers have been increasing.  Therefore, if one



considers that man's progress is correlated with urbanization



and increased breaking of the soil for new roadways, sub-



divisions, and farms, then the increase of ragweed and hay



fever is a problem resulting from this progress.

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2.  EFFECTS



    2.1  Effects on Humans



    2.1.1  Natural Effects



            The major effects of aeroallergens  on human health



    are allergic rhinitis and bronchial  asthma.  Acute allergic



    dermatitis also sometimes occurs.



            Allergic rhinitis is characterized  by a profuse,



    clear,  watery nasal discharge,  sneezing,  and itching of the



    nose, eyes, roof of the mouth,  and posterior pharynx.  If



    these symptoms occur during a particular  season of the year,



    the rhinitis is commonly called hay  fever or rose fever and



    is likely to be caused by plant pollen or mold spores.  If



    the symptoms occur randomly or  nonseasonally, they may



    result from such materials as house  dust  and animal danders,



    or nonairborne allergens.



            Bronchial asthma is a syndrome characterized by



    recurrent, periodic paroxysms of wheezing that are frequently



    associated with dyspnea, choking,  and coughing due to ob-



    struction of expiratory air flow.  The patient is free of



    symptoms during periods between attacks.  Aeroallergens are



    associated with bronchial asthma.



            Some authorities divide bronchial asthma into two



    groups:  extrinsic and intrinsic.20   Others consider the



    syndrome too complex and do not attempt any special group-



    ings.^2  Extrinsic bronchial asthma  is thought to be an

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atopic condition (i.e., a state of clinical hypersensitivity



associated with a family history of allergy.)   It seems



to occur primarily in individuals before the age of 45.



Skin tests with known allergens are positive,  and the in-



dividual has had other clinically determined allergies.



Infection of the respiratory tract is not a factor in the



symptomatology, although it might complicate the condition



later.  The symptoms are brought on by exposure to pollens,



molds, occupational or house dusts, animal danders, foods,



etc.  Intrinsic bronchial asthma is not associated with any



demonstrable evidence of atopy or family history of allergy,



and skin tests are negative.  It is more common after age



45, and is associated with infection in the upper respira-



tory tract.



        Allergic dermatitis occurs less commonly than rhinitis



or asthma.  It is characterized by hives, eczema, or con-



junctivitis.



        The potential aeroallergens present in nature are



numerous.  The ease with which humans can be sensitized to



these materials varies greatly.  Approximately 15 percent



of the population in this country is sensitized easily to



many materials.  These sensitivities often appear at an



early age and are severe enough to be obvious, producing



symptoms of hay fever, asthma, and eczema.  Another 25 to



30 percent of the population is progressively less easily

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sensitized, and their symptoms are subtle.   Most of the



remaining 55 to 60 percent appear never to  become sensitized.



The family history for obvious allergy is usually strongly



positive in the first group, less so in the second, and



absent in the third.  However, on the basis of the physiolog-



ical processes-whereby allergies are manifested, individuals



of the third group could also become sensitized, given the



proper condition.^^



        In addition to the specific syndromes elicited by



the aeroallergens, an additional health problem exists in



that further complications may appear in time.  There is



evidence indicating that sensitivity to one allergen pre-



disposes a sensitivity to other allergens.   In addition, some



hay fever sufferers develop bronchial asthma, which may be



complicated by intractable asthma, pulmonary emphysema,



bronchitis, and pneumonitis.^S  in a continuing community



survey of Tecumseh, Mich., begun in 1957, hay fever and



asthma were separately diagnosed in approximately 10 percent



of the 9,800 population, and both hay fever and asthma in



approximately 2 percent of the population.10  Of those



persons with a history of asthma, the first attack commonly



occurred under the age of 5 years; hay fever frequently began



at a later age.  The analysis also indicated that 5 to 10



percent of persons subject to hay fever will develop asthma



if the disease runs unchecked.11

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        Of the common allergens listed in Table 1, Appendix,



the most important natural sources of air pollution concerned



with inhalant allergy are the wind-pollinated plants (Table



2, Appendix).  Their seasonal and geographical occurrence



are shown in Table 3, Appendix.  Plants of the ragweed



family (Ambrosiaceae) are the most frequent cause of hay



fever in North America; grass pollens seem to cause most of



the cases in Europe; and in the Scandinavian countries blue



grass (Poa pratensis), timothy (Phleum pratense), orchard



grass (Dactylis glomerata), rye grass (Lolium), and rye



(Secale cereale) are the chief offenders.*^  in addition,



trees are an important source of pollen, causing hay fever



throughout North America in the early spring of the year.60



The ubiquitous allergenic saprophytic molds are found both



outdoors and indoors.  A list of the most common aeroaller-



genic molds is presented in Table 4, Appendix.



2.1.2  Dose-Effect Relationship



        Some efforts have been made to establish a quanti-



tative dose-effect relationship for pollens.  Blumstein8



and Tuft et aJL,.^0^ dipped a toothpick into pollen and held



it under the patient's nose as the patient inhaled vigorously.



Using different pollens, they were able with this method to



determine to which pollen the patient was sensitive.  The



disadvantages of this technique were these:  (l) quantitation



of the pollen dosage (through dilution with talc)was crude;

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(2)  a clinical episode of hay fever could be produced by



one or two inhalations, thus suggesting that the dosages



were massive as compared to ordinary environmental exposure;



and (3) the administration of hazardous dosages was possible.



Although the quantity of pollen constituting an overdose is



unknown, there is evidence which suggests that an overdose



is not a fixed amount but probably varies from patient to



patient, and even varies in the same patient under differing



conditions.16



        Solomon1^ hag reported data which suggest that in




persons with allergic rhinitis, prior exposure, body position,



and breathing patterns may be important modifying factors



in nasal responses.  He has initiated studies, the results



of which have not yet been reported, utilizing a large test



chamber in which the temperature, relative humidity, and



duration of extraseasonal pollen exposure of test subjects



is possible.1(^



        Connell16'1^'1^ has devised an apparatus for exposing



subjects to quantitative dosages more analogous to environ-



mental exposure levels.  The apparatus consists of a 22-liter



flask in which the pollen is kept in suspension.  The patient



applies a face mask connected to the flask and inhales



through his nose.  Sensitive individuals respond with the



clinical signs and symptoms of hay fever:  itching of the



eyes, ears, nose, and throat; nasal discharge; etc.  Dosages

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                                                           8
can be controlled so that the total accumulative nasal



exposure (challenge) in an hour is from 100 to 300 pollen



grains.  It was found that hay fever symptoms were frequently



produced by as little as 100 grains in a 1-hour challenge.



Nonallergic individuals have showed no response to as much



as 50,000 grains in 30 minutes.  Connell's studies showed



that with daily ragweed pollen challenges, smaller and



smaller doses were required each succeeding day to cause the



same or greater degree of hay fever.  He called this increased



sensitivity the "priming effect" and concluded that environ-



mental exposure during the ragweed pollinating season



similarly caused priming of the nasal membranes.  The priming



was reversible, and the time period (days to weeks) for



priming was dependent upon the degree of exposure to the



pollen.  When pollen was administered only to one nostril,



unilateral priming and allergic rhinitis occurred only in



that nostril, and resistance of the other nostril remained



unchanged.  In the case of one individual, after 2 weeks



of daily priming of one nostril, response was obtained to a



total dose of 30 ragweed pollen grains, but the non-primed



nostril showed no response to 303 grains.  It took additional



exposure to pollen (priming and challenge) before the latter



nostril also responded.  Connell further observed that in



subjects sensitive to more than one pollen, priming could be



accomplished with one pollen (such as sorrel) and the hay

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fever symptoms produced by challenge with another pollen



(such as ragweed)-



2.1.3  Svnercfistic Responses and Effects of Unknown Substances



        Naturally occurring aeroallergens such as pollens



have been known and studied for several decades.  However,



the ability of .other air pollutants to cause a potentiating



or synergistic response with the natural allergens has become



a new area of study in recent years.



        Schoettlin and Landau^4 studied 137 asthmatic



patients in the Los Angeles area for 98 days (September 3 to



December 9, 1956).  This period roughly corresponded to the



smog season.  They studied the number of asthmatic attacks



in relationship to air pollution as measured by total atmo-



spheric oxidants, particulates, carbon monoxide, relative



humidity and temperature, and plant damage.  Low positive



correlations were found between chemical measures of air



pollution and the number of persons suffering attacks.  Low



correlations were also noted for temperature, relative



humidity, and water vapor pressure.  A significantly greater



number of persons had attacks on days with oxidant values



high enough to cause eye irritation or cause plant damage



than on other days.  Sterling et jal^P-06 aiso in Los Angeles,



observed an increase in hospital admission rates for allergic



disorders as well as a number of other respiratory syndromes



on days of high air pollution (Table 5, Appendix).

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                                                          10
        Zeidberg et ^.l17 reported on an extensive air



sampling and medical evaluation study of Nashville, Tenn.



They observed a group of 84 patients with bronchial asthma—



49 adults and 35 children—for a 10-month period (October



1958 to July 1959).  A total of 3,647 asthmatic attacks



occurred during 27,440 person-days of observation.   An



overall attack rate of 0.133 per person-day was reported.



In adults, the asthmatic attack rate varied directly with



the sulfate level in their residential environment and was



three times as high for those living in the high pollution



area as for those in the low.  This correlation could not



be demonstrated with the children.



        Following a 2-year epidemiological study of asthma



in children in Philadelphia, Girsh et aJL.40 concluded that



the occurrence of stable weather conditions with stagnant



air seemed to correlate with peak incidences of bronchial



asthma.  They had observed a total of 1,346 patients during



the two-year period; the average normal 24-hour incidence



of asthma was 2.5 patients (standard deviation of +. <2).



The incidence was considered high when five or more patients



were seen in 24 hours; and on 70 days of the 676 days, the



incidence was 5 to 14 per 24 hours.  The incidence of bronchial



asthma was three times greater during days of high air



pollution, and there was a fourfold increase during days with



high barometric pressure.  When both air conditions coincided,

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                                                          11
there was approximately a ninefold increase in bronchial



asthma.  The authors concluded that the greater incidences



were not due to ragweed pollen, but speculated that some



unknown pollutant was present which was not being measured.



        Booth et al.9 studied the records of asthma emer-



gency visits for 10 hospitals in seven cities during 1960.



For most of the hospitals, noticeable peak months for



asthma occurred in the autumn of the year, as shown in



Table 6, Appendix.  No single cause could be established;  the



investigators' conclusions were that multiple causes probably



had existed.



        Greenburg and co-workers have analyzed records of



emergency clinic visits for asthma and for other respiratory



illnesses in New York City hospitals for a number of years



in relationship to air pollution episodes.  Although they



observed an increase in upper respiratory infections during



the November 1953 air pollution incident, there was no



associated increase in asthma clinic visits.4^  Continuing



their analysis, Greenburg .et §JL.47'48'49 observed that peak



rates of asthma clinic visits occurred in September for the



years 1957, 1961, 1962, 1964, and 1965 (Tables 7 and 8,



Appendix), but that there was no correlation of the visits



with air concentrations of pollens, molds, or other air



pollutants.  The best correlation seemed to be with the



onset of cold weather.  They speculated either that the

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                                                          12
first onset of cold weather was the "triggering" factor,  or
that the first indoor home heating following the summer
months stirred up dormant allergenic dust and mold particulates,
or that something was present in the air which was not being
measured.
        The monthly admissions for asthma and asthmatic
bronchitis to the Brisbane Children's Hospital in Australia
for three yearsr July 1955 to June 1958 (Table 9, and Figures
1 and 2, Appendix)f showed a minor increase during the
spring months and a major wave in autumn and early winter.23
It was concluded by Derrick et al. that the seasonal occur-
rences of these attacks were not correlated with atmospheric
pressure, temperature, humidity, rainfalls, hours of sun-
shine, or wind velocity or direction.  They suspected the
cause of the seasonal peaks was an unidentified pollen or
pollens.
        New Orleans has been a subject of study since 1958
when an outbreak of asthma occurred resulting in approximately
100 cases and 3 deaths.  This pattern reoccurred in following
years.  In earlier reports, asthmatic attacks have been
associated with certain local wind conditions.68  At first,
spontaneous underground combustion in abandoned city dumps
seemed to be responsible.  However, a recent study suggests
that there is more than one air pollution source causing  an
asthmatic-type disease in New Orleans, and that there are

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                                                          13
probably multiple sources.



        Kantor et al..63 reported on an aerobiological survey



conducted concurrently with a study of 56 clinically deter-



mined asthmatics who lived for a period of one year in the



new Judean desert town of Arad.  The pollen and mold con-



centrations in the air of Arad were about one-third and one-



half, respectively, of those observed at the same time of



day in the central coastal community of Beilinson, Israel,



where the humidity was higher and more vegetation existed.



(See Table 10, Appendix).  Excellent to moderate clinical



responses were observed in 84 percent of the patients during



their stay in Arad, followed by relapse on their return



home to Beilinson.  The speculation was that the mold and



pollen concentrations represented a subthreshold level for



the patients, which they could tolerate without any adverse



effects.



2.1.4  Effects of Vegetable Dusts



        The harmful effect of grain dust on health has been



recognized for many years.  However, there was little interest



in the scientific literature concerning this subject until



Duke29 described four cases of bronchial asthma among flour



mill workers, who experienced asthmatic attacks on exposure



to the dust from the first cleaning of the wheat grain.



Other reports have since appeared describing adverse health



effects caused by grain dust among workers loading and

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                                                          14
unloading grain30'54 and mill workers.101  Williams et al.13-6



surveyed 502 country grain elevator agents in Saskatchewan.



Of these, 54 percent had a history of one or more asthmatic



symptoms, including attacks associated with exposure to



grain dust (oats, wheat, barley, and rye), with barley re-



ported as the most responsible.  There was some evidence



indicating that the allergic response was due to mold spores



(Aspergillus, Penicillium, Mucor,  and Rhizopus) present in



the grain.



        Industrial plants that handle, process, and mill



cereal grains have been suspected of emitting allergenic



dust into the surrounding atmosphere.  The University of



Minnesota campus in Minneapolis is surrounded by such storage



and processing plants.  Outbreaks of asthma have occurred



from time to time among students at the University, and it



was considered that these outbreaks were due to some pollu-



tants from the plants.41  However, although air samples taken



in the vicinity revealed pollens (dependent upon seasonal



release), fungi, plant hair, and starch grains from the



nearby mills, no correlation was observed between their



presence and the incidence of asthmatic attacks.  Coppers



and Paulus43 collected air samples from a similar grain



milling plant area.  Two substances (compounds A and B)



were extracted from the collected particles that could cause



allergic reactions in persons susceptible to bronchial asthma

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                                                          15
and hay fever.  The investigators extracted a number  of



grains, seeds, plants, and weed pollens in the same manner



and were able to isolate one of the two substances (compound



A) from each material examined.  Their results are presented



in Table 11, Appendix.



        Allergic reactions to the castor bean and its products



are well known to the allergist.  Apen et al.7 have reviewed



the literature on the health aspects of castor bean dust up



to 1967.  Castor pomace is the residue that remains after



the castor oil has been removed from the beans of the castor



plant, Ricinus communis.  This pomace contains one of the



most potent allergens known, which, in a fine, light  powder



form, is readily carried by the wind from the processing



plant and shipping areas into the surrounding community. A



number of well-documented outbreaks of illnesses have been



traced to this aeroallergen.  For example, Ordman77 reported



an outbreak of bronchial asthma in South Africa caused by



inhalation of castor bean dust.  The highly allergenic dust



affected 200 persons in a castor oil processing plant.



        The pressed castor bean has been used increasingly



in recent years as a fertilizer.  Small100 reported two



patients with bronchial asthma and two with hay fever caused



by inhalation of castor bean pomace used as a fertilizer-



Additional patients showed positive skin test reactions  to



the castor bean extract.  This investigator anticipated  that

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                                                          16
more cases will occur as the use of the pomace as fertilizer



increases.



        Panzani and Layton80 recorded 478 cases of castor



bean dust allergy in Marseilles and the neighboring country-



side during the period 1951 to 1962.  Pollinosis caused by



the pollen of the castor bean plant also can occur.  Linden-



baum   observed a patient who had symptoms of hay fever



complicated by asthma.  The many castor bean plants growing



in the vicinity of the patient's home were the source of



the pollen.  Layton et al.67 concluded that apparently



castor pollen and castor pomace share common antigens.  Re-



actions to the pollen are milder than to the pomace,  but



the pollen can induce sensitivity to the pomace.



2.1.5  Effects of Molds



        The allergies discussed thus far are considered to



be atopic allergies13—that is, usually characterized by



hereditary predisposition and high and immediate sensitivity.



Desensitization is difficult and usually only partial at



best, skin reactions are marked and specific, and considerable



amounts of antibody are demonstrable in the serum.  However,



in nonatopic allergy, the converse is true; i.e., sensitivity



is low and delayed, desensitization is usually successful,



skin reactions are weak and nonspecific, and few antibodies



are demonstrable in the serum.   Nonatopic allergy can also



be associated with an infection.  Authorities differ on the

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                                                          17
significance of this classification and whether distinction
between atopic and nonatopic allergy is quantitative rather
than qualitative.  However, most agree that the distinction
has clinical utility.  This section primarily discusses
nonatopic allergy.
        Watkins-Pitchford112 reviewed the occurrences of
"farmer's lung" in Great Britain, Europe, and the United
States.  Pulmonary disability among agricultural workers
handling moldy hay and composts has been known for genera-
tions, and many of the early descriptions of such cases un-
doubtedly were what is presently called "farmer's lung."
Some hours after exposure to the hay, acute attacks begin
that are characterized by shortness of breath, fever, and
cough generally followed by recovery.  If the acute attack
has been severe or if exposure is continued, the illness
progresses to a severe dyspnea and cough which become  pro-
gressively worse.  It is generally agreed that "farmer's
lung" is an inhalant allergic response to Thermopolvspora
polyspora growing in the hay.24  Although the true incidence
of the disease can only be conjectural, it has been estimated
that approximately 1,000 cases occur annually in Great
Britain.
        Sakula92 reported respiratory symptoms resembling
those seen in farmer's lung in workers at mushroom-growing
farms.  The process of composting on these farms favors the

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                                                          18
growth of fungi, and the inhalation of their spores is re-
sponsible for allergic response of certain mushroom workers.
Four cases were reported in Sussex, where 50 percent of the
mushrooms in England are commercially grown, and 16 cases
among immigrant Puerto Ricans working in the Chester County
area of Pennsylvania, where 90 percent of the mushrooms con-
sumed in the United States are cultivated.  Similar allergic
responses occur in "bagassosis," caused by inhalation of the
moldy dust of the bagasse fiber (the sugar cane residue left
after removal of the sugar).  Asthma symptoms have developed
in some individuals following exposure to specific wood
dusts. 0'  Byssinosis, or cotton lung resulting from inhala-
tion of cotton dust, has been included in this group in the
past,24 but gome recent studies indicate that it has a re-
versible chemical component resembling that of metal fume
fever and a chronic component.58
        Itkins and Dennis^^ tested 81 patients for sensitivity
to the common fungus Candida albleans.  Forty percent gave
positive bronchial reactions to inhalation of the fungus.
These results are presented in Table 12 (Appendix).
2.2  Effects on Animals
2.2.1  Commercial and Domestic Animals
        Commercial and domestic animals generally are not
affected by inhalation of aeroallergens.

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                                                         19
2.2.2  Experimental Animals



        Experimental animals generally are not affected by



inhalation of aeroallergens.  However, animals are used in



allergy studies.  Dogs sometimes show spontaneous sensitivity


                  1 82
to ragweed pollen. '    Also, researchers at the National  In-



stitutes of Health have begun to work with inbred strains  of



guinea pigs to study the inheritance of allergic tendencies.



2.3  Effects on Plants



        The physiological response of allergy cannot occur in



plants, and, therefore, no allergic effects of aeroallergens



on plants are possible.  However, as an anomalous effect,



growers of genetically pure seeds for crops and flowers are



faced with the problem of airborne cross pollination with


                   ft 7
undesirable plants.    The disease-producing effects of molds,



and other microorganisms on plants is discussed in the com-



panion report on Biological Aerosols.



2.4  Effects on Materials



        As in the case of plants, aeroallergens do not



produce allergic response on materials.  The effects of the



growth of molds and other microorganisms on materials are



discussed in the companion report on Biological Aerosols.



2.5  Environmental Air Standards



        Insufficient information exists on which to establish



environmental air standards for the aeroallergens.  However,

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                                                          20
a pollen count, from which a pollen index is derived,  is



determined daily in many local areas and is compared to



previously observed indexes in the same area.  The comparison



is used as an indication of the incidence of pollinosis to



be expected in the susceptible population.  Generally,



although the values vary from locality to locality, an index



of 5 to 15 is considered moderate, and at this level acute



hay fever symptoms generally last only a few days.  An



index above 15 is indicative of heavy pollen concentrations,



and 25 or more on any given day will usually cause severe



symptoms of hay fever in most of the susceptible population.^



However, individual sensitivities to the pollen vary,  and an



extremely sensitive individual may suffer intensely with a



low index of 10 while another less sensitive individual may



show no response to a count of 100.  This situation may even



reverse itself in another geographical area, where the prev-



alence of certain pollens may be different.



        The pollen index should be considered only as a



guideline rather than as a standard.38  The counts are ob-



tained on rooftops, and the actual exposure value at street



level may be greater or less than this.  Also, the counts



represent integrated values for the 24-hour period and do



not reveal surges of high concentrations which exist for



only short periods of time.  In addition, small clouds of



pollen could occur in certain areas due to local point

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                                                         21
sources or micrometeorological conditions (i.e.,  local  air



calms, turbulences, etc.) and yet not be reflected in the



count obtained short distances away.  Shapiro and Rooks,96



in 1951, placed pollen samplers near known stands of ragweed



in residential areas of Iowa City.  They noted higher values



and marked variation of the pollen counts as compared to



the counts obtained with the standard procedure of a central



sampler on a rooftop.



        Regardless of their limitations as a standard,  the



pollen counts do serve a useful purpose.  The more data of



this type which are accumulated and analyzed with related



meteorological, medical, and control data, the better will



be the total understanding of the problem.  In addition,



the pollen counts aid both the practitioner and the patient.



The symptoms of pollinosis can be better diagnosed and



understood when there is reported evidence that a certain



pollen is prevalent in a given area.  After a period of



time some individuals may even be able to anticipate an



attack of pollinosis on the basis of the daily pollen count



and take some precautionary steps, such as avoiding areas



high in vegetation and taking medication before the onset



of symptoms.

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                                                             22
3.   SOURCES



    3.1  Natural Occurrence



           The aeroallergens  encompass a wide variety of materi-



    als, as shown  in Table 1  (Appendix), but the pollens are the



    most important member of this group.  The plants which pro-



    duce allergenic pollens are widely distributed geographically,



    and their  distribution and seasonal growth characteristics



    in part determine  their importance as aeroallergens.  The



    most prevalent pollen-producing plants have been listed in



    Table  2 (Appendix),  and the most  important pollens of each



    State  and  their seasonal occurrence are presented in Table



    3 (Appendix).



           Ragweed has  been found in all 50 States; it produces



    large  quantities of  pollen, and the pollen grains are especially



    adapted for aerial dissemination  by virtue of their size (20



    |j), shape, and density.60  In addition, because of its aller-



    genic  property for a large percent of the population, it has



    been the most  studied.



           Approximately 40 species  of ragweed are known at the



    present time,  the  majority of which occur in North America.83



    Ragweed is most prevalent  in the  North Central and North-



    eastern States followed by the Southern, Great Plains, Inter-



    mountain,  and  Pacific Coast States.  The weed has also been



    introduced into Hawaii.91  Six species are sufficiently



    widespread and abundant within the United States to be of

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                                                          23
importance as pollen sources on the State or national level.



These ragweed species characteristically establish themselves



quickly in freshly turned soil, and their special ability to



grow as weeds enables them to flourish in cultivated grain



fields, where they are most abundant today.   Pollination is



by aerial dissemination.  Because pollination by wind is very



inefficient, many thousands of pollen grains must be liberated



and disseminated for effective pollination.



        Ragweed tends to be crowded out by other vegetation



if the soil is not disturbed.  Therefore, the plowing of



fields, especially cereal grain fields, is responsible for



the growth of a major portion of the ragweed in many areas



of the country.  The ragweed seedlings develop during the



ripening stage of the cereal grain and grow rapidly in the



stubble after the grain has been harvested in late summer.



An estimated one-third of the 60 million acres of wheat



stubble is infested with ragweed.39  In fields of winter



wheat, oats, and barley, with no cover crop, about 172,000



ragweed plants per acre have been observed,  which is more



than 300 times the plant density in pastures.55  Ragweed is



found also in urban areas where soil has been disturbed.



For example, in a new subdivision where the  soil had been



overturned but untouched during the spring and summer, rag-



weed concentration amounted to 56,500 plants per acre.55



Along railroads, one count gave 13,000 plants per acre.55

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                                                          24
Gorlin44 has estimated that an acre of giant ragweed may



produce as much as 50 pounds of pollen during a single



season.  A ragweed survey was conducted within parts of the



Ann Arbor and Superior Townships of Washtenau County, Mich.



Land along selected roads and highways was classified by



land use/ and the density of ragweed was then determined.



The results of the survey showed that the general pattern



of ragweed distribution prevailed:  high density in crop-



lands and low density in marshes and woodlands, as shown in



Table 13, Appendix.



        The fungi (molds and yeasts) are an important group



of aeroallergens; the most common ones are listed in Table



3, Appendix.  These microorganisms are ubiquitous and sapro-



phytic in nature.  Their usual habitats are soil and dust, and



they become airborne by means of local air disturbances.  Their



concentration in the air is dependent upon the magnitude of



the source, their death rate in the air, humidity, temperature,



and other factors.  The largest percent of the airborne fungi



are found in the air up to 5,000 feet, and their number de-



creases rapidly above that level.  However, viable mold



spores have been recovered up to 90,000 feet.12



        Danders, which include feathers of fowl and hair of



animals (including humans), are found in the air close to



their source point.  Their concentration in the air is limited,



and they are allergenic to humans when the source is in close



proximity to the susceptible individuals.

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                                                          25
        House dust consists of the small particulate organic



materials (fragments of animal hair, wool, cotton linters.



Kapok, feathers, pollen which has seeped in from outdoors,



et;c.) found in the home.  Molds are also frequently found in



the home—especially in old, damp dwellings (e.g., basements)



—and can be included in house dust.  The house dust which



is of concern in this connection does not include sand, soil,



and powdered rock, which are not allergenic.



        Voorhorst et al_.HO concluded from their studies that



the mite Dermatophagoides pteronysissimus was the allergenic



agent in house dust.  They were able to isolate the mite



from such dust, and skin reactions to extracts of this mite



in sensitive individuals were both quantitatively and quali-



tatively indistinguishable from those obtained with house



dust.  In addition, the mite has a worldwide distribution, as



well as seasonal variation of occurrence similar to the fre-



quency of house dust allergenicity (peak in the autumn).



        Although a nonspecific house dust extract is available



for skin testing, some clinicians believe that house dust is



specific for each home.  Because it is found in every indoor



environment, house dust is probably the most common aero-



allergen after pollens.52  Jaggi and Viswanathan61 skin-tested

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                                                          26
patients with seasonal and perennial rhinitis and asthma with



extracts of pollen, house dust, and fungi.   Of the 462 patients



tested, 61.9 percent reacted positively to  house dust.  Of



these positive subjects (286), 17.5 percent were positive to



house dust alone, and the rest were positive to one or more



of the other allergens beside house dust.



        The miscellaneous aeroallergens—which include



vegetable fibers and dust,  cosmetics,  paints, and varnishes



—are limited in the air and affect susceptible individuals



only when in close proximity to the source.  They constitute



a minor problem in terms of the total number of people



affected.



3.2  Production Sources



        The occurrence of certain aeroallergens in the air



has been due to production sources.  A survey of country



grain elevator agents in Saskatchewan showed a greater than



50 percent prevalence of asthmatic and related symptoms in



these individuals.116  Grain dusts produced in flour-milling



plants similarly have been the(cause of asthmatic and other



symptoms in mill workers29'101 and workers  loading and un-



loading grain.30*54  There has been some evidence that flour



mills can emit the allergenic dust into the surrounding



atmosphere.  Coppers and Paulus43 in 1967 were able to isolate



both from the air near a flour mill and from grains an un-



identified compound which could cause allergic reactions in

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                                                          27
persons  susceptible to bronchial asthma and hay fever.
         Inhalation of castor bean dust in castor oil process~
ing plants has been the cause of outbreaks of bronchial
asthma.7  In addition, the use of the pressed castor bean
pomace as fertilizer has caused bronchial asthma attacks in
individuals.7
        The syndrome of farmer's lung has been observed in
farm workers handling moldy hay and compost.U-2  similar
symptoms have occurred among workers on mushroom-growing
farms, 2 and workers inhaling the dust from bagasse fiber
(sugar cane residue) and cotton dust.^4»52
3.3  Product Sources
        There are many materials which are aeroallergenic to
sensitized individuals.  Since some of these materials are
incorporated into other items, their presence cannot always
be recognized.  For example, pillows often contain allergenic
chicken, duck, or goose feathers, or kapok; and stuffed toys
may contain kapok or cat hair-  Table 14, Appendix, presents
a list of some common items which may be aeroallergenic
because of their contents.  '
3.4  Environmental Air Concentrations
        The concentration of aeroallergens present in the
air at any given time is a function of many factors:  source
strength, distance from source, humidity, temperature, and
sunlight.  Of the aeroallergens, pollens have been studied
most; and of the pollens,  ragweed,  the major cause of

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                                                          28
pollinosis in this country, has been studied the most ex-
tensively.  Table 15, Appendix, lists a number of pollens and
dispersal data.
        It has been observed that emission of pollen from
ragweed plants is not continuous but is dependent upon-
the time of day, temperature, and humidity.^5  Pollen re-
lease from the ragweed flower is accomplished in the several
hours after sunrise, as shown in Figures 3, 4, and 8,
Appendix.  The release is triggered by the drop in humidity
resulting from the warmth of the sun.  Usually, only a small
percentage of the pollen becomes directly airborne.  The
great majority of the grains fall on nearby vegetation and
soil but may become airborne at a later time.14  it has
been estimated that only approximately 6 percent of the
released pollen becomes airborne.99
        Once airborne, the dispersal of the pollen is depen-
dent upon the horizontal and vertical air movements.  Raynor
and Ogden®^ in 1965 sampled the air from a cultivated ragweed
source 180 feet in diameter and found that if the horizontal
air flow is relatively slow  (a wind of about 1 m/sec) and
the vertical flow is small, the airborne pollen concentration
becomes negligible within a short distance.  The pollen
concentration had dropped to 1/100 of the source concentration
at 500 feet from the source.  The patterns observed at three
different sampling heights of a similar 90-foot-diameter
ragweed source are presented in Figure 5, Appendix.  Day-to-day

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                                                          29
concentration patterns varied primarily because of changes



in wind direction, speed, and turbulence.  A composite of



these daily patterns showed the seasonal average concentra-



tion around the source, sample at a 5-foot height (Figure 6,



Appendix).  Although the wind was primarily in one direction,



significant amounts of pollen spread in all directions.



        Allessio and Rowley4 in 1956 took a daily pollen



count for 7 months at two sites on the University of Massa-



chusetts campus, while looking for and tabulating 42 differ-



ent pollens plus fern, moss, and fungal spores.  Although



they could not rule out long-distance dispersal by their



results, the presence of all materials observed could be



accounted for by local vegetation.



        If there is sufficient upward air flow due to warmed



air and other processes, the ragweed pollen may be carried



aloft to high elevations.  Pollen grains at 40,000 feet have



been reported, and not infrequently there may be a higher



concentration of pollen at an altitude of 4,000 to 6,000



feet than nearer the surface of the ground.60  Although



some grains may fall to the ground, horizontal dispersal over



long distances can occur.  As local air movement and turbulence



diminish during the day, the pollen grains fall back to earth



at a rate of 3 to 10 ft/min,14'60 and a minimum surface air



concentration occurs just prior to sunrise.  Figures 7 and 8



in the Appendix show this diurnal ragweed pollen emission cycle


     ^                                                   75
        Other pollens also exhibit a diurnal periodicity.

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                                                          30
Acalypha pollen has been reported to reach a maximum between
6 and  7 a.m.,79 Corvlus at 1 p.m., Artemisia at 9 a.m., and
Pinus  at 3 to 5 p.m.50  The patterns for timothy, corn, and
castor bean plant are presented in Figure 9.
        The ability of rain to cleanse the air of pollen has
been investigated.27  Pollen showed a rapid decrease in the
air during heavy rain, with a partial recovery between
separated showers.  There was a possibility that a more or
less continuous replenishment of the airborne pollen con-
tamination into the storm was necessary in order for pollen
to be  found in the rain that fell more than a few minutes
after  the beginning of the rainfall.
        During the ragweed season, daily pollen concentrations
over much of the Eastern and Central United States may reach
350 to 1,000 grainsper cubic meter of air.  During the peak
day of the 1966 season, the pollen count in Ann Arbor, Mich.,
exceeded 4,400 grains per cubic meter.^3  Durham-^ in 1947
estimated that during the ragweed pollen season in the
District of Columbia, an individual would inhale approximately
4,000  ragweed grains in 24 hours.   However, during periods
of maximum air contamination, the grass and ragweed pollen
concentration may reach millions of grains per cubic meter.
Raynor and Ogden^7 (1965) reported that the general back-
ground ragweed pollen concentration at the Brookhaven
National Laboratory, New York,  ranged between 10 and 150
grains per cubic meter.  The area  within a mile or two of

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                                                          31
the Laboratory is relatively ragweed-free,  but at greater



distances ragweed grows in normal profusion in all directions.



        Table 16, Appendix* summarizes the monthly average



counts obtained by the gravity slide method for the most



common pollens found in the St. Louis area from March



through September in both 1963 and 1964.36  The pollen



season begins in early spring with tree pollination, and



continues through the summer with grass pollination, and



into the fall with ragweed pollination.  The weather condi-



tions under which pollen usually was emitted were light



winds, clear to partly cloudy skies, and at least moderate



convection, all of which contribute to vertical movement of



air.



        Hornedo and Tillman57 (1959) reported the results of



a 2-year pollen survey in El Paso, Tex.  Using the gravity



slide method, they observed that weed pollens (Russian



thistle, careless weed, and ragweed) predominated in early



fall; tree pollens (elm, cedar, cottonwood, ash, pine, and



mulberry) in late winter and'early spring; and mulberry,



pine, and oak in late spring.  Peaks in pollen counts



occurred in the fall (about September 30) and in February



and April; and lows were in January, June, and November.



The highest daily count (89 grains per cubic centimeter)



occurred October 4, 1957 (see Figure 10, Appendix).



        Many fungi are common in the air since their spores

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                                                          32
are adapted for aerial dissemination.  Production of vast

numbers of spores in periodic waves is characteristic of

many fungi.  Allergy to fungi may occur seasonally, depending

upon climate and geography.  Hormodendrum and Alternaria are

especially abundant during May to September in the Central

States.20

        Morrow et al.    have summarized the most frequently

isolatedmolds from 41 sampling stations across the country.

No two stations had the same lists/ but a basic group of

dominant genera appeared to occur.  These were:

                Alternaria       Trichoderma
                Homodendrum      Fusarium
                Aspergillus      Helmintho sporium
                Penicillxum      Cryptococcus
                Pullularia       Rhodotorula
                Phoma

        Similar genera of fungi were observed in Tucson,

Ariz.,35 in Phoenix, Ariz.,42 in Albuquerque, New Mex..31

and in Los Angeles, Calif.97

        The airborne concentration of fungi changes from

season to season, from day to day, and even from hour to

hour.  Table 17, Appendix, illustrates the hourly fluctua-

tions of Alternaria spores observed by Pathak and Pady.

Some fungi appear to have a diurnal periodicity.79'81  One

explanation offered for the latter fact is that—as, for

example, in Cladosporum—a single crop of spores is pro-

duced per 24-hour period, maturing at night and ready to be

released just before daylight.  Morning turbulence produces

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                                                          33
a midmorning peak by carrying the spores into the air in



large numbers/(for example, 100 per cubic foot (3,500 per



cubic meter)).  Decreasing air turbulence later in the day



produces a late afternoon or early evening peak.88  pady^S



found fungi spores present in the atmosphere at an elevation



of 150 feet throughout the year in Kansas, with peaks in



July and August.  In summer their number varied from 50 to



700 per cubic foot (1,765 to 24,700 per cubic meter), while



in winter they ranged from 5 to 20 per cubic foot (175 to



700 per cubic meter).

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                                                             34
4.   ABATEMENT
            The abatement and control of  aeroallergens have been
    primarily directed at the control of  ragweed.  Ragweed grows
    very quickly in areas where the  soil  has been  disturbed; it
    is not found in areas shaded by  trees or in heavy growth of
    grass, shrubs, or ferns.  If soil is not disturbed, grass will
    eventually crowd out ragweed. Although some ragweeds are
    annual plants,  the seeds can remain viable for many years
    and ready to germinate once the  soil  is disturbed.  Therefore,
    control of ragweed by pulling it up is not satisfactory,
    since the soil  is thus disturbed and  the growths may be
    heavier the following year.  Control  by cutting is satisfac-
    tory only if done prior to flowering; otherwise, flower
    heads will continue to develop and pollinate.  Soil steri-
    lants are toxic to most other plants  as well as to ragweed;
    and if successful,  their use leads to the problem of soil
    erosion.
            Herbicides such as 2,4-D (dichlorophenoxyacetic
    acid salts) have been used successfully where  applicable.
    Since 2,4-D is  lethal to most broadleaf plants (including
    ragweed) and to vegetables, flowers,  and some  grasses, its
    use on crop acreage is limited.
            The recommended application of 2,4-D is one-half
    pound per acre, although 0.1 pound per acre has also yielded
    good results.39  It is best used when diluted  in water (5
    to 100 gallons  per acre).   The spraying program for ragweed

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                                                          35
should be initiated just before the flower buds open.   The

herbicide 2,4,5-T (trichlorophenoxyacetic acid),  a compound

similar to 2,4-D, has also been used.  The potential side

effects and toxicity of herbicides are discussed in a sepa-

rate report of this series, "The Air Pollution Aspects of

Pesticides."

        Considerable money and effort have been expended by

several municipalities in attempts to reduce or eliminate

ragweed within their boundaries, but generally these attempts

have not significantly changed pollen concentrations.^^

Walzer and Siegle^-H reported on the effectiveness of a

ragweed eradication program in New York City.  After the

program was initiated in 1946, a 50 percent reduction in

ragweed plants within the city was observed at the end of 4

years.  However, the program did not produce any further de-

cline during the next 5 years.  Also, during this 9-year

period there was no change in the pollen count from 30

stations in and around New York City (see Figure 11, Appendix).
                             i
During certain seasons/ pollen counted on a lightship in

New York harbor, at a point 9 miles from the nearest land,

amounted to as much as 45 to 60 percent of the pollen collected

in New York City, and on some days exceeded those of the city.

The data indicated that the city probably received as much

windborn pollen from areas to the west as was generated

locally.  Furthermore, no differences were found on comparing

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                                                          36
the New York City counts and those from neighboring New



Jersey and Connecticut, where no eradication programs were



underway.  These authors concluded that the elimination of



ragweed pollen in the air cannot be accomplished on a local



level, but that any program must be developed on a regional



basis.  Obviously, if the pollen entering a city from outside



is enough to cause pollinosis in all susceptible individuals,



local control is inadequate.  On the other hand, if the



background pollen reaching a given local area is low—that



is, insufficient to cause symptoms in the population—



eradication of any local sources would be a beneficial



preventive measure which could be undertaken by a local



authority.  The latter situation prevailed in Detroit, where



a ragweed control program was accomplished.  Nearly all of



the pollen observed in sampling counts was produced in the



immediate vicinity.  Local eradication of ragweed plants,



therefore, was beneficial in controlling the pollen concen-



tration.39



        Some aeroallergens are troublesome only in close



proximity to susceptible individuals.  Eradication of the



nearby source often can reduce the daily exposure to such



a degree that the individual's threshold sensitivity is



decreased and he will not experience severe symptoms with



normal background levels.  In particular* the allergenic



effects traceable to trees,  plants/ and flowers can be

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                                                     37
avoided by not growing them in the yard.  Similarly,  some


allergies can be prevented by not keeping pets that produce


danders and by avoiding certain cosmetics.



        Molds, like pollens, are found in the outside air,



and to some extent in the home.  The concentration found in



the home is associated with house dust which may also be



allergenic to susceptible individuals.  Many of the allergies



due to molds arise from specific molds found in damp  places



in the home (e.g., basements).  These can be reduced  in


concentration or eliminated by the use of disinfectants.


            21
Criep et al.    have recommended the use of inexpensive



Roccal (benzalkonium chloride) at 1:1,000 to 1:10,000



dilution as a spray, or 1:1,000 dilution of trioxymethylene



(crystalline paraformaldehyde) as a wash solution or  vapor



for treating musty houses.  Table 18, Appendix, lists other



germicidal substances which can be used.


        Another approach to the control of pollinosis has



been for sensitive individuals to avoid contact  with the



pollen by remaining indoors as much as possible during the



hay fever season.  Since 67 percent of the ragweed pollen



collected during 24 hours was found between 9 a.m. and 1 p.m.,



Smith and Rooks102 (1954) have recommended that sensitive


individuals seek "shelter" during those hours.  However,


Dingle26 (1957) has demonstrated that ragweed and other



pollens can penetrate the cracks around doorways and  windows

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                                                          38
of the average house.  The amount of penetration was dependent
upon the meteorological conditions.  Air-conditioned buildings
give some relief to susceptible individuals,  but most air
systems are not designed for aeroallergen control.   This is
especially true of home systems where house dust and/or
molds are involved.  The air ducts harbor large quantities of
dust and molds, and the filters usually are not adequate to
remove them from the air stream.  However, present-day
technology is capable of adequately designing air systems
which benefit sensitive individuals.22
        Control of pollinosis is often accomplished by
temporary or permanent departure of the individual from a
given area to one which is free of the specific pollen to
which he is sensitive.  A map of the United States showing
&reas relatively free of ragweed pollen is shown in Figure
12, Appendix.  Although moving has been of benefit to many,
some individuals after moving may develop a sensitivity to
a new local pollen, or a plant to whose pollen he was already
sensitized may be introduced into the area as an ornamental
plant.
        Many susceptible individuals can be treated so that
exposure to the aeroallergen does not produce symptoms or
the symptoms are reduced in severity.  Extracts of various
allergens are available which can be injected in small
amounts into susceptible individuals to desensitize them

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                                                          39
temporarily to the allergen.  Some persons routinely undergo
a preseasonal series of desensitizing injections and later
develop few symptoms during the aeroallergen season.  The
antihistaminic drugs have been used quite extensively to
alleviate the acute symptoms of hay fever.  However, anti-
histamines hav.e not been a cure-all, and some individuals
react adversely to these medications.

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                                                             40
5.   ECONOMICS
            Ridker90 has stated that because  in many cases there
    are either insufficient or no  data  concerning the number of
    persons with an illness and very little information concern-
    ing the cost of treatment, the economic loss due to the health
    effects of air-pollutants is most difficult to  estimate.
    This applies equally to the economics  of  aeroallergens.
    Ridker has attempted to place  a conservative dollar value
    estimate on some diseases (Table 19, Appendix), with diseases
    other than asthma listed for comparative  purposes.  The
    costs of prescribed medicine for asthma and/or  hay fever as
    reported by the National Health Survey are presented in Table
    20 (Appendix)-
            Because of this difficulty  in  estimating the cost of
    effects on health, another approach is to consider the in-
    cidence and prevalence of the  illnesses.  The Allergy Founda-
    tion of America has estimated  that  8 to 9 million people in
    the United States are adversely affected  by seasonal hay
    fever."  The addition of nonseasonal  sufferers would increase
    this figure. • The National Health Survey  has reported that
    the number of asthma and/or hay fever  sufferers was over
    14,000,000 in 1964.3  It has been estimated that 5 to 10
    percent of the untreated patients with hay fever develop
    bronchial asthma.10  Some estimates place the number of work
    days lost each year due to hay fever at 25 million.

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                                                          41
        Data pertaining to the "limitation in activity" by



asthma and/or hay fever sufferers as reported by the National



Health Survey are shown in Tables 21 and 22 in the Appendix.



        Besides the direct cost and discomfort associated



with hay fever and bronchial asthma, there are secondary



economic effects:  the ensuing lack of sleep and fatigue



lowers efficiency at work.  Additional secondary economic



consequences are the sedative effect of medications, the



danger of a sneezing attack while driving an automobile or



while operating some mechanical device which could be hazard-



ous to others, and the fact that swollen respiratory passages



are prone to bacterial overgrowth and infection which may



continue far beyond the hay fever "season."



        The death rate due to asthma for 1964-1966 compared



to other selected causes of death is presented" in Table 23,



Appendix.



        Of the aeroallergens, the pollens are the worst



offenders, and ragweed pollen specifically is responsible



for greater than 90 percent of the pollinosis in the United



States.  Some costs are available for the control of ragweed.



Freedman39 (1967) has reported that in New Hampshire in 1948,



ragweed control cost about $2.00 for one mile of highway on



both sides.  This figure included labor, cost of the chemical,



and truck mileage.  In 1967, a similar control program along



roadsides and rights-of-way, using a combination of chemical

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                                                          42
and mechanical measures,  probably would have cost  in  the



range of $5 to $10 per acre.39  in open areas,  such as



pasture and wheat fields, the weed could have been controlled



with 2,4-D at an annual cost of $1.50 per acre;  in congested



areas the cost of control by mechanical techniques or in



combination with sprays would have been approximately $25



to $50 per acre.39

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                                                             43
6.  METHODS OP ANALYSIS



            The methods used for the analysis of aeroallergen



    pollution are based primarily  on microscopic observations of



    collected samples from the air.   Basically, these procedures



    are qualitative, but a relative degree of quantitation  is



    introduced by standardization of the procedure.  Some



    quantitative procedures are used that sample a given volume



    of air, and the results are expressed in terms of a count



    per unit volume of air.  Most of the procedures have been



    concerned primarily with pollen and molds;  little attempt



    has been made to sample for the other aeroallergens.



    6.1  Qualitative Methods



            The "gravity slide" method for pollen sampling, first



    used by Durham^S in 1946, was accepted as the standard  pro-



    cedure by the Pollen Survey Committee of the American Academy



    of Allergy8^ in the same year.   This standard air sampling



    device consists of two circular parallel planes of polished



    steel 9 inches in diameter and 3 inches apart, with a slide



    holder raised 1 inch above the lower plane.  It is supported



    by a 30-inch metal rod on a tripod laboratory stand.  A



    petrolatum-coated slide is placed in the slide holder and



    exposed to the air on an unobstructed seven- to eight-story



    rooftop for 24 hours.  The entire exposed area of the slide



    (4.84 cm2) is examined microscopically and  the pollen counted.



    The count is divided by 4.84 and expressed  as a count per

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                                                          44
square centimeter, or simply as a number-   The count can be



converted into short ragweed pollen grains per cubic yard



by multiplying by the factor of 3.6; to giant ragweed by



3.87; to timothy by 1.14; to corn by 0.17; and so forth.32



        The pollen count is determined (by many local authori-



ties) by daily exposing a series of these Durham gravity



slides at various sites in and about an area.  Some slides



may be exposed at ground level. The number of particles



trapped on the slide is dependent upon wind conditions during



the sampling period, and therefore, it is difficult to relate



the counts to actual concentrations in the air.  However, the



pollen counts thus obtained, after several years, show a



pattern of pollen concentration increase and decrease and



correlate to some degree with the general incidence of hay



fever in a given local area.



        The gravity slide method for pollen determination



has a number of limitations. In particular, the sampling is



for a 24-hour period and does not give any indication of



pealc concentrations which might have existed at any time



during the sampling period.  Also, Ogden and Raynor74



demonstrated that slides placed parallel to the airflow



collect much more pollen than those placed at right angles,



and this difference becomes greater at the higher wind



speeds prevalent at greater heights.  An increase of 3 to 4



miles per hour in wind speed may result in a 50 percent

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                                                          45
increase in the amount of pollen trapped.2  Therefore,  the
gravity slide method yields values which may not be entirely
comparable to neighboring sampling sites because sampling
heights and wind speed and direction cannot be standardized.
6.2  Quantitative Methods
        Several volumetric devices are available for drawing
a measured amount of air (using a vacuum pump) into a
sampler.  The intent here is to determine as accurately as
possible the actual concentration present in the air at any
given time.  A photoelectric, continuous-recording particle
sampler has been used by Smith and Rooks (1954) for studying
the diurnal fluctuations of airborne ragweed pollen. ^2
Raynor®6 made use of a membrane filter device with an attached
timer and measured air intake to obtain a series of sequen-
tial pollen samples.  The membrane filters were then viewed
through a microscope and the counts determined.  The Hirst
Spore Trap draws a measured amount of air through an orifice,
and the pollen is impacted on a microscope slide moved past
the orifice at a rate of 2 mm/hr by a clock mechanism.   A
24-hour sample thus can be obtained, but the deposition has
been spaced in time along the slide.56
        Volumetric sampling devices have presented the problem
of isokinetic sampling.  That is, with volumetric samplers,
the intake opening must be continuously oriented into the
wind, and the airflow through the sampler must be equal at

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                                                          46
all times to the wind speed in the free air approaching the
intake.  If these conditions are not met, a true representa-
tive sample cannot be obtained for particles the size of
pollens. ^3
        Because the use of volumetric samplers is too difficult
in routine pollen sampling, the simple Durham gravity slide
method has remained the standard technique in spite of its
deficiencies.  Although it is inaccurate for short-term (1
day or less) measurements, it has been satisfactory for de-
termining seasonal patterns.  However, a number of devices
have been devised which attempt to retain simplicity but yet
improve upon the gravity slide method.  The simplest sampler
has been a vertically-oriented wire of about 1 mm in diameter
which is placed in the air stream containing pollen.  The
air can go around the wire but the pollen is impacted on the
surface.  The wire is then examined through a microscope
and the pollen grains counted.  An improvement upon this has
been the flag sampler.^1  It consists of an ordinary house-
hold pin set in a glass bearing in which it moves freely.  It
has a flag of transparent tape wound about it that works like
a weather vane to keep the coated leading edge of the pin
facing into the wind.  Particles unable to follow the air
stream around the curved surface of the pin impact upon it
and are counted by means of a microscope.  A similar device
uses a larger wind vane to keep the edge of a microscope

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                                                          47
slide facing into the wind to act as the trapping surface.



Such samplers are inexpensive and are suitable for use when



a large number of samples are to be taken.   However/  their



disadvantages are that they are efficient only when there is



some wind (at least 5 m.p.h.), and the impaction surfaces



are quickly covered and require frequent changes.  Also,  the



wind velocity and fluctuations need to be known, which re-



quires the use of a separate recording anemometer.



        Another approach to impaction sampling has been to



mechanically move an adhesive-coated surface through the air



to be sampled.  The rotorod sampler-^ consists of two



vertical rods (plastic or metal) rotated about a vertical



axis approximately 2 inches away at a speed of about 2,000



rpm.  The coated collecting surface of the rods moves at a



tangential speed of approximately 25 mph, which is higher



than most air velocities sampled and thereby has a relatively



high collection efficiency independent of wind speed.  The



rods are examined through the microscope and pollen counts



made.  Modifications of this'device have been the rotobar,51



with a bar-shaped surface used instead of a rod, and the



rotoslide,74 which uses a microscope slide.  The main dis-



advantage of these samplers has been that a high concentra-



tion of pollen can build up on the impaction surface in a



short time (an hour or less) and, therefore, frequent changing



is required when continuous sampling is desired.  To obviate

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                                                          48
this difficulty, the rotodisk sampler15 has been devised,
which substitutes disks for the rods.  The vertical edge of
the disk is covered except for a small slit, and a timing
mechanism automatically shifts the slit to expose a fresh
sampling surface.
        Fungi have also been sampled by the methods given
above.  As the pollen are being counted, some investigators
may also count mold spores.  However, fungi lend themselves
to other sampling procedures that utilize growth of the or-
ganisms as a means of measurement.  The basic methods are:
        (l)  Sedimentation:8"In this simple method of
sampling airborne organisms, the suspended particulates are
allowed to settle on plain surfaces or on surfaces coated
with a nutrient growth medium.  This method yields informa-
tion on the total number of viable particles that have
settled out during the given sampling period of time.
        (2)  Impingement into liquids:19'37'45'72  Air is
drawn through a small jet and is directed against a liquid
surface, the suspended fungi being collected in the liquid.
Because of the agitation of the particles in the collecting
liquid, aggregates are likely to be broken up.  Therefore,
the counts obtained by this method tend to reflect the total
number of individual cells in the air and are higher than the
value obtained by other methods.
        (3)  Impaction onto solid surfaces:6'28  Air is

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                                                          49
drawn through a small jet(s) and the particles are deposited
on dry or coated solid surfaces, or on an agar nutrient.
This method has been used to determine total cellular numbers,
size distribution, total viable numbers,  and variation in
concentration per unit of time during a long sampling period.
        (4)  Filtration:70'73,95,108 T^ particulates are
collected by passage of the air through a filter which can
be cellulose-asbestos paper, glass wool,  cotton, alginate
wool, gelatin foam, or membrane material.  The particulates
are washed from the filters and assayed by appropriate
microbiological techniques.  Since the viability of the
organisms can be detrimentally affected by dehydration in
the air stream, the results may be biased in this method.
        (5)  Centrifugation:93,114  The particulates are
propelled by centrifugal force onto the collecting surface,
which can be glass or an agar nutrient.  Particulate size
and particulate concentration can be obtained by this
method.
        (6)  Electrostatic precipitation:66  Particles are
collected by drawing air at a measured rate over an electri-
cally charged surface of glass, liquid, or agar.  The
number of particles or viable number is then determined.
        (7)  Thermal precipitation:65  The organisms are
collected on surfaces by means of thermal gradients.  The
design is based on the principle that airborne particles

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                                                          50
are repelled by hot surfaces and are deposited on  colder



surfaces by forces proportional to the temperature gradient.



The particle size distribution can then be determined.

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                                                             51
7.   SUMMARY AND CONCLUSIONS
            Aeroallergens are airborne materials which elicit
    a hypersensitivity or allergic  response  in  susceptible in-
    dividuals.   The major effects of  aeroallergens on human
    health are  the production of allergic  rhinitis and bronchial
    asthma.  If the symptoms of allergic rhinitis occur during
    a particular season of the year,  it is commonly called hay
    fever.  It  has been estimated that there are 10 to 15 million
    hay fever sufferers in the United States and that 5 to 10
    percent of  the untreated patients will develop bronchial
    a sthma.
            The common aeroallergens  affecting  human health are
    pollens of  wind-pollinated plants/ molds, house dust, and a
    miscellaneous group of vegetable  fibers, cosmetics, paints,
    and others.  The pollens are the  most  important of the entire
    list,  and ragweed provides the  most common  of the pollens.
    More than 90 percent of the pollinosis occurring in this
    country is  due to ragweed pollen.
            Laboratory animals are  used routinely in allergy
    studies, but exposure is usually  by injection? most animals
    do not exhibit allergenic reaction to  inhalation of aero-
    allergens.   There is no evidence  that  aeroallergens have
    adverse effects on plants or materials.
            Insufficient information  exists  to  establish environ-
    mental air  standards for the aeroallergens.  Daily pollen
    counts are  taken and pollen indexes derived in many local

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                                                          52
areas of the country by the use of a standardized procedure.
However, because many variables are involved,  these values
are used more as guidelines than as standards.  Generally,
indexes of 5 to 15 are considered moderate,  and acute hay
fever symptoms last only a few days.  Indexes  above 15
are indicative of heavy pollen concentrations, and an index
of 25 or more on any given day will usually cause severe
symptoms of hay fever in most of the susceptible population.
These values are relative, however, and may vary considerably
between local areas.
        Ragweed establishes itself readily in  freshly turned
soil, and therefore is found in abundance both in farmlands
and in urban areas in most parts of North America.  Of the
other aeroallergens, the molds are ubiquitous; their usual
habitat is the soil and dust, and they become  airborne
through local air disturbances.  House dust consists of
small organic particulates.  Because it is found in every
indoor environment, house dust is probably the most common
aeroallergen after pollens.  Danders and other similar aero-
allergens are found in the air close to their  source, and
their concentration in the air is therefore limited.  They
are allergenic to humans when the source is in close proximity
to the susceptible individual.
        The potential of other air pollutants  to act syner-
gistically with the natural allergens has become a new area

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                                                          53
of study in recent years.  Several investigators have ob-
served an increase in hospital admissions for bronchial
asthma on days of high air pollution.
        There are many materials which are aeroallergenic
to sensitized individuals.  However, some of the allergens
are incorporated into products in such a way that their
presence cannot always be recognized.  Stuffing in pillows,
mattresses, and toys may be of feathers, kapok, or other
materials that can be highly allergenic.
        The emission and dispersal of ragweed pollen have
been studied in much detail.  It has been found that pollen
release occurs primarily in the early morning, and once the
pollen is airborne, its dispersal is dependent upon horizontal
and vertical air movements.  If there is little air movement,
dispersal of the pollen from a given source may be negligible.
However, upward air flow can carry pollen up to high eleva-
tions, whereas horizontal air movements can carry the pollen
great distances in all directions.  During the ragweed
season, daily pollen concentrations over much of the Eastern
and Central United States commonly reach 350 to 1,000 grains
per cubic meter of air.
        The abatement and control of aeroallergens have been
concentrated on ragweed.  Considerable money and effort have
been expended by local municipalities in attempting to
reduce the pollen concentration in the air by reducing the

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                                                          54
ragweed plant density.  Herbicides such as 2,4-D have been
used extensively for this purpose.  However,  many of the
eradication programs have had little success, primarily be-
cause windborne pollen from outside the control area usually
has entered the city in sufficient quantities to cause
pollinosis in the local susceptible population.
        The economic costs incurred by the effects of and the
control of aeroallergens cannot be adequately estimated.
Insufficient data are available regarding the costs of allergic
illnesses, and there are no estimates for the cost of abatement
on the regional scale that would be required for adequate
control.
        The standard procedure for the analysis of pollens
recommended by the Pollen Survey Committee of the American
Academy of Allergy is the gravity slide method.  Basically,
the procedure involves exposing an adhesive-coated slide to
the air for 24 hours, following which it is examined micro-
scopically and a count made of the particles deposited.
        Based on the material presented in this report,
further studies are suggested in the following areas:
        (1)  Additional investigations are needed concerning
the cause of periodic peak occurrences of bronchial asthma
and associated illnesses.
        (2)  There is need for a relatively inexpensive
automatic device for both research and routine sampling and

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                                                          55
and counting of pollen and other aeroallergens.



        (3)  A cost benefit analysis of regional versus local



ragweed control programs is warranted.



        (4)  Better estimates are needed of economic costs



associated with the illnesses caused by aeroallergens.



        (5)  The design of air systems, especially for homes/



should be evaluated and improved for indoor control of



aeroallergens.

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                                                               •56
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                                                                57
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                                                                59
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                                                               60
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-------
                                                               61
64.  Kern, R. A., Environment in Relation to Allergic Disease,
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-------
                                                                62
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 77.  Ordman, D., An Outbreak of Bronchial Asthma in South Africa,
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 78.  Pady, S. M., Quantitative Studies of Fungus Spores in the  Air,
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                                                                63
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                                                               64
102.   Smith,  R.  D., and R. Rooks,  The Diurnal Variation of Airborne
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103.   Solomon, W. R., Comparative Effects of Transient Body Surface
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104.   Solomon, W. R., Air Pollution by Ragweed Pollen.  IV.  Aspects
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107.   Susman, A. J., et al., Hyper sensitivity to Wood Dust, J..
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108.   Thomas, D. J., Fibrous Filters for Fine Particle Filtration,
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109.   Tuft, L.,  G. I. Blumstein,  and Y.  M. Hecks, Pollen Tolerance
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110.   Voorhorst, R., F. T. M. Spieksma,  H. Varekamp,  M. J.  Leupen,
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111.   Walzer,  M., and B. B. Siegle,  The Effectiveness of the Rag-
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112.   Watkins-Pitchform, J., Farmer's Lung:  A Review,  Brit. J.  Ind.
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113.   Watson,  H. H., Errors due to Anisokinetic Sampling of Aerosols,
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                                                               65
114.   Wells,  W. F.,  Apparatus for Study of Bacterial Behavior of
      Air,  Am. J. Public Health 23:58 (1933).

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116.   Williams, N.,  A. Skoulas, and J. E. Merriman, Exposure to Grain
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117-   Zeid.berg, L.  D., R. A. Prindle, and. E. Landau, The Nashville
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      A Preliminary Report, Am. Rev. Respirat. Diseases 84:489 (1961),

-------
APPENDIX

-------
                                                             67
                       Asthma
               Ul
               C
               o
               E
               •0
               o
               6
Asthmatic Bronchitis
                     JULY  OCT.   JAN.  APR.  JULY
                           FIGURE 1

         Seasonal  Variations in Respiratory Illness.
   Monthly admissions  to the Brisbane Children's Hospital  in
   Australia,  1955-58,  for asthma, asthmatic bronchitis, and
                        bronchitis.
                        1955-56
                     JULY OCT.  JAN.   APR.  JULY
                            FIGURE 2

Seasonal Fluctuations  in  Respiratory Illness for Several Years
Monthly admissions  for asthma (including asthmatic bronchitis)
to the Brisbane  Children's  Hospital in Australia for each year
of the study.  The  spring and autumn waves occurred in each
year, with some  variation in height and      ^

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                                                                68
4 h
                               O
                                             Morning
                                            (0830-1030)
                                                Afternoon
                                               (1330-1530)
  19  20  21  22 23  24  25  26 27  28  29  30 31  01  02 03  04  05  06
  AUG                                       SEPT
                          FIGURE 3
 Ragweed  Pollen Concentrations During the 19
           Season at  and near Ann Arbor/ Mich.
Ragweed

-------
  14000
  12000  -
  10000  -
   8000  -
.E

(/>
c
   6000 —
    4000 —
    2000 —
      0626
                            I         I

0650      0714     0738     0802      0826


                           Hours ( E.S.T. )
                                                           0850
0914
0938
1002
                                                                                                    <^
                                                                                                    vO
                                       FIGURE  4


  Variations  in Ragweed  Pollen  Concentrations Close to the  Pollen  Source.

-------
                                                              70
       5 FEET
                              10 FEET
                                      F£CT 0 20 40 60 8O IOO
                                                     is FEET
                             FIGURE 5

      Ragweed Pollen Concentration Patterns  (grains/mr' ) .
These patterns were observed  at heights of 5,  10,  and 15 ft in
            an east field  plot',  August 10-11,  196187
                                             100
                                             100
                                             100
                                     0  50  100
                                       FEET
                             FIGURE 6
   Seasonal  Average Concentration of Ragweed Pollen  (grains/m3)
         Sampled at 5 ft, east  field plot, in-season,  1961.»

-------
o
_>
§ 10
          Q>
          O
             o
                          0600
                                   BACKGROUND RAGWEED
                                                            1961-64
                              1200          1800
                               Time ( E.S.T. )
                                2400
                                          FIGURE 7

                    Diurnal Ragweed  Pollen Concentration Patterns  at  a
                          Location Distant from a Local Source
5 20
                  PRESEASON
                               RAGWEED

                                1961-63
                           IN-SEASON
                0600
                1200
1800
 2400    0
Time  ( E.S.T. )
                                                         0600
1200
1800
2400
                                          FIGURE 8

                Diurnal Pollen Emission Patterns  from Fields of Ragweed
                                                                 76

-------
   40
   20
>

S  20
Q

«-^
o

-------
Pollen Count
   60
   50
    40
    30
    20
     10
                                                                                   I    I    I
i	I
         SEPT. OCT. NOV. DEC. JAN. FEB. MAR. APR. MAY. JUNE JULY AUG. SEPT. OCT. NOV. DEC. JAN.  FEB. MAR.APR, MAY JUNE JULY AUG.
         1957                               1958                                        1959
                                                      FIGURE  10
                                 Weekly Average Pollen Counts,  El Paso, Tex.
                                                                                   57

-------
                                                      74
   55


   50

   45


   40

V  35
   30
o
0
£  25
   20

   15
    1946
1948
1950
1952
1954
                    FIGURE  11

  Pollen Count for New York. City, 1946-1954
                                  111

-------
LEGEND
      * Excellent
      + Good
      ® Fairly Good

All solid discs represent by smaller or larger
size the varying pollen pollution in different
area] or in specific communities.
                                                    FIGURE  12
                           Ragweed  Pollen  Refuges  in  the  United States
                                                                                         34
                                                                                                                                  - 1
                                                                                                                                  (J1

-------
APPENDIX
                                                          76
                         TABLE 1

                 COMMON AEROALLERGENS52'64
Aeroallergens
             Source
Pollens


Molds


Danders
House dust
Mi scellaneous
  Vegetable fibers
    and dusts
  Cosmetics
  Insecticides
Wind-pollinated plants, grasses
  weeds,  and trees

Usually saprophytic, prevalence
  depending upon humidity

Feathers of chickens, geese,
  ducks;  and hair of cats, dogs,
  horses, sheep, cattle, labo-
  ratory animals,'and humans

A composite of all dusts found
  about the home
Cotton, kapok, flax, hemp, jute,
  straw, castor bean, coffee
  bean, oris root, rye, wheat

Wave set lotions, talcs, per-
  fumes, hair tonics

Insecticides containing pyre-
  thrum as a common ingredient
  Paints, varnishes,
    and glues
Linseed oil and organic solvents

-------
APPENDIX
                                                              77
                             TABLE 2

                  COMMON WIND-POLLINATED
Common Name
   Botanical Name
Diameter
(microns)
                                                          Specific
Giant ragweed
Burweed marsh elder
Short ragweed
False ragweed
Marsh elder
Southern ragweed
Western ragweed
Cocklebur

Russian thistle
Palmer's amaranth
Western water hemp
Mexican fireweed

Annual sage
Tall wormwood
Sagebrush

Nettle
Red sorrel
Hemp
English plantain

Bluegrass
Bluegrass
Bermuda grass
Orchard grass
Timothy
Rye
Corn

Sycamore
Mountain cedar
Hazelnut
Birch
Alder
Ash
Cottonwood
Elm
Bur oak
Shingle oak
Ambrosia trifida          19.25
Iva xanthifolia           19.3
Ambrosia elatior          20.0
Franseria a.canthicarpa    22.0
Iva ciliata               23.0
Ambrosia bidentata        23.0
Ambrosia psilostachya     26.4
Xanthium commune          27.0

Salsola pestifer          23.6
Amaranthus palmeri        25.8
Acnida tamariscina        27.5
Kochia scoparia           32.7

Artemisia annua           20.4
Artemisia caudata         21.0
Artemisia tridentata      25.85

Urtica qracilis           14.0
Rumex acetosella          21.45
Cannabi^ sativa           25.0
Plantago lanceolata       27.5

Ppa pratensis             28.0
Poa pratensis             30.0
Capriola dactvlon         28.5
Dactylis glomerata        34.0
Phleum pratense           34.0
Secale cereale            49.5
Zea mays                  90.0
Platanus occidental's     22.22
Juniperus sabinoides      22.8
Corylus. americana         23.6
Betula niqra              24.6
Alnus glutinosa           26.0
Fraxinus_ americana        27.1
Populus virainiana        30.0
Ulmust americana          31.2
Quercus macrocarpa        32.3
Quercus imbricaria,        33.1
              0.52
              0.79
              0.55*
              0.75
              0.58
              0.50
              0.57
              0.45

              0.90
              1.02
              1.01
              0.97

              1.02
              1.04
              1.03

              0.77
              0.78
              0.82
              0.97

              0.90
              0.90
              1.01
              0.91
              0.90
              0.98
              1.00

              0.92
              1,
              1,
  ,08
  ,09
0.94
0.97
0.90
0.79
1.00
1.04
1.04
                                                       (continued)

-------
                                                             78
APPENDIX
                       TABLE 2  (Continued)


                 COMMON WIND-POLLINATED PLANTS11
Common Name
Walnut
Beech
HicTcory
Scotch pine
Bull pine
Botanical Name
Juqlans niqra
Faqus grandifolia
Carya ovata
Pinus svlvestris
Pinus pondero sa
Diameter
(microns)
35.75
44.0
45.0
52.0
60.0
Specific
Gravity
0.93
0.94
0.79
0.45
0.45

-------
                  TABLE 3
POLLEN SEASONS THROUGHOUT THE UNITED STATES
                                            60
Location
Alabama
Montgomery
Arizona
Phoenix
, Kingman
Arkansas
Little
Rock
California
North
Western
Southern
Area
San Fran-
cisco Bay
Area
Tree
1/15
6/7
2/1
5/1

2/7
5/7
2/1
7/1
1/15
3/1
2/15
6/15
Grass
4/1
10/1
4/1
11/1

5/15
10/1
4/1
8/7
3/1
12/1
4/1
1/1
Rag-
weed
9/1
10/7
9/15
11/1
8/1
10/15
8/15
10/15
7/1
11/15
7/7
11/1
6/22
11/1
Ama-
ranth

5/15
12/22





Russian
thistle


6/15
10/1


7/1
11/1

Salt
bush


6/15
10/1

4/1
9/22


Saqe




7/1
11/15
7/7
11/1
6/22
11/1
Chene-
pod




4/1
9/22


Dock






5/1
9/1
Plan-
tain






5/1
9/1
Kochia







Hemp







Elm







                                                               (continued)

-------
            TABLE 3 (Continued)



POLLEN SEASONS THROUGHOUT THE UNITED STATES
Location
Colorado
Denver
(Connecticut
(Delaware
1
District of
Couiribia
Washing-
ton
Florida
Miami
Tampa
teorgia
Atlanta
Tree
3/15
4/15
3/15
5/22
3/1
5/15
2/1
5/15
2/1
4/1
2/7
5/15
1/15
5/7
Grass
6/7
7/15
5/15
7/15
5/15
7/7
5/15
7/7
3/1
6/1
1/1
12/31
5/1
9/15
Rag-
weed
8/1
10/1
8/15
9/15
8/15
10/1
8/15
10/1
5/15
9/15
8/7
12/1
8/15
10/1
Ama-
ranth







Russian
thistle
7/7
9/15






Salt
bush







Saqe
8/15
10/1






Chene-
pod







Dock







Plan-
tain







Kochia
7/7
9/15






Hemp



i



Elm







                                                                (continued)
                                                                         oo
                                                                         o

-------
            TABLE 3  (Continued)




POLLEN SEASONS THROUGHOUT THE UNITED  STATES
Location
Idaho
Southern
Area
Illinois
Chicago
Indiana
Indiana-
polis
Iowa
Ames
Kansas
Wichita
Kentucky
Louisville
(Louisiana
1 New
1 Orleans
Tree
3/15
5/1
3/15
6/1
3/15
6/7
3/15
5/15
3/1
6/1
3/1
6/1
1/1
4/1
Grass
5/1
8/7
5/22
7/15
5/15
7/7
5/15
9/1
5/1
6/15
5/15
7/1
4/1
12/1
Rag-
weed
8/15
9/22
8/15
10/1
8/15
10/1
8/15
10/1
8/15
10/1
8/15
10/1
8/15
10/22
Ama-
ranth




7/15
10/1


Russian
thistle
7/7
10/1



7/15
10/1


Salt
bush
7/7
10/1






Sage
7/22
10/15






Chene-
pod







Dock







Plan-
tain







Kochia







Hemp







Elm







                                                                 (continued)
                                                                          oo

-------
            TABLE 3 (Continued)



POLLEN SEASONS THROUGHOUT THE UNITED STATES
Location
[Maine
(Maryland
1 Baltimore
Massachu-
setts
Boston
Michigan
Detroit
Minnesota
Minneap-
olis
Mississippi
Vicksburg
Missouri
St. Louis
Kansas
City
Tree
4/1
6/1
3/1
5/7
4/1
6/1
3/1
6/1
4/1
6/1
2/1
5/1
3/1
6/1
Grass
5/15
7/7
5/7
7/1
5/15
7/15
5/15
7/15
5/22
7/7
5/7
10/1
5/15
7/7
Rag-
weed
8/7
9/22
8/15
10/1
8/15
10/1
8/15
10/1
8/15
9/22
9/1
10/7
8/7
10/1
Ama-
ranth




6/1
10/1


Russian
thistle







Salt
bush







Saqe







Chene-
pod




6/1
10/1


Dock







Plan-
tain







Kochia







Hemp







Eln







                                                                (continued)
                                                                          oo
                                                                          to

-------
           TABLE 3 (Continued)


POLLEN SEASONS THROUGHOUT THE UNITED STATES
1
Location
Montana
Miles City
iebraska
Omaha
evada
Reno
New Hamp-
shire
New
Jersey
JNew Mexico
1 Ro swell
New YorTc
1 New York
North Caro-
lina
Raleigh
Tree
4/1
6/1
3/1
6/1
4/1
6/1
4/1
6/1
3/15
6/1
3/15
5/1
3/15
6/1
2/1
6/1
Grass
5/1
9/22
5/15
7/15
6/1
8/1
5/15
7/22
5/15
7/15
5/15
10/15
5/15
7/15
5/15
7/15
Rag-
weed
8/1
10/7
8/15
9/22
8/22
10/1
8/15
10/1
8/15
10/1
8/22
10/22
8/15
10/1
8/15
10/1
Ama-
ranth





7/1
10/1


Russian
thistle
7/7
9/7
7/7
9/7
7/1
10/1


,


Salt
bush


7/1
10/1


7/1
10/1


Saqe
8/1
10/7

8/1
10/7


8/22
10/22


Chene-
pod








Dock








Plan-
tain








Kochia








Hemp

8/1
9/7






Elrr








                                                               (continued)
                                                                          oo
                                                                          00

-------
            TABLE 3  (Continued




POLLEN SEASONS THROUGHOUT THE UNITED STATES
jocation
forth Dakota
Fargo
Dhio
Cleveland
Oklahoma
Oklahoma
City
)regon
Portland
Area East of
1 Cascade
1 Mountains
Pennsylvania
Rhode
1 Island
Tree
4/1
6/1
3/15
6/15
2/15
6/15
2/22
5/1
3/15
4/15
3/15
5/15
3/15
6/1
Grass
6/1
8/1
6/1
7/15
5/1
10/1
4/22
9/1
5/7
7/1
5/7
7/15
5/22
8/1
Rag-
weed
7/22
9/7
8/15
9/22
8/22
10/7

8/15
9/15
8/15
10/7
8/15
10/1
Ama-
ranth


7/1
10/7




Russian
thistle
7/1
9/15



7/7
10/1


Salt
bush




7/7
10/1


Saqe
8/15
9/22



8/22
10/1


Chene-
pod







Dock



5/1
10/1



Plan-
tain



5/1
10/1



Kochia







Hemp







Elm







                                                                 (continued)
                                                                           CO

-------
            TABLE 3 (Continued)




POLLEN SEASONS THROUGHOUT THE UNITED  STATES
Location
South Caro-
lina
Charles-
ton
South
'Dakota
Tennessee
Nashville
Texas
Dallas
Utah
Salt Lake
City
Vermont
Virginia
Richmond
Tree
2/15
5/22
3/1
5/1
2/22
5/22
12/15
5/1
4/1
5/22
4/1
6/1
2/1
6/15
Grass
5/15
8/7
5/15
7/15
5/1
9/7
4/1
10/1
5/7
7/22
5/22
7/15
5/15
7/15
Rag-
weed
8/15
10/7
7/22
10/1
8/22
10/7
9/1
10/1
8/15
10/1
8/15
9/22
8/15
10/1
Ama-
ranth







Russian
thistle

7/1
10/1


7/15
9/15


Salt
bush







Saqe

8/22
10/1
9/7
10/1

9/7
10/11


Chene-
pod







Dock







Plan-
tain







Kochia







Hemp







Elm


7/7
10/1
8/22
10/1



                                                                 (continued)
                                                                           CO

-------
            TABLE 3 (Continued)




POLLEN SEASONS THROUGHOUT THE UNITED STATES
Location
Washington
Seattle
Eastern
Area
West
Virginia
Wisconsin
Madison
Wyoming
Trees
2/22
5/1
3/15
4/15
3/15
6/15
4/1
6/1
3/22
5/1
Grass
4/22
10/15
4/22
7/7
5/22
7/15
6/1
7/22
6/7
8/1
Rag-
weed

8/15
9/22
8/15
9/15
8/15
9/22
7/7
9/15
Ama-
ranth





Russian
thistle

7/15
10/1


7/1
9/15
Salt
bush

7/15
10/1



Saqe

8/22
10/1


8/15
10/15
Chene-
pod





Dock
5/1
10/15




Plan-
tain
5/1
10/15




Kochia





Hemp





Elm





                                                                         00

-------
APPENDIX
                                                           87
                          TABLE  4


           MOST COMMDN AEROALLERGEN1C  FUNGI20'52



                  Alternaria


                  Aspergillus


                  Botrytis


                  Cladosporium


                  Curyularia


                  Epicoccuiu


                  Fusarium


                  Helm intho spor i urn


                  Hormodendrum


                  Macro spor ium


                  Penicillium


                  Phoma


                  Pullalaria


                  Spondvlocladium


                  Stemphyllum

-------
                                           TABLE  5
SUMMARY OF DIFFERENCES BETWEEN HOSPITAL ADMISSIO^ RATES
FOR DAYS OF HIGH AND DAYS OF LOW AIR POLLUTION 106
Oxi-
Disease Groupings dants CO SO-? NO?
Allergic disorders x x
Acute upper respira- - x +
tory infections
Pneumonia x + x
Bronchitis 4- + x
Oxides
of
NO Nitrogen
x
+ X

X X
Oxi-
dant
Precur-
Ozone sor
-
X. X
X X
— X
Partic-
ulate Temper-
Matter ature
x x
x x.
x +
X
Humid-
ity
+

—
x.
Diseases of tonsils
  and adenoids
                                                            x
Other diseases of
  respiratory system
x
x
                                                            X
        *Differences of 5 percent or less are not shown.
         Differences of 6-10 percent indicated by +  or  —.
         Differences of 11 percent or more indicated by x or  x. (if negative).
                                                                                                 oo
                                                                                                 oo

-------
APPENDIX
                                             Table 6




                    PERCENTAGE DISTRIBUTION OP EMERGENCY VISITS BY MONTH,  1960
9
Hospital
and City
Harlem
New York
Metropol it an
New York
D.C. General
Washington
General
Philadelphia
Cook County
Chicago
St. Francis
Evanston
Thomas Memorial
Charleston
Michael Reese
Chicago
De Paul
Norfolk
Charity
New Orleans
Month
Jan.
8.45
7.49
9.60
7.98
5.95
3.70
7.76
7.23
6.22
5.94
Feb.
7.58
7.33
6.33
7.38
4.93
2.47
5.94
6.54
6.73
5.12
Marcli
7.82
7.93
8.25
5.57
4.64
7.41
5.02
5.22
2.85
4.08
April
5.82
5.96
6.44
5.65
5.53
3.70
8.68
6.41
5.96
5.22
May
6.29
6.55
5.99
7.55
7.09
7.41
3.20
6.60
8.81
8.18
June
5.89
5.95
5.31
6.91
7.72
8.64
5.02
6.22
7.77
5.89
July
5.12
5.58
6.55
7.81
6.30
8.64
4.57
7.10
6.48
8.00
Aug.
7.11
5.73
4.97
8.20
7.46
9.88
9.13
8.04
4.40
6.64
Sept.
8.09
7.47
7.91
7.51
10.30
17.28
15.07
7.92
9.07
7.93
Oct.
12.85
14.76
14.69
13.29
17.34
18.52
10.50
15.65
14.77
15.01
Nov.
13.40
14.41
13.67
13.72
12.78
6.17
13.24
12.51
14.51
17.10
Dec.
11.58
10.82
10.28
8.42
9.96
6.17
11.87
10.56
12.43
10.89
Total
No. of
Cases
10,684
5,432
885
2,317
3,795
81
219
1,591
386
8,163
Percent
100
100
100
100
100
100
100
100
100
100
                                                                                                   00

-------
APPENDIX                                                   90
                         TABLE 7

         AVERAGE DAILY NUMBER OF EMERGENCY CLINIC
           VISITS FOR ASTHMA, 1962, 1961, 195747
                                    Hospital
Year	Bellevue	Metropolitan    Harlem

1962

  September 1-18          12.0           23.2         39.7
  September 19-26*        21.0           42.4         60.3
  September 27-30         12.3           30.3         57.0

1961

  September 1-15           7.4           27.3         14.5
  September 16-19*        30.5           60.8         44.8
  September 20-30         12.9           40.2         24.1

1957

  September 16-25          6.6            9.0         25.0
  September 26-30*        16.2           24.0         52.4
  October 1-15             9.7           11.2         26.3

        *Critical period.

-------
APPENDIX
                                                          91
                         TABLE 8

             AVERAGE DAILY NUMBER OF EMERGENCY
     CLINIC VISITS FOR ASTHMA, SEPTEMBER, 1964, 196549
Location and Time
1964
Bellevue Hospital
September 1-12
September 13-16*
Harlem Hospital
September 1-12
September 13-16*
Metropolitan Hospital
September 1-12
September 13-16
1965
Bellevue Hospital
September 1-24
September 25-30*
Harlem Hospital
September 1-24
September 25-30*
Metropolitan Hospital
September 1-24
September 25-30*
Average
Visits Percent
Per Day Increase Probability

13.3
28.0 +110.5% <0.01
37.9
55.5 + 45.4% <0.01
16.8
37.0 +120.2% <0.01

12.1
35.7 +195.0% <0.01
32.5
66.7 +105.2% <0.01
16.6
47.2 +134.3% <0.01
        *Critical period

-------
                                                          92
APPENDIX
                         TABLE 9

     ANNUAL ADMISSIONS, BRISBANE CHILDREN'S HOSPITAL23



Group	1955-1956    1956-1957    1957-1958   Total

Asthma             115           98          107       320

Asthmatic
  bronchitis        71           93           72       236

      Total        186          191          179       556

-------
APPENDIX
93
                         TABLE 10

          COMPARISON OF MOLD AND POLLEN COUNTS63
                          (Israel)
Town
Arad
Beilinson
Mold Counta
Monthly Mean per Plate
6.5-15.5
20.0-61.5
Pollen
Monthly Mean
per can2
962
1,934
Countb
Mean per cm2
530
1,222
         a9  cm sediment plate exposed  J
         ^Durham gravity slide sampler,

-------
APPENDIX
                                                         94
                         TABLE 11

     MAXIMUM DESITOMETRIC READINGS FOR COMPOUND A*43
Grains
Botanical Name
                                         Readings
Stem   Husk
Bran   Grain
Wheat
Corn
Oats
Barley
Rye
Triticum vulqare
Zea mays
A vena sativa
Hordeum vulqare
Secale cereale

58

53
47
15
94
88
84
78
22
86
85
82
79
10
12
14
13
9
4
 Seed
Group
    Botanical Name
                                Readings
         Husk
        Seed
Flax
False Flax
Rape
Turnip Rape
   Linum usitatissum
   Camelina sativa
   Brassica napus
   Brassica rapa
          94
          93
          95
          82
         95
         94
         95
         92
Pollen
 Group
          Botanical Name
                   Readings
Short ragweed
Oak
Cottonvrood
Maple
Elm
Alfalfa
      Ambrosia, artemisiifolia
      Quercus alba
      Populus trichocarpa
      Acer saccharum
      Ulmus americana
      Medicago sativa
                      96
                      95
                      92
                      87
                      63
                      38
        *A high reading represents a high relative concentra-
tion of Compound A.

-------
APPENDIX
95
                         TABLE  12

   DERMAL AND BRONCHIAL REACTIVITY TO  CANDIDA ALBICANS59
                       (81  Patients)
No. of
Patients
19
23
16
23
Total
Immediate
Immediate Delayed Bronchial
Skin Skin Reaction
Response Response Only
+ + 8
+ - 4
+ 3
3
18
Immediate
& Delayed
Bronchial
Reaction
4
8
1
2
15
Negative
Bronchial
Reaction
7
11
12
18
48

-------
APPENDIX                                                   -_
                                                           96
                          TABLE 13

  ABUNDANCE OF RAGWEEDS ACCORDING TO LAND USE CATEGORIES98
                             Percent of Areas Represented in
                               Each Abundance Scale Rating3
Land Use Category
Cropland — corn
Cropland — wheat
Cropland — oats
Alfalfa meadow
Pasture
Grass Meadow
Park lands
Woods
Marshes
Roadsides
Residence property
Soybeans
Clover and clover mixtures
Timothy and timothy mixtures
One to three-year abandonment
Summer-fallowed fields
0
8.5%
0
0
75
81
91
90
100
100
87
100
0
40
100
0
50
1
50%
0
0
25
0
0
10
0
0
10
0
50
0
0
44
50
2
33%
0
33
0
9.5
0
0
0
0
1
0
50
40
0
14
0
3
8.5%
33
33
0
0
9
0
0
0
2
0
0
20
0
28
0
4
0%
67
33
0
9.5b
0
0
0
0
0
0
0
0
0
14
0
        aAbundance Scale Rating:

           0—No ragweeds observed.
           1—Ragweeds present, but averaging less than 0.5
              plants per square meter.
           2—Ragweed density averaging from 0.5 to 1 plant
              per square meter.
           3—Ragweed density averaging from 1 to 10 plants
              per square meter.
           4—Ragweed density averaging over 10 plants per
              square meter.
           is value is based on observations of pastures
used exclusively by swine at the time the study was made

-------
APPENDIX
                                                          97
                          TABLE 14

               COMMON ALLERGEN1C PRODUCTS52'64
Product
Pillows


Stuffing in mattresses
  and toys

Rugs

Fabrics


Brushes


Purs


Wigs

Cosmetics  (wave  set lotions,
  hair tonics, talcs, and
  perfumes)

Insecticides
Paints and  varnishes
Fertilizer
Chicken, duck, and geese
feathers, kapok

Kapok; cat, cattle, and horse
hair

Cattle and horse hair,  wool

Goat, cattle, and horse hair;
wool

Cattle and horse hair,  hog
bristles

Sensitivities exist to  indi-
dual furs

Human and horse hair

Many contain orris root or
flaxseed
Many contain pyrethrum derived
from dried flowers of the
chrysanthemum family, which is
related to ragweed

Flaxseed is used to produce
linseed oil, an ingredient in
many paints and varnishes

Castor bean pomace

-------
APPENDIX
                                            TABLE 15



                                        10LLEN DISPERSAL5
Species (Means of Dispersion)
Horizontal Distances and Units Dispersed

lAgropvron cristatum (Wind)
JA. intermedium (Wind)
r
JBeta sp. (Wind)
JBromus sp. (Wind)
1
Cedrus atlantica (Wind)

b. libani (Wind)
jDactvlis sp. (Wind)
[Fraxinus sp. (Wind)

Juglans regia (Air currents)

Lolium SP. (Wind)

Malus pumila (Wind)

3ryza sativa (Dehiscense and
1 Wind)
Rods from field
Pollen grains
Rods from field
Pollen grains
Meters from seed fields
Pollen grains/cm2
Rods from field
Pollen grains
Feet from source tree
Pollen grains
Feet from source tree
Pollen grains
Meters from field
Pollen grains/cm2
Feet from source tree
Pollen grains
Feet from pollen source
Pollen grains/mm2y24 hr
Meters from ryegrass field
Pollen grains/cm3
Feet from source tree
Pollen grains
Centimeters from pollen source
Pollen grains
5
72
5
44
0
5
146
40
189
15
127
0
3,096
25
2,545
60
4
0
0
13
25
22
15
29
12
17
300
15
41
120
116
75
62
200
447
50
1,008
150
2.8
200
165
2
50
9
25
10
25
4
500
25
21
240
71
135
37
400
172
150
141
500
1.4
500
330
0.9
100
3


800
40
10
325
51
195
22
600
120
400
29
1,000
0.6
700

150
1



60
4
700
0.1

800
86

1,600
0
900

200
0.4
                                                                                      (continued)
                                                                                                 00

-------
TABLE 15 (Continued)




  POLLEN DISPERSAL
Species (Means of Dispersion)

[Panicum virqatum (Wind)
1
IParthenium arqentatum (Wind)
iPenniselum qlaucum (Wind)
iPhleum pralense (Wind)
iPicea sp. (Wind)
r
JP. cembroides (Wind)
L
Populus sp. (Wind)

P. deltoides

Secale cereale (Wind)
•
Ulmus sp. (Wind)

Zea mays (Wind)
Horizontal Distances and Units Dispersed

Rods from field
Pollen qrains
Yards from guayule plants
Pollen qra ins/in2
Yards from release point
Pollen, percent
Meters from timothy field
Pollen qra ins/cm3
Feet from source tree
Pollen qrains
Feet from source tree
Pollen qrains
Feet from source tree
Pollen qrains
Feet from source tree
Pollen qrains
Rods from rye field
Pollen grains
Meters from rye field
Pollen qra ins/cm2
Feet from source tree
Pollen qrains
Rods from field
. Pollen qrains
Feet from pollen source
Pollen grains
5
27
100
4
100. C
0
0
9.7
10
8,479
50
107
25
115
5
453
100
4,181
500
115
5
18
10
7,330
15
7
400
50
8.9
100
165
0.1
75
462
500
86
250
62
15
232
300
2,579
L,100
152
15
6
30
341
25
4
850
200
0.8
200
330
0.7
150
86
1,400
76
500
46
25
124
500
1,834
2,700
12
25
3
50
121
40
2
1,200
400
0.4
300

225
38
3,200
69
1,550
20
40
52
700
1,343
5,500
8
40
2
70
30
60
0.5


500

300
52
4,200
66
3,550
0.3
60
11


60
0.8


-------
                 TABLE 16
POLLEN COUNTS, ST. LOUIS SITE,  1963-196436

        (In Grains per Cubic Yard*)

Month Year
March 1963
1964
Upril 1963
1964
May 1963
1964
IJune 1963
1964
July '1963
Aug. 1963
1964
Sept. 1963
- r
Average Count Per Month
Elm
ackberrv
134.9
40.4
1.7
12.8
0.2
0.9
0
0
0
0
0
0
0
0
Poplar
Cottonwood
30.5
0
7.5
10.0
0
1.4
0
0
0
0
0
0
0
0
Maple
4.9
1.1
0.4
2.1
0
0
0
0
0
0
0
0
0
Oak
17.5
0
254.3
117.2
6.2
78.2
0
0
0
0
0
0
0
0
Sycamore
4.2
0
92.1
78.6
0.3
9.4
0
4.4
0
0
0
0
0
0
Hickory
Walnut
2.5
0
13.4
3.3
12.3
39.3
0.3
0.2
0
0
0
0
0
0
Grass
0
'0
0.2
0
5.0
4.6
7.6
4.3
2.8
0.8
0
0
0
0
Plantain
0
0
0
0
0
0
1.2
2.8
1.9
1.3
1.6
0
0
0
Goosefoot
0
0
0
0
0
0
0
0.5
0.7
0.9
21.9
11.2
31.4
30.7
Ragweed
0
0
0
0
0
0
0.2
0.2
1.0
0.8
86.4
47.8
112.7
144.3
*Multiply by 1.3 = grains per cubic meter.
                                                                        o
                                                                        o

-------
                                                       101
APPENDIX
                  TABLE 17

NUMBER OP ALTERNARIA SPORES PER CUBIC FOOT81
Time
5 a.m.
6 a.m.
7 a.m.
9 a.m.
11 a.m.
1 p.m.
2 p.m.
5 p.m.
6 p.m.
Number per Cubic Foot
12
7
9
13
16
17
13
16
19

-------
APPENDIX
102
                         TABLE 18

         RECOMMENDED CONDITIONS FOR USE OP COMMON
     GERMICIDAL SUBSTANCES  (AT ROOM TEMPERATURE, 25°C)62
                         FOR FUNGI
Germicide
Phenol
Lysol
Hypo chlorite + 1% wet-
ting agent (Naccanol,
etc.)
Caustic sodium hydroxide
Formalin (37% HCHO)
Steam formaldehyde vapor
in closed area
-Propiolactone
Ethyl ene oxide gas
Concentration
5%
3%
2,000 ppm
10%
5% solution
1 ml/ft3 in air with
Rh* above 80%
200 mg/ft3 in air with
Rh* above 80%
300 mg/liter
Exposure
Time
15 min
15 min
10 min
30 min
10 min
30 min
30 min
8-16 hr
         *Rh  =  relative humidity.

-------
                         TABLE 19


RESOURCE COSTS OF DISEASES ASSOCIATED WITH AIR POLLUTION57
Basis
of
Cost
Premature
death
Premature
burial
Treatment
Absenteeism
Annual Costs Associated With Selected Diseases,* Millions of Dollars
Cancer
of the
Respiratory
Svstem
518
15
35
112
Chronic
Bronchitis .
18
0.7
89
52
Acute
Bronchitis
6
0.2


Common
Cold
-

200
131
Pneumonia
329
13
73
75
Emphysema
62
2


Asthma
59
2
138
60
*Using a discount rate of 5 percent.
                                                                                O
                                                                                CO

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APPENDIX                                                    104
                         TABLE 20

          ASTHMA-HAY FEVER PURCHASED ACQUISITION
       OF PRESCRIBED MEDICINE, JULY 1964-JUNE 19543
Number of conditions                            14,375,000

Percent of total prescribed medicine
  acquisitions                                     2.7%

Number of purchased acquisitions of
  prescribed medicine                           21,194,000

Percent of total number purchased
  acquisitions of prescribed medicine:

      Cost under $2.00                            26.0%
                  2.00-2.99                       25.2%
                  3.00-4.99                       33.4%
                  5.00-6.99                        9.3%
                  7.00

Average cost per purchase of prescribed
  medicine                                        $3.30

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APPENDIX                                                 105
                         TABLE 21

           SIX MOST FREQUENT CAUSES OF NON-MAJOR
        ACTIVITY LIMITATION, JULY 1963-JUNE 196525
Cause	Percent of Total

Arthritis and rheumatism                         11.9

Heart condition                                  10.7

Impairment of back or  spine                       7.7

Mental and nervous conditions                     7.0

Asthma-hay fever                                  6.8

Impairment of lower  extremities and hip           6.3

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                                                                106
APPENDIX
                               TABLE  22

       AVERAGE NUMBER OF PERSONS REPORTED AS LIMITED IN ACTIVITY
       DUE TO SELECTED  CHRONIC CONDITIONS, JULY 1961-JUNE 1963115


                                 Usual Activity Status
                     (Average Number  of Persons with Conditions X 1,000)
                        All     Usually Keeping
Cause of Limitation  Activities Working  House  Retired (Age; 17+ yrs)

All conditions        22,275     6,384   7,525   4,668       2,257

Asthma-hay fever        1,118        242     281     190         117

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APPENDIX
107
                               TABLE 23

         DEATH RATE  (1950 TO 1966) AND DEATHS  (1965 AND 1966)
                          FROM SELECTED CAUSES*05

Cause
All Causes
Tuberculosis
(all forms)
Meningococcal
infection
Asthma
Influenza and
pneumonia
(except pneu-
monia of
newborn )
Influenza
Pneumonia
Bronchitis
Deaths per 100,000 Population
1950
963.8
22.5
0.6
2.9
31.3
4.4
26.9
2.0
1955
930.4
9.1
0.6
3.6
27.1
1.7
25.4
1.9
1960
954.7
6.1
0.4
3.0
37.3
4.4
32.9
2.4
1964
939.6
4.3
0.4
2.3
31.1
0.9
30.2
2.8
1965
943.2
4.1
0.5
2.3
31.9
1.2
30.8
3.0
1966
951.3
3.9
0.4
2.2
32.5
1.4
31.0-
3.1
Deaths
1965 1966
1,828,136 1,863,149
7,934 7,625
850 876
4,520 4,324
61,903 63,615
2,295 2,830
59,608 60,785
5,772 6,151

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