DOC
United Stales National Oceanic and Atmospheric Administration
Department of Environmental Research Laboratories
Commerce Seattle WA 98115
United States Office of Environmental Engineering and EPA-600 7-79-213
Environmental Protection Technology November 1979
Agency Washington DC 20460
Research and Development
The Strait of
Juan de Fuca
Intertidal and
Subtidal Benthos
Interagency
Energy/Environment
R&D Program
Report
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RESEARCH REPORTING SERIES
Research reports of the Office of Research and Development, U.S. Environmental
Protection Agency, have been grouped into nine series. These nine broad cate-
gories were established to facilitate further development and application of en-
vironmental technology. Elimination of traditional grouping was consciously
planned to foster technology transfer and a maximum interface in related fields.
The nine series are:
1. Environmental Health Effects Research
2. Environmental Protection Technology
3. Ecological Research
4. Environmental Monitoring
5. Socioeconomic Environmental Studies
6. Scientific and Technical Assessment Reports (STAR)
7. Interagency Energy-Environment Research and Development
8. "Special" Reports
9. Miscellaneous Reports
This report has been assigned to the INTERAGENCY ENERGY-ENVIRONMENT
RESEARCH AND DEVELOPMENT series. Reports in this series result from the
effort funded under the 17-agency Federal Energy/Environment Research and
Development Program. These studies relate to EPA's mission to protect the public
health and welfare from adverse effects of pollutants associated with energy sys-
tems. The goal of the Program is to assure the rapid development of domestic
energy supplies in an environmentally-compatible manner by providing the nec-
essary environmental data and control technology. Investigations include analy-
ses of the transport of energy-related pollutants and their health and ecological
effects; assessments of, and development of, control technologies for energy
systems; and integrated assessments of a wide range of energy-related environ-
mental issues.
This document is available to the public through the National Technical Informa-
tion Service, Springfield, Virginia 22161.
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THE STRAIT OF JUAN DE FUCA INTERTIDAL AND SUBTIDAL BENTHOS
Second Annual Report
Spring 1977 - Winter 1978
by
Carl F. Nyblade
University of Washington
Friday Harbor Laboratories
Friday Harbor, Washington 98250
Prepared for the MESA (Marine Ecosystems Analysis) Puget Sound
Project, Seattle, Washington in partial fulfillment of
EPA Interagency Agreement No. D6-E693-EN
Program Element No. EHE625-A
EPA Project Officer: Clinton W. Hall (EPA/Washington, D.C.)
NOAA Project Officer: Howard S. Harris (NOAA/Seattle, WA)
This study was conducted
as part of the Federal
Interagency Energy/Environment
Research and Development Program
Prepared for
OFFICE OF ENERGY, MINERALS, AND INDUSTRY
OFFICE OF RESEARCH AND DEVELOPMENT
U.S. ENVIRONMENTAL PROTECTION AGENCY
WASHINGTON, D.C. 20460
March 1979
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Completion Report Submitted to
PUGET SOUND ENERGY-RELATED RESEARCH PROJECT
MARINE ECOSYSTEMS ANALYSIS PROGRAM
ENVIRONMENTAL RESEARCH LABORATORIES
by
Friday Harbor Laboratories
University of Washington
Friday Harbor, Washington 98250
This work is the result of research sponsored by the Environmental
Protection Agency and administered by the Environmental Research
Laboratories of the National Oceanic and Atmospheric Administration.
The Environmental Research Laboratories do not approve, recommend,
or endorse any proprietary product or proprietary material mentioned
in this publication. No reference shall be made to the Environmental
Research Laboratories or to this publication furnished by the Environmental
Research Laboratories in any advertising or sales promotion which would
indicate or imply that the Environmental Research Laboratories approve,
recommendj or endorse any proprietary product or proprietary material
mentioned herein, or which has as its purpose an intent to cause directly
or indirectly the advertised product to be used or purchased because of
this Environmental Research Laboratories publication.
ii
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FOREWORD
Substantially increased petroleum tanker traffic and refining
operations are anticipated in the region of northern Puget Sound and the
Strait of Juan de Fuea as Alaskan crude oil production increases and as
pipeline deliveries of crude from Canada to the region are terminated.
This increased transport and refining activity will increase the opportu-
nities for spills and leaks of crude oil and refined products into the
marine environment. Recognizing the need for environmental information in
the region, the U.S. Environmental Protection Agency has supported the
Puget Sound Energy-Related Project under which studies involving biological
characterizations, physical oceanography, trajectory modeling, pollutant
monitoring, and fate and effects of oil have been implemented. This report
has been administered by NOAA's Marine Ecosystems Analysis (MESA) Puget
Sound Project office. A major part of the Project has involved a variety
of biological studies intended to provide information on the character-
istics of biological communities at risk to oil pollution in the region.
This report presents the results of a two year field study of intertidal
and shallow subtidal communities found at ten sites representative of
common shoreline habitats along the Strait of Juan de Fuca. This study was
conducted in consultation with representatives of the Washington State
Department of Ecology.
iii
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ABSTRACT
Because of the threat of oil pollution from large scale oil shipment
through the Strait of Juan de Fuca, this study was undertaken to document
the pre-pollution communities in the shallow water zones along the Washing-
ton coast of the Strait of Juan de Fuca. The study objectives were to
adequately describe the distribution, abundance, seasonal variations and
annual variation of the intertidal and shallow subtidal benthos of this
coastal area.
During the first year ten sites, representative of the range of
habitats present, were sampled quarterly. Stratified random replicates
were collected at each area. Strata used were high (+6'), mid (+3'),
low (+0') intertidal and -5 m and -10 m. Distribution sampling at inter-
mediate strata was conducted once. During the second year seven of the
same sites were again sampled quarterly in the intertidal, while the
remaining three intertidal sites and all subtidal strata were sampled once.
Over 1,000 different plant and animal species were collected during
these two years, 176 of which had not previously been recorded in standard
taxonomic keys for our region. The dominant groups were algae, molluscs,
polychaete annelids, and crustaceans. In the intertidal, rock habitats
were the richest in terms of number of species, density, and biomass (and
probably productivity), followed by cobble, protected soft sediment,
exposed sand, and exposed gravel habitats. Species richness values ranged
from 177.5 (Tongue Pt. +0') to 1 (Dungeness Spit +6'); diversity 3.24
(North Beach Cobble +0') to 0.17 (Dungeness Spit +6'); density 56,874/m2
(Jamestown +6') to 44/m2 (Twin Rivers +6'); biomass 11,375 g/m2 (Pillar
Pt. +0') to < 2 g/m2 (Dungeness +6'). Strong intertidal vertical zonation
was found at all but the most exposed gravel and sand sites. Subtidal
study sites were consistently rich. Community comparisons of the areas
and levels sampled during this study validated the type habitat approach
and the selection of strata to be sampled. Patchiness of organisms in the
communities sampled generally obscured seasonal patterns in populations of
component species. However, summed over all levels and areas, summer was
most often the peak for species richness, abundance, and biomass and
winter most often the low. Year to year community similarity was high in
the rock, cobble, and protected soft sediment areas.
An important generalization from this mass of data stood out. The
rich, complex, productive communities are to be found in rock and protected
soft-sediment sites. These communities were the least variable over time
and are the most vulnerable to long term damage from an oil spill. These
data will be useful in assessment of damage from an oil spill, especially
the measures of community similarity, species richness, and diversity,
since the values of these parameters change with oil pollution effects. In
addition these data should have broad use in coastal zone management
decisions.
iv
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CONTENTS
Page
FOREWORD iii
ABSTRACT iv
FIGURES vi
TABLES vii
LIST OF APPENDICES ix
ACKNOWLEDGMENTS ........ . xi
I. INTRODUCTION 1
II. CONCLUSIONS 3
III. RECOMMENDATIONS .......... 5
IV. METHODS AND MATERIALS 7
IV-A. Field and Laboratory Procedures .... 7
IV-B. Data Analysis 10
V. RESULTS 13
Tongue Point 15
Pillar Point 28
North Beach Cobble 40
Morse Creek 47
Beckett Point ¦ 57
Dungeness Spit 68
Twin Rivers 74
North Beach Sand 80
Kydaka Beach 88
Jamestown 94
VI. DISCUSSION 105
IV-A. Study Area Comparisons 105
IV-B. Seasonal Changes 110
IV-C. Annual Changes . 114
VII. REFERENCES/BIBLIOGRAPHY ... 123
APPENDIX I. Complete Data Sets 130
APPENDIX II. Complete Species Lists ........ 378
APPENDIX III. Physical Parameters 472
v
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FIGURES
Number Page
1 Map of study sites . 9
2 Change in species richness compared with mean species
richness 119
3 Change in diversity compared with mean diversity 120
4 Mean species richness compared with sediment particle
size 121
5 Year to year community similarity compared with mean
species richness 122
vi
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TABLES
Number - Page
1 Listing of study areas , 8
2a Tongue Point +6 16
b +3 18
c +0 22
d Subtidal . 25
3a Pillar Point +6 ........... 29
b +3 31
c +0 34
d Subtidal 38
4a North Beach Cobble +6 . 41
b +3 . . . 42
c +0 44
5a Morse Creek +6 48
b ^*3..... ...... ........... 49
c +0 51
d Subtidal 55
6a Beckett Point +6 . , . 58
b +3 59
c +0 62
d Subtidal 66
7a Dungeness Spit +6 69
b +3 70
c +0 71
d Subtidal ........... 72
8a Twin Rivers +6 . . 75
b +3 .......... 76
c +0 77
d Subtidal ..... 78
9a North Beach Sand +6 81
b +2 ................. 82
0 +0 ...... .83
d Subtidal ...... 85
vii
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Number Page
10a Kydaka Beach +6 89
b +3 . 90
c +0 . 91
d Subtidal 92
11a Jamestown +6 95
b +1.4 96
c +0 98
d Subtidal . 102
12 Intertidal Summary 106
13 Subtidal Summary , 108
14 Seasonal Change: Species Richness Ill
15 Seasonal Change: Density 112
16 Seasonal Change: Biomass 113
17 First Year/Second Year Summary 115
18 Year One/Year Two Community Similarity 117
viii
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LIST OF APPENDICES
Page
APPENDIX I. COMPLETE DATA SETS 130
Table la Tongue Point +6 131
b +3 140
c +0 155
d Subtidal 174
2a Pillar Point +6 192
b +3 196
c +0 205
d Subtidal 215
3a North Beach Cobble +6 224
b +3 226
c +0 230
4a Morse Creek +6 240
b +3 242
c +0 250
d Subtidal 264
5a Beckett Point +6 270
b +3 273
c +0 281
d Subtidal 292
6a Dungeness Spit +6 298
b +3 299
c +0 ......... 300
d Subtidal 301
7a Twin Rivers +6 307
b +3 308
c +0 309
d Subtidal 310
8a North Beach Sand +6 312
b +2 314
c +0 317
d Subtidal 320
9a Kydaka Beach +6 337
b +3 338
c +0 339
d Subtidal 341
10a Jamestown +6 346
b +1.4 348
c +0 354
d Subtidal 363
ix
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Page
APPENDIX II. COMPLETE SPECIES LISTS 378
Table 1 Tongue Point 379
2 Pillar Point 398
3 North Beach Cobble ........ 412
4 Morse Creek 419
5 Beckett Point 430
6 Dungeness Spit 438
7 Twin livers ..... 447
8 North Beach Sand . 452
9 Kydaka Beach 460
10 Jamestown 464
APPENDIX III. PHYSICAL PARAMETERS 472
Table 1 , Salinity, Water Temperature,
Weather, Sampling Data ..... 473
2 Sediment Type by Study Area -
Tide Height 475
Figure 1 Kydaka Beach Study Area 476
2 Pillar Point Study Ares ...... 477
3 Twin Rivers Study Area ...... 478
4 Tongue Point Study Area ...... 479
5 Morse Creek Study Area 480
6 Dungeness Spit Study Area ..... 481
7 Jamestown Study Area 482
8 Beckett Point Study Area 483
9 North Beach Study Areas ...... 484
x
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ACKNOWLEDGMENTS
The author wishes to acknowledge the generous taxonomic and support
assistance of : K. Banse, P. Illg, E. Kozloff, R. Norris, R. T. Paine,
R. Shimek, and D- Willows.
The diligent efforts of Louisa Norris, Tony Roth and the additional
technical staff are gratefully acknowledged.
The following generously allowed access through or to their property
during the course of this study: Clallam County Commissioners; Crown
Zellerbach Corporation and C. B. Paulson, Forest Manager; Four Seasons
Maintenance Commission; C. W. Gunstone, Jr.; D. B. Mclnnes; Merrill and
Ring Corporation and J. Vadnais, Managing Forester; D. Moriarity;
Nisqually National Wildlife Refuge, W. B. Hesselbart, Refuge Manager;
M. Schiefelhein; C. Swain; Twin Rivers Investment Club; A. Webster;
G. Wood.
xi
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Xii
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SECTION I
INTRODUCTION
This report presents the second year results of an intertidal and
shallow subtldal benthlc sampling program at ten sites along the Washington
State coast of the Strait of Juan de Fuca. The purpose of this program
was to provide a quantitative characterization of the marine, shallow-water,
bottom communities of the Strait.
In the past five years the greater Puget Sound region has seen a
dramatic increase in marine crude and refined oil transport with the
replacement of overland pipeline transport by large tanker transport. The
possibility of Alaskan crude oil transshipment through this region to the
Midwest means additional increases are possible in the near future. This
tanker traffic increase also increases the risk of catastrophic and chronic
oil pollution of the marine environment in this region.
Previous to 1974 virtually no quantitative data existed on the inter-
tidal and shallow subtidal benthos of this region. The seriousness of
this absence of information was two-fold. First the communities of these
organisms have great recreational, ecological, and economic importance.
This is especially true for the greater Puget Sound region. Hundreds of
thousands of people live on or very near the shore and make use of its
varied recreational activities from yachting to clam digging. These same
areas are also highly productive of eelgrass and macro-algae, plants which
are responsible for a very large percentage of greater Puget Sound primary
production. After the economic importance of recreational uses of these
communities, there is a large direct economic value based on commercial
fisheries. Virtually all juvenile salmonids are dependent on intertidal
and shallow water communities for food. These communities are also highly
productive of commercial bottom fish and of shellfish.
The second reason an absence of quantitative data on the intertidal
and shallow subtidal benthic communities of this region was so serious
is the widely documented high susceptibility of these communities to
damage from spilled oil (National Academy of Sciences, 1975; Halins, 1977;
Wolfe, 1978). Put simply much or most oil floats and in the virtually
closed greater Puget Sound basin most floating oil will come ashore (Oil
on Puget Sound, 1972).
To respond to this oil pollution threat, in 1974 the Washington State
Department of Ecology initiated field work for their Oil Baseline Studies,
This work was largely confined to the San Juan Islands and the Rosario
Strait mainland areas. In 1976 the federal government initiated field
sampling along the Strait of Juan de Fuca under the EPA's Puget Sound
1
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Energy-Related Research Project administered through NOAA's Marine
Ecosystem Analysis Puget Sound Project office. This project was designed
to identify the potential ecological consequences of increased petroleum
transport and transfer activities anticipated for the greater Puget Sound
region.
First year research components of the study reported on previously
(Nyblade, 1978) consisted of defining the habitat types present along the
Strait of Juan de Fuca, largely according to substratum/exposure, selec-
tion of ten sites along the length of the Strait representative of these
habitat types, and quarterly determination of the vertical distribution of
the organisms found at each.
In order to verify and amplify trends observed the first year and to
document natural year to year variation, a second year of sampling was
undertaken. Second year sampling consisted of continued quarterly
determination of community composition at seven sites in the intertidal
and annually at three sites intertidally and all sites subtidally. These
components have permitted documentation of both seasonal and annual
changes in the communities sampled.
This information is critical? to develop an understanding of the
biology of the shallow water marine communities of this region, to
evaluate the regional ecological importance of the various habitats, to
determine the economic value of the various types of communities, and to
enable any careful assessment of the impact of man's activities along the
Strait, especially in assessing damage to this environment from oil
pollution. Also, uses of this data base may very well involve activities
quite unrelated to oil pollution, eg. damage from other pollutants,
tideland utilization, siting studies—informed coastal zone management.
2
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SECTION II
CONCLUSIONS
With the completion of this two year program of sampling along
Washington's coast of the Strait of Juan de Fuca, a quantitative data set
on the composition of the Strait's intertidal and shallow subtidal marine
benthic communities has been established prior to any major perturbation
such as a large oil spill from a tanker or submarine pipeline.
Exposed intertidal sand and gravel habitats contained relatively
sparse, simple, low diversity communities dominated by worms and small
crustaceans. Protected soft-sediment habitats exhibited dense, very
diverse infaunal communities dominated by a vast array of polychaete
species, small and large bivalves, and small and large crustaceans.
Cobble and rock areas contained the richest communities with the largest
standing crop biomass. Cobble and rock communities were dominated by
macro-algae, herbivorous gastropods, barnacles, mussels, large and small
crustaceans. Subtidal rock areas were equally rich. Communities there
contained a large variety of algae, gastropods, small crustaceans, and the
dominant algal grazers, sea urchins. Subtidal soft sediment areas were
also species rich, but standing crop was much lower. Communities in these
areas contained literally hundreds of species of polychaetes as well as a
great variety of small bivalves and crustaceans.
Over 1,000 different plant and animal species were collected during
the study. Second year species richness values ranged from 177.5 (Tongue
Point +0') to 1 (Dungeness Spit +6'); diversity from 3.24 (North Beach
Cobble +0') to 0.17 (Dungeness Spit +6'); density 56,874 /m^ (Jamestown
+6') to 44 /m2 (Twin Rivers +6'); biomass 11,375 g/m^ (Pillar Point +0')
to < 2 g/m^ (Dungeness +6'). Strong intertidal vertical zonation was
found at all but the most exposed gravel and sand sites.
Little seasonal change in communities was documented during either of
the two years of study. Surprisingly little annual change in communities
was found comparing the first and second year data sets. It is clear that
data variability may very well have masked real seasonal and annual
changes. On the other hand major changes should be reflected in the
summary parameters measured such as mean species richness, diversity, and
community similarity. But little evidence was found among these parameters
for seasonal or annual changes. Sanders (1978) argues forcefully for the
utilization of such synthetic parameters in examining community change,
rather than traditional species lists and the vast, unmanageable data set
on individual species populations.
What is there specifically to be concluded about the response of the
3
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communities studied to perturbations such as an oil spill? The following
summarizes these conclusions. Life span/recruitment rates are the author's
best estimate from direct experience.
Increasing
Damage/
Increasing
Recovery
Time
Number of Life Span/
Macro-Species Recruitment Rates
Habitat/Study Areas in Community of Dominants
Gravel - Dungeness Spit
Twin Rivers few to 10's «1 yr.
Sand - Kydaka Beach
North Beach Sand 10's to 20's <1 yr. to 1 yr.
Mud - Jamestown
Mixed - Beckett Point
50 to 100
1 yr. to
several years
Cobble - North Beach
Morse Creek
100's
several yrs.
to decades
f
Rock - Pillar Point
Tongue Point
There finally has begun to develop a literature to support such
conclusions concerning longterm damage. Torrey Canyon recovery took from
5 to 10 years and by some measures remains incomplete (Southward and
Southward, 1978). Other studies of other spills for shorter periods
continued to show damage after 2h to 3 years (Sanders, 1978; Hampson and
Moul, 1978) with no prediction on when recovery might be complete.
Hampson and Moul have documented severe salt marsh erosion following the
killing of marsh grass by oil erosion of areas that may have taken
hundreds of years or longer to accumulate. The rich, complex communities
of rock and protected soft-sediment areas are the most vulnerable to long
term damage because they are dominated by long-lived, irregularly
recruiting species. Recovery of these communities from an oil spill could
take decades here.
4
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SECTION III
RECOMMENDATIONS
A number of recommendations for further studies can be made based on
the two year Strait of Juan de Fuca sampling program:
1. Further quantitative monitoring of communities at the study sites
should be undertaken. Sampling methodology must be identical or
compatible with previous work in order to make comparisons with
previous data possible. However, for the more complex communities
only a subset of the dominant species need be fully processed, the
remainder of the sample being processed for long term storage
only. The need for further monitoring is two-fold. First, more
information is needed on normal year to year variation of these
marine communities. Second, only by updating will the validity
of the original data base be maintained for post-perturbation
comparisons. Should an oil spill occur in 1987, use of a ten year
old data base for damage assessment would be weak unless validated
by long-term monitoring.
2. Studies of community recruitment dynamics, especially of the key,
long-lived species of the complex rock and protected soft-bottom
communities, should be undertaken. Almost nothing is known about
recruitment dynamics of regional marine species, particularly the
long-lived species which apparently are highly irregular in
recruitment. Without a better understanding of recruitment, there
can be no basis for specific predictions on the recovery of their
communities from a perturbation such as an oil spill which
destroys key elements of the communities.
3. A general analysis and synthesis of the greater Puget Sound
shallow water marine benthos data base should be undertaken.
Although sampling methodology has been similar in studies during
the past five years undertaken for METRO, the Washington State
Department of Ecology, and the MESA Puget Sound office, technical
report data presentation has not been compatible and generally the
original data has not been presented.
4. Following data synthesis a field test should be undertaken to
determine the degree to which the data base can be extrapolated to
areas not previously sampled.
5. Additional regional taxonomic work is desperately needed on a
number of important groups; gammarid amphipods, oligochaetes,
free-living nematodes, ostracods, and cumaceans - in descending
order of importance. The lumping of the species of these groups
into higher taxa creates great problems In data analysis and
comparison. When dominant species (eg. amphipods and oligochaetes
in exposed gravel areas) are not identified to species, nothing
can be said about long term community change or a potential
change caused by pollution.
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6. Samples processed and stored must be carefully curated as long as
there Is any use for the data based on them. The science of
taxonomy is undergoing constant change. Without recourse to the
original samples no use can be made of current taxonomic informa-
tion.
6
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SECTION IV
METHODS AND MATERIALS
IV-A. Field and Laboratory Procedures
Second year procedures followed directly those of the first year
(Nyblade, 1978) to guarantee comparable data sets. The purpose of this
methodology was to provide data to best document the abundance, distribu-
tion, seasonal and annual variation of organisms and populations in each
major habitat type present and to do this with a finite set of resources.
All ten study sites sampled during the first year were resampled
during the second (Table 1, Figure 1, site maps in Appendix III). Because
of problems with irregularly shifting sediment (North Beach Cobble and
Twin Rivers) or limited area available to sample (Pillar Point), these
three areas were only sampled once during the second year. Since during
the first year subtidal areas were fully processed for only one quarter
they were sampled only once during the second year.
Because tidal height/water depth has an over-riding influence on
community type present within a given habitat type, the strata marked and
sampled during the first year were resampled during the second to obtain
comparable data. Three strata were selected for sampling in the inter-
tidal (high +6'; mid, generally +3'; low +0') and two in the subtidal
(-5m, -10m). First year distributional sampling (Nyblade, 1978), showed
these levels gave full coverage of the range of organisms present over the
entire tidal and shallow water range at study areas.
The number, size, and type of randomly located replicates taken at
each stratum in general was identical during the second year to the first
year.
They were as follows:
Rock - intertidal (Tongue Point and Pillar Point): Four 0.25 m2
quadrats, each consisting of five 0.01 m2 subsection scrapes and
the residual 0.2 m2 scrape.
Cobble - intertidal (North Beach and Morse Creek): Four 0.25 m2
quadrats, each consisting of five 0.01 m2 subsection scrapes,
the residual 0.2 m2 scrape, and the under-cobble 0.05 m2 x 15 cm
sediment core fixed and dead-sieved through 1 mm mesh; four
0.25 m? x 30 cm deep quadrats live-sieved through 12.5 mm mesh.
Exposed Gravel and Sand - intertidal (Dungeness Spit, Twin Rivers,
North Beach, and Kydaka Point): Five quadrats of 0.05 m2 x 15 cm
deep sediment cores fixed and dead-sieved through 1 mm mesh; five
of 0.25 m2 x 30 cm live-sieved through 12.5 mm mesh.
7
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TABLE 1. LISTING OF STUDY AREA NAMES, LOCATIONS (COORDINATES), AND
INTERTIDAL 1977-78 SAMPLING DATES
Study Area- Iritertidal
(intertidal habitat type) Location Sampling Dates
1
Kydaka Point -
(exposed sand)
124°
48°
22'
16'
20"
14"
W
N
Apr., Jun.,
Oct., Feb.
2
Pillar Point -
(exposed rock)
124°
48°
06'
12'
03"
51"
W
N
May
3
Twin Rivers -
(exposed gravel)
123°
48°
56'
09'
57"
55"
W
N
May
4
Tongue Point -
(exposed rock)
123°
48°
41'
09'
42"
57"
W
N
May, Jun.,
Oct., Jan.
5
Morse Creek -
(exposed cobble)
123°
48°
20'
07'
48"
09"
W
N
May, Jul.,
Nov., Feb.
6
Dungeness Spit -
(exposed gravel)
48°
123°
08'
11'
47"
12"
N
W
May, Jul.,
Nov., Jan.
7
Jamestown -
(protected sand)
48°
123°
07*
05'
51"
11"
N
W
Apr., Jun.,
Oct., Jan.
8
Beckett Point -
(protected gravel/sand)
48°
122°
04'
52'
37"
56"
N
W
Apr., Jul.,
Oct., Jan.
9
North Beach -
(exposed cobble)
48°
122°
08'
46*
36"
59"
N
W
Apr.
10
North Beach -
(exposed sand)
48°
122°
08'
46'
35"
51"
N
W
May, Jul.,
Nov., Jan.
Protected Sand and Mixed - intp.rtidal (Jamestown and Beckett Point) :
Two quadrats, each 0.05 x 15 cm deep, divided into 0.025m2 x
15 cm cores, fixed and dead-sieved through 1 mm mesh; two 0.25 m
x 30 cm deep cores live-sieved through 12.5 mm mesh.
Rock - subtidal (Tongue Point): four quadrats, each a 0.25 m2 scrape.
Soft-sediment - subtidal (all sites except Tongue Point): Two quadrats
each a 0.1 m2 Van Veen grab sample partitioned on the boat into
equal halves.
8
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124c 123°
VANCOUVER
ISLAND
CAPE
Ju*N ncr s^rt5
k FUca
C!all an*
48
OLYMPIC
PENINSULA
Po\t Seattle
Jefferson j J
!24
123
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In an effort to increase replicate number and hopefully to decrease
sample variance at Beckett Point, Jamestown, and all soft bottom subtidal
sites, the first year quadrat size was halved in the second year by sample
partitioning. Instead of three replicates, four half size replicates were
taken.
All samples collected during the second year program were completely
processed and are stored in the Washington State Baseline Sample Repository
of the University of Washington Friday Harbor Laboratories.
Highly detailed methodology descriptions, site maps, criteria for
study area selection, explanations of the maps, driving directions, tidal
reference point data, and details on access permissions from private and
public agency land owners were presented in the first year report (Nyblade,
1978).
There are many potential sources of error in any field sample
collection and their processing in the laboratory. Field errors include
improperly determining quadrat boundaries, collecting the sample under
water, failure to accurately count small organisms not removed. Although
these mistakes generally cannot be detected once the sample has been
collected, a carefully supervised and experienced field crew reduced these
problems to a minimum.
Potential laboratory processing errors include improper sieving of
samples, mis-counting, mis-weighting, mis-identifications, and clerical
errors each time the data are transcribed. A carefully supervised, experi-
enced staff and frequent referral to the original sample were used to
minimize these sources of sampling error.
IV-B. Data Analysis
The Data Management Plan for the Puget Sound Energy-Related Research
Project and MESA Puget Sound Project received from the Seattle-based project
Office dated 15 January 1976 with subsequent revisions has been followed in
this study. Raw data in the form of keypunched cards have been submitted
to the Project Office on a quarter-by-quarter basis for ultimate archival
in the United States Environmental Data and Information Service. All study
data are available through the E.D.I.S.
A variety of data analyses have been used in this report. Each is
listed and described below. Taken together they provide a clear descriptive
summary of the communities studied and enable community comparisons.
IV-B-1. Species Richness:
As in the first year report, here species richness was the total
number of identification categories/study area stratum/sampling period, thus
summed over the replicates. The identification categories for plants and
animals include both species and higher taxonomic categories where
identification to species was not possible. This total number of species
10
-------
found is a useful figure which summarizes one kind of community complexity
but says nothing about relative abundance or numerical dominance.
IV-B-2.
Diversity - H'
The following formula for diversity has been used (see Pielou, 1975,
for a description of its use and calculation):
This weighs both species number and the evenness of their occurrence. It
thus provides a better measure of community complexity than species richness
because it weighs the relative abundance or the evenness of constituent
species spread through their community This index does not differenti-
ate between low diversity due to low species richness or due to the over-
whelming dominance of one or a very small number of species. In order to
combine plants (biomass) and animals (individuals) into a single H', plant
biomass was converted to individuals (0.1 g = 1). (Algae of <0.1 g were
considered 0.1 g for this index.)
IV-b-3. Total Number:
Total number is the summed number of individuals per identification
categories where individuals are counted, eg. most animals but not plants
which are weighed and not counted. Especially where community biomass
is low, total number gives the most useful information on overall community
abundance. This is the only measure which excludes plants.
IV-b-4. Total Biomass:
The total biomass is the biomass of categories where 0.1 g or more was
present. When the "<0.1 g"'s seemed significant compared to the other
biomass, they were totaled, added to the small real weight, and this new
sum entered as a "less than" weight. This made it very clear when no mean-
ingful community biomass information had been obtained. However, for
algal rich communities, total biomass is the only realistic measure of
overall community abundance. Biomass is also the first step in understand-
ing community productivity.
IV-B-5. Similarity Index - D:
The key analysis in this second year report was the comparison of the
first and second year communities at each stratum sampled. For this
comparison the similarity index D (Schoener, 1968) was used.
s
H* pi log pi
i = 1
n
D- 1-hl
1-1 P.
x,i
- P
y,i
li
-------
This index gives weight to both species (identification) categories present
and their abundance. The values of D range from 0.00 to 1.00, where 0.00
represents total dissimilarity from first to second year, while 1.00 means
completely identical. Abundance values used (weight for algae, number for
animals) were annual means or means for a comparable number of seasons.
Because evolution of taxonomic knowledge from year to year may result in the
same organism being called different names, artificially decreasing similar-
ity, every effort was made to make the identification categories comparable
from year one and year two. The index uses relative abundance of a species
in the community from one year to the next. It is insensitive to absolute
changes in abundance, as long as percentage community composition remains
the same. Abundance changes, if uniform across the spectrum of community
constituents, do not reflect a change in community structure. However,
such changes may be of interest.
12
-------
SECTION V
RESULTS
Replicate samples were successfully collected all four seasons at the
seven seasonally sampled study areas and at all intertidal strata except
+0' Tongue Point in the fall and winter and +0' Morse Creek in the winter.
Spring sampling at Pillar Point, North Beach Cobble, Twin Rivers and all
subtidal areas was successfully completed with the exception of -5m Morse
Creek and -10m Twin Rivers. Bad weather/wave conditions were responsible
for sample collection failures. All samples collected were completely
processed and are presently stored at the Washington State Baseline Sample
Repository, Friday Harbor Laboratories.
Approximately one thousand species of plants and animals were identi-
fied to species during the course of this study. Crustaceans were most
numerous (280+), followed by algae (245+), annelids (220+), and molluscs
(135+).
As found in the first year in general, although each stratum of each
study area had a unique community, distinct substratum/exposure-associated
communities were recognizable and these communities persisted from the
first through the second year of sampling. The results are presented
below by study area. These are arranged in increasing substratum fineness
and where habitat types were paired, the eastern then western site.
The data are given largely in tabular format. The tables in the
Results section are normalized to 1 m2 surface area and abridged to include
only the communi ty species which are dominant by virtue of their high
biomass, numerical abundance, or trophic importance. This determination of
dominants is subjective. For a full listing see Appendix I. The values
for species richness, diversity, total number (normalized to 1 m2), and
total biomass (per 1 m2) were taken from the complete data sets given in
Appendix I. Cobble and rock methodology precluded adding the 0.01 m2
subsamples with the residual 0.2m2 scrape for normalizing. In this case
values for both sizes were normalized to 1.0 m2 and species by species the
value was selected for the table which would give the best measure of the
true value in the quadrat. In general for small organisms the 0.01 m2
normalized value was taken, while for large organisms the 0.2 m2 normalized
value was used. Cobble infaunal organisms were normalized to 1.0m2 and
added to epifaunal scrape organisms. Because large infaunal organisms were
generally found only at Beckett and Jamestown, live sieve data are usually
not commented on.
Special attention should be given to the fact that rock and cobble
sampling methodology changed during the first year (Nyblade, 1978).
13
-------
Original sampling methodology (first three quarters) involved first remov-
ing all large organisms from the 0.25 m2 area, then taking the subsaraples.
This allowed recombination of normalized subsamples with the 0.25 m2 large
organism scrape, resulting in a single set of numbers for each quadrat
location. Methodology for the last five sampling quarters involved remov-
ing the subsamples first; then the large organisms were removed from the
remaining 0.2 m2 area. The resulting sets of numbers are not objectively
additive. To generate a single set of values for the organisms collected
at each quadrat location, a species by species determination (subjective)
of which subsampling value represents the best value would be necessary.
This was done in order to compute first and second year community simi-
larity. However, Appendix I gives the complete data set.
The Appendix I unabridged tables give number of replicates, means, and
standard deviations in the sampled quadrat size; e.g., 0.05 m2 x 15 cm or
0.25 m2 (rock scrape). Cobble and rock entries are given in two or three
data columns: the 0.01 m2 scrape subsamples, the residual scrape (0.2m2),
and for cobble the sediment sample (0.05 m2 x 15 cm deep). In Appendix I
tables / means present but not quantified,
A Note of caution: The tables in the "Results" section are abridged,
and the values are means normalized to 1 m2 surface area. Appendix I
tables must be consulted for the complete data. All statements concerning
patchiness are based on the replicate variance seen in Appendix I table
standard deviations.
Results of the physical field parameters measured {first year (Nyblade,
1978) sediment analysis summary, water temperature, salinity, and weather}
are presented in Appendix II. Where noteworthy, they are mentioned in the
study area results presented below.
14
-------
Tongue Point (rock)
The substratum at Tongue Point over the tide heights and the depths
sampled was solid rock. This rock was relatively smooth and flat with few
pools or large crevices, and it sloped rather uniformly from +6* to 0'.
This site was strongly exposed to local wave action from the north. How-
ever oceanic swells generally were fairly well damped this far into the
Strait. No change in the physical environment was observed from year one
to year two.
Table 2a presents a summary of the biological community over the four
quarters of sampling at +6'. This community was dominated by a few species
of red algae, herbivorous gastropods, and planktivorous barnacles and
mussels. Algal biomass was highest in spring/summer declining in the fall
and winter. Seasonal patterns in the herbivores were obscured by large
winter recruitment of Littorina sitkana. As in the first year barnacles
showed a major recruitment between spring and summer quarter sampling.
Especially noteworthy was the massive recruitment of Muscuius pygmaeus
between winter 1977 and spring 1977 (5200 to 68,770 / m2) and its subsequent
decline through the year.
Alaria was present in some quantity in the spring, but vanished in the
following quarters. It probably burned off during summer low tides. The
other algae showed no consistent pattern of seasonal change, while the
herbivores increased in number and biomass from spring through summer
declining in the fall/winter. Barnacle number peaked in the summer and
then declined.
The massive barnacle and Musculus recruitment dominates the seasonal
changes in diversity, total number, and biomass. Although species richness
decreased from spring to summer (79 to 66), diversity increased because of
the numerical dominance of Musculus in the spring.
Examination of the means and standard deviations in the Tongue Point
+6' Appendix I table illustrates the spatial patchiness of the organisms in
the +6' community. Still, dramatic population changes such as that of the
barnacles and mussels showed through this variance. Despite the Musculus
spring numerical dominance, overall community similarity between year one
and two was high. (See Table 18 for all similarity values.)
Table 2b gives the abridged results for Tongue Point +3'. This commu-
nity was structurally dominated by the brown alga Alaria, articulated
corraline algae (Corallina and Bossiella), mussels, and barnacles.
Associated with these were organisms which ate them—the herbivorous
chiton (Cyanoplax) and gastropods (Collisella, Notoacmea, and Qnehidella)
and the carnivorous Thais and Leptasterias—and small organisms which
inhabited the structure they provided—nematodes, polychaetes, oligochaetes,
tanaids, isopods, amphipods, insect larvae, and the small Cucumaria. With
the exception of the massive Musculus recruitment little consistent
seasonal change in populations appeared, largely because of the patchiness
of the major structural dominants. Corallina is a long-lived perennial
alga which occurred in descrete patches of very dense algal turf.
15
-------
Table ?a. Tongue Point (rock) dominant benthic organisms from the high (+6^ intertidal zone
collected spring 1977 through winter 1978, expressed as number and/or biomass (g)
per m . For complete data set see Appendix I.
Spr 77
# wt
Sum 77
# wt
Fall 77
# wt
Win 78
# wt
Phaeophyta
Alaria
spp.
Rhodophyta
Endocladia
muricata
Gigartina
papillata
Halosaccion
glandiform
Iridaea
spp.
Porphyra
spp.
Mollusca
Gastropoda
Collisella
digitalis
C.
strigatella
Littorina
sitkana
Siphonaria
thersites
Bivalvia
Musculus
pygmaeus
Mytilus
spp.
17.0
130.0
217.5
23.5
17.5
<10.0
260.0
130.0
570.0
440.0
48.5
40.0
10.0
50.0
60.0
96.5
84.5
302.5
3.5
370.0 45.0
210.0 5.0
2650.0 50.0
610.0 20.0
68770.0 230.0 2040.0 10.0
304.0 21.0 60.0 2.0
10.0
23.5
4,5
63.0
0
290.0 44.5
180.0 3.5
750.0 10.0
70.0 <10.0
320.0 <10.0
20.0 <10.0
<10.0
10.0
1.5
0.5
2,0
3.0
290.0 29.5
70.0 0.5
9990.0 *10.0
0
10.0 <10.0
20.0 <10.0
-------
Spr 77
# wt
Crustacea
Cirripedia
Balanus.
spp.
Chthamalus
dalll
Tanaidacea
Pancolus
calif ornierisis
Isopoda
Dynamenella
CI isheareri
Amphipoda
Camnaridea
spp.
Insecta
Dipteran
larvae spp.
Species Richness
Diversity,
Total Number
Total Biomass (p)
5*6.0 70.5
3980.0 70.0
2090.0 <10.0
3680.0 10.0
320.0 10.0
390.0 <20.0
79
1.50
89,216
3,466
Table 2a, (cont.)
Sum 77 Fall 77 Win 78
# wt # wt # wt
16790.0 641.5
4310.0 70.0
420.0 <10.0
670.0 <10.0
330.0 50,0
330.0 <10.0
66
2,79
29,800
1,619
4620.0 144.0
2270.0 50.0
510.0 <10.0
3590,0 40.0
370.0 10.0
360.0 <30.0
56
2.75
14,880
446
5480,0 415.0
3610 60.0
250.0 <10.0
1990,0 <10.0
40 <10.0
150.0 <10.0
44
2.54
13,220
374
-------
Table ?h, Tongue Point (rock) dominant benthic organisms from the mid (+3^ intertida1 zone,
collected spring 1977 through winter 1978, expressed as number and/or biomass (g)
per . For complete data set see Appendix I.
Spr 77
// wt
Sum 77
wt
Fall 77
wt
Win 7B
# wt
Phaeophyta
Alaria
spp.
Rhodophyta
Bossiella
plumosa
Corallina
Vancouvertensis
Halosaccion
glandiforme
Cnidaria
Anthozoa
Anthopleura
elegantissima
Nematoda spp,
Moiluaca
Aniphineura
Cyanoplax
dentiens
Gastropoda
Barleeia
haliotiphila
Collisella
pelta
Notoacmaea
scutum
1173.5
140.0
970.0
6.0
899.5
120.0
1050.0
55.0
30.0 <10.0 30.0 <10.0
1460.0 <10.0 3720.0 <10.0
30.0 < 10,0 50.0 0.5
1500.0 <10.0 0
230.0 100.0 10.0 10.0
0 10.0 <10.0
1208.0
220.0
1060.0
969.5
270.0 40.0
1300.0 <10.0
160.0 10.0
4250.0 <10.0
20.0 91.5
0
617.5
90.0
250.0
227.5
340.0 50.0
270.0 <10.0
50.0 <10.0
4360,0 <10.0
40.0 30.0
70.0 4.5
-------
Table 2b. (cont.)
Spr 77 sun 77
# wt # wt
Fall 77
# wt
Win 78
# wt
Mollusca (cont.)
Gastropoda
Onchldella
borealIs
Tha Is
spp •
Blvalvla
Musculus
pygmaeus
Hytllus
spp.
Annelida
Polychaeta
Syllldae
spp.
Ollgochaeta
spp.
Crustacea
Clrripedia
Balanus
spp.
TanaIdacea
AnatanaIs
normanl
Pancolus
callforniensls
Isopoda
Dynamenella
shearer!
0
0
1810.0 <10.0
270.0 3.0
3110.0 <40.0
1480.0 <10.0
440.0 30.0
270.0 <10.0
640.0 <10.0
4970.0 10.0
0
50.0 5.0
48380.0 160.0
15.0 6.0
2360.0*20.0
6310,0 <10.0
2400.0 20.0
3320.0 <10.0
1090.0 <10.0
4050.0 10.0
370.0 190.0
20.0 <20.0
300.0 10.0
200 61.0
6840.0 <30.0
2200 <10.0
480.0 30.0
2490.0 <10.0
2260.0 <10.0
880.0 <10.0
220.0 <10.0
20.0 <20.0
1080.0 <10.0
80.0 1.0
1490.0 <20.0
590.0 <10.0
810.0 590.0
80.0 <10.0
770.0 <10.0
2000.0 <10.0
-------
Table 2b. (cont.)
Mollusca (cent.)
Isopoda
Idotea
spp.
Amphipoda
Gaimnarldea
spp.
Decapoda
Pagurus
hirsutiusculus
Insecta
Dipteran
larvae spp.
Echinodermata
Asteroidea
Leptasterlas
hexactis
Holothuroldea
Cucumaria
pseudocurata
Species Richness
Diversity, H^"
Total Number
Total Biomass (g)
Spr 77
# wt
114
3.15
30,750.0
2984.5
Sum 77
# wt
Fall 77
wt
27.5 1.0 20.0 6.5 54.0 8.0
20.0 1.0 10.0 <.10.0 40.0 1.0
1260.0 <10.0 5410.0 <10.0 410.0 <10.0
136
2.79
97,740.0
3711.5
126
3.08
45,071.5
4807.0
Win 78
# wt
19.0 2.0
2390.0 3.0 5800.0 30.0 11250.0 70.0 2780.0 30.0
0
200.0 <-20.0
10.0 <10.0 50.0 <10.0 20.0 <10.0
7120.0 150.0 6520.0 170.0 3150.0 140.0 1250.0 50.0
97
3.14
20,500.5
2279.5
-------
Species richness was fairly constant over the year. Muscuius recruit-
ment was mainly responsible for the total number peaking in the summer.
Biomass peaked in the fall, largely because Maria, large Corallina, and
Halosaccion patches were sampled. Year one/year two community similarity
was high (see Table 18).
Table 2c presents the summary results of Tongue Point +0'. Wave
conditions prevented sampling at this tide height in the fall and winter.
This community was structured by the brown algae Alaria and Hedophyllum, the
seagrass Phyllospadix, the boring clam Hiatella arctica, and the large
barnacles Balanus cariosus and B_. nub ills. Important herbivores were the
chitons. Lacuna, and the spider crab Pugettia gracilis and carnivores,
Cancer oregonesis and Leptasterias. Small organisms associated with the
strucutral organisms included the polychaetes, tanaids, isopods, and
amphipods. No consistent seasonal population changes were detectable
because of the over-riding patchiness of the major structural organisms of
this community and because surf conditions prevented fall and winter
sampling.
Species richness, diversity, total number, and biomass were high both
quarters. The large patch of Phyllospadix with its associated worm fauna
was responsible for the lower year one/year two similarity compared to +6'
and +3'.
Table 2d gives the abridged results at Tongue Point -5 m and -10 m.
The community at Tongue Point -5 m was dominated by algae and the urchins,
Strongylocentrotus. Grazers besides the urchins included chitons, Acmaea
mitra, Calliostoma, Lirularia, Margarites and Pugettia gracilis. The
grazers exerted obvious strong pressure on this community. The only algae
present in quantity have thwarted herbivores by chemical noxia (Desmarestia)
or are structurally unpalatable (the calcareous alga Calliarthron).
Numerical dominance at this level was by small organisms associated with
Calliarthron—Granulina, the isopods, and amphopods. The community at -10 m
was also dominated by grazers (chitons, Acmaea, Lirularia, Strongylo-
centrotus spp.). However, suspension feeders made an appearance
(Calyptraea and Spirorbis). Low -5 m year one/year two similarity is due
to the larger patch of Calliarthron sampled in year two with its associated
rich worm fauna. Low - 10m first and second year similarity was caused
by much richer flora the second year. At both levels low similarity
probably reflected patchiness, rather than major community change.
21
-------
Table 2c. Tongue Point (rock) dominant benthic organisms from the low (W) intertidal zone,
collected spring 1977 through summer 1977, expressed as number and/or biomass (g)
per in". For complete data set see Appendix I.
Spring 1977
f wt
Summer 1977
* wt
NJ
NJ
Phaeophyta
Alaria
spp.
Hedophyllum
sessile
Rhodophyta
Bossiella
plumosa
Iridaea
cordata
Odonthalia
flocossa
Spermatophyta
Phyllospadix
scouleri
Nematoda
spp.
Mollusca
Amphineura
Katharina
tunicata
Tonlcella
lineata
Gastropoda
Lacuna
variegata
Velutina
laevigata
3773.5
343.5
580.0
204.0
50.0
927.0
700.0 <-10,0
6.5 <0.5
10.0 22.5
20.0 <-10.0
0
3104.0
60.0
10.0
26.0
30.0
4359.5
2030.0 <-10.0
20.0
30.0
90.0
30.0
140.0
4.0
<10.0
C10.0
-------
Table 2c. (cont.)
Mollusca (cont.)
Bivalvia
Hiatella
arctica
Annelida
Polychaeta
Capltellldae
Capltella
capitata
Clrratulidae
Clrratulus
ciTratus
Nereldae
Nereis
spp.
Sabellidae
spp.
Syllldae
spp.
Terebellldae
spp.
Ollgochaeta
spp.
Crustacea
Clrrlpedla
Balanus
carlosus
B.
nubllus
Spring 1977
# wt
50.0
50.0
280.0
1510.0
1060.0
80.0
40.0
10.0
<10.0
J
420.0 <10.0
620.0 <80.0
< 70.0
190.0 <10.0
203.0
130.5
Sumner 1977
# wt
20.0
40.0
1660.0
2500.0
1600.0
2830.0
<10.0
1100.0 10.0
< 10.0
590.0 <30.0
< 50.0
< 70.0
< 30.0
< 10.0
210.0 159.5
0
-------
Crustacea (cont.)
Tana Ida cea
Ana tana is
nonnani
Isopoda
Limnoria
a1parum
Amphipoda
GammarIdea
spp.
Decapoda
Cancer
oregonensis
Oedignathus
inermis
Pugettia
gracilis
Echinodermata
Asteroidea
Leptasterias
hexactis
Species Richness
1
Diversity, H
Total Number
Total Biomass (g)
Table 2c. (cont.)
Spring 1977 Summer 1977
# wt # wt
300.0 <10.0 780.0 <10.0
90.0 <10.0 80.0 <10.0
1320.0 <10.0 3600.0 20.0
10.0 10.0 10.0 40.0
20.0 40.0 0
170.0 10.0 670.0 20.0
10.0 <10.0
180
2.33
10,470.0
6,506.0
10.0 <10.0
175
2.17
28,279.5
8,229.8
-------
Table 2d. Tongue Point (rock) dominant benthic organisms from the shallow (-5m, -10 m) subtldal
zone, collected spring 1977, expressed as number and/or biomass (g) per m2. For
complete data set see Appendix I,
-5 m
# wt
-10 m
# wt
NS
Ul
Phaeophyta
Desmarestia
viridis
Rhodophyta
Calliarthron
tubereulosum
Mollusca
Amphineura
Tonicella
lineata
Gastropoda
Acmaea
mitra
Amphissa
Columbiana
Calliostoma
ligatum
Calyptraea
fastigiata
Fusitriton
oregonensis
Granulina
margaritula
Lirularia
lirulata
Margarites
pupillus
Ocenebra
lurida
154.8
10.8
4487.2
59.2
7.2 2.0
315.2 13.6
524.0 117.2
0
0
540.0 4.0
159.2 1.2
228.0 20.0
53.2 7.6
49.2
5.2
7.2
2.0
17.2
6.8
33.6
1.6
1.2
0.0
2.0 <0.4
1.2 86.4
0
0
0
0
-------
Annelida
Polychaeta
Nereidae
Nereis
spp.
Platyrtereis
bicanalieulata
Serpulidae
Spirorbis
spp.
Crustacea
Isopoda
Ianiropsis
spp.
Munna
spp,
Amphipoda
Gammarldea
spp.
Decapoda
Cancer
oregonensis
Paguridae
spp.
Pupettia
gracilis
Echirtoderma ta
Echinoidea
Strongylocentrotus
droeba chiens is
2d, (cont.)
-5 m
# wt
-10 m
# wt
160,8 *0,4
2153.2 <0.4
63.2 <0.4
377.2 0.0
265.2 0.0
9348.0 12.8
28.0 18.4
429.2 15.6
960.0 38.0
25.2 1116.0
7.2 40.4
62.0 40.4
2.0 <0.4
7.2 <0.4
0
120.0 1.2
0
0
4.0 <0.4
16.0 701.2
-------
Table 2d.
Echlnodermata
Echinoldea
Strongylocentrotus
franclscanus
Species Richness
Species Diversity,
Total Number
Total Blomass (g)
(corit.)
-5 m -10 n
# wt # wt
9.2 2786.4 5.2 1723.2
209 94
1.45 3.00
18,634 834
9,160 2,988
-------
Pillar Point (rock, intertidal; sand, subtidal)
The Intertidal at Pillar Point from +0' to +6' was solid rock. Unlike
Tongue Point, the rock was an irregular conglomerate, not smooth at all.
The slope varied from 45° to 90 . There were no large crevices, and no
pools were sampled. Seasonally, sand scouring at +01 might be significant
as the rock at that level was adjacent to a sandy bottom. Because of this
fact and the very limited area available for sampling on the rock outcrop,
this area was sampled only once during the second year, in the spring. This
site was exposed to both extreme wave action from the north and to rather
continuous oceanic swells. Subtidally, the substratum was medium and fine
sand at -5m and fine sand at -10m. Salinity in winter quarter of the first
year showed some slight freshwater influence from the Pysht River, likely
of no biological consequence. No major changes in the physical environment
were observed over the course of the study.
Table 3a presents the abridged results for Pillar Point +6'. The
community at this level was relatively simple and was dominated by grazers
(Collisella digitalis and Littorina spp.) and by the planktivorous
barnacles (Balanus glandula and Chthamalus dalli). Associated with this
algal barnacle matrix were small crustaceans and dipteran larvae. The
similarity between the first and second year community was very high.
Table 3b presents a summary of the +3' Pillar Point data set. The
community at this level was structurally dominated by algae (Alaria,
Hedophyllum, and to a lesser extent Corallina, Gigartina, Odonthalia, and
Iridaea), Mytilus spp., and barnacles (Balanus spp.). Two sets of organisms
were associated with these structuring components. There were those which
eat them, the herbivores (limpets and chitons) and the carnivores (Thais
spp.). And there were small organisms intimately dependent on the physical
structuring of the dominants: nematodes, polychaetes, oligochaetes,
tanaids, isopods, amphipods, and insect larvae.
Year one/year two community similarity was quite high despite consider-
ably lower mussel and barnacle numbers.
An abridged data set for Pillar Point +0' appears in Table 3c. Two,
things stood out in this community. The community was totally structurally
dominated by plants (Alaria, Egregia, Iridaea, and especially Phyllospadix),
and large barnacles were absent. Sand scouring may have been responsible
for the latter. Herbivores (chitons, Lacuna, Idotea, and Pugettia) and
small plant-associated animals (polychaetes, oligochaetes, tanaids, isopods,
and amphipods) were also abundant. Cancer oregonensis was the only major
carnivore collected.
The similarity in data sets between year one and year two was low due
to the massive amounts of Phyllospadix and its associated worm fauna
collected the second year. This represents patchiness, not community
change.
28
-------
Table 3a. Pillar Point (rock) dominant benthic organisms from the high (+6') intertidal zone,
collected Spring I977t expressed as number and/or biomass(g) per nr. For complete
data set see Appendix I.
vD
# Wt
Rhodophyta
Gigartina 10.0
papillata
Mollusca
Collisella 120.0 41.0
digitalis
Littorina 210.0 0.5
scutulata
L. 1090.0 20.0
sitkana
Crustacea
Cirripedia
Balanus 1770.0 70.0
glandula
Chthamalus 7460.0 190.0
da 111
Isopoda
Idotea 31.5 2,0
montercyensis
Amphipoda
Gammaridea 70.0 <10.0
spp.
Insecta
Diptera 170.0 <10.0
larvae spp.
-------
Species Richness
Diversity, H1
Total Number
Total Biomass
(cont,)
1t wt
42
i .85
11 ,620
480
-------
Table 3b, Pillar Point (rock) dominant benthic organisms from the mid(+3^ intertidal
collected spring 1977, expressed as number and/or biomass (g) per m . For
data set see Appendix I.
zone,
complete
# wt
Phaeophyta
Alaria 2131.0
sp.
Hedophyllum 1432,0
sessile
Rhodophyta
Corallina 20.0
vancouveriensis
Cigartlna 33,0
papillate
Halosaccion 10.0
w glandiforme
Iridaea 784.0
cordata
Odonthalia 220.0
floccosa
Hemertea 150.0 10.0
spp.
Nematoda 980.0 10.0
spp,
Mollusca
Amphineura
Cyanoplax 1.5 <0.5
dentiens
Katharina 19.0 298.0
tunicata
-------
Mollusca (cont.)
Gastropoda
Thais
spp.
Bivalvla
Hiatella
arctica
Mytilus
spp.
Annelida
Oligochaeta
spp.
Polychaeta
Sabellidae
spp.
Syllidae
spp.
Pycnogonida
spp.
Crustacea
Cirripedia
Balanus
spp.
Tanaidacea
Leptochelia
dubia
Table 3b. (cont.)
_# wt
20.0 <20.0
40.0 2.0
102.5 0.5
140 <.10.0
530.0 -450.0
370.0 <40.0
70.0 <50.0
421.5
2160.0
863.5
<10.0
-------
Crustacea (cont.)
Isopoda
Dynamenella
sheareri
Muttna
c hr oma tocepha la
Amphipoda
Caprellidea
Cercops
compactus
Ganmarldea
spp.
Insecta
Dlptera
larvae app.
Species Richness
1
Diversity, H
Total Number
Total Bloaass
Table 3b. (cont.)
#
wt
550.0
790.0
290.0
100.0
130.0
131
2.50
8,994
6,317
<10.0
<10.0
<10.0
<10.0
<10.0
-------
Table 3c. Pillar Point (rock) dominant benthic organisms from the low (+0) i^tertidal zone,
collected spring 1977, expressed as number and/or biomass(g) per m For complete
data set see Appendix 1.
Phaeophyta
Alaria
spp.
Egregia
menziesii
Rhodophyta
Iridaea
cordata
w
¦e-
Spermatophyta
Phyllospadix
scouleri
Nematode
spp.
Hollusca
Amphineura
Tonicella
lineata
Gastropoda
Lacuna
variegata
Bivalvia
Nytilus
sp. (Juv.)
Annelida
Oligochaeta
spp.
wt
2593.5
1621.5
261.0
6119.5
8200.0
<10.0
2.5
<0.5
20.0
<10.0
20.0
<10.0
1630.0
<10.0
-------
Annelida (cont.)
Polychaeta
Arenlcolidae
spp.
LumbrinerIdae
Lumbrlneris
spp.
Spionidae
spp.
Syllidae
spp.
Crustacea
Tanaldacea
Antanais
normani
Isopoda
Idotea
spp.
Atnphipoda
Caprellidea
Caprella
spp.
Gaonaridea
spp.
Decapoda
Cancer
oregcmensis
Pugettla
gracilis
Table 3c. (cont)
wt
2050.0
<10.0
480.0
<10.0
930.0
<50.0
280.0
<40.0
40.0
<10.0
230.0
<20.0
10.0
<10.0
510.0
2.0
2.5
10.0
1.5
13.0
-------
Table 3c. (cont.)
Species Richness
Diversity, H1
Total Number
Total Blomass (g)
124
2.04
15,642
11,375
-------
Table 3d gives a summary of the subtidal Pillar Point data set. The
communities at -5m and -10m were similar. They were dominated by epifaunal
and infaunal deposit feeders. Species richness and diversity were high at
both levels. Neither density nor measured biomass were particularly high
at either level. These communities run energetically on imported organics
and an unknown amount fixed at the levels by the largely microscopic
epiflora. Year one, year two similarity was surprisingly high considering
the crudeness of grab sampling.
37
-------
OJ
00
Table 3d, Pillar Point (fine-medium sand) dominant benthic organisms from the shallow (-5 m.slO m)
subtidal zone, collected spring 1977, expressed as number and/or biomass (g) per m .
For complete data set see Appendix 1.
-5 io -10 m
# wt # wt
Mollusca
Bivalvia
Macoma 62,0 <2.0 12.0 <2,0
spp.
My sella 400.0 <2.0 96.0 <2.0
tumida
Psephidia 476.0 2.0 216.0 2.0
lordi
Tellina 46.0 <2.0 52.0 <2.0
spp.
Tresus 70.0 <2.0 36.0 <2.0
capax
Annelida
Polychaeta
Capitellidae
Capitella 870.0 <2.0 66.0 <2.0
capitata
Mediomastus 1830.0 <2.0 380.0 <2.0
sp.
Cirratulidae 320.0 <2.0 392.0 <2.0
spp.
Orbiniidae
Scoloplos 200.0 <2.0 76,0 <2.0
sp.
Spionidae
Prionospio
steenstrupi
3000 <2.0 770.0 <2.0
-------
Table 3d.
Crustacea
Cumacea
spp.
Tanaidacea
Leptochelia
dub la
Anphipoda
Gammaridea
spp.
Species Richness
Diversity,
Total Number
Total Biomass (g)
(cont.)
-5 n -10 in
# wt # wt
282.0 <2.0 12.0 <2.0
366.0 <2.0 110.0 <2.0
2290.0 2.0 700.0 <2.0
77 86
2.82 3.26
13,814 4,42*
<156 <172
-------
North Beach Cobble (cobble)
This area was selected in 1976 as a cobble habitat although the +6'
substratum consisted of coarse sand. During the first year the +0' cobble
was buried in sand, and cobble was uncovered at +6'. This sediment
instability made this area unsuitable for baseline population monitoring.
Because of this the area was sampled only once during the second year.
The beach had a fairly gentle slope and an offshore kelp bed doubtless
moderated this area's exposure. There were no ocean swells at this end of
the Strait and given the prevailing winds wave activity was probably fairly
moderated.
North Beach subtidal will be discussed with North Beach Sand.
Table 4a presents summary data for North Beach Cobble +6'. The
community present at this level was fairly simple, dominated by a grazer
(Littorina scutulata), a surface detritivore (Exosphaeroma), an infauna
detritivore (oligochaetes), and barnacles. Low similarity between year one
and year two resulted from sediment instability. Spring year one found the
area sand covered, while spring year two found the cobble exposed.
Table 4b presents an abridged data set for North Beach Cobble +3'.
This community was dominated by grazers (Collisella spp., Littorina spp.),
planktivores (Balanus spp.), under-rock detritivores (Exosphaeroma,
Gnorimosphaeroma, and Hemigrapsus), and predatory gastropods (Thais spp.).
Macroalgae were a very minor constituent of this community, as was the
infauna.
As would be expected given the nature of the substratum in this area,
the expected rock organism patchiness was even more extreme. The variance
among replicates was very high (Appendix I). Year one/year two similarity
was lower than for the Morse Creek +3' cobble community. A major factor in
this was the much lower isopod number the second year.
Abridged results from North Beach Cobble +0' are given in Table 4c.
The rich spring algal flora was dominated by Ulva, Alaria, Nereocystis.
Iridaea and Pterosiphonia. Major epifaunal constituents were grazers
(chitons, Lacuna, Notoacmea, and Idotea), planktivores (Balanus spp.),
predators (Thais spp., Cancer spp.,) and under-rock detritivores
(Exosphaeroma, gammarids). The abundant worm infauna consisted of an active
predator (Hemipodus), tube-building algal grazers (Nereidae spp., Onuphis),
and detritivores (oligochaetes, Malacoceros).
The richer second year algal flora was primarily responsible for the
fairly low year one/year two community similarity.
40
-------
Table 4a, North Beach Cobble (cobble over sand) dominant benthic organisms from the high (+fc7
inter tidal zone, collecteJ spring 1977, expressed as number and/or biomass (g) per at1.
For complete data set see Appendix I.
# wt.
Mollusca
Gastropoda
Littorina 720.0 34,0
scutulata
Annelida
Oligochaeta 140.0 <2.0
spp.
Folychaeta
Nereidae
Nereis 30.0 <-2.0
sp.
Crustacea
Cirripedia
Balanus 320.0 18.0
glandula
Isopoda
Exosphaeroma 366.0 <2.0
media
Amphipoda
Gammaridea 6,0
-------
Table 4b. North Beach Cobble (cobble over sand) dominant benthic organisms from the mid (+3^
intertidal zone, collected spring 1977, expressed as number and/or biomass (g) per m .
For complete data set see Appendix I.
# wt
Cnidaria
Anthozoa
Anthopleura 8.5 3.0
elegantissima
Mollusca
Gastropoda
Collisella 50.0 0.0
pelta
Collisella 50.0 0.5
strigatella
Littorina 252.0 12„G
scutulata
to
L. 1066.0 52.0
sitkana
Thais 96,0 7,5
spp.
Annelida
Oligochaeta 76.0 <2.0
spp.
Polychaeta
Nereidae
Nereis 56.0 <2,0
vexillosa
Spionidae
Polydora 76.0 <2,0
proboscidea
Syllidae
Syllis 126.0 <2.0
spp.
-------
Table 4b. (cont.)
Crustacea
Clrripedia
Balanus
spp.
Isopoda
Exosphaeroma
media
Gnorlmosphaeroma
oregonense
Decapoda
Hemigrapsus
nudus
Insecta
Dlptera
larvae spp.
Species Richness
1
Diversity, H
Total Number
Total Blomass
#
wt
126.0
107.0
380.0
10.0
20.0
38
2.48
3,802
128
14.0
<1.0
<2.0
23.0
<10.0
-------
Table 4c. North Beach Cobble (cobble over sand) dominant benthlc organisms from the low (40*)
intertidal zone, collected spring 1977, expressed as number and/or biomass(g)per m^.
For complete data set see Appendix I,
# wt
Chlorophyta
tllva 90.0
spp.
Phqeophyta
Alaria 713,0
spp.
Nereocystis 156.5
luetkeana
Rhodophyta
Gigartina 45.0
papillate
Iridaea 409.5
•c- cordata
Pterosiphonia 174.0
bipinnata
Mollusea
Amphineura 10.0 <2.0
spp.
Gastropoda
Lacuna 886.0 <10.0
variegata
Notoacmea 16.0 <-10.0
scutum
Annelida
Oligochaeta 326.0 <10.0
spp.
Folychaeta
Glyceridae
Hemipodus 120.0 <2.0
borealls
-------
Table 4c. (cont.)
wt
Annelida
Polychaeta
Nereldae 50.0 <10.0
spp.
Onuphldae
Onuphls 100.0 ^2.0
stlgmatis
Splonldae
Malococeros 270.0 <10.0
glutaeus
Crustacea
Clrrlpedla
Balanus 10.0 <10.0
carlosus
Isopoda
Exosphaeroma 30.0 <10,0
ampllcauda
E. 430.0 <10.0
media
Idotea 1186.0 0.5
spp.
Amphlpoda
Gammaridea 317.5 <2.0
spp.
Decapoda
Cancer 10.0 <10.0
spp.
Species Richness 125
-------
Diversity, H1
Total Number
Total Biomass
Table 4c. (cont.)
3,24
6,724
1,862
-------
Morse Creek (cobble, intertidal; gravel, subtidal)
Although +6' at this area, as at North Beach Cobble, also consisted of
sandy gravel over buried cobble, this area was selected for a cobble habitat.
During the course of the year, the +6' cobble was never uncovered, although
live barnacles were recovered from buried cobble in several quadrats. Mid
and low intertidal zone sediments consisted of cobble over sand. The
subtidal sediment at -10 m consisted of gravel. No samples were collected
at -5 m because small cobble prevented the grab from operating.
This beach had a fairly gentle slope and like North Beach only a
moderate exposure to wave activity. No major changes in the physical
environment were observed between the first and second year.
Table 5a presents the abridged results for Morse Creek +6'. This
community was very simple, consisting of detritus feeding oligochaetes,
isopods, and gammarid amphipods. The barnacles were buried in the gravel,
and the Littorina were probably drift. The oligochaetes showed no seasonal
pattern. However, the amphipods showed a peak in summer quarter. Diversity
and species richness were uniformly low. Total number generally followed
the amphipods. Biomass was insignificant.
Areas with sparse fauna generally show extreme patchiness, and this
level was no exception (see variances, Appendix I). However, despite this
patchiness, year one and year two community similarity was very high.
Abridged results for Morse Creek +3* are presented in Table 5b. The
rock community at this area and level was basically two dimensional, with
no structural dominants such as at Tongue and Pillar. Algal species
richness was low and what algae occurred regularly (Fucus and Gigartina)
were very patchy. Gastropod grazers, Idotea, and barnacles dominated the
epifaunal community. Hemigrapsus and Pagurus, detritivores, dominated
under-rock. The infaunal community was dominated by the detritivores
Capitella, Malacoceros, Corophium. and dipteran larvae.
Species richness, diversity, and biomass were fairly constant.
Numbers peaked in summer/fall. Intense barnacle recruitment occurred
between spring and summer quarter sampling. The second year community was
very similar to the first year.
Table 5c gives the summary results from Morse Creek +0*. Algae
(Alaria, Hedophyllum, and Iridaea) were the structural dominants of the
epi-community. Herbivore associates included Lacuna, Notoacmea and
Pugettia. The infaunal community was dominated by detritivores (nematodes,
Abarenieola, Capitella, Cirratulus, Armandia, spionids, Leptochella and
some gammarids). However, a herbivore (Nereis) and suspension feeders
(Protothaca, Tresus, and sabellids) were also abundant. Important
predators in this community were Thais spp. and Cancer spp.
47
-------
Table 5a. Morse Creek (sandy gravel) dominant benthic organisms from the high (+6) intertidal
zone, collected spring 1977 through winter 1978, expressed as number and/or biomass(g)
per m . For complete data set see Appendix I.
Mollusca
Gastropoda
Littorina
sitkana
Annelida
Oligochaeta
spp.
Crustacea
Cirripedia
Balanus
spp.
Isopoda
Gnorimosphaeroma
oregonense
Amphipoda
Gammaridea
spp.
Species Richness
Diversity,
Total Number
Total Biomass (g)
#
Spr 77
60.0
6.0
6,0
wt
<2.0
<2 .0
<2.0
110.0 <<2.0
8
1.31
206
<16
Sum 77
# wt
Fall 77
#
wt
40.0 <2.0
326.0 4.0
2028.0 4.0
16
0.76
2,458
<36
10.0
<2.0
110.0
6.0
10.0 <2.0
1106.0 <4.0
7
0.48
1,257
<20
Win 78
80.0
wt
8,0
20.0 <2.0
406.0 12.0
116.0 <2.0
126.0 <2.0
15
1.72
844
<46
-------
Table 5b. Morse Creek (cobble over sand) dominant benthic organisms from the mid (+35 ineertidal
zone, collected spring 1977 through winter 1978, expressed as number and/or biotnass (g)
per n>2. For complete data set see Appendix I.
Fhaeophyta
Fucus
distichus
Rhodophyta
Gigartina
papillata
Platyhelminthes
Turbe1laria
spp.
Nemertea
spp.
Mollusca
Gastropoda
Collisella
pelta
C.
strigatelia
Lacuna
variegata
Littorina
sitkana
Bivalvia
Hytilus
spp.
Annelida
Polyehaeta
Capitellidae
Capitella
capitata
Spr 77
# wt
Sura 77
# wt
Fall 77
wt
Win 78
# wt
-------
Table 5b, (cont.)
Annelida (cont.)
Spionidae
Malacoceros
g|utaeus
Crustacea
Cirrtpedia
Balanus
glandula
8,
spp, (juv.)
Isopoda
Idotea
wosnesenskii
Amphlpoda
Gamma r idea
spp.
Coropblum
spp.
Decapoda
Hemigrapsus
spp.
Fagurus
spp.
Insecta
Dlpteran
larvae spp.
Species Richness
Diversity, H3-
Total Nunber
Total Biomass (g)
Spr 77
# wt
0
0
90.0
14.0
20.0
36.0
2.0
7.0
6.0 <2.0
10.0 <10.0
60.5
18.0
59
2.34
6,452
653
Sum 77
# wt
Fall 77
# wt
320.0 <2.0
1830,0 <2.0
620.0 28.0
18,930.0 516.0 50.0 <10.0
36.0 20.5
66.0 <12.0
1290.0^24.0
76
2.04
29,420
1,047
136.0 20.5
476.0 0.5
17,590.0 10.0
116.0 184.5 180.0 43.5
96.0 27,0 46.0 1.0
120.0 <10.0 2,136.0 <-12.0 2l,200<12.0
70
1.97
33,278
916
Win 78
# wt
60.0 <2.0
5430.0 616.0 5246.0 660.0
0
21.5 10.5
100.0 <12.0
2836.0 <12.0
157.0 110.0
286.0 <27.0
400.0 <12.0
64
2.50
13,514
1,088
-------
Table 5c. Morse Creek (cobble over sand) dominant benthic organism from Che low (+0$
1ntert1daI zone,, collected spring 1977 through fall 1977, expressed as number
and/or biomass (g) per m , For complete data set see Appendix I.
Spr 77 Sum 77 Win 78
# wt # wt # wt
Clorophyta
Ulva 13.5 117.5 10.0
«pp.
Phaeophyta
Alarla 702.5 503.0 1473.5
spp.
Hedophyllum 57.5 0 0
sessile
Rhodophyt*
Gigartina 39.5 142.5 118.0
papillata complex
Iridaea 2,397.5 1,975.0 468.0
cordata
Cnidarla
Anthozoa
Anthopleura 130.0 5.0 50.0 2.5 16.0 10.0
elegant is sitna
Nemertea 0 360.0 4.0 250.0 <12.0
spp.
Nematoda 490.0 <12.0 536.0 <12.0 114.0 <12.0
spp.
Mollusca
Gastropoda
Lacuna 126.0 0.5 1000.0 12.0 834.0 12.0
variegata
Rotoacnea 26.0 5.5 62.0 7.5 46.0 20.0
spp.
-------
Table 5c. (cont.)
Mollusca (conC.)
Gastropoda
Thais
spp.
Blvalvia
Mytilus
spp.
Protothaca
staminea
Tresus
capax
Annelida
Polychaeta
Arenicolidae
Abarenicola
spp.
Capitellidae
Capitella
capitata
Cirratulidae
Cirratulus
cirratus
Nereldae
spp.
Opheliidae
Armandia
brevis
Sabellidae
spp.
5pr 77 Sum 77 Fall 77
# wt # wt # wt
3.1 0.5
190.0 <10.0
6.0 <2.0
10.0 102.0
10.0 <10.0
0
30.0 12.0
0
0
6.0 <2.0
416.0 <12.0
0
1826.0 <12.0
766.0 <34.0
0
11416.0 <62.0
62.0 <2.0
1100.0 <12.0
1810.0 82.0
362.0 64.0
60.0 <12.0
7816.0 <70.0
166.0 <2.0
3556.0 <12.0
1380.0 <2.0
166.0 <28.0
214.0 <2.0
76.0 <30.0
-------
Spr 77
# wt
Annelida (cont.)
Polychaeta
Spionidae
Malacoceros 16.0 <2.0
glutaeus
Crustacea
Tanaidacea
Leptochelia 36.0 <12.0
dubia
Amphlpoda
Gatranaridea 916.0 12.0
spp.
Decapoda
Cancer 12.5 15.5
spp.
Pugettia 40.0 <12.0
gracilis
Species Richness 112
1
Diversity, H 2.14
Total Number 17,843
Total Biotftass (g) 3,936
5c. (cont.)
Sum 77 Fall 77
# wt # wt
736.0 <12.0
2286.0 <2.0
36.0
<12.0
14.0 <2.0
1176.0 <148.0
1120.0 10.0
46.0 35.0
436.0 10.0
6.0
56.0
4.0
2.0
117
2.55
17,450
3,176
82
2.29
10,360
2,216
-------
Species richness, total numbers, total bioraass, and populations of most
component species showed a spring/summer maximum. However, as in all rock
areas, the community components were very patchy. Despite this, a compari-
son of the first and second year data sets showed high similarity.
Table 5d presents the subtidal abridged data set for Morse Creek. The
-10 m community was dominated by detritus feeders: Macoma, capitellids,
maldanids, Armandia, spionids, Exogone, most gammarids, and ophiuroids.
Suspension feeders (Calyptraea, Crenella, Mysella, and Leptochelia) were
also abundant at -10 m.
Species richness and diversity were very high. However the total
biomass was fairly low. Year one/year two community similarity was high.
54
-------
Table 5d. Morse Creek (gravel) dominant benthic organisqis from the shallow (-10 m) subtidal
zone, collected spring 1977* expressed as number and/or biomass (g) per m^. For
complete data set see Appendix I.
Annelida
Ui
-10 m
wt
Phaeophyta
Desmarestia ligulata <2.0
Mollusca
Gastropoda
Araphissa columbiana 250.0 16.0
Calyptraea fastigiata 60*0 2.0
Blvalvia
Crenella decussata 6.0 <2.0
Macoma spp. 86.0 0.0
Mysella tuodda 396.0 2.0
Polychaeta
Capitellidae
Mediomastus sp. 510.0 *2.0
Maldanidae spp. 156.0 <2.0
Nereidae
Platynereis bicanaliculata 70.0 <2.0
Opheliidae
Armandia brevis 36.0 <2.0
Spionidae
Malacoceros glutaeus 0
Prionospio cirrifera 20.0 <2.0
P. steenstrupi 106.0 <2.0
Syllidae
Exogone spp. 416.0 <2.0
-------
Crustacea
TanaIdacea
Leptochelia dubia
Amphipoda
Gamroaridea spp.
Decapoda
Cancer productus
Pagurldae spp.
Echinodermata
Ophiuroidea spp.
Species Richness
1
Species Diversity, B
Total Number
Total Blomass (g)
Table 5d. (cont.)
-10 m
# wt
4806,0
2.0
2030.0
4.0
20.0
116.0
12.0
8.0
166.0
127
2.72
11,654
<290
2.0
-------
Beckett Point (protected sand-gravel, intertidal; sand, subtidal)
The sediment at Beckett Point was a sandy gravel at +61, a gravel-sand
mix at +3', a mdeium-fine sand with gravel at +0', fine sand at -5m, and
medium to fine sand at -10m. The study area was completely protected
inside Discovery Bay. The beach slope was fairly steep, and tidal action
probably was mainly responsible for the fairly coarse sediment at the study
site in the intertidal. The salinity showed no freshwater influence.
Major physical environment changes from year one to year two were not
observed.
Table 6a presents the summary data from +6' at Beckett Point. The
community had two components, planktivorous epifaunal Balanus, responsible
for most of the total biomass, and detritus feeding worms (nematodes,
syllids, and oligochaetes) and isopods. The isopods and oligochaetes both
showed peak numbers in fall quarter both years, perhaps correlated with fall
plant die back. Year one, year two community similarity was quite high.
An abridged data set for Beckett Point +3' is presented in Table 6b.
The community at this level was dominated by suspension feeding bivalves
(Mysella. Mytilus, Protothaca, Transennella, and Tresus), detritus feeders
(capitellids, oweniids, spionids, syllids, oligochaetes, isopods, amphipods,
Dendraster. and Leptosynapta), and a couple of carnivores (nemerteans and
Hemigodus). The bivalves were very patchy in distribution. Seasonal
pattern showed a fall peak for species richness, diversity, and density.
Year one/year two similarity was low, probably reflecting the patchy nature
of the community rather than community change.
Table 6c gives the abridged results for Beckett Point +0'. This rich
diverse community was dominated by suspension feeding bivalves
(Cllnocardium. Mysella, Protothaca, Transennella. and Tresus); deposit/
detritus feeding worms, crustaceans, and echinoderms; and a number of
carnivores (nemerteans, Nassarius, Hemigodus, Glycinde, hesionids,
nephtyids, phyllodocids, polynoids, Cancer, and Crangon). The majority of
species and the total number peaked strongly in the fall. Species richness
and diversity remained fairly constant through the year. Year one/year two
community similarity was high.
Abridged subtidal results for Beckett Point are presented in Table 6d.
The very rich, diverse communities at -5 m and -10 m at Beckett were very
similar. The communities were dominated by deposit feeders: nematodes,
Hacoma, Telllna, oweniids, spionids, tanaids, and amphipods. There were a
few suspension feeders (Mysella, chaetopterids) and carnivores (Nassarius,
hesionids, and phyllodocids). The Van Veen grab operates best in fine
sediments such as at Beckett. It is not surprising that Beckett subtidal
communities showed the greatest year one/year two similarity of all
subtidal communities sampled since grab-caused error would be at a minimum.
57
-------
Table 6a.
Beckett Point (sandy gravel) dominant benthic organisms from the high (+6)
intertidal zone, collected sprang 1977 through winter 1978, expressed as
number an/or biomass (g) per m . For complete data set see Appendix I.
Spr 77 Sum 77 Fall 77 Win 78
, # wt # wt # wt # wt
Nematoda 570.0 <2.0 172.0 <4.0 272.0 <4.0 420.0 <4.0
spp.
Annelida
Oligochaeta 580.0 <2.0 92.0 <4.0 832.0 <4.0 780.0 <4.0
spp.
Polychaeta
Nereidae 40.0 <2.0 12.0 44.0 <4.0 24.0 <8.0
spp.
Syllidae
Syllis 340.0 <2.0 140.0 <4.0 60.0 <4.0 140.0 <4.0
SPP-
Crustacea
Girripedia
Balanus 0 112.0 28.0 480.0 168.0 432.0 168.0
glandula
Isopods
Exosphaeroma 10.0 <2.0 0 2452.0 16.0 80.0 <4.0
media
Species Richness 10 13 15 12
Diversity, Hi 1.43 2.04 1.44 1.69
Total Number 1630 632 4,404 1904
Total Biomass <20 <68 <232 <212
-------
Table 6b» Beckett Point (gravel-sand) dominant benthic organisms from the mid(+3$ intertldal zone,
collected spring 1977 through winter 1978, expressed as number and/or biomass (g) per m^.
For complete data sec see Appendix I.
Nenertea
spp.
Mollusca
Gastropoda
Lacuna
variegata
Bivalvla
Mysella
tumlda
Mytilus
edulis
Protothaca
staminea
Transennella
tantilia
Tresus
sp.
Annelida
Oligochaeta
spp.
Polychaeta
Capitellidae
Notomastus
tenuis
Glyceridae
Hemipodus
borealis
Oweniidae
Owenia
fusiformis
Spr 77
# wt
Sum 77
# wt
Fall 77
# wt
Win 78
# wt
30.0
<2.0
100.0 «».0
212.0 <4.0
100.0
0
20.0 <2.0
110.0
90 <2.0
60.0 <2.0
0
20.0 <2.0
0
1370.0 s/
190.0 <2.0
172.0 0.0
492 4.0
172.0 4.0
52.0 <4.0
132.0 0.0
6.0 292.0
100.0 <4.0
340.0 <4.0
1680.0 y
280.0 >/
3092.0 12.0
492 4.0
9012.0 20.0
172.0 16.0
380.0 8.0
90.0 26.0
120.0 <4.0
132.0 <4.0
3152.0 J
8292.0 V
0
152.0 <4.0
460.0 <4.0
32.0 <4.0
20.0 <4.0
10.0 40.0
152.0 <4.0
312.0 <4.0
1212.0
-------
Table 6b,
Spr 77
# wt
Annelida
Polyehaeta
Spionidae
Pygospio 0
elegans
Spio 0
filicornis
Spiophanes 0
bombyx
Syllidae
Syllls 20.0 <2.0
spp.
Crustacea
Cirripedla
Balanus 60.0 16.0
glandula
Isopoda
Exosphaeroma 460.0 <2.0
spp,
Amphipoda
Gsmmaridea 40.0 <2.0
sp.
Echinodermata
Echlnoldea
Dendiaster 0
excentrlcus
Holothuroidea
Leptosyriapta 0
clarki
(cont.)
Sum 77
# wt
Fall 77
wt
Win 78
# wt
100.0 <4.0
52.0 <4.0
0
188.0 <4.0
572.0 <4.0
60.0 <4.0
0
0
0
40.0 <4.0 12.0 <4.0
4712.0 96.0 792.0 136.0 40.0 0.0
432.0 <8.0
1600.0 12.0
12.0 <4.0
1172.0 4.0 400.0 <4.0
32.0 <4.0
40.0 <4.0
660.0 <4.0
0
12.0 <4.0 80.0 4.0
-------
Table fib. (cont.)
Spr 77
Sum 77
Fall 77
Win 78
Species Richness
23.0
55.0
75.0
29.0
Diversity,
1.89
2.20
2.73
2.15
Total Nuriaer
3,450
11,068
48,228
3,964
Total Blomass
4.54
<836
<532
<184
-------
Table 6c.
Beckett Point (medium-fine sand) dominant benthic organisms from the
zone, collected spring 1977 through winter 1978, expressed as nunber
per m^. For complete data set see Appendix I.
low (+0) intertidal
and/or biotnass (g)
Spr 77 Sum 77 Fall 77 Win 78
# wt # vt # wt # wt
Hemertea 530.0 <2.0 860.0 <~ 352.0 <4.0 480.0 <4.0
spp.
Nematoda 900.0 <2.0 292.0 <4.0 8832.0 <4.0 1652.0 <4.0
spp.
Mollusca
Gastropoda
Lacuna 80.0 <2.0 652.0 4.0 152.0 <4.0 32.0 <4.0
variegata
Nassarius 30.0 18.0 20.0 12.0 20.0 <4.0 0
tnendicus
Bivalvia
Clinocardium 180.0 18.0 12.0 4.0 580.0 8.0 972.0 8.0
nuttallii
Macoma 60.0 < 2.0 36.0 48.0 64.0 4.0 184.0 12.0
spp.
Mysella 4250.0 18.0 9772.0 16.0 19500.0 32.0 8812.0 16.0
tumida
Protothaca 180.0 2.0 32.0 12.0 292.0 36.0 280.0 4.0
staininea
Transennella 490.0 2.0 1120.0 4.0 232.0 4.0 480.0 <4.0
tantilla
Tresus 0 572.0 64.0 172.0 32.0 72.0 44.0
capax
Annelida
Oligochaeta 220 42.0 52.0 <4.0 1240.0 <4.0 140.0 <4.0
spp.
Polychaeta
Capitellidae 680.0 <8.0 824.0 <8.0 1712.0 <16.0 948 <16>0
spp.
-------
Annelida
Folychaeta
Glyceridae
Hemipodus
borealis
Goniadidae
Glycinde
picta
Hesionidae
spp.
Nephtyidae
spp.
Nereidae
Platynereii
bicanalieulata
Opheliidae
Armandia
brevis
Oweniidae
Owenia
fusiformis
Phyllodocidae
spp.
Polynoidae
spp.
Spionidae
spp.
Table 6c. (cont.)
Spr 77 Sum 77 Fall 77 Win 78
# wt # wt # wt # wt
940.0 <2.0
180.0 <2.0
186.0 <4.0
0
250.0 <2.0
20.0 <2.0
310.0 <2.0
290.0 <10.0
50.0 <6.0
645.0 <10.0
692.0 J
40 <4.0
232.0 <8.0
0
300.0 y
60.0 <0.0
1032.0 >/
484.0 <16.0
52.0 <8.0
15236.0<8.0
480.0 <8.0
0
292.0
24.0 4.0
1692.0 %/
1720.0 <4.0
1392.0 <4.0
1160.0 <20.0
64.0 8.0
2232.0 <28.0
480.0 <4.0
200.0 <4.0
0
20.0 <4.0
1512.0 <4.0
740.0 <4.0
872.0 <4.0
396.0 <16.0
64.0 <12.0
644.0
-------
Tab le 6c. ( cont.)
Spr 77 Sum 77 Fall 77 Win 78
# wt # wt # wt # wt
Annelida (cont.)
Polychaeta
Syllidae
Exogone 720.0 <4.0 8920 <4.0 3104.0 <8.0 1600.0 <8.0
spp.
Crustacea
Cumacea
Cutnella 510.0 <2.0 52.0 <4.0 160.0 <4.0 32.0 <4.0
vulgaris
Tanaidacea
Leptochelia 9670.0 2.0 1798.0 8.0 44632.0 24.0 20372.0 8.0
dubia
Atnphipoda
Ganraaridea 1410.0 <2.0 3532.0 32.0 880.0 <4.0 820.0 4.0
spp.
Decapoda
Cancer 0 112.0 <8.0 12.0 <4.0 0
spp.
Crangon 0 12.0 <4.0 0 0
nigricauda
Pagurus 0 52.0 <4.0 0 0
spp.
Echinodermata
Echinoidea
Dendraster 94.0 2.0 912.0 76.0 992.0 16.0 480.0 12.0
excentricus
Holothuroidea
Leptosymapta 110.0 2.0 112.0 4.0 140.0 <4.0 80.0
clarki
-------
S£T 77
Species Richness 68
Diversity, H* 2.52
Total Number 24,830
Total Biotnass 4.1,226
&
Ln
Table 6c. (cont.)
Sum 77
Fall 77
Win 78
83 93 79
2.51 2.13 2.30
58,584 97,112 45,784
2,780 < 3,080 <2,784
-------
Table 6d. Beckett Point (medium-fine sand) dominant benthie organisms from the shallow (-5 m,
-10 bi) subtidal zone, collected spring 1977, expressed as number and/or biomass (g)
per m2. For complete data set see Appendix I.
On
-5 m -10 m
£_ wt # wt
Nemertea spp. 260.0 < 2.0 400.0 <2.0
Nematoda spp. 160.0 < 2.0 360.0 <2.0
Mollusca
Gastropoda
Alvlnia sp. 316.0 <2.0 340.0 6.0
Lacuna variegata 190.0 < 2.0 66.0 <2.0
Mitre11a tuberosa 50.0 2.0 326.0 14.0
Nassarius mendicus 0 20.0 0.0
Bivalvia
Macoma spp. 200.0 4.0 206.0 2.0
Mysella tumida 1290.0 2.0 3076.0 4.0
Tellina sp. 376.0 10.0 146.0 12.0
Annelida
Polychaeta
Chaetopteridae
Mesochaetopterus 36.0 <2.0 280.0 < 2.0
taylori
Phyllochaetopterus 0 826.0 <2.0
prolifica
Spiochaetopterus 110.0 <2.0 160.0 <2.0
costarum
Hesionidae
Hicropodarke dubia 566.0 <2.0 950.0 <2.0
Nereidae
Platynereis 2096.0 >/ 1360.0 <2.0
bicanaliculata
Owentidae
Owenia fusiforrais 130.0 <2.0 6.0 <2.0
-------
Table
Annelida (cont.)
Polychaeta
Phyllodocidae
Eulalla sanguines
Phyllodoce spp.
Spionidae
Polydora socialis
Prionospio steenstrupi
Spiophanes berkeleyorum
Crustacea
TanaIdacea
Leptochelia dubia
Antphlpoda
Gammaridea spp.
Decapoda
Pagurldae spp.
Species Richness
Diversity,
Total Number
Total Biomass
6d. (cont.)
-5 m -10 m
# wt # wt
156.0 <2.0 250.0 <2.0
122.0 <2.0 912.0 <2.0
76.0 <2.0 1080.0 <2.0
1630.0 <2.0 730.0 <2.0
10.0 <2.0 0
1260.0 <2.0 316.0 <2.0
2976.0 2.0 3026.0 6.0
36.0 <2.0 280.0 <2.0
83 90
2.77 3.05
13,416 17,542
<170 <214
-------
Dungeness Spit (sand-gravel)
The sediment at Dungeness Spit was sandy gravel at +6', gravel at +31 ,
fine sand with gravel at +0', medium to fine sand with gravel at -5 m and
medium sand with gravel at -10 m. The intertidal had a moderate slope and
was extremely exposed to severe wave action. It was the only area east of
Port Angeles which proved difficult to sample because of surf conditions.
No noteworthy differences in the physical environment were observed between
the first and second year.
Abridged results for Dungeness Spit +6', +31, and +0' are given in
Tables 7a-c respectively. All three levels had a very species-poor commu-
nity. In fact at +6' fall quarter absolutely no organisms were found. The
only organisms found with any regularity at the three levels were deposit/
detritus feeding oligochaetes and amphipods. No consistent seasonal
pattern in populations appeared. The community similarity between the first
and second years was high at +3, and low at +6T and 0', because of fewer
oligochaetes and more amphipods found the second year. As expected with
such a sparse fauna, patchiness was extreme (see Appendix I).
Subtidal summary results for Dungeness Spit are presented on Table 7d.
Fauna was still extremely sparse at -5m. The community at -10 m, although
of low biomass, was fairly rich in both species and total number. The
community was composed of small bivalves and deposit feeding polychaetes
(capitellids, dorvilleids, spionid, and syllids, and gammarids). Year one
and year two subtidal community similarity was low at both levels. This
probably reflects sampling inadequacy rather than community change.
68
-------
Table 7a. Dungeness Spit (sandy gravel) dominant benthic organisms from the high (+6') intertidal
zone collected spring 1977 (through winter 1978)» expressed as number and/or biomass (g)
per For complete data set see Appendix I,
Spr 77 Sum 77 Fall 77 Win 78
# wt # wt # wt # wt
Annelida
Oligochaeta
spp.
4.0 <2.0
0
0
4.0
Crustacea
Amphipoda
Gammaridea
SPP»
4.0 <2.0
332.0 <2,0
0
0
Species richness
2
1
0
1
Diversity - H»
0.69
0.00
0
0.00
Total number
8
332
0
4
Total biomaas (g)
<2
< 2
0
<2
-------
Table 7b. Dungeness Spit (sandy gravel) dominant benthic organisms from the mid (+3*) intertidal
zone collected spring 1977 (through winter 1978), expressed as number and/or btomass (g)
per m^. For complete data set see Appendix I,
Spr 77 Sum 77 Fall 77 Win 78
# wt # wt # wt # wt
Annelida
Oligochaeta
4.0 <2.0
0
0
0
spp-
Crustacea
Amphipoda
Gammaridea
272.0 0.0
4.0 <2.0
16.0 <2,0
572.0
spp.
Species richness
2
2
1
1
Diversity - H*
0.08
0.56
0.00
0.00
Total number
276
16
16
572
Total biomass (g)
< 2
< 4
< 2
< 2
-------
Table 7c. Dungeness Spit (sandy gravel) dominant benthic organisms from the low (+0*) intertidal
zone collected spring 1977 (through winter 1978), expressed as number and/or biomass (g)
per m . For complete data set see Appendix I.
Spr 77 Sum 77 Fall 77 Win 78
# wt # wt # wt # wt
Annelida
Oligochaeta
spp.
8.0 < 2.0
0
0
0
Crustacea
Amphlpoda
Garamaridea
spp.
40.0 < 2.0
140.0 <2.0
492.0 <2.0
68,0
Species richness
3
2
5
1
Diversity - H*
0.69
0.13
0.27
0.00
Total number
52
144
520
68
Total biomass (g)
< 6
< 4
< 10
< 2
-------
Table 7d. Dungeness Spit (medium-fine sand with gravel) dominant benthic organisms from the shallow
subtidal zone collected spring 1977, expressed as number and/or biomass (g) per For
complete data set see Appendix I.
-5 m
wt
-10 m
# wt
fs3
Nematoda spp.
Mollusca
Bivalvia
Crenella decu-sftata
Ma coma spp.
Mysella tumida
Psephidia lordi
Annelida
Capitellidae
Mediomastus sp.
Dorvilleidae
Protodorvillea gracilis
Hesionidae
Micropodarke dubia
Spionidae
Prionospio steenstrupi
Spiophanes bombyx
Syllidae
Exogone spp.
Crustacea
Tanaidacea
Leptochelia dubia
30.0 <2.0
0
30.0 <2.0
AO.0 <2.0
70.0
0
0
<2.0
6.0
<2.0
876.0
66.0
52.0
70.0
10.0
1060.0
456.0
220.0
146.0
6.0
508.0
240.0
<2.0
<2.0
<2.0
<2.0
<2.0
0.0
<2.0
<2.0
<2.0
<2.0
<2.0
<2.0
-------
Table 7d. (Cent.)
Amphipoda
Gamut-idea spp.
Species richness
Diversity - H*
local number
Total biomasa (g)
*¦-4
U3
-5 m
# vt
-10 D
# wt
380.0 <2.0 560.0 <2.0
28 136
2.43 3.82
858 6764
<56 <322
-------
Twin Rivers (.sand-gravel)
The sediment at Twin Rivers was sandy gravel at +6', gravel at +3',
gravel with fine sand at +Ol, gravel at -5 m. The beach had a fairly steep
slope and was very exposed to both waves and ocean swells. Wave conditions
prevented sampling at -10 m during the second year.
Tables 8a-c present abridged results for Twin Rivers +6', +3', and +0'
respectively for the single quarter sampled. At all levels species richness,
diversity, and biomass were low. The communities, such as they are, were
primarily composed of deposit feeding worms and gammarid amphipods. As
expected with a sparse fauna, it was extremely patchy spatially. Year one,
year two similarity was high at +6' and low at +3' and +0'.
Twin Rivers -5 m subtidal abridged results are given in Table 8d.
Weather conditions prevented -10 m sampling. The community was dominated
by deposit feeding annelids, and crustaceans. As with most of the Van Veen
samples, variance among replicates was very high and might not reflect
organism patchiness. In addition there was very little similarity between
first and second year samples. First year species richness was 139 compared
to 28 the second year. This is most likely a reflection of poor sampling,
not year to year change.
74
-------
fable 8a. Twin Rivers (sand, gravel) dominant benthic organisms from Che high (46') intertidal zone
collected spring 1977, expressed as number and/or biomasa (g) per For complete data
set see Appendix I.
"•si
U1
ipr 77
~ wt
Annelida
Oligochaeta 32.0 <2.0
»PP.
Crustacea
Amphipoda
Gammaridea 8.0 <2.0
* '
Species richness 3
Diversity - H' 0.76
Total number 44
Total biomass (g) ^6
-------
Table 8b* Twin Rivers (sand, gravel) dominant benthic organisms from Che mid (+3*) intertidaL zone
collected spring 1977, expressed as number and/or biomass (g) per m*. For complete data
set see Appendix I.
Spr 77
# wt
CT>
Annelida
Oligochaeta 28.0 <2,0
spp.
Crustacea
Isopoda
Gnorimosphaeroma 32.0 <2.0
oregonense
Amphipoda
Gammaridea 576.0 <2.0
spp.
Species richness 4
Diversity - H* 0.63
Total number 692
Total biomass (g)
<8
-------
fable 8c. Twin Rivera (sand, graveI) dominant benthie organisms from the lov f+01) intertidal zone
collected spring 1977, expressed as mnaber and/or biomass (g) par » . For complete data
set aee Appendix I.
Spr 77
# vt
-%J
-"J
Annelida
Polychaeta
Capitellidae
Capitella 168.0 <2.0
capitata
Spionidae
Malacocerus 660.0 <2.0
glutaeua
Crustacea
Amphipoda
Gamaridea 480.0 <2.0
•pp.
Species richness IS
Diversity - H' 1.62
Total rnuaber 1448
Total bioauisa (g)
<30
-------
Tabic 8d. Twin Elvers (gravel) dominant benthic organisms from the shallow subtidal zone collected
spring 1977, expressed as number and/or biomass (g) per m2. For complete data set see
Appendix 1.
-5m
vt
Mollusca
Bivalvia
Hysella tumida 6.0 <2.0
Frotothaca statninea 10.0 6.0
Annelida
Oligochaeta spp. 6.0 <2.0
Polychaeta
oo Capitellidae
Mediomastus sp. 410.0 <2.0
Cirratulidae
Tharyx multifilis 5190.0 /
Goniadidae
Clycinde picta 70.0 <2.0
Nereidae
Platynereis blcanaliculata 16.0 <2.0
Oweniidae
Ovenia fusilorols 670.0 <2.0
Spionidae
Prionospio steenstrupl 150,0 <2.0
Crustacea
Cumacea
Oiastvloptl* 1166.0 2.G
-------
Table 8d. (Cont.)
Isopoda
Gnorimosphaeroma oregonense
Amphipoda
Gamnaridea spp.
S Species richness
Diversity - H'
Total number
Total bioma88 (g)
-5 m
# wt
230.0
1486.0
2.0
2.0
28
1.69
8852
<56
-------
North Beach. Sand Csand)
The sediment at +6' was sand with gravel, at +2' medium to fine sand
with gravel, at +0' medium to very fine sand, at -5 m medium to coarse sand,
and at -10 m sand and gravel. The mid-tide height of +2r was selected
instead of +3' to stay out of the more gravelly upper intertidal. North
Beach Sand had a moderately sloped beach and moderate exposure, as North
Beach Cobble. No major changes in the physical environment were observed
between the first and second year.
Abridged results for North Beach Sand +6' are given in Table 9a. The
community was very low in species richness, diversity, total number of
organisms, and total biomass. It was composed of deposit feeding worms and
crustaceans. Species richness and total number appeared particularly
depressed in the winter. The sparse fauna was of course very patchy in
spatial distribution. First and second year community similarity was low.
Table 9b gives a summary data set for North Beach Sand +2'. Major
components of this community were all detrital/deposit feeders (Paraonella.
Eohaustorius. and Paraphoxus). The community was dominated by the fossorial
amphipod Eohaustorius. Its populations peaked in the summer. Species
richness, diversity, and total biomass were low and exhibited little season-
al pattern. The community was extremely similar from year one to year two.
Table 9c presents the abridged results for North Beach Sand +0r. This
low diversity, low biomass community was comprised almost totally of deposit/
detrital feeding polychaetes and crustaceans plus carnivorous nemerteans and
was dominated by three species: the paraonid polychaete Paraonella. the
mysid Archaeomysis, and the amphipod Eohaustorius. Paraonella and
Eohaustorius populations peaked in the spring. Archaeomysis showed a severe
population decline in the fall/winter. This may merely indicate migration
of the species out of the littoral zone during this period. As at +2' this
community showed great similarity from year one to year two.
Abridged subtidal data are given in Table 9d for North Beach. Except
for fewer plants at -10 m, the communities at -5 m and -10 m were quite
similar. They were composed of herbivores (Lacuna and Onuphls). suspension-
feeding bivalves (Clinocardium. Crenella, Mysella, Psephldia), many deposit
feeding annelids and small crustaceans, and carnivores (Natica, Micropod-
arke, and Cancer at -5 m). Species richness, diversity, total number, and
biomass were high at both levels. Year to year similarity was low, again
likely due to poor sampling methodology.
80
-------
Table 9a. North Beach Sand (medium-fine sand with gravel) dominant benthic organisms from the
high (46') intertldal zone collected spring 1977 (through winter 1978), expressed as
number and/or biomass (g) per m^. For complete data set see Appendix I.
Spr 77 Sum 77 Fall 77 Win 78
# Vt # lit # wt # wt
Annelida
Oligochaeta 24.0 <2.0 0 28.0 <2.0 8.0 <2.0
•PP.
Crustacea
Anphipoda
Gammaridea 4.0 <2.0 0 0 8.0 <2.0
spp.
Species richness
10
8
6
5
Diversity - H*
2.02
1.48
1.35
1.55
Total number
92
156
48
28
Total biomass (g)
<20
<18
<12
<10
-------
Table 9b. North Beach Sand (medium-fine sand with gravel) dominant benthic organisms from the mid
(+2') intertidal zone collected spring 1977 (through winter 1978), expressed as number
and/or biomass (g) per For complete data set see Appendix I.
Annelida
Polychaeta
Paraonidae
Paraonella
platybranchia
Spionidae
Pygospio
elegans
Crustacea
Isopoda
Exosphaeroma
media
Amphipoda
Eohaustorius
spp.
Paraphoxus
spp.
Species richness
Diversity - H*
Total number
Total biomass (g)
Spr 77 Sum 77 Fall 77 Win 78
# wt # vt •# wt # wt
156.0 <2.0
132.0 <2.0
0
12.0 <2.0
36.0 <2.0
0
0 0 0 2.6 <2.0
2408.0 6.0 6072.0 8.0 1524.0 4.0 1988.0 2.0
0 8.0 <2.0 156.0 <2.0 200.0 <2.0
10 15 10 ¦ 10
0.33 0.26 0.70 0.66
2600 6372 1824 2352
<24 <32 <28 <22
-------
fable 9c. North Beach Send (medium-fine sand with gravel) dominant benthic organisms from the
low (+0*) intertidal zone collected spring 1977 (through winter 1978), expressed as
number and/or biomass (g) per ra2. For complete data set see Appendix 1,
Spr 77 Sum 77 Fall 77 Win 78
# wt # wt # vt # wt
Hemertea spp. 16.0 <2.0 20.0 <2.0 0 12.0 <2.0
Annelida
Polychaeta
Arenicolidae
Abarenicola 64.0 <2.0 4.0 <2.0 24.0 <2.0 4.0 <2.0
claparedi
oceanica
Orbiniidae
Scoloplos sp. 112.0 y 132.0 ~ 32.0 <2.0 112.0 *2.0
Faraonidae
Paraonella 2168.0 S 1984.0 J 480.0 <2.0 1448.0 <2.0
platybranchia
Spionidae spp. 92.0 <2.0 56.0 <2.0 124.0 <2.0 456.0 <2.0
Syllidae
Syllis spp. 116.0 <2.0 124.0 <2.0 0 196.0 <2.0
Crustacea
Hyaidacea
Archaeomysis 16.0 <2.0 1188.0 4.0 4.0 <2.0 0
grebnitzkii
Amphipoda
Kohaustorius 2940.0 6.0 992.0 0.0 2792.0 0.0 2796.0 2.0
vashingtonianus
-------
Tabic 9c. (Cont.)
Spr 77
# vt
Amphipoda (cont.)
Paraphoxus 32.0 <2.
•pp.
Species richness 16
Diversity - H' I.11
Total number 5608
Total biomass (g) <28
Sum 77
# wt
Fall 77
# wt
Win 78
# wt
16.0 <2.0 4.0 <2.0 0
21 18 19
1.56 1.02 1.36
3692 3792 5296
<34 <36 <38
-------
fable 9d. North Beach (coarse aand-gravel) dominant benthic organisna from the ahallov aubtidal
cone collected spring 1977, expressed as number and/or blomass (g) per m2. For
complete data set see Appendix 1.
00
Ul
-5 at -10 a
~ wt # vt
Phaeophyta
Laminar!a saccharine 1655.0 0
Pterygophora californica 9163.0 0
Shodophyta
Odonthalia vashingtoniensls 0 0
Opuntiella californica 700.0 0
Nematode spp. 330,0 <1.0 260.0 <2.0
Kollusca
Gaatropoda
Lacuna variegata 940.0 5.0 40.0 <2.0
Margaritas pupillus 120.0 <1.0 0
Natica clause 5.0 <1.0 6.0 26.0
Bivalvia
Clinocardlun nuttallii 1375.0 1.0 30.0 0.0
Crenella decusaata 15.0 *1.0 646.0 2.0
Macoma ap. 0 10.0 <2.0
Hysella tumlda 100.0 <1,0 0
Psephidia lord! 105.0 <1.0 46.0 0.0
Annelida
Oligochaeta app. 175.0 <1.0 1250.0 <2.0
-------
Table 94. (Cont.)
Polychaeta
Capitellidaa
Mediomaatui ap.
Oorvilleidac
Protodorvlllea gracilla
Hesionidae
Mlcropodarke dttbla
Omiphiidae
S Onuphls spp.
Sabellidae «pp.
Splonldae
Polydora hamata
P. pygldlalla
Prlonosplo atcenatrupl
Splo fllicornla
Syllidae
Exogone spp.
Archiannelida
PolygordiLdae
Polygordius ap.
Cruatacca
Tanaldacea
Leptochelia dubia
-5 a
#
-10 m
# vt
35.0
290.0
540.0
285.0
25.0
420.0
275.0
130.0
90.0
395.0
520.0
175.0
<1.0
<1.0
<1.0
<1.0
<1.0
<1.0
<1.0
<1.0
<1.0
<1.0
<1.0
<1.0
6.0
30.0
416.0
10.0
0
0
0
30.0
270.0
166.0
560.0
176.0
<2.0
<2.0
<2.0
<2.0
<2.0
<2.0
<2.0
<2.0
<2.0
-------
Table 9d. (Cont.)
Isopoda
Exosphaerotna spp.
Aophipoda
Garamaridea spp.
Dccapoda
Cancer spp.
Paguridae spp.
Species richness
Diversity - H'
Total number
Total biomass (g)
-5 m
# wt
-10 in
# vt
655.0 0.0 136.0 <2.0
4275.0 6.0 630.0 2.0
30.0 9.0 0
130.0 5.0 36.0 <2.0
201 151
1.44 3.86
14103 7442
12021 1474
-------
Kydaka Beach (sand)
The sediment at Kydaka Beach at +6% +3', and +0' was uniformly very
coarse to fine sand, at -5 m medium to fine sand, and at ^10 m fine sand.
The beach slope was moderately steep, and the area was exposed to extremely
violent wave and ocean swell action. Salinity during winter quarter
sampling was quite low (20.1 /oo). No major difference was observed in the
physical environment between the first and second year.
Tables lOa-c present abridged results for Kydaka Beach +6', +3', and
+0' respectively. Th$ very low diversity, low total number, low biomass
communities were fairly similar at all three levels. Deposit/detrital
feeders dominate (oligochaetes, gammarlds, Archaeomysis). A carnivore
(Nephtys) occurred at +0'. No clear seasonal patterns emerged, .probably
because of the difficulty of accurately sampling such a sparse fauna.
Winter did appear to be a depressed time for total number. Similarity in
communities between year one and two was low at all three levels, doubtless
due to the problems in sampling such a sparse fauna.
The summary subtidal results for Kydaka Beach appear in Table lOd.
Major constituents were dfeposit feeding bivalves (Tellina), small suspension
feeding bivalves (Mysella), deposit feeding polychaetes (capitellids and
spionids) and small crustaceans (ostracods, cumaceans, and gammarid
amphipods). Year one to year two community similarity was high given the
limitations of grab sampling noted earlier.
88
-------
fable 10a, Kydaka Beach (coarse-fine sand) dominant benthic organisms from the high (+6*) intertidal
zone collected spring 1977 (through winter 1978), expressed as number and/or biomass (g)
per m . For complete data set see Appendix I.
Spr 77 Sua 77 Fall 77 Win 78
# Vt # lit # wt # ttt
Annelida
Oligochaeta
•pp.
8.0 <2.0
68.0 <2.0
12.0 <2.0
8.0
Crustacea
Anphipoda
Gammaridea
spp.
0
44.0 <4.0
76.0 <2.0
0
Species richness
2
6
4
2
Diversity - H*
0.64
1.23
0.71
0.64
Total number
12
144
96
12
Total biomasa (g)
<4
*12
*8
<4
-------
Table 10b. Kydaka Beach (coarse-fine sand) dominant benthic organisms from the mid (+3') intertidal
zone collected spring 1977 (through winter 1978), expressed as number and/or biomass (g)
per m^. For complete data set see Appendix I.
Spr 77
# wt
Sum 77
# wt
Fall 77
# wt
Win 78
wt
Annelida
Oligochaeta
¦PP.
Crustacea
Mysidacea
Archaeomysis 4.0 <£2.0 212.0 <2.0 40.0 <2.0 0
grebnitzkii
Species richness
2
5
2
1
Diversity - H*
0.6°
0.87
0.45
0.00
Total number
8
328
48
12
Total biomass (g)
<4
*10
<4
<2
-------
Table 10 c. Kydaka Beach (coarse-fine sand) dominant benthic organisms from the low (-<-0') intertidal
zone collected spring 1977 (through winter 1978), expressed as number and/or biomass (g)
per m . For complete data set see Appendix I.
VO
Spr 77 Sum 77 Fall 77 Win 78
# wt # wt # wt # wt
Annelida
Polychaeta
Nephtyidae
Nephtys 0 0 8.0 <2.0 0
spp.
Paraonidae
Paraonella 0 24.0 <2.0 8.0 <2.0 0
platybranchia
Crustacea
Ifysidacea
Archaeomysis 4.0 <2.0 16.0 <2.0 0 0
grebnitzkii
Anphipoda
Gamnaridea spp. 268.0 <8.0 64.0 <4.0 0 0
Species richness 6 5 7 1
Diversity - H* 0.61 1.25 1.77 0.00
total number 272 112 56 8
Total biomass (g> <12 <10 <14 <2
-------
Table lOd. Kydaka Beach (medium-fine sand) dominant benthic organisms from the shallow subtidal
tone collected spring 1977, expressed as number and/or biomass (g) per m?. For
complete data set see Appendix 1.
-5 m -10 m
# vt 4 vt
Mollusca
Bivalvia
Mysella tumida 610.0 2.0 1656.0 4.0
Psephidia lordi 0 1316.0 8.0
Tel Una sp. 26.0 2.0 56.0 2.0
Annelida
vo Polychaeta
Capitellidae
Mediomastus sp. 256.0 <2.0 1670.0 0.0
Orbiniidae
Scolop1os spp. 110.0 <2.0 50.0 0.0
Oweniidae
Owenia fusiformis 150.0 <2.0 220.0 0.0
Spionidae
Polydora socialis 60.0 <2.0 116.0 0.0
Prionospio steenstrupi 740.0 <2.0 3850.0 0.0
Crustacea
Ostracoda spp. 196.0 2.0 1560.0 2.0
Cumacea
Diastylis sp. 0 46.0 2.0
Diastylopsis sp. 476.0 2.0 6.0 0.0
-------
Table lOd. (Cont.)
Isopoda
Edotea subllttorallt
Aophipoda
Gammaridea spp.
Echinodermata
Holothuroidea
Leptosynapta clarki
Species richness
Diversity - H1
Total number
Total bionass (g)
-5 m
# »t
-10 m
# vt
20.0 2.0
2076.0 4.0
336.0 6.0
54
2.55
6230
<114
0
2576.0 6.0
0
96
2.83
17,410
<222
-------
Jamestown (protected sand)
The Jamestown sediment was sandy gravel at +6*, fine sand at +1.4',
medium sand at +01*, coarse to medium sand at -5 m, and coarse to fine sand
at -10 m. The +1.41 level was selected to avoid the upper intertidal
gravel. The beach was fairly well protected by Dungeness Spit. Its slope
was very gradual and was the widest beach sampled. No major changes in the
physical environment were noted between the first and second year.
Table 11a presents the summary results of Jamestown +6'. This low
diversity community w^s composed predominantly of deposit feeders (nematodes,
oligochaetes and gammarid amphipods). Lowest species richness occurred in
the summer- fall, highest total number in winter. The patchiness of the
fauna was very great (Appendix I). However, because of the overwhelming
numerical dominance of oligochaetes, year one and year two similarity was
total.
Abridged results for Jamestown +1.4' are presented in Table lib. The
major components of this community were a small suspension-feeding bivalve
(Transennella), a deposit feeding bivalve (Macoma), other deposit feeders
(nematodes, arenicolids, capitellids, paraonids, spionids, oligochaetes,
gammarids, and Leptosynapta), and carnivores (Nephtys, Eteone). No clear
seasonal patterns appeared in the results* although spring appears to have
lowest species richness and number. First and second year community
similarity was high, but densities were greater in the second year.
Table 11c presents an abridged data set for Jamestown +0*. This high
density, high biomass community was quite complex. Tube-building poly-
chaetes and crustaceans and Upogebia provided major structuring elements
and dominated the community. Most of the worms, Macoma, small crustaceans,
and Leptosynapta were deposit/detritus feeders. Carnivores included
nemerteans, hesionids, phyllodocids, and polynoids. A small, suspension-
feeding bivalve (Transennella) was also abundant. No clear community
seasonal changes appeared in the results. However, many polychaetes
(e.g. Platynereis) showed peak numbers in fall/winter, evidence of nectult-
ment. Community similarity between year one and two was Very high.
Subtidal results for Jamestown are presented in abridged form in
Table lid. The communities of these two levels, -5 m, and -10 m, were
quite similar. Both were very species rich, diverse, and had a high total
number. Over ten species of suspension feeding bivalves and one deposit
feeder were present. Herbivores included Margarites/Lirularia. Platynereis,
Onuphis, and Pugettia. Among carnivores were nemerteans, Nassarius.
Natica. hesionids, and polynoids. In addition there were many deposit/,
detritus feeding polychaetes and small crustaceans. Replicate variance was
fairly low at this area and year one to,year two similarity rather high
for subtidal grab-sampled communities.
94
-------
Table lis.
Jamestown (sandy gravel) dominant benthlc organisms from the high (+6') intertidal
zone collected spring 1977 (through winter 1978), expressed as number and/or biomass
(g) per m2. For complete data set see Appendix I.
Nematode
spp.
Spr 77
# wt
60.6 <2.0
Svun 77
# wt
Fall 77
# wt
Win 78
# wt
72.0 <4.0
60.0 <4.0
20.0 <4.0
Annelida
Oligochaeta 148,830.0 <2.0 44,332.0 <4.0 30,640.0 <4.0 1,612.0 <4.0
spp.
Crustacea
Amphipoda
Caanaridea
spp.
650.0 <2.0
140.0 <4.0
80.0 <4.0
Species richness
Diversity - H*
Total number
Total biomass (g)
8
0.07
150,220
<14
4
0.02
44,428
<16
4
0.07
30,972
<16
14
0.73
1876
<52
-------
Table lib. James town (fine sand) dominant benthic organisms from the mid (+1.4') intertidal
zone collected spring 1977 (through winter 1978), expressed as number and/or biomass
(g) per m^. For complete data set see Appendix I.
Spr 77
# wt
Nematode
spp.
Mollusc*
Bivalvia
Macoma
nasuta
Transene11a
tantilla
Annelida
Folychaeta
Arenicolidae
sp.
Capitellidae
Capitalla
capitata
Paraonidae
Paraonella
platybranchia
Phyllodocidae
£teone
longa
Fhyllodoce
maculate
60.0 156.0
1270.0 8.0
Sum 77
# wt
Fall 77
# wt
20.0 148.0
760.0 <4.0
380.0 <-2.0
20.0 <2.0
17,520.0 <4.0
12.0 <4.0
Win 78
# wt
40.0 <"2.0 412.0 <4.0 252.0 <4.0 472.0 <4.0
72.0 340.0 12.0 80.0
7232.0 32.0 5280.0 <4.0
50.0 <2.0 700.0 <4.0 7052.0 <4.0 2132.0 <4.0
870.0 <2.0 3632.0 <4.0 3012.0 4.0 1140.0 <4.0
60.0 <2.0 332.0 <4.0 112.0 <4.0 52,0 <4.0
712.0 <4.0 1352.0 <4.0
320.0 < 4.0 340.0 <4.0
-------
TabI* lib. (Cont.)
Splonldae
Malacoceros
glutaeus
Polydora kempl
japonica
Pygospio
elegans
Crustacea
Tanaidacea
Leptochelia
dubla
Amphlpoda
Garaoaridea
spp.
Echinodermata
Holothuroldea
Leptosynapta
clarki
Species richness
Diversity - H*
Total nuaber
Total blonass (g)
Spr 77
# wt
5750.0 <2.0
80.0 <2.0
820.0 <2.0
10.0 <2.0
Sum 77
# wt
820.0 <4.0
1880.0 <4.0
51,512.0 <4.0
40.0 <4.0
906,0 < 2.0 1 048.0 <20.0
10.0 <2.0
33
I.87
II,536
<224
38
1.19
68,244
<340
Fall 77
# vt
Win 78
# vt
11,892.0 y
2940.0 y
2740.0 <4.0
600.0 <4.0
1868.0 <16.0
12.0 <4.0
48
2.02
61,368
<704
13,040.0 <4.0
272.0 <4.0
1980.0 <4.0
1252.0 <4.0
24.0 <8.0
172.0 <4.0
38
1.90
28,632
<220
-------
Table 11c. Jamestown (medium sand) dominant benthic organisms from the low (+0') lntertidal zone
collected spring 1977 (through winter 1978), expressed as number and/or biomass (g)
per m^. For complete data set see Appendix 1.
Spr 77 Sum 77 Fall 77 Win 78
# vt # wt # wt # wt
Nemertea 50.0 <2.0 32.0 <4.0 120.0 <4.0 10010 <4.0
spp.
Nematode 0 12.0 <4.0 60.0 <4.0 340.0 <4.0
spp.
Mollusca
Bivalvia
Macoma 320.0 <2.0 32.0 24.0 20.0 4.0 12.0 4.0
spp.
Transennella 780.0 4.0 540.0 <4.0 2560.0 12.0 1572.0 *4.0
tantilla
Annelida
Oligochaeta 11,690.0 <2.0 9532.0 <4.0 19,820.0 <4.0 27,072.0
-------
Tabic 11c. (Cont.)
Spr 77
# vt
Dorvilleldae
Dorvlllca 230.0 <2.0
rudolphi
Heslonidae
Ophiodrooua 60.0 <2.0
pugettensis
Lumbrlnaridae
Luabrlneria 1210.0 <2.0
»PP-
Maldanldae 550.0 <2.0
»P« 0«v')
Hcraldac
Platynereia 10.0 <2.0
bicanaliculata
Ophclildaa
Armandia 90.0 <2.0
brevls
Orbinlidae
Naineris 560.0 <2.0
spp.
Phyllodocldae
Eteone 40.0 <2.0
longa
Sun 77
# vt
180.0 <4.0
92.0 <4.0
1112.0 <4.0
312.0 <4.0
0
20.0 <4.0
452.0 <4.0
20.0 <4.0
Fall 77
# vt
380.0 <4.0
120.0 <4.0
2044.0 76.0
2300.0 4.0
820.0 0.0
212.0 <4.0
692.0 <4.0
12.0 <4.0
Win 78
# vt
192.0 4.0
160.0 <4.0
ii72.o y
2172.0 y/
332.0 <4.0
280.0 <4.0
716.0 y
40.0 <4.0
-------
Tabic lie. (Cont.)
Spr 77
# vt
Phyilodocidae (cont.)
Phyllodoce 40.0 <2.0
maculata
Polynoidae
Harmothoe 50.0 <2.0
imbricata
Splonidae
Malacoceros 650.0 <2.0
glutaeus
Syllidae
Exogone 5220.0 <4.0
lourei
Terebellidae
Plata 560.0 /
brevlbranchiata
Crustacea
Tanaidacea
Leptochelia 810.0 <2.0
dubia
Amphlpoda
Canmaridea 590.0 <2.0
•pp.
Sum 77
# vt
Fall 77
# vt
Win 78
# vt
52.0 <4.0 120.0 <4.0 252.0 <4.0
380.0 <4.0 492.0 8.0 52.0 J
600.0 <4.0 2692.0 < 4.0 4052.0
y
2560.0 <4.0 5180.0 <4.0 3432.0 <4.0
512.0 <4.0 340.0 20.0 492.0 >/
1492.0 <4.0 1760.0 <4.0 2080.0 <4.0
452.0 <4.0 218.0 16.0 660.0 ^4.0
-------
fable 11c. (Cont.)
Spr 77
# wt
Dccapoda
Plnnlxa 50.0 4.0
•pp.
Upogebla 120.0 40.0
pugettensis
Echinodermata
Holothuroldea
Leptosynapta 220.0 <2.0
clarki
Species richness 47
Diversity - H' 2.36
Total number 25,910
Total biomass (g) 302
Sum 77
# wt
Fall 77
# wt
Win 78
# wt
24.0 0.0
660.0 60.0
112.0 <4.0
57
2.42
24,216
252
140.0 <4.0
220.0 308.0
20.0 <4.0
78
2.48
50,412
1084
52.0 <4.0
352.0 <4.0
100.0 <4.0
62
2.10
51,644
20
-------
Table lid, Jamestown (coarse-fine sand) dominant benthic organisms from the shallow subtidal
zone collected spring 1977, expressed as number and/or biomass (g) per m*. For
complete data set see Appendix I.
-5 in -10 m
# wt # Wt
Nemertea spp. 96.0 <2.0 170.0 <2.0
Nematode spp. 1336.0 <2.0 426.0 <2.0
Molluscs
Gastropoda
Alvinia sp. 410.0 <2.0 26.0 <2.0
£ Margarites/Lirularia spp. 196.0 <2.0 226.0 <2.0
nj Nassarius mendicus 0 6.0 <2.0
Natica clausa 0 6.0 <2.0
Bivalvia
Cardita ventricosa 0 0
Clinocardium nuttallii 210.0 2.0 20.0 <2.0
Crenella decussata 1366.0 2,0 86.0 <2,0
Macoma spp. 126.0 4.0 452.0 34.0
Mysella tumida 1450.0 2.0 280.0 <2.0
Psephidia lordi 210.0 <2.0 156.0 <2.0
Annelida
Ampharetidae
Ampharete arctica 0 240.0 <2.0
Capitellidae
Mediomastus sp. 426.0 <2.0 796.0 <2.0
Chaetopteridae
Phy1lochaetop t erus 280.0 <2.0 80.0 <2.0
prolifica
-------
Table 114. (Coat *)
Clrratulldaa app.
Tharyx nultlfills
Dorvilleidaa
Frotodorvillaa gracilis
Healonldae
Mlcropodarkc dubia
Maldanidaa app.
Nicoaacha paraonata
Neraidaa1
Platynaraia blcanallculata
Omiphidaa
Onuphia ap.
Ovanlldaa
Omnia fusiform!a
Paraoaldaa
Arlcldaa ap.
Polynotdaa
Haraothoa imbrlcata
Sabcllidac
Sabella media
-5 a
# nt
-10 m
vt
50.0
76.0
236.0
406.0
392.0
400.0
326.0
10.0
130.0
36.0
416.0
-£2.0
*2.0
<2.0
<2.0
<2.0
<2.0
*2.0
<2.0
<1.0
<2.0
<2.0
646.0
136.0
56.0
636.0
290.0
40.0
176.0
66.0
50.0
26.0
200.0
<2.0
<2.0
<2.0
<2.0
<2.0
<2.0
<2.0
<2.0
<2.0
<2.0
<2.0
270.0
<2,0
-------
Table Lid. (Cont.)
Spionidae
Malacoceros glutaeus
Polydora socialis
Prionospio steenstrupi
Spiophanes boobyx
Syllidae
Exogone sp.
Sphaerosyllis pirifera
Oligochaeta spp.
Crustacea
Ctmacea
Diastylis ap.
Tanaidacea
Leptochelia dubla
Aaphipoda
Gaamaridea app.
Decapoda
Pugettia gracilis
Species richness
Diversity - H'
Total number
Total blauta
-5 IB
# wt
-10 n
# wt
10.0
26.0
146.0
50.0
*2.0
<2.0
<2.0
<2.0
16.0
46.0
166.0
20.0
<2.0
<2.0
<2.0
<2.0
1940.0
70.0
<2.0
<2.0
270.0
<2.0
826.0
20.0
<2.0
<2.0
620.0
<2.0
6.0 <2.0
2646.0 <2.0
1660.0 4.0
146.0 <2.0
136
3.49
17,364
782
50.0 <2.0
1370.0 <2.0
1536.0 2.0
30.0 <2.0
156
3.81
12,864
76
-------
SECTION VI
DISCUSSION
Prior to the initiation of these studies in the spring 1976 absolutely
no quantitative information existed on the shallow water benthic communities
along the Washington coast of the Strait of Juan de Fuca. With the comple-
tion of this second year of sampling and sample analysis a vast storehouse
of quantitative data on the full range of intertidal and shallow subtidal
communities has been accumulated. Strict, clear, replicable sampling has
been done. Literally hundreds of thousands of organisms attributed to over
one thousand species have been processed. All processed organisms have been
retained in a long-term storage repository to enable future reference.
In the following discussion first the study areas will be compared.
Then seasonal and annual changes will be considered.
VI-A. Study Area Comparisons
Summary information on species richness, species diversity, density,
and biomass for the intertidal and subtidal are presented on Tables 12 and
13 respectively. The intertidal values were averaged over the four sample
periods for the seven areas sampled quarterly.
Second year data confirmed first year data trends. Intertidal species
richness and diversity were highest in the rock sites (Tongue Point, Pillar
Point), followed by cobble (North Beach, Morse Creek) and protected sediment
areas (Beckett, Jamestown). They were lowest in the exposed sand (North
Beach, Kydaka) and gravel (Dungeness Spit, Twin Rivers) areas.
Intertidal species richness increased with decreasing tide height in
all haMi-fli-s exceot at Pillar Point where sand scouring reduced richness at
+0' (Nyblade, 1978). In the severely exposed gravel (Dungeness Spit, Twin
Rivers) and sand (Kydaka) areas the very low richness showed little tidal
height difference. Species diversity showed a less distinct pattern.
Except in species poor areas lowest diversity was found in the high inter-
tidal.
High diversity and species richness were found in the subtidal.
Patterns among the areas or between the two depths were difficult to
discern. Generally, species richness was less in the most exposed areas
(Kydaka, Twin Rivers —5m, Dungeness —5m). Still, all subtidal areas were
species rich compared to gravel and sand intertidal habitats. At the
subtidal rock site (Tongue Point) intense grazing by sea urchins was
observed at both -5m and -10m. Species richness was much lower at -10m
both years reflecting the decline in algal species number with depth in
-------
Table 12. Intertidal summary of second year community parameters by study area - tide height.
Pillar Point, North Beach Cobble, and Twin Rivers are spring 1977 data only, other areas means
of four seasons (spring 1977 - winter 1978) (Tongue Point +0', spring-summer only; Morse Creek
+0', spring - fall only)
Study Area
Mean Species
Richness
Mean
Diversity
Mean Total
Density (#/m )
Mean Total^
Biomaas (g/m )
Tongue Point
W
61.3
2.40
36,779
1476
(rock)
+3'
118.3
3.04
48,516
3446
+0'
177.5
2.25
19,375
7368
Pillar Point
+6'
42
1.85
11,620
480
(rock)
+3/
131
2.50
8,994
6317
+0'
124
2.04
15,642
11375
North Beach Cobble
+6'
16
1.99
2148
78
(cobble over sand)
+3'
38
2.48
3802
128
-K)/
125
3.24
6724
1862
Morse Creek
+6'
11.5
1.07
1191
<30
(cobble over sand)
+3/
67.3
2.21
20,666
926
+0'
103.7
2.33
15,218
3109
Beckett Point
+6'
12.5
1.65
2143
*133
(protected
+3'
45.3
2.24
16,678
•c402
gravel-sand)
40/
80.8
2.37
56,578
<2468
Dungeness Spit
1
0.17
86
<2
(exposed gravel-
+3/
1.5
0.16
220
<3
sand)
40 '
2.8
0.27
196
<6
Twin Rivera
+6/
3
0,76
44
<6
(exposed gravel-
+3'
4
0.63
692
<8
sand)
+0 /
15
1.62
1448
^30
North Beach Sand
+6 '
7.3
1.60
81
*15
(semi-exposed
+2'
10.5
0.49
3287
•*27
sand)
+0 '
18.5
1.26
4597
<34
-------
Table 12. (Cont.)
Mean Species
Study Area Richness
Kydaka Beach +6' 3.5
(exposed sand) +3' 2.5
+0' 4.8
Jamestown +6' 7.5
(protected sand) +1.4' 38.2
40' 60.3
Mean
Diversity
Mean Total-
Density (#/m )
Mean Total
Biomass (g/m )
0.81
66
<7
0.50
99
*-5
0.91
112
<10
0.22
56,874
<25
1.75
42 ,445
<372
2.34
38,046
<115
-------
Table 13. Subtidal summary of second year community parameters by study area - depth. All
parameters are for June 1977.
Species Diversity Total Density Total Biomass
Study Area Richness Hj (#/m2) (g/ro )
Tongue Point
.5
in
209
1.45
18,635
9160
(rock)
-10
m
94
3.00
834
2988
Pillar Point
-5
m
77
2.82
13,814
<156
(sand)
-10
m
86
3.26
4426
<172
Morse Creek
-10
m
127
2.72
11,654
<290
(cobble-gravel)
Beckett Point
-5
m
83
2.77
13,416
*-170
(sand)
-10
m
90
3.05
17,542
<214
Dungeness Spit
-5
m
28
2.43
858
*56
(sand-gravel)
-10
m
136
3.82
6764
<322
Twin Rivers
-5
m
28
1.69
8852
<56
(gravel)
North Beach
-5
m
201
1.44
14,103
12,021
(cobble-grave1-
-10
m
151
3.86
7442
1474
sand)
Kydaka Beach
-5
m
54
2.55
6230
<114
(sand)
-10
m
96
2.83
17,410
<222
Jamestown
-5
m
136
3.49
17,364
782
(sand)
-10
m
156
3.81
12,864
76
-------
part but also a decline in polychaete and small crustacean species, This
latter would imply that more or larger sediment pockets were sampled at -5m
compared to -10m*
The above illustrates the difficulty of determining causes for observed
patterns from the data alone. A very large number of factors, both real and
sampling induced, could produce the observed patterns in species richness.
Fortunately poor sampling is more likely to obscure patterns than create them.
Of the patterns mentioned above it is clear that two factors are primarily
responsible. Pirst is the type of substratum. Rock supports the richest
marine communities in this region. Second and virtually of equal importance
is exposure to wave and current activity. Exposed rock communities are much
richer than those from protected rock areas. The opposite exposure effect
is found for soft sediment areas. Exposed soft sediment areas support very
species poor communities, while the communities found in protected soft
sediment areas may approach the richness of protected rock communities.
(See Figure 4.)
Intertidal abundance and biomass followed fairly closely species
richness patterns with highest values at the rock, cobble, and protected
soft sediment areas. Patterns at these areas relative to tidal height were
more complex. Biomass increased with decreasing tide height. No clear
pattern in abundance was observed. In gravel and sand areas, low biomass
precluded determination of tide height patterns. Density was usually
lowest at the highest tide height.
For subtidal areas lowest densities occurred at exposed Dungeness -5m
and Tongue Point -10m. In general, biomass decreased with increasing
depth, due largely to decreased plant biomass with depth.
It is also possible to rank all study sites by biomass or standing
crop and from this to make inferences about productivity arid energy flow in
the community. Rock habitats usually had by far the greatest standing crop
with as much as 11.4 kg/m2 found in this study. The 12,0. kg/m^ found at
North Beach -5m was mostly due to a single large Pterogophora: very large
algae are not adquately sampled by the methodology employed in this study.
Cobble areas were next in standing.crop, although protected sediment areas
at some levels where large bivalves and crustaceans were abundant also had
a large standing crop. In rock and cobble areas a large percentage of the
standing crop was benthic macro-algae (and some eelgrass), the major
primary producers in these communities. Therefore, areas with little or no
macro-algae such as the gravel and sand habitats would have low productivity.
Energy flow in these communities would be based on importation from drift
or the plankton. Although turnover rates are unknown, it is hard to imagine
they are high enough to raise the energy flow and indirect productivity of
these sparsely populated gravel and sand communities to the level of those
of rock, cobble, or fine sediment communities. Ranking energy flow and net
productivity in the rock, cobble and protected sediment systems is impossi-
ble without detailed rate studies.
The first year report (Nyblade, 1978) gave a detailed analysis of
study area and tidal level similarities. Second year data confirmed first
year patterns. Rock, cobble, and gravel intertidal areas were similar to
-------
each other, Subtidal areas of similar sediment type showed high community
similarity. Low similarity was found between communities at high, mid, and
low tidal height at given study sites * Appendix II Tables 1-10 give the
cumulative two year species list for each study site with the tidal height/
depth range indicated for each species. Examination of these tables
reinforces the patterns observed above,
In last years report a brief comparison was made between the Strait
study sites and some San Juan Island DOE study sites. It would be very
desirable to be able to expand on that to compare all study sites sampled
by comparable methodology in the greater Puget Sound basin in the past
five years. It would be valuable to know if regional faunal/floral trends
exist within this larger area.
The number of sites sampled is now quite large with METRO sites from
Puget Sound, DOE sites from San Juan Island and the Rosario Strait mainland
area, and NOAA sites from Whidbey Island. To obtain the necessary original
data and put it into a uniform format would be both very time consuming and
expensive, putting regional comparisons beyond the scope of this report.
(See Recommendations.)
Site to site trends in replicate variability would be of interest and
would be possible using a similarity index. This would require extensive
computer analysis and proved to be beyond the scope of this report.
VI-B. Seasonal Changes
Tables 14, 15, and 16 summarize seasonal changes many of which were
noted area by area in the Results. Table 14 gives a summary of species
richness seasonal changes. In areas with a high species richness, species
richness remained high through the year, with some areas showing a slight
decline in richness in the winter (Tongue +3, +0; Pillar +0). There was
also often a winter depression in species richness at gravel and sand sites,
species poor areas. Such winter depressions may be due to annual algal
die-off and storm disturbance.
A summary of seasonal change in community density is presented in Table
15. Seasonal patterns were not particulary clear. However, more areas had
their highest densities in the summer and lowest in the winter than during
the other three quarters.
Seasonal change in biomass (Table 16) was not obvious. No meaningful
biomass information was obtained from the low biomass areas, a majority of
sites, giving a much shorter table than Table 14 and 15. Algae dominated
the biomass at higher biomass areas. The patchiness of large algae distri-
bution masked any seasonal trends.
Except for species which had a discrete period of massive recruitment
(Balanus, Muscuius, Platynereis), seasonal population changes of individual
species were often not apparent. Reaspns for this may be several. First,
there may have been little population change over the year. The species may
be long-lived or the individuals of a species may be replaced at the same
rate as mortality. Inadequacy of the sampling methodology to adequately
-------
Table 14. Second year (1977-78) seasonal change
Asterisk indicates seasonal peak.
Study Area Spring
Tongue Point +6' 79*
(rock) +3' 114
40' 180
Morse Creek +6* 8
(cobble over +3' 59
sand) +0' 112
Beckett Point +6' 10
(protected +3' 23
gravel-sand) 40' 68
Dungeness Spit +6' 2*
(exposed gravel- +3' 2*
sand) +0' 3
North Beach Sand 46' 10*
(semi-exposed +2' 9
sand) +0' 16
Kydaka Beach 46' 2
(exposed sand) 43' 2
40 6
Jamestown +6' 8
(protected sand) 4-1.4' 33
40' 47
in species richness by study area -
Sumner Fall
tide height.
Winter
66
136*
175
16*
76*
in*
13
55
83
1
2*
2
8
13*
21*
6*
5*
5
4
36
57
44
97
15
64
12
29
79
1
1
1
5
10
19
2
1
1
14*
38
61
56
126
7
70
82
15*
74*
93*
0
1
5*
6
10
18
4
2
7*
4
46*
76*
-------
Table 15. Second year (1977-78) seasonal change in density by study area - tide height.
Asterisk indicates seasonal peak.
Study Area Spring Summer Fall Winter
Tongue Point
+6'
89,216*
29,800
14,880
13,220
(rock)
+3'
30,750
97,740*
45,072
20,501
+0'
10,470
28,280
—
Horse Creek
+6'
206
2,458*
1,257
844
(cobble over
+3'
6,452
29,420
33,278*
13,514
sand)
40/
17,843*
17,450
10,360
_
Beckett Point
+6'
1,630
632
4,404*
1,904
(protected
+3/
3,450
11,068
48,228*
3,964
gravel-sand)
+0/
24,830
58,584
97,112*
45,784
Dungeness Spit
+6'
8
332 *
mm
0
4
(exposed gravel-
+3'
276
16
16
572*
sand)
40'
52
144
520*
68
North Beach Sand
+6'
92
156*
48
28
(semi-exposed
+2'
2,600
6,372*
1,824
2,352
sand)
40'
5,608*
3,692
3,792
5,296
Kydaka Beach
+6 /
12
144*
96
12
(exposed sand)
+3/
8
328*
48
12
+o'
272*
112
56
8
Jamestown
+6 '
150,220*
44,428
30,972
1,876
(protected sand)
+1.4'
11,536
68,244*
61,368
28,632
+0'
25,910
24,216
50,412
51,644 *
-------
Table 16. Second year (1977-78) seasonal change in biomass by study area - tide height where
significant biomass was measured. Asterisk indicates seasonal peak.
Study Area . Spring Summer Fall Winter
Tongue Point
(rock)
+6'
+3'
4-0 '
3466*
2985
6506
1619
3712
8230
446
4807*
374
2280
Morse Creek
(cobble over
sand)
43 '
40'
653
3936*
1047
3176
916
2216
1088*
Jamestown 40'
(protected sand)
302
252
1084*
20
-------
reflect real population change may be another reason. The methodology
used was designed to optimally describe the communities present, not
necessarily to provide population information on component species useful
for examining seasonal population change.
It is satisfying to note that the general composition or structure of
the communities sampled remained stable at most areas sampled in terms of
the dominant species and their general order of abundance. Community
composition remained stable over the seasons sampled and between the first
year and second, as discussed in the next section.
VI-C. Annual Changes
An essential goal of the second year program was to document changes
from first year results. Observed changes may be due to real or extrinsic
factors or may be the result of poor sampling design. Once changes are
clearly determined to be due to extrinsic factors, speculation on what
factor or factors produced the change becomes possible. One factor may be
only the normal biological variation in an otherwise stable community, its
biological noise. The community may not be stable. It may be undergoing
slow directional change, because of its early successional stage or long-
term changes in the physical environment. It may not be a stable community
at all, representing just a single time frame of random change.
First year/second year comparisons of species richness, diversity,
abundance, and biomass are presented in summary form for all sites in
Table 17. Species richness and diversity showed very little year to year
change especially in the richer intertidal sites. Figures 2 and 3 present
graphically the relationship between richness/diversity and year one/year
two percentage change in these parameters. The more complex the communities,
the lower the year to year change in these parameters. Larger subtidal
changes may be only a product of sampling error. Abundances also show
surprisingly little change, generally much less than a factor of ten, and
biomass changes are even less. An exception is North Beach -5m where much
algae was collected the second year, probably a sampling problem caused by
collecting larger cobble in the second year in this apparently heterogeneous
area.
Table 18 presents a summary of the values for community similarity
between the first and second year. These were mentioned in the Results.
Community similarity in the intertidal from the first to second year was
greatest at the rock, stable cobble, and protected soft bottom areas, all
areas with rich, complex communities. This year to year constancy in
overall community composition was an especially noteworthy result when one
considers sampling problems due to organism! patchiness arid the irregular
recruitment of some important individual component species. Figures 4 and
5 graphically present the relationship between sediment type, community
complexity, and year to year community constancy. In general, species
richness increases with increasing sediment particle size. The complex
communities associated with rock/cobble and protected sediments show less
year to year community change than the- simpler exposed gravel and sand
communities, with the exception of a few simple communities where taxonomic
lumping may create artificially high community constancy.
114
-------
Table 17. Comparison of first and second year summary community parameters. Asterisk: Pillar Point,
North Beach Cobble, Twin livers comparison of spring 1976 with spring 1977; Tongue Point +0'
SDrini?—snitnffiAf 1Q76 w-ffh — in-*-*- « ^ ~~
m
Study Area
Stratum
Mean
Species Richness
'76-'77
'77-*78
Tongue Point
+6*
47.3
61.3
+3'
103.3
118.3
*4-0*
150.3
177.5
-5m
133
209
-10m
59
94
Pillar Point
*+6'
26
42
+3'
169
131
40*
123
124
-5m
92
77
-10m
91
86
North Beach
*+6'
12
16
Cobble
+3'
49
38
+0'
122
125
Morse Creek
+6*
8.5
11.5
+3'
56.8
67.3
*+0'
101.8
103.7
-5m
74
-10m
149
127
Beckett Pt.
+6'
17.3
12.5
+3*
40.0
45.3
+0'
80.3
80.8
-5m
101
83
-10m
132
90
Dungeness Spit
+6*
4.0
1.0
+3*
3.0
1.5
+0»
3.3
2.8
-5m
30
28
~10m
90
136
Mean
Diversity
H'
Mean
Total Abundance
(#/m2)
Mean
Total Biomass*
(g/m2)
76-'77
'77-'78
'76-f77
"-J
00
'76-'77
'77-'78
2.2
2.4
18,109
36,779
928
1,476
2.6
3.0
28,371
48,516
2,575
3,446
2.5
2.3
4,474
19,375
5,262
7,368
2.6
1.5
6,004
18,635
8,539
9,160
2.1
3.0
604
834
778
2,988
0.9
1.9
27,282
11,620
640
480
3.1
2.5
11,540
8,994
11,766
6,317
1.8
2.0
729
15,642
7,548
11,375
3.1
2.8
6,123
13,814
_
3.3
3.3
4,715
4,426
_
_
1.5
2.0
283
2,148
—
78
1.6
2.5
9,077
3,802
353
128
3,5
3.2
5,512
6,724
908
1,862
1> ^ 1
1,1
3,131
1,191
_
1.8
2.2
21,388
20,666
791
926
2.6
2.3
12,794
15,218
1,683
3,109
3.0
1,495
—
2.8
2.7
8,863
11,654
_
1.8
1.7
1,713
2,143
_
2.0
2.2
4,716
16,678
2.1
2.4
34,102
56,578
MM*
_
2.9
2.8
15,522
13,416
*-
3.1
3.1
18,122
17,542
1.0
0.2
60
86
0.7
0.2
150
220
—
0.8
0.3
97
196
_
2.4
2.4
283
858
L
3.0
3.8
3,828
6,764
-
—
-------
Table 17. (Cont.)
Mean
Study Area Stratum Species Richness
Mean
Diversity
Mean
Total Abundance
(#/m2)
Mean
Total Biomass*
(g/m2)
'76-'77
»77-»78
*76-'77
00
r*.
r*-
*76-'77
'77-'78
'76-'77
00
1 !
r**
Twin Rivers
*+6'
3
3
0.3
0.8
184
44
_
_
+3'
5
4
0.6
0.6
332
692
_
-
+0'
10
15
0.9
1.6
420
1,448
-
-
-5m
139
28
3.5
1.7
5,282
8,852
-
-
-10m
65
-
2.1
-
6,093
-
-
-
North Beach
+6*
7.8
7.3
1.1
1.6
221
81
-
-
Sand
+2*
10.8
10.5
0.7
0.5
3,521
3,287
_
-
+0'
14.8
18.5
1.0
1.3
4,223
4,597
-
-5m
163
201
3.6
1.4
9,303
14,103
390
12,021
-10m
109
151
3.1
3.9
5,881
7,442
62
1,474
Kydaka Beach
+6*
3.3
3.5
0.8
0.8
175
66
_
-
+3'
4.8
2.5
0.5
0.5
443
99
-
_
+0'
6.3
4.8
1.5
0.9
42
112
-
-
-5m
51
54
3.0
2.6
2,300
6,230
-
-
-10m
53
96
2.9
2.8
4,568
17,410
-
Jamestown
+6*
6.8
7.5
0.1
0.2
22,319
56,874
-
_
+1.4'
30.0
38.2
1.9
1.8
11,579
42,445
-
+0'
64.0
60.3
2.2
2.3
56,076
38,046
_
-
-5m
174
136
3.5
3.5
21,712
17,364
266
782
-10m
144
156
3.2
3.8
20,747
12,864
106
76
* Blanks occur where community biomass for first, second or both years was low and uncertain due to the
large numbers of <0.1 g. weights.
-------
Table 18. Community similarity between year one and year two by study area - tide height. Similarity
values range from 0.00, no similarity^ to 1.00, total similarity. Blanks oceur where two years of
data were not obtained. Values for Pillar Point, North Beach Cobble, Twin Rivers are based on
spring 1976 and spring 1977 only; Tongue Point +0* spring-summer 1976 and spring-summer 1977; Horse
Creek +0* spring-fall 1976 with spring-fall 1977 only.
Study Area Habitat Type Tide Height
+6*
+3'
+0'
—5m
-10m
Tongue Point
rock
0.56
0.61
0.49
0.31
0.08
Pillar Point
rock intertidal; sand subtidal
0.80
0.67
0.36
0.57
0.54
North Beach Cobble
cobble over sand
0.17
0.47
0.33
-
-
Morse Creek
cobble over sand
0.79
0.68
0,52
-
0.61
Beckett Point
protected gravel-sand
0.76
0.37
0.56
0.58
0.65
Dungeness Spit
exposed gravel-sand
0.24
0.83
0.21
0.30
0.34
Twin Rivers
exposed gravel-sand
0.79
0.09
0.12
0.21
-
North Beach Sand
semi-exposed sand intertidalj
sand-gravel-cobble-subtidal
0.28
(+2')
0.96
0.79
0.08
0.42
Kydaka Beach
exposed sand
0.12
0.26
0.32
0.55
0.39
Jamestown
protected sand
1.00
(+1.4')
0.61
0.79
0.42
0.48
-------
Generalizations in the subtidal are more of a problem because of the
obvious inadequacy of the Van Veen grab sampling -methodology. In sediment
where the grab worked the best, year to year similarity was high. It is
possible that most subtidal communities would show the community constancy
found intertidally in rock and protected fine sediment areas if adequately
sampled.
Comments on annual changes of dominant species in the various
communities studied were made in the Results Section. In the exposed
gravel and sand habitats populations changed greatly from year one to year
two, i.e., community similarity was low. This might be only a sampling
artifact due to the difficulty of sampling a sparse often aggregated fauna.
However, great population oscillations may as likely be a real characteris-
tic of these areas. In the habitats with more complex communities, some
species populations changed greatly from year to year. However, the
majority remained fairly stable from year one to year two.
In summary then, should one of the study sites be visited five or ten
years from now, the same basic community should be found. Species composi-
tion, richness, and diversity should remain very similar, with perhaps
slightly greater changes in overall abundance and biomass.
118
-------
M.FAN SPECIES RICHNESS
S3 U>
O U3
Figure 2. Intertidal species richness (mean of the first and second years) by study area - tide
height compared with the percentage change in species richness between the first and second years.
BP6 = Beckett Point +6', BP3 = Beckett Point +3', BPO = Beckett Point +0'; DS6, DS3, DSO =
Dungeness Spit +6', +3', +0'; JT6, JT3, JTO = Jamestown +6', +1.4*, +0": KB6, KB3, KBO = Kydaka
Beach +6', +3', +0'; MC6, MC3, MGO » Morse Creek +6', +3'» +0'; NBC6, NBC3, NBC0 = North Beach
Cobble +6', +3', +0'; NBS6, NBS3, NBS0 = North Beach Sand +6', +2', +0'; PP6, PP3, PP0 = Pillar
Point +6', +3', +0'; IP6, TP3, TP0 = Tongue Point +6' , +3', +0'; TR6, TR3, TR0 = Twin Rivers +6',
+3', +0 *.
-------
MIAN DIVERSITY
^ ®w
woo
ON hrf
Figure 3. Intertidal diversity (mean of the first and second years) by study area - tide height
compared with the percentage change in diversity between the first and second years. For study
area abbreviations (e.g., BP6), see Figure 2.
-------
MEAN SPECIES RICHNESS
to
Ui
l_n
O
Ui
o
o
ro
Ln
Ln
O
Ln
(-1
H
Co
• • •
L35 J 3
ddW bd
cnco m
ovjj o
S
m ^
•• td
. • O
trt
Co
W
Tt
O
•
C-i
H
O
L •••••
I
HONOON
I &
OG\
• c/v
3N-3
» fxJ
t(U>
CO
• •
o 3 2
ca n 53
o n
ON
w
TJ
u>
2!
dd
O
U>
irJ
-3\
o
u>
>-d
a\
•
X
o
o
hd
u>
25
W
o
o
"d
TJ
O
;-d
CO
O
Figure 4. Intertidal species richness (mean of the first and second years) by study area- tide
height compared with sediment particle size. For study area abbreviations (e.g., BP6) , see
Figure 2. FS = fine sand, FMS = fine-medium sand, MS = medium sand, FCS = fine-coarse sand,
SG = sand and gravel, CS = cobble and sand, CSG = cobble, sand and gravel, R = rock.
-------
MFAN SPECIES RICHNESS
Figure 5. Intertidal species richness (mean of the first and second years) by study area*- tide
height compared with the similarity of communities between the first and second years. Values
for similarities range from 0.0 (no similarity) to 1.0 (total similarity). For study area
abbreviations (e.g., BP6), see Figure 2.
-------
VII, REFERENCES/BIBLIOGRAPHY
In addition to those references cited in the text, the following list
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-------
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124
-------
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¦ ¦» • —— «v ^—« -
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125
-------
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126 —
-------
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127
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129
GPO 697-148
-------
APPENDIX I
COMPLETE DATA SKTS
130
-------
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biottM* < g p*r <•! "ii '•»».'• r. 2 < *»-41 ;'••!•>•
Sprint
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•0
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-0. I 0
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0 0 0 0 0. ? -". I
•P- •n.',
Pi MTiclill# 0.1 <0.1 0 0 0-1
o IfArMchli fQ.2
' iodor idw
r^l#iLa 0 0 o.l -o.i y o.l 'O.i
*•">.) '}.2
-------
Point
S^r ing '11 r
0.01 m7 0.2 «2 0.01 .' 0.
9 wt t wi 9 vt 9
Km1«11* 0.1 <0.1 0 - 0
rO. 2
B«1«I I* 0 0 >},2 *n. j o
•»> -A.*
pp. 0 0 0. ^ '-OS fl
* I. )
FikrtcU 0.1 <0. S 0 *.2 'Q, i 0
arn—ici tO. 7 • ? .*
Orloptli 0 0 1.7 -0.5 0
~2.6
kktu^MclU 0 0 0.2 '0.1 a
rO.S
s*r?«l Ite
Iflnrtll 0 0 0 0
2.4 <0 i 0 J.S
•V) »V 4
0 0 0
0 0 0
0 0 0. i
•o :
0. i ^0.1 0 0
!«*•«¦$I +Q. 2
i#M»rwylM- 0 I <0.S n 0 ?
fti1l«r« *0 5 »0.")
0.0: m4- 0.1 m2 il.OJ «' W. . ft"
' w r $ wt & *r t ¦wi
0
0 >; O ,i
2. S OS u 0
h b
l;.<* -si i .> n 2 . i) ; y
t , <» • l' 4
•7 f; U n
OS -(i \ 0 o a
0 5> 0.1 <0,1 u
'0.
1.4. .<»_ ! r. 0.6 <0.1 O
fe. 7 - 1. «
f>.2 <0 1 m '>. 1 - (,i. ! I
O. % ' L>. 7
2 'i 0 -o.l <> 3 h
0 " ' 0 i i: 0 'i
0 ?
0 2 ' : ' 1 m
5
<5 r a
-------
T n»-jr P»Snf * S
S$r 1 Ig *n '
_D-°L"i. _ o.i k o.oi ,2 0
t wt if w*__ fl wi #
Irliu 0.1 <0.1 0 0 <">
milt—at mil *0. )
Mill* ya.% «o. i o 2). , . o
•*. Ml. J
TvrckflU^w
«f. O.J '0.1 0 0.1 • a. s 0
O.t -0.2
Hflfc Uflti 0 0 0.6 r 0
c Irrtt 1 'I..'
tlcoiM 0 0 D,} '0.1 0
lacttrlcoli *0 6
$lpw»c«U
C 0 0. i <0. * 0
tift'.i ?0.2
0,1 <0.1 D ! , | 10-i n
¦ *0 k <1.5
0.2 <0.1 0 0, I *0.1 0
drUt*-
*0.7 *fV 2
mivrnem 0 0 Q 0
•»«
fWiiekiU^lia 0. 1 <0. 10 0 G
t#m&rmwm ?0.5
0 0 oa -o.j n
r0.3
Crwltcti
Ovtmedi
0 4 <0 1 0 9 1
? i.b *\S .Q
S SI
M . f*
' '1.$
0. I
•0. >
U 8
*2 1.*
I , «
- / I
1,0
' \.h
•J
¦ 1
•0 t
•0 i
0.2
<0.5
0
0. 1
»o - 2
a
14. 1
*37.6
0
7 5 r . !
* U.0
-------
Tongu* Pf»int *}
Spring
' 77
SuMM
T *71
0.
,01 «2
Q,
.2 m2
0.
.01 ¦_2__
0.2 mZ
i
wt
#
wt
#
i 1
[rriHU
kalMtaa
D.J
0. 1
0
l.ft
0. i
0
to.*
*1,1
**. *
*0.2
htlm»
0
0
0
0
|Wi1j
1*1 a«M
<0, 1
0
22.4
0. J
0
«*-
>«.S
•44,9
*0.2
Cfctfctaalna
0.4
0. 1
0
C
0
<1*111
~1.1
tO.O
rsiitciptf
0
D
o
0
jolymrM
tMlatill
2» 7
*0. i
0
•C. 1
0
¦ami}
I 3.0
U*« 0
hirfwdi
*t?
?0.J
?3.«
~0. i
.'6.2 '0
fTj llll Bill ¦! Ill iia
4f, 7
0.1
0
*U. 5
0.1
0
«fc*ar*rl
to.2
*5?. J
*0. i
t«nn>hirf<>ii
0. 3
<0. 1
0
0
0
mf 1 lct«4«
:I,3
€seTiaee?ii«*rcma
0.1
0
0. t
<0. i
0
gfipmnii
tfl.5
*o.s
Uattapiii
i.7
<0.1
0
0. J
'0, \
0
kiacMi4i
<3.3
f ], 1
tMlroyi'a
0, i
<0. t
Q
0
0
to. 3
liblM
3, t
<0,1
5.5
0.2
i.#
a. i
4.0 1
VOADMfR'-hl 1
?0 .2
»
u.o
*04
?6. 5
'0.5
*4,7 ~ 1
IS?
4,;
12.2
0. I
*0.3
0
e,4
"0.9
0, 1
*1 , I
'0,j
0, 3
•0 5
0
0
0
u- 1
0. ^
Q. I
~0. !
0.6
l.l
17-3
0. i
• 'i. ft
0
0,6
: V I
9.0
* n. <¦}
* s
*8. «<
0
0
24*
i
I. %
*4.4
22.3
42, ^
-n, i
'0.1
< f.M
0,8
» t. •
0
7. 7
?«. 2
1.0
• 3. i
*.8
4
0
0
J .8
~S. 5
0
O.I
* I. i
n.o
MS {
0.3
*'1.0
~a.2
o
G
0
0
C:
•J
! U.S
» j > . ?
, .6
~i.*
(J
20.0
*29 *
a
a. i
-3.2
n.a
~2.3
'0, I
0. I
*0.2
J.%
¦ s. %
-------
T--»r,a<*r F 'lr»r * J
Spring '71 Sdftcr ' 77
0.CM a2 0,2 m2 0.0} 0.
t
lfc
?2,*
0. i
*0. S
0.9
a
o. i
~0. 5
0
117.
•104.
0. ?
* 0. B
41-8
•(SB. ?
27.8 0.1
~12.9 *Q.7
. 0 0.
. 7 • 0
0. S
*0.2
0
0- 1
~ 0.2
0. i
~0.2
-------
T.-ntij* P ' I n r ~ j
Spring
2
Rf*J«
irwrflcoriii
enlIformica
¦ r*W
lalc«U
QiLpchlB«
Il^tl
far^lrailM
Sfvcteildlia
rtctlpalM
C«rt4«* 0
*p.
m 0. t
hlrMtt«cali« t0.2
«0.2
Fi^tt 11« 0
gfac 11 U
tMKU
Cfclrwmi<» 0
ColMftira 0
Dlpt«r«a U. 6
L«rw
Bojicbo^odid** 0
»r
lt+>rdr idar 0
0,0
'0. i
O.J
*0.5
0
0, 1
~0. 5
0 2
•0.4
0.0
*0. S
0. i
t0.2
0
0. i
?Q. )
0. J
~0.6
0
iO
%h
* I 12
~0. !
'0. I
<0. 1
'0. I
f\ I
'0 , 1
<0. I
0.0
«o,a
0
o
0. ^
*1,0
0
0 I
•0.1
a *
* W{
0.0
*0.0
0 i
3 *
<2. i
0. J
Mi 2
-------
TonjiuP P• > i n~ * <
Sprli>t '77 SuHfr ' ' J
_ 0_.pi »2 0, 2 j#' _ 0.01 «? _ 0-
# vt # ft J wr f
Irrom * 0 «
M-
tcblw4ff«t«
4»c«rol4*«
*f. o o o.i -o.i n
~o. 1
0.1 o Q.s «0.1 o i o
(*»*'< 1« ?0.2 *Q.fl •1.0
Bei0t&eret4*A
•y. 0 0 0,1 '0.; 0
*0, 7
CaCMirli 0 0 9 0
¦laiita
CrncmtmrU M.3 i.5 i.O 0.5 65.2 1.7 0
>mlBC
Dlvtrilt)! ¦' ).i" 2. ?9
?ot*l ?, ?«W.« 2«», Wv). t
T«t*l llawa 799.4 9#» V 2
f i s. '. r
o. 2
-------
Table 1c. Tongue Point (rock) benthic organisms from the
spring and smnner 1977, exnresst-J is number anu/nr bi
'N = -) m-1 +- stini'ard deviation.
, Spr 7 7
0.01 a 0,2 m'
# vt # wt
Chlorophyt*
HoMstroma <0.1 0
ap.
Rhlzoclonlua <0.1 0
rlpariura
Spongoaarpha <0.1 0
coalita
Spoogoaorpha <0.1 0
¦plnescens
Ulva 0.0 0
8p• +0,1
Bacilldi'lophyta
Petmalee <0.1 0
spp.
Phaeophyta
Alaria 57.6 ?54.7
¦p. 1110.9 1521.5
Co#taria 0.1 0.7
castata ±0.2 ±0.8
Deesarestia 0 2.2
ligulata ±4.5
Hedophyilua 55.9 68.7
sessile 159.9 ' 30.2
LAffiinarla 0 0.4
saccharina ±0.7
La1® ilia r Lai e s 0.2 0
sp. *1.0
1 55
c w (.+0 ') intertidal zone collected
>nwis:i (t>) per 0 ,!) i (N-20) ,ind
Sum 7 7
0.01 m
# wt
0.2 in
wt
0
0
0
0
: 0,1
0
0
0
0
0
<0.1
65.0
:118 . 6
0
0
0
0
0
620.8
±752.8
0
0
12.0
*16.7
4.9
+ 8.0
0.0
±0.1
-------
Tongue Point
2Spr 77
0.01 is 0.2 n
f wt # wt
Rhcx.'ophjrta
Acrochaetlua <0.1 0
«P
-------
Tongue Point
, Spr
77
0.01 m
0.2 in
# vt
# wt
Cryptosiphonia
<0.1
0
wood i 1
Delesscrlaceae
<0.1
0
sp.
Endocladia
<0.1
0
auricata
Cigartina
0.0
0
papiilata
±0.2
CracliarLa
0
<0.1
sp.
Raloaacclcm
<0.1
0
glandIforme
Halymenla
0
1.3
cocclnaa
+ 2.7
Hateroalptaonla
<0.1
_ Q
sp.
Hildenfarandla
r'
0
occldcntale
Hj»enena
0.3
3.7
flabclligera
±0. 9
±4.4
Bynencfia
0.2
0.8
sp.
±0.3
±1.0
Iridaea
1.5
40.8
cordata
?2.9
±79.1
Iridaea
0.0
0.0
sp.
±0.1
± 0.1
Laurencla
0.0
0
spectabt*
±0.1
Melobesia
0
<0.1
aedlocrls
157
+0
Sum 7 7
0.01 m-
# vt
0. 2 m
# wt
0
0
0
0
0
0
0
0
0
0
0.7
±0.3
2.1
±4.2
0
0
<0.1
0
0
0
0
0
0
0
0
0
0
0
5.2
±10. 5
0
0
0
-------
Tongue Point
Spr 77
•}
0.01 as
0,2 m
# «rt
# wt
Meabranoptera
<0.1
0
sp.
Mesophyllua
0
/
conchatun
sophy Hum
/'
0
Sp.
Microcladia
0.0
0
bcrealls
±0.0
Heoptilota
0.0
0
hypnoidea
±0.0
Neoptiloti
<0.1
0
Sp- (Juv.)
Odonthalls
0.5
0.1
f loccosa
±1.0
±0.3
0d on thai la
<0.1
0
kaatschatica
(Mont ha 11a
<0.1
0
sp.
Petrocells
¥
0
¦lddeadorfit
Peyssonelt«
<0.1
0
sp.
Plocaaiua
<0.1
0
tenue
PolysIphonia
<0.1
G
hendryi
Polystphonla
<0.1
0
pacifies
Porphyra
<0.1
0
•p.
151
+0
Sura 77
0.01 m
# wt
0.2 ci-
# wt
0
¦
1
0
<0.1
0
0
0
0
0
0
0.3
±1.0
0
0
0
o
<0.1
<0,1
0.1
±0.2
0
0
0
0
0
0
0
0
0
-------
Tongue Point
»PP
2 Spr " ,
0.01 m 0.2 m2
} wt # wt
Pterosiphonia <0.1 0
biplanata
Pteroalphonla <0.1 0
dendroIdea
Pc llothaniopsls <0.1 0
lejollsea
Rhodochorton 0 0
«P-
Rhodymenia 0.2 0.6
palaata ±0.5 ±1.2
Rhodyaenla <0,1 0
sp.
9permatophyta
Phylloapadlx 0.9 185.4
•coulerl ±3.4 ±370.9
Porlf era
•pp. *' 0
Halichondrla / 0
paclf lea
Ophlitaspongia 0 0
peonata
Cnidarla
Aothozoa 0.1 <0.1 0
+ 0.5
Anthopleura 0 0
elegantlssiaa
Hydrozoa
app. >'
159
+0
Sum 77
0.01 m
_# wt
<0.1
0
0
<0.1
0
0
0.2 m
_# wt_
0
0
0
0
0
0
34.6 871.9
±58.9 ±1030.6
0 /
0 0
0.
+ 1.
0,
+ 0.
0.1
1 °'3
0.0
+ 0.1
0.8
+ 1.5
<0.1
-------
aittlta
eschrlchtii
Calllost
-------
Tongue Point
Neaertea
»PP
Meaacoda
•pp.
Molluaca
Aaphlnaura
Cyanoplax
dentlens
Sacbari&a
tunicata
Nopal la
sp. K
Tonicell*
1 in eats
Ga#tropoda
AfiBSU
¦dtra
Acaacldae
up. (juv)
Acteocina
¦p.
Alvinia
Bp.
Rarlaela
haliotiphila
Spr 77
0.01 a2 0.2 m-
# wt # wt
1.4
±1.5
7.0
4- 9.9
<0.1
0.2
±0.4
0
0.1
tO. 3
0.0
tO.O
0.0
±0.1
0.5 0.1
±0.6 ±0.1
1.3 <0.1
±1.3
0
1.5 4.5
±0.6 ±4.5
0
0.2
tO.6
0.3
tO.4
<0.1
<0.1
0
0
0
160
+0
Sun 77
0.01 m
# wt
0.2 nf
# wt
2.2
±4.0
<0.1
20.3
34.4
<0.1
0.1
±0.2
0.2
±0.4
0.3
±0.6
0.0
±0.1
1.4
±4.3
0.0
±0.1
0
0
0.3 1.2
±0.5 ±2,3
1.3 0,8
±1.5 ±1.6
0.1
±0.2
0.1
±0.2
0.3
±0.9
2.4
±9,8
0.7
±0.6
0.5
±2.1
<0.1
<0.1
<0.1
<0.1
0
0
0
-------
Thais
can&liculata
Thais
esarginata
Thais
•p. (Juv.)
Trlchotropla
caocellata
Velutina
laevigata
Blvalvla
Hlatella
arctlea
Lasaea
clscula
Lyonnla
callfornlca
Modiolus
rectus
Muscuius
pygmaeus
Mysella
turn Ida
MytHus
californianus
Mytllus
edulla
Mytilus
•p. (Juv.)
Protothaca
ataainea
Tongue Point
Spr 77
0.01 m
# wt
0.2 m
# wt
0
0
0
0.1
±0.2
0
<0.1
0
0
0
0
0
0.5
±1.2
0.2
±0.7
0
0.7
~1.6
0
0
0.7
±0.9
0.0
±0.0
0.1
±0.4
<0.1
<0.1
<0.1
<0.1
0.8
±1.5
0
0
0
0
0
0
0
0
0
0.4
±0.8
162
+0
Sura 77
0.01 m
# wt
0.2 m
# wt
0.1
±0.2
0.2
±0.7
0.1
±0.2
0
0.3
±0.7
<0.1
<0.1
<0.1
<0.1
0
0
0
0
0
0.2
±0.4
0
0
0.2
±3.8
0.1
±0.2
0.1
+ 0.2
0.1
±0.2
0.1
±0.2
0.2
±0.5
<0.1
<0
<0
<0
<0
<0
<0,1
0
0
0
0.1
±0.2
0.1
±0.2
0
0
0
0
0
0
0.0
±0.1
0.1
±0.2
-------
Tongue Point
2 SPr 77 2
0.01 m 0.2 ra
f Wt # Wt
TransennelLa 0 0
cantilia
Zirfaea 0.1 0.0 0
pilsbryi *0.5 ±0.0
Annelida
Oligochaeta 1.9 <0.1 0
¦pp. ±4.5
Polychaata
Arenicolidae 0.1 <0.1 0
¦pp. ±0.2
Branctxloaaldane 0.1 <0.1 0
vlscentl ±0.4
Caplccllldae
Capitella 0.5 <0. 0
capitata ±1.1
Chrysopetalldae
Paleoootua 0.2 <0.1 0
belli* -0.4
CirraCulidae 0.1 <0.1 0
spp. ±0.2
Cirratulue 2.8 / 0
cirratus ±6.3
Eunicoidea 0.4 <0.1 0
spp. ±0.9
Euclcldae
Eunice 0.2 / 0
valena ±0.4
Luabrlneridae
Luabrineria 0.1 <0.1 0
inflate ±0.2
163
+0
Sum 77
0.01 ra^
# wt
0.2 n'
# wt
0.7
*1.5
0
0.0
±0.1
0
28.3
±68.0
0
3.5
17.6
11.1
±24.8
0.5
±0.9
1.7
±3.1
0.4
±1.3
0.2
±0.5
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
0
0
0
0
0
0
0
-------
Tongue Point
2 SPr 77 2
0.01 ¦ 0.2 m
# wt # wt
Ltabrlncrla 1.2 -"0.1 0
•p. ±2.0
Haldanidae 0 0
¦p. (Jirv. )
Hereidac
sp. (Juv. ) 0 0
Nerets 4.2 <0.1 0
sp. ±4.9
Ncreia 0 0
pelagica
Piatjrnercsis 1.1 <0.1 0
blcanallculata ±3.0
Onupbldae
Oouphia 0.4 <0.1 0
stlgut Is tl. 3
Oauphls 1.1 <0.1 0
sp. ±3.0
Orbinildae
Nalneris 0.1 <0.1 0
dendritlea "0.2
Hainerla 0.1 <0.1 0
quodriccuSf ia ±0.4
Malnerla 0.4 <0.1 0
sp. 1.6
Scoloploa 0.1 <0.1 0
sp. ±0.4
Phyllodocidae
Eujalla 0 0
btllneat a
Eulalia 0.2 <0.1 0
quadrloculata ±0.5
164
Sua 77
0.01 m2 0.
# wt #
3.2 <0.1 0
±6.4
0.1 <0.1 0
±0.2
4.5 <0.1 0
1.3 <0.1 0
±1.7
0.1 <0.1 0
±0.2
1.2 <0.1 0
+2.6
11.7 <0.1 0
24.9
0 0
7 0 <0.1 0
14.9
0 u
2.2 <0.1 0
±5.5
0 0
0.1 <0.1 0
±0.2
0.6 <0.1 0
±1.2
-------
Eulall*
•P-
Polynoicue
¦P-
Halosydna
brevisetosa
Sabellidae
app.
Chone
ecautls ta
Eudiatylta
Vancouveri
Fabricla
oregonica
Fabricinae
¦p.
Orlopsis
«p.
Sabella
media
Sabellastarte
¦p.
Schixobranchla
insignia
Sabellarildae
Idanthyrsua
anutut
Sarpulldae
Spirorbis
ap.
Spbaerodoridae
Sphaerodorople
mlnutfc
Tongue Point
0.01 m
# wt
Spr 77
0.2 n
i wt
0.1
±0.2
0
0.1
±0.2
3.2
±5.4
0
U.l
±0.2
0.1
±0.2
1.1
±4.5
1.1
±4.7
0.2
±0.4
0.2
±0.5
0.2
±0.7
<0.1
<0.1
<0.1
0
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
0
0
0
c
0
0
0
0
0
0
1.5
±2.8
0.3
±0.8
<0.1
<0.1
165
+0
. Sum 77
0.01 m 0.2
# wt # wt
0.1
±0.2
0.3
±0. 7
0
<0.1
<0.1
0
0
0
2,4
±2.9
0.8
±1.0
0
0
12.6
±27.]
0. o
± 1.2
0.2
±0.7
G
<0.1
<0.1
<0.1
<0.1
<0.1
0
0
0
0
0
0
0
0
0
±0.3
2.7
±6.6
± ">.9
<0.1
<0,1
'0.1
-------
Tongue Point
U. U1 D . i. HI
# wt # wt
Spionidae
Polydora
0.1
<0.1
0
Columbiana
±0.3
Polydora
0.3
<0.1
0
pygldlalla
±0.8
Polydora
0
0
•pouglcola
Polydcra
0.4
<0.1
0
ap.
±1.4
lldac
Brania
0.4
<0.1
0
brevlpharyngea
±0.0
Exogcrae
1.4
<0.1
0
gnmlfera
±2,1
Exogone
0.5
<0.1
0
lourei
±1.2
Exogone
01
<0.1
0
ap.
±0.3
Odontosyllls
0.8
<0.1
0
•P.
±1.9
Sphacroayllla
0.4
<0.10
brandboratl
±1.3
SphaeroeyllIs
0.4
<0.10
0
plrlfera
±0.8
Sphaeroayllia
0
0
sp.
/
r
Syllia
11.1
0
ap, i
:25. 2
166
+0
Sum 77
0. 01 m
# wt
0.2 ®
# wt
0.2
±0.4
2.3
±7.5
0.2
±0.5
0.9
±1.5
0.2
+0. 5
0.7
±1.0
2.2
±5.5
0
0.9
±2.1
0
0.6
±1.0
0.7
±2.0
19.7
±29.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
0
0
0
0
0
0
0
0
0
0
0
0
0
-------
Tongue Point
2 SPr 77 2
0.01 m 0.2 m
# wt # vt
Terebellidae 4.1 / 0
¦pp. ±8.7
Nicolea 6.5 <0.1 0
roatericola ±28.1
Polycirrus 0 0
ap.
Sipunculida
Fhascolososa 0.1 <0.1 0
agaaaizii ±0.2
Pyccofonid*
Halosoma 0.1 <0.1 0
coapactua ±0.2
Cruatacaa
Ostracoda
epp. 0.4 <0.1 0
Cirripadia i
lalanua 0.8 <0.1 7.3 40.6
cariosua ±1.3 ±14.5 ±81.2
Balanua 0.1 <0.1 0
giandula ±0.2
Balanua 0.4 17.2 1.5 2fc.l
oubilia il.l ±58.8 ±1.9 141.0
Badanua 0.7 0.0 0
«rp. (Juv.) ±3.1 ±0.1
Chchaaalua 0 0
d&lli
cea
Cunclla 0 0
vulgaris
167
0.01
# wt
Sum 77
0.2 m"
# wt
0.1
±0.2
12.7
20.4
3.2
±8.7
<0.1
<0.1
<0.1
0
0
0
0.2
t0.5
<0.1
0.1
±0.2
0.0
^0.1
0.1
±0.2
<0.1
3.4
6.4
2.1
~8.0
0
<0.1
2.0
±8.4
0
2.0 31.9
>2.3 ±42.7
0
2.6
±6.5
0.1
±0.2
0.4
±1.4
0.0
±0.1
<0.1
<0.1
0
0
0
-------
Tongue Point
Taoaidacea
Anatanais
no man i
Leptochella
dub la
Pancolus
californlensis
Isopoda
Cirolana
narfordi
DynaMaella
glabra
Dynaaenella
sheareri
Dynamenella
Bp. (JUV.)
Exosphaerotaa
media
Exosphaeroaa
rboaburum
Exoaphaerosa
sp. (Juv.)
lanlropsls
klncaidi
laoiropsls
trldens
lanlropsls
sp.
Idotea
aontereveneis
Idotea
stenopa
0. 01 m
# %rt
Spr 7 7
0.2
# vt
3.0
±5.1
4.9
*13.0
0.1
±0.2
0.1
±0.2
0.8
±1.5
1.5
±3.2
0.1
+ 0.3
0
5.7
:14. 7
0.4
±1.6
0.2
±0.5
0
0
0
0
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
0.0
±0.1
<0.1
<0.1
0
0
0
0
0
0.1 <0.1
±0.4
0
0
0
0
0,1 0.0
±0.4 ±0.1
0.1 0.0
±0.2 ±0.1
163
4-0
Sum 77
0.01 m'
t vt
0.2 n.
# wt
7.8
±12.8
2.2
±5.6
1.0
±4.3
0
o.t
±1.3
0.5
±1.2
0
0.1
±0.2
2.6
±4.3
0
1.4
±3.2
0.1
±0.3
1.9
'5.0
0
0
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
'0,1
^0.1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
-------
Tongue Point
2 Spr 77 2
0.01 m 0.2 m
f vt # wt
Idotea 0.2 0.1
wosnesenskii *0.5 +0.2
Idotea 0.5 <0.1
sp. (juv.) i0.9
Jaeropais 0.1 <0.1
dubia ±0.2
Jaeropais 0
lobaCa
Jaeropais 0
sp.
L tumor ia 0.9 <0.1
algarum ±2.5
Lisioria 0.4 <0.1
1 lgnorum ±1.3
Muona 0.4 <0.1
chrooatocephala *0.8
Munna 0
atephenseni
Munna 0
ubiquita
Paranthura 0
elegans
Sjmidotea 0
pettiboneae
Synidotea 0
sp. A.
Anphipoda
Ampithoe
sp. C.
Aapithoidae
spp.
169
Sum 7 7 _
0.01 m2 0.2 m
# wt # wt
0
0.1
0.2
0
0.8
2.2
0.1
0.2
0.8
2.0
0
0.1
0.2
0. 3
0.8
0.2
0.7
0, 2
0.4
O.z
0.7
1.1
3.9
1.6
4.6
2.5
4.9
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
0. 5
to.6
0
0
0
0
0
0
0
0
0
0
0
0.0
to. 1
0.8
:1.5
0.1
:0. 2
-------
Tongue Point
Aorodea
columbiae
Atylus
leviiensis
Caprella
greenleyi
Caprella
verrucosa
Ceropa
compactus
Cyaadusa
uncinata
Gaaaaridea
ep-p,
GaMUropsls
tht»psoni
Hyale
frequens
laehjrroeerus
anguipes
Jaasa
falcata
Lepldepecremn
(gurjanovae)
Lepldepecreum
sp. A
"ellta
callfornlca
Met'opa
c istella
Ha jna
consilionna
Spr 77
o.ni m£
* wt
0,2 m
f wt
±0
0
to
0
to
<0.1
<0.1
<0.1
13.2
±9. 6
<0.1
0
0
0
1.0
tl,2
0.0
~0.1
170
7 Sura 7 7
0.01 m" "j.2 m'*
# wt # wt
5.8
6. 7
0.5
¦0.9
0
0
0.1
:0, 2
0.4
1.1
<0.1
<0.1
<,.1
0.0
~0.1
0
0.3
~0.5
0
0
7.5
'15.0
<0.1
0,5
'1.1
0.3
0.8
1.1
2.4
1.0
1.9
0.2
.0.5
0.1
0.5
0.5
2.0
1.7
3.5
0.1
0.2
0.2
¦0.7
<0
<0
<0
<0
<0
<0
<0
<0
<0
0
0
0
0
0
0
0.3
-0,5
0
<0.1
-------
Tongue Point
0.01 n2 0.2 m2
# wt # wt
Najaa
sp.
Oligochlnus
light!
Parapleustes
naut Hue
Parapleustes
pugettensis
Photia -
bifareata
Photlfl
brevipes
Photia
Sp. A
Photia
•pp.
Polycheria
osborai
Pontogeneia
intermedia
Tritella 0.1 <0.1 0
pilimana +0,2
Deeapoda
Cancer
0.1
0.1
0.1
0.2
orego&enais
±0.3
±0.4
±0.3
±0.2
Cancer
0.1
0.0
0
sp. (juv.)
±0.3
±0.1
Cryptolithodes
0
0
•itcheasis
Oedignathus
0.2
0.4
0.1
0.6
inermis
i0.4
±1.4
±0.3
±0.6
171
Sum 7 7
0.01 sn
# wt
0.2 ii
# wt
1.3 <0.1
'4.3
0.1 <0,1
tO.2
1.4 <0.1
~2.0
1.0 <0.1
2.5
0.4 <0.1
0.9
0.2 <0.1
0.9
5.4 <0. 1
9.4
5.2 <0.1
8.9
0.1 <0.1
0.2
0.4 <0.1
0.8
0.3 <0.1
0.6
0.1 0.4
0.3 ±1.6
0
0.5 0.4
1.0 ifi.8
0
2.8 <0.1
±3.8
0
0.5 <0.1
±1.0
0
0
0
2.0 <0.1
±4.0
0
0
0
0
0.5 0.0
±0.6 tO.l
0
0.3 0.0
±0.5 ±0.1
0
-------
Tongue Point
Spr 77
0.01 a
# wt
0.2
Pagurus 0 0
birsutiusculus
Pagurus 0.7 0.2 0.8 0.1
•p. (juv.) *3.1 ±0.7 ±1.5 ±0.2
Pinnotheres 0 0
pugettensls
Plxmotherldae 0.1 <0,1 0
spp. ±0.2
Pugetcia 1.7 0.1 0.5 0.3
gracilis ±3.5 ±0.4 ±1.0 ±0.6
InsecCa
Coleoptera 0.1 <0.1 0
app. ±0.2
Diptera 0.3 <0,1 0
sf-p. ±0.7
Bryozoa
8pp.
Echlnoderwata
Ophluroidea 0.1 <0.1 0
sp. (Juv.) ±0.3
Aateroidea
Benrlcla 0.1 0.0 0.1 1.0
leviuscula ±0.3 ±0,1 ±0.3 ±0.1
Leptaaterlas 0.1 <0.1 0
hezacdb ±0.4
Holothuroldea
Cucmurla 0.1 <0.1 0
pseudocuratJ ±0.2
-K)
0. 01 m
f wt
_ Sum 77
2 0.2 in2
# wt
0
±0
0
±0
0
±0
0
<0.1
<0.1
0.0
±0.1
0
0
0
0
6.
18.
0.2
±0.4
1.0 1.8
±5.5 ±2.3
0
0. .
±0.2
<0.1
0
0
C.l
±0.2
<0.1
0.2
±0.7
0.1
±0.3
0.2
±0.5
0.2
±1.0
<0.1
0.0
±0.1
0
0
-------
Tongue Point
Spr 77
0.01 m2 0.2 m2
f # vt
Chordats
Aacldacea
¦ p.
Species Richness 178
Diversity - 2.33
local Nuaber 2617.5
Total Bloaasa (g) 1626.5
173
+G
Sum 77
0.01 ra"
# wt
0.2 tn
# wt
170
2.17
6183.8
2057.5
-------
Table Id, Tongue Point (rock) oenthic organisms fron the subtidal collected
I? June 1977, expressed as number and/or biomass (3) per 0.25 + standard
deviation (N=4).
Chlorophyta
Chaetotaorpha
ap.
Derbesia
marina
Monostroms
¦ p.
Rhizocloniua
riparium
Spongooorpha
coalita
Spongoaorpha
spinescena
Ulvaria
sp.
Baclliariophyta
Pennales
spp.
Phaeophyta
Costaria
costaca
Desmarestia
llgulata
Desaarestla
virldis
Desaarestia
sp. (Juv.)
Ec tocarpaceae
-5 m
vt
-10
wt
0.0
to. 1
0.1
±0. 1
0.0
3-0. 1
0.0
30. 1
0.0
±0. 1
0
0.0
30. 1
0
0.0
±0. 1
0.0
±0. 1
0
0
0.0
~0.1
0
0.0
10
. 1
0
0
0
±0
1
2
7
*2
1
0
0
±0
1
0
0
-0
1
0.0
~0.1
3.6
±7,3
0,8
±1.7
4.3
±8.5
0
0
-------
Glffordla
ovata
Laminariales
ap.
Pterogophora
callfornica
Pylalella
tenella
Syringoderasa
abysaicola
Rhodophy ta
Acrochaetima
ap.
Aopllaiphonla
paclfica
Antithamrtion
defectum
Antithanmlon
kylinii
Boss lei1 a
pluaosa
Bonneaaiaonia
r.oockana
Cal1iarthron
tuberctilosura
Callithamnion
blseriatum
Callithamnion
pikeanum
Tongue Point
Sub tidal
-5 is
wt
-10 m
wt
0.0
±u. 1
2,3
+ 3.9
0
0.0
tO. 1
0.0
±0. 1
0.0
to, 1
0.0
±0. 1
0. 1
±0.0
0,0
+0. 1
0.4
i 0.4
0
1121.8
~688.5
0.0
±0.1
0.0
rO.l
0
0
2,7
±5.3
0
0.0
±0. I
0
0
0.0
+0. 1
0,0
zO. 1
0
0.9
±1.9
1 . 7
±3.4
0
0
175
-------
Ca 11 i thansnion /
Pleo.iOBporiusa
Callophyllis
flabellulata
CallophyllIs
haenophy11a
Ceramiun
callfornicum
Ceramiurn
sp.
Choreocolax
polysiphoniae
Corallinaceae
Cruoria
profunda
Cryptonemla
ovalifolia
Delesseriac^ae
Fauchea
laciniata
Cri ffithsia
pacifica
Herposiphonia
pluaula
Herposiphonia plumula
var. plunula
Hoi1enbergia
subulata
Tongue Point
Sub tidal
-5 tn
wt
-10 m
wt
0.1
~0. 1
0
0
0.0
+ 0,1
0.0
1
0. 1
iO.O
0.0
to. I
0. 1
±0. 1
0
0.0
±0,1
2.9
±2.6
0
33. 5
±27.7
0.0
±0.1
0.0
to. 1
0.0
±0. 1
0.2
10.2
0.0
±0.1
0.1
±0.1
0
0.0
±0.1
0.0
10. 1
0
0. 1
±0.2
0
0.0
±0.1
0.3
±0.5
0. 1
±0.2
0
0.0
±0.1
176
-------
Tongue Point
Sub tidal
-5 m
-10 m
# wt
t) ut
Hymenena
4.3
0
Sp,
±4,2
Iridaea
0.0
0
sp.
±0.1
Mesophyllura
0.0
0
conchatini
±0.1
Mesophyllun
0.0
0
sp.
±0. 1
Microcladia
0.1
0
borealia
±0.1
Microclad la
0.0
0
sp.
±0.1
Opuntiella
0
0.0
callfornica
10. 1
Peyssonelia
0.0
0
pacifica
±0. 1
Peyssonelia
0.0
0
sp.
±0.1
Piatythamnlon
0.0
0.2
pectlnat um
±0.1
±0.2
Platythamnion
0
0.0
villosum
±0.1
Pleonosporium
0.0
0
Vancouverlanun
=0.1
Polyslphonla
0. J
0
hendryl
+ 0. 1
Polysiphonia
0,1
0.0
pacifica
±0. 1
±0. 1
Polysiphonia
0.1
0.0
paniculata
±0.0
+0.1
17 7
-------
Polysiphonia
ap.
Polysiphonia
seopulorusi
vlliosun
Pterosiphonia
bipinnata
Pterosiphonia
dendroidea
Pterosiphonia
gardtieri
Pterosiphonia
gracilis
Ptilothanmiops1s
lejollsea
Rhodoptilum
plumosum
Rhodymenia
sp.
Porifera
sop.
Cnidaria
Hydrozoa
Aglaophenia
Plumularia
sp.
Tongue Point
Subtidal
-5 m -10 m
# wt 9 vt
00
0
to. 1
0.0
0.0
to. 1
±0.1
0. 1
0.0
±0.1
*0.1
0.1
0.0
to. 1
±0. 1
0. 1
0.1
+ 0. J
±0. 1
0.0
0.0
±0.1
to. 1
0.1
0.0
±0.0
±0.1
0
0.0
±0.1
0. 1
0
±0. 1
0.0
0
±0. 1
*
r
0
0.1
±0.2
0
2.3
±4,6
178
-------
Anthoroa
Actiniaria
spp.
Platyhelminthes
spp.
Nemertea
spp.
Neisetoda
spp.
Molluscs
Aaphlneura
ap. (juv.
Cyanoplax
dentleu8
Mopalla
sp.
Tonice.la
llneata
Gastropoda
Atraaea
mltra
Acmaeidae
Aeolldldae
sp.
Tongue Point
Sub tidal
-5 a
wt
10 n
wt
0.6
<0. 1
4.7
t8. 1
<0. 1
15.3
±9.9
<0, 1
0.5
±0.6
<0.1
31.0
;43. 3
<0. 1
0.3
±0.5
<0. 1
6.0
!n . U
0
0.7
+ 0.6
38.7
±31.9
0.0
i0. 1
0. 1
±0.2
14.8
10.9
0
0.3
±0, 1
0
12.3
±13.7
<0.1
8.A
10.8
1.8
±2.i
0.3
±0.5
0.5
±1.0
0.5
±0. 7
<0. 1
<0.1
1.3
±2.5
0
2.5
±3.8
0.'*
±0.8
<0. 1
179
-------
Tongue Point
Subtidal
-5
m
-10 m
#
wt
P
wt
Alvinia
23.0
0. 1
-.5
<0.1
sp.
±22.3
±0. 1
±0.6
Amphissa
78.8
3.4
1.8
0. 3
colon*} iana
±54, 3
±3. 1
±2.2
±0. 3
Balels
0.3
<0.:
0
nontereyensls
tO.5
Bittium
2.0
0. 7
0
eschrichtl1
8
tO. 9
Bittlun
0.9
0.3
0.0
spp.
±0,9
±0.5
±0. 1
Calllostoma
29.3
0.5
0.0
ligation
±29.5
+ 0.6
±0. 1
Calyptraea
a
0.5
<0. 1
fsstigiata
±1.0
Ceratoacoina
0
0.8
3.4
follatum
±1.0
±4.5
Cerithiopsis
13.5
0.4
0.3
<0.1
sp.
±15. 3
±0.5
±0.5
Colllsella
0
0.8
0. 1
onhracea
±1.5
±0. 1
Crepidula
6.3
0.3
0.3
<0.1
sp.
±5.7
±0.4
±0.5
Crepipatella
0.3
0.1
0
lingulata
±0.5
±0.1
Cryptobranchia
0.5
<0.1
0
concentrica
±1.0
Fusltrlton
0
0.3
21.6
oregonensis
±0.5
±43.2
Cranulina
135.3
1.0
0
aargaritula
t186,9
±1.4
180
-------
Tongue Point
Sub t idal
-5 is
wt
-10 10
wt
Lacuna
24.3
0.0
0.5
•¦ariegata
~31. 3
±0. 1
+ 1.0
Lirularla
39.8
0. 3
0
lirulata
±41.3
:0.4
MargarItea
57.0
5.0
0
pupillus
*50,9
±4.8
Kargarites/
2.0
0.0
0.3
Lirularia
+4.0
±0. 1
±0.5
Mitrell:
9.3
1.4
0
carinata
±9. 1
±1.5
Hoelleria
0. 3
<0. 1
0
quad re
+0.5
Nassarlus
0.5
0.2
0
¦endicus
±0.6
±0.2
Ocenebra
13. 3
1.9
0
lurida
~12.0
±2.4
Odostomia
0.3
<0. I
0
«P-
±0.5
Oenopota
0.3
<0, 1
0
tabulata
±0.5
Retusa
0. 3
<0.1
0
sp.
±0.5
Searlesia
0.3
0.5
0
dlra
±0.5
±1.0
Tegula
0.3
<0. 1
0
sp. (juv.)
iQ. 5
Trichotropis
0.5
0. I
0. 3
cancel lata
±1.0
+0.2
±0.5
Turridae
0.3
<0, 1
0
±0.5
181
-------
Velutina
laevigata
Bivalvia
Clamys
hastata
Eotodesma
saxicola
Hiatella
arct i; \
Annelida
Oligochaeca
spp.
Polychaeta
Ampharetidae
SPP-
Cirratulidae
spp.
Dodecaceria
cencharum
tunicidae
Eunice
valens
Hesionidae
Micropodarke
dubla
Ophlodromus
pugettensis
Lutnbrineridae
Luaibrinerls
spp.
Tongue Poiat
Subtidal
li-Zb
$ wt
u. j <0.1
±1.0
0
0.3 <0. 1
±0.6
0
# wt
0
0.3 0.0
+0.5 ±0.i
0
0.3 <0.1
10.5
0.8
±1.5
0.3
±0.5
13.0
r 11 - 2
5.8
±4.8
0.3
±0.5
0.3
tO. 5
0.3
+0,5
13.0
: 1 5. 0
<0. 1
<0.1
<0. 1
<0. 1
<0. I
<0. 1
<0.1
<0.1
0.5
±1.0
0
<0. 1
-------
S'ere idae
sp.
Nereis
app.
Platynerels
bicanalicuiata
Onuphidae
Oru'ph is
spp.
Opheliidae
Arm-india
brevis
Phyllod>-»c Idae
Eulalia
quadrloculata
Phyllodoce
sp.
Pilargidae
Slqarnbra
teritaculata
Polynoidae
spp.
Arc tone
vi t tat;
Harmothoe
imbricata
Polyodoniidae
Peisidiee
as pera
Tongue Point
Subtidal
-5 ra
wt
•10 m
wt
22.3
il9.7
17.9
H5.9
538.3
±387.8
0.8
±1.5
2.5
±5.0
0
5.3
i 7.4
1.5
±2.4
0.3
~0.5
5.5
±5.6
0
1.8
13. 5
0.3
+0.5
183
<0.1
<0. 1
<0. I
<0.1
<0.1
<0. I
<0.1
<0. J
<0. 1
<0.1
<0. 1
0
1.8
±2.4
15.5
±19.4
0
0
0.3
±0.5
0.3
±0.5
0.3
±0.5
0
0.8
±1.0
0.3
±0.5
0
0
<0.1
<0. 1
-------
Sabellariidae
Idanthvrsus
arraatus
Sabellaria
cementar iun
Sabel1idae
spp.
Chone
ecaudata
Oriopsls
minuta
Serpulidae
spp,
Spirorbts
spp-
Stgalionidae
spp,
Sphaerodoridae
Sphaerodoropsis
minuta
Spionidae
Mai ococeros
glutaeus
Polydora
sp,
Prionospio
s teenstrupi
PrIonospio
sp.
Tongue Point
Sub tidal
— > i-
wc
¦10 ra
wt
2.3
: 3. 0
3.0
±5.4
4, 3
±6,1
0. 3
zO. 5
1.8
~2.1
15.8
r 31. 5
0.3
:0. 3
8.8
+ 13.1
2.5
t3. 3
8.5
-8.7
2.0
±4.0
1. 3
r 1 .9
<0, 1
. i
1.0
0
0
0.5
tl .0
0
0
0
0
'0. 1
<0. 1
-0.1
18-'«
-------
Splo
fillcornis
Syllidae
SPP-
Irania
brevipharyngea
Evogone
gctsri fera
Exogone
lourei
Odontosvllis
sp.
Sphaerosy11 is
piri fera
SylI is
sop,
Terebellidae
spp.
Nicolea
zostericola
Pis ta
brevibranchlata
Polyc irrus
spp.
ArchlannelIda
Polygordlldae
spp,
Tongue Point
Subtida1
¦ 5 in
wt
-10 m
wt
i. 3
±1.5
7.5
112.5
4.5
zl, 7
38. 3
i64.2
23.5
~27.3
9.5
±15.1
1.3
±1.9
31.0
~26,4
5.0
t8. 7
4.8
±5.6
2.0
~ 4.0
4.5
16. 1
0.3
rO. 5
<¦0. I
<-0. 1
<0.
<0. 1
-------
SIpunculida
Phascolosoraa
agassizii
Pvcnogonida
Achelia
chelata
Crustacea
Os trac oda
spp,
Cumacea
Cmnel la
vulgar is
Curael1 a
sp.
Diastvlopsis
rp.
Lampropidae
sp.
Leptoctraa
sp.
Tanaldarc a
Ana tana is
no man!
Leptochelia
diibia
I sopoda
Anthuridae
sp.
Tongut Point
Subr ida1
10 m
wt
¦ 5 ffi
Wt
2.8
-4. 1
<0. 1
0. 3
"0.5
< 0. 1
12.0
'16,6
5. 3
: 10. 5
3. 5
~2.9
0. 3
'0.3
0. 3
*0.5
2.8
•4 .9
1. 3
-2.k
3.8
~5.3
0,3
rO. 5
'0. I
<0. 1
<0. 1
'0.1
- 0. I
<0. 1
-0. 1
-0. 1
<0. 1
0
0
0
0
0
0
0. 5
1.0
0
<0,
-------
Tongue Point
Sub tidal
Colanth».ra
squamoslsslaa
Dyrwmenelia
dllltata
DyTiaMe«wlla
glabra
Epicaridea
sp.
Exosphaereina
r "buroH
Iai.^^opcis
analoga
Ianiropsis
klncaidi
Ianiropsis
trldens
Ianiropsis
sp.
Idotea
sp. (J uv.)
Jaeropsis
dubia
Jaeropals
lobata
Munaa
chromatocephala
Munoa
ublqulta
Munna
sp.
•5 m
wt
¦10
wt
25.3 <0.1
±30.5
0.3 <0.1
±0.5
0.5 «0.I
11.0
0.3 <0.1
+0.3
0.J <0.1
10.5
75.5 0.0
±57.1 ^0.1
11.8 <0.1
±12.9
1.5 <0.1
±3.0
5.5 <0.1
*7.6
1.3 <0,1
1. 7
4.0 <0.1
* 3,9
0
J.5 <0.1
±1.9
26.5 <0.1
±31,7
38.3 0.0
±46.2 -0.1
187
0
0
0
0
0
0
0
0
0
3.8 0.1
±7.5
0.3 0.1
±0.5
0.3 0.1
±0.5
0
1.6 0.1
±2.1
0
-------
Parathura
elegans
Anphlpoda
Caprella
cali fornica
Caprella
sendax
Caprella
sp.
Caprelltdea
sp. (Juv.)
Gasmarldea
spp.
Metacaprella
anoaala
Metacaprella
kennerly i
Metacaprella
sp,
Etecapoda
Cancer
oregonensis
Cancer
sp, (juv.)
Caridea
sp. (juv.)
Crypto11thodes
sitehens is
Crypto11thodes
typicus
Tongue F'oint
Sub tidal
-O O
wt
-10 m
wt
1.5 <0.1
±1.9
2.0 <0.1
+ 4.0
0.8 <0.1
-1.5
0
0.8 <0.1
11.5
2337.0 3.2
+ l<»91.4 3.7
0.8 <0.1
±1.5
1.5 <0.1
i 3.0
0.5 <0.1
±1.0
7.0 4.6
±5.7 ±4,9
0,3 <0.1
±0.5
29.8 -0.1
±31.5
1.3 0-5
±2.5 ±0.9
1,3 0.2
±1.9 ±0.3
0.3 <0.1
± 0.5
0
0
0.3 <0.1
±0.5
93.5 <0.1
±183.0
30.0 0.3
±36.2 0.6
0
19.5 <0.1
±37.7 *0. 1
0
0
0
0
0
0
138
-------
Cryptolithodcs
sp. (juv.)
Discorsopagurus
schmltti
Elassochirus
tenuimanus
Hlppolytidae
sp. (juv.)
Lebbeus
sp. (juv.)
Majidae
sp. (juv.)
Mlmulus
foliacus
Oregonia
gracilis
Pagurus
berlnganus
Pagurus
dalll
Pagurus
kennerlyi
PagurIdae
sp. (J uv,)
Petrolisthes
erloraerus
Phy1 lollthodes
papillosus
Puget tia
gracilis
Tongue Point
Subt idaI
-5 m
wt
10 m
ut
10.8
-9.5
3.0
±3.2
0
15.0
0
i 1
92
r 70. 1
1.8
±2. 1
, 5
,6
.5
0
±0
6
±11.7
13. 5
119.8
4 . 3
±7.9
83.0
±61.6
5.0
16.0
2.3
t 3. 3
240.0
•235.3
0. 1
±0. 1
0. 1
±0. 1
<0. 1
0. 1
0.2
±0.2
0.6
*0. 7
0,
*•0,
!,
±2,
0,
1 I,
0,
±0,
1,
~ l,
0,
*0.2
<0. I
9.5
10. I
0
0
0,3
"0. 5
0
0
0
0
0
0
0
0. 3
±0. 5
2.0
±2.7
0
0
I .0
±1.4
0. 1
~0. 2
0.2
>0. 3
0. 1
•0. 1
<0. 1
189
-------
Spirontocaris
pr ianota
SpirontocarIs
sp.
Insecta
Diptera
larvae spp.
Bryosoa
spp,
Heteropora
sp,
Echlnodermata
Ophluroidea
(j uv.)
Anphlpholis
squamat a
Asteroldea
sp.
E.~hinoidea
St rongylocentrotus
droebachiensis
St rongylocentrotus
franc iscanus
Srrongylocent rot us
purpuratus
St rongylocenrroc us
sp. (juv.)
Tongue Point
Subt idal
-5 m
wt
- 10 m
wt
1. 3
~2.5
64.8
-79-5
0, 1
~0.2
0.3
•0.4
0
0
3.0
3.8
<0.1
19.9
39,8
o
0
2.8
~5.6
11.8
12.2
4.0
*8.0
0. 3
~0.5
u. 0
:0. 1
: 0. I
-0. 1
1 . 3
~1.9
0,8
-1.5
0
<0. 1
'0. 1
6. 3
±4.8
2.3
r 2.6
0.3
±0.5
2.5
±2.9
2 79.0
±206.5
696.6
±804.6
32.4
•64.8
4. 7
~7.4
4,0
: 6.2
1
~ 1.
175.3
~319.2
430.8
~391.3
1 90
-------
labia 2a. Pillar Point (rocit) benthic organisms f ros the high (+6*) intertidal zone
collected spring 1S77, expressed a= number an.J/or bion.nss < g) per 0.01 (N=20)
and 0,2 (X=-i) m- + standard deviation.
0. Olra"
wt
0.2m
wt
Chlorophyta
Cladophoraceae
RhizocIonium
riparium
Ulva
Urospora
sp.
sp.
ap,
<0.1
<0.1
<0.1
<0.1
0
0
Bacillailophyta
Peniiales
Phaeophyta
Anallpus
j aponlcus
Fucus
distichus
spp.
<0.1
0.2
±0.5
0.3
±1.1
0
2.2
±4.4
Rhodcphyta
Antitharanion
dendro ideuB
Endocladi a
muricata
Grigartica
papillata
Halosaccion
glandiforme
Hildenbrandia
sp.
Iridfea
heterocarpa
<0.1
0.1
±0.2
0.1
±0.2
±0.2
»'
0.1
±0.0
0
0
0.7
±0.9
±0.2
0.0
±0.2
192
-------
Kolothuroidea
sp.
Eupencacta
sp»
Parasc ichopus
calif orn iensis
Chorodata/
Ascldiacea
Species Richness
Diversity - H'
Total Number
Algal Biomass
Animal Blomass
Total b" ¦ ¦",?!
Tongue Point
Sub tIda 1
-5 31 -10 m
ut if w r.
6.0
i9 »4
0.3
^0.5
0
2,9
t4.8
-0. 1
2.0
t4»0
0.4
r0.8
0.Z
:0.5
0
<0.1
0.3
to.5
85.3
±170.6
0
209
1,45
4,658.7
1.169.4
1.120.5
2,289.9
94
3.00
208.5
15.0
731.9
746.9
191
-------
Pecrocelis
sp,
Porphyra
sp.
Pteroaiphonla
Spematophyta
Phyllospadi*
scouleri
Neaatoda
sp.
Mollusca
Gastropoda
Acsaeidae
sp. (juv.)
Golllsella
digitalis
C.
pelta
C.
strifacella
Littorina
scutulata
L.
sitkana
Nitoacaaea
persona
N.
scutua
Thais
emarginata
Pillar Poln t
+6
0.01m
wt
0,2m
0.1
0.5
±1.4
1.2
±1.9
0.9
0.1
*0.2
2.1
-1.8
10.9
r 9.2
±0.7
0.1
±0.2
*
0.0
tO.O
n.i
o
o
<0.1
<0.1
0.7
±1.2
2.1
<0.1
0.0
±0.1
0.2
i 0. 3
0
±0.5
0.1
±0.3
11.3
±6-1
2.4
2.3
±4.5
0.3
to.5
0
0.5
±0.6
±1 . '+
0.5
±1.0
wt
0
0
0
0
1.1
±2.2
0
0
8.2
+ 4.0
2.4
0.5
±1.0
0.1
±0,1
0.4
±0.6
il.l
0.6
±1.2
193
-------
Pillar Point
Blvalvla
Mytilus
californianus
M.
edulis
H.
(spp. (Juv.)
Annelida
Ollgochaeta
ap.
Polychaeca
S/llidatf
Syllls
adamantea
S.
spp.
Crustacea
Clrripedia
Balanua
carlosus
B.
glaodula
8.
pp. (juv. )
Ciith^Aal us
dalli
Tanaldacea
Pancolus
californiens is
194
+6
O.Olts
0. 2m
~ 0.4
wt
wt
0.1
±0.2
4.0
±6.5
0.0
±0.1
0.1
±0. 3
3.3
+6.5
0
0
0
1.6
±3.2
0.3
-0.3
<0.1
0.2
±0.4
cO.l
0.
1.
17.
"8.
0.
±0.
74.
±47.1
0.
0.
0.
±0.
<0,
1.
±24
2.3
: 3.9
0
0
0
12.8
±18. 3
0.1
.2
<0.1
-------
Pillar Point
Isopoda
Cryptothir
belanl
Idotea
¦ontereyenels
Aophipoda
CaaoarIdea
sp.
Insecta
Dlptera larvae
Species richness 42
Diversity - H* 1.85
Total number 2905.0
Total blonass 120.0
195
+6
0.Oiffl*
0. 2nf
wt
wt
0.1
0.4
0
<0.1
0
6.3
:12 . 5
0.4
tO. 7
0. 7
1.5
<0.1
1.8
*3.5
<0.1
1.7
2.1
<0.1
0
-------
Table 2b. Pillar Point (rock) ben chic organisms f r»>m the nld (-«-3') intertid.il zone col !<•«.*ted
spring 1977 , expressed as nuaihf r and/or bioraass (g) pt-r 0.01 (N»20) -ir»d 0.2 (N=4) m- -f-
standard deviation,
O.Olm2 0.2m2
if V # wt
Chlorophyta
Chaetoaorpha <0.1 ;i
sp.
Cladophoraceae <0.1 0
sp.
Elite rotsorpha <0.1 0
I. in* a
Rhitoclonium <0.1 >
ripariun
Spongoaorpha 0.0 0
con U3 *0,1
S. -0,1 0
sp.
Ulva 0.1 0.5
sp. '0,2 *0.6
Baclllariophyta
Females <-0.1 0
spp. ii, I 0
Phaeophyta
Alaria
17.5
-+26.2
sp.
'37.6
-772.1
Ectocarpales
0.0
sp.
-0.0
Ectocarpus
0.0
0
sp.
•0.1
Egregia
0.2
nenzl»sli
t0. 7
Hedophy11 am
9.1
286.4
sessile
r 37. ?
¦ 365.7
Lamlnariales
0.0
0,0
sp. (juv. )
iO.i
'0.1
196
-------
Pillar Point
Ralfsiaceae
sp.
Spkacelarla
raceraosa
S,
sp.
Rhodophyta
Acrochaetium
sp.
Ahnfeltia
plicata
Antitham.iion
defectum
Anti thatanionella
Daciflea var. pacifica
A.
glandulifera
Bossieila
pi ifflosa
Call 1thamalon
pleonosporlum
C.
pitceanoni
Cerami ales
sp.
ChoreocoJ ax
t-c! ys lphoni.ee
Coraliina
Vancouverlens Is
Corallinaceae
sp.
Cryp tos iphonia
woodii
1"
2
0.2m
wt '/ wt
~ 0
0.0 0
0.1
i 0.1 0
<0.1 0
< 0. i 0
0.0 0
<0. 1
<1.1 c
<0.1 0
0.0 0
"0.1
<0.1 0
<0.1
<0.1 0
<0.1 0
0.2 0
10. s
-------
Pillar Point
0,Gin"
wt
0.,
~*t
Delessariaceae 0
sp.
Delessaria 0.0 0
decipiens tO.O
Gigartinaceae 0.2 0
sp. t0.3
Glgarcina 0.3 6.6
pap 11lata -0.7 *6.1
Halosacclon 0.1 0.2
glandiform ^0¦1 +0.4
Iridaea 7.1 156.8
cordata -10.9 ±161.5
C. 1 0.1
sp. >0.2 . 2
Mesbranoptera 0.0 0
nultiramosa *0.1
<0,1 0
sp.
Hlcro-ladla 0.9 0
borealis -1.0
Neoptllota 0.8 0
asplenioldes i2.6
Odonthalia 2.2 0
f loccosa . 9
Petrocel is 0
niddendcr f 1 i
Phvcodrys <0,1 0
isabellae
Platvthanmlon <0.1 0
heteroaorphum
P. «-0.1 0
VlllOSUB
P- <-0.1 0
sp.
198
-------
Pillar
Plocami ua
tenue
Polyslphonla
hendryi
P.
pacifica
Porphyra
sp.
r'terosiphonia
bipinnata
Ptilota
filicina
Ptllothaanionopsis
lejoliaea
Rhodyaenia
palmaca
Scagelia
occidentale
Porifera
spp.
Platyhelminthes
Turbellaria
spp.
Nem«rtea
spp,
Hematoda
spp.
Mollusca
Aaphlnera
Cyanoplax
dent lens
K-atharina
tunlcata
0.0io2 0.2m2
" Wt * Wt
0,5 0
*2.2
<0,1 0
0.1 0
to. 1
0,1 0,5
to. I *0.b
<0.1 0
0.8 0
i 2.0
<0.1 0
0.0 0.1
}0.1 i 0. 3
< 0.1 0
0
0.1 <0.1
0.2
1.5 <0.1
• 2.5
0
0
9.8 <0.1 0
- 19.6
0 0.3 <-0.1
0.5
0.1 1.3 3.8 59.6
,0.4 , 5.8 +5.0 162.9
199
-------
Mopalia
sp.
Gastropoda
Acaae 1 dae
sp. (juv.)
Barleela
haliotiphila
Thais
canaliculata
T.
sp.
Bivalvia
Adula
californiensia
Hiatella
artlca
Mitaculus
p yg*aeus
Mytilus
edulis
M.
sp. (juv.)
Prothaca
staalnea
Tresus
capax
AnnelIda
Oligochaeta
spp.
Polychaete
^renicolldae
3P-
Pillar Point
0. 01m
+ 3
2
wt
0.2m'
wt
0. 3
±0.5
2.2
•4.4
0. 3
*0.4
0.5
t2.0
0.1
*0.4
<0. 1
<0.1
<0.1
0
0
0
:0. 3
0.8
t2. 3
0.4
~C.8
0.2
f0.2
0
0
10.0
*23.8
0.1
-0.2
0.1
»0 4
0
0.0
+0.1
<0.1
0.0
tO.l
<0. 1
cO. 1
0.3
±0. 5
1.0
~2.0
0
0.5
il.O
0
0
0
C.l
±0.1
0.4
*0.7
0.1
-0.2
1.4
•2.4
1.0
>2,6
<0.1
00
-------
Capitellidae
Capltell£
Capitata
Eunlcoidea
sp.
Lissbrlneridae
Lmbrlnerls
ap.
Nereidae
sp.
Nereis
sp.
Orblnildae
sp.
Nalneris
dendrlcica
Phyllodocldae
Eulalla
quadrioculata
E.
sp.
Polynoidae
Halosydna
brevisetosa
Sabel1ldae
spp.
Fabricia
oregonlca
Pillar Point +3
0.01 in~
It WC
0,2
rO. 7
0.2
*0.4
0,
>0.
0,
tl.
0.
±0,
0.
±0.
0.
iO.
-------
Schlzobranch ia
insignis
Orlopsis
ainuta
Serpul idae
spp.
Slgalionidae
Pholoe
ainuta
Spionidae
spp.
Polydora
coluabiana
P.
sp.
Syllidae
Exogone
gem&lfera
E.
sp.
Odontosyllis
sp.
Syllis
spp.
Pycnogonida
Achelia
graci1lpes
A.
nudluscula
Kalosoraa
coopaciua
Pillar Point
O.OtnT
0.2m*
wt
WT
0. 3
±1-1
0.7
tl.2
0.1 tQ.2
0.9
+ 2.4
0.1
jG.2
0.1
.0.2
0.7
±1.4
. 1
<0.1
<0. 1
<0.1
<0.1
<0.1
<0.1
0
0
0
0
0
0
0
0.1 <0.1 0
±0. 3
0.1 <0.1 0
*0.2
0.3 <0.1 0
tl. 1
3.2 <0.1 0
i 5. 7
0.2 <0.1 0
~0.7
±0.2
0.1 <0.1 0
i0.2
-------
H.
virldintestitiale
Nyaphon
pixellae
Crustacea
Ostracoda
SPP-
Cirripedla
Balanus
carlosus
B,
nubilIs
B.
Sp.(Juv)
CiMcea
Cuaella
wlgaris
C.
sp.
Tanaidacea
Leptochella
dubia
Pancolus
callforniensis
Isopoda
GiroIana
harfordi
Dyaauwnella
shearl
Exosphaerona
aaplicauda
E.
aedia
Pillar Point
O.Olcf
0. 2m
wl
-0
0
~0
0.1
<0.1
0
0
0.2
: 0 . 5
2.1
14.0
0
0.5
~1.1
0.2
>0.5
0.1
-0.3
<0.1
14.6 13.3
±33.6 t21.5
0.8
±1.5
<0.1 0
<0.1
<0.1
130.4
±211.1
42. 3
±84.6
21.6
~ 82.2
0.9
<0.1
<0.1
0
0
0.2
4-0.5
5.5
±9.5
0.4
*1.1
0.3
t0.6
<0.1
<0.1
<0.1
<0.1
0.8
¦1.5
0
0
0
0.1
*0.2
203
-------
Pillar Point
#
laairopsis 1.1
kincaidi 0.2
Idotea 0.1
wosneseaskl *0.2
I. 2.9
sp. (j uv.) - 5.8
Hunna 7.9
chroaatocephala ±30.0
Aaphlpoda
Cercops 2,9
coapactus *4,8
Gamaaridea 1.0
spp. ±2.2
Irssecta
Diptera larvae 1.3
spp. ±1.8
Bryozoa
spp.
Echinoderraata
Eupentacta 0.5
qulnquesemita *2.0
Leptasterias 0
ftexactis
Ophi uroidea 0.1
sp. *0,2
Chcrdata
Ascidacea
Species richness 131
Diversity - H* 2.50
Total Number 2243.4
Total blonass 1579.3
+3
0.01m„
wt
<0.1
<0,1
<0,1
<0.1
0
0
0.2m
wt
<0.1
<0.1
<0.1
<0,1
<0.1
0.3
*0.5
0
0.2
+ 0.3
/
»
0
-------
Tab 1 e 2c. Pillar Point (rock) btnthic . • ri',m it.s from the low (+0') intertidal zone
collected spring 1977, expressed as number and/or binrnass (g) per 0.01 (N=20)
and 0,2 (N»M b-1 + standard deviation.
0.01»2 0.2m2
spp.
Phaeophyta
8?p.
Bp
Bcclllarlophyta
Pennales <0.1
f wt * wt
Chlorophyta
Chaeto»orpha <0.1 0
ap.
Cladophoraceae *0.1 0
sp.
Entero«orpha <0.1 0
1 lnza
P. <0.1 0
sp.
Rhlzoclanlua <0.1 0
rlparlua
Spongoaorpha +0.0 0
coallta -0.0
Ulothrix <0.1 0
sp.
"lva <0.1 .0.4
*0.9
Alarte +24.0 +518.7
-78.8 -410.2
Pctocarpua <0.1 0
sp.
ERregia 5.3 + "526-1
menzlenii -23.7 -648.5
Feldmamla <0,1 0
simplex
Hedopny1li» 0 ^15.5
sessile -30.9
20%
-------
Pillar Point
Lanlsarla
aetchel111
Laalnarlales
¦P-
Phaestrophion
irregulars
Spbacelarla
raeewosa
S.
ap.
Stietyosiphon
Cot!lis
Khodephyca
Acrochaetlua
•pp.
Ant 1 than ion
defectu*
A.
dendroldeu*
Rosa leila
pltaoaa
Calllthaanlon
plkeamTn
Callithanlcm
Pleonoaporlua
Ceraaina
eaton 1anuv
Choreoeolax
po 1 ys lphon lae
Coral Una
vancouverlcnala
Coral1laneeae
ap.
20ft
+ n
O.Olt
0.2m
wt
+ 1.6
-7.0
+ 1.8
-16.1
*0.1
<0.1
<0.1
<0.1
wt
0
n
o
o
o
o
o
to.o
<0.1
<0.1
+0.1
-0.1
0.0
0.0
<0.1
<0.1
<0.1
<0.1
o.o
.0
4
^0.0
0
0
0
0
0
0
0
0
0
0
-------
Pillar Point
Cryptoslphonia
woodli
Deleaaerlaceae
•p.
Endoeladia
¦urieata
Cigartlna
paplllata
Clgartlnactae
¦p. (Jot.)
Gracllarla
andcraonll
Cracllarlophlla
orytoidea
Gywnogcmgrus
leptophyllua
Haloaacclon
ftlandiforae
Irldaea
cordata
Irldaea
sp.
Meabranoptera
wulelraaoaa
M.
•p. (juv.)
Mlcrocladf.a
boreal is
Odonthalla
floccoaa
PlaryChasnlon
sp.
PlocaaltH
sp. (J«v.)
20 7
01m
0.2m
vt
<0.1
vt
0
;0.1
0.0
-0.0
0.1
-0.2
+o.i
io.i
0.0
-0.1
<0.1
+0.2
-0.5
<0.1
+5.1
-7.9
+oa
HI. 3
<0.1
<0,1
<0.1
0.0
-0.1
<0.1
0
0
0
0
0
0
0
0
52.2
-64.6
<0.1
0
0
0
0
0
<0.1
0
-------
Pillar Point
Polyaiphenia
hendryi
P.
pacifies
P.
•P-
Porphyra
•P-
Ptaroaiphonia
bipinnata
Pterosiphonla
Bp. CjuV.)
Ptilota
filiclna
KhcHJymrr.ia
•p.
Sp«Tmatophyf a
Phylloapadi*
•couleri
Pcrifera
•pp.
Onldarla
Hydroioa
•pp.
Neaertea
•PP.
208
+ 0
o.nit
0.2a
wt
•¦0.1
<0.1
<0.1
<0.1
<0.1
<0.1
+Q.3
-a,6
<0.1
wt
0
0
0
0
0
n
0
0
+2*.l
-57.3
~
1223.9
-1685.2
2.1 <0.1
+ 5.4
0
-------
Pillar Point
Weaatoda
ipp.
Molluaca
Aaphinetira
Tonicella
llneaca
Gastropoda
Acmaeldae
¦p. (Juv.)
Lacuna
variegata
Thais
laselloaa
Blvalvla
Ad ul a
calfornienala
Muaculus
pypueus
Mysella
tuaida
Mytil ua
¦P- (1uv.)
Annelida
Ollgochaeta
•pp.
209
+ 0
0.01a
P2.0
±203.6
vt
<0.1
0.2a
#
0
wt
0.5
±0.1
0.1 <0.1 o
n.2
-------
Pillar Point
Polychaeta
spp,
Aren'colidae
Bp.
Branchiomalaane
»incenti
Cap1 tellidae
Capltella
capitata
Mediouatua
sp.
Cirratulidae
Cirratulus
cirratus
Eunleoldea
ap.
lAiabrinerldae
Luasbriiieris
sp,
Nereidae
Hmrei*
pelaglca
H.
•p.
Orbiniidae
ap,
Nalnerls
dendritica
Fhyllodocidae
Eteooe
looga
210
+ a
0.01m"
wt
0.2
±0.4
20.4
47.5
0.1
±0,1
0.4
±0.8
0.2
±0.7
0.2
±0.5
0.1
±0.2
4.8
±R. 6
0.1
±0.2
0.2
±0.5
0.2
r0.5
1.6
±3.9
0.1
±0.3
<0
<0
<0
<0
<0
<0
<0
<0
<0
<0
<0
<0
<0.1
-------
Pillar Point
Eulalia
quadrioculata
Phyllodoce
uaculata
Polytioldae
Halosydua
brevlsetoaa
Sabell ?riit»ae
sp,
SabellIdas
sp.
Schiro' -anchia
ir.aignls
Splonidae
MalMocerus
gluteus
Polydora
eolu*biana
P.
liairola
P.
sp.
Pygcspio
elegans
Syl1ldae
Branla
brevlpharvriRea
Exogone
lourcl
E,
sp.
Syllls
spp.
2 1 1
~ 0
O.Oli
0. 2m*
1.2
*3.1
0.2
±0.7
0,1
±0.2
0.1
+ 0,2
t8:J
0,1
±0.3
o.i
±0.2
1.7
±5.1
1.3
±4.1
5.4
±15.1
0.8
±2,1
0.1
±0.2
0.1
±0.3
0.1
±0.2
2.5
±4.7
wt
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
• 0,1
<0.1
<0.1
#
0
wt
0
0
0
0
0
0
0
0
0
0
0
-------
Pillar Point
Terebe111 due
sp.
The 1 epus
cr1spus
PTCnofconida
H«losoma
compact u»
Crustacea
Ost ricod*
spp.
Cirr ip€dia
Balanus
sp, (Juv.)
CiMcet
Cunel1 a
vulgaris
Tana * daces
At .t anal s
nonaan i
Leptoche1i a
dubia
Pancolua
cal lfom 1 ensi '
Isopoda
F.xosphaerowa
acipl icauda
t
¦edia
rtioaburus
0.01m*"
* wt_ ut
0.1 <0.1
~0,4 n
0.2 <0.1 0
*0,6
0.2 <0.1 0
~0.7
0.2
tn.f>
o.i
t 0.2
0.7
*2.2
0.4
- 1 .6
0.2
*0.5
0.2
to. S
0.2
-~0.4
0.1
~ 0.2
n.i
to.3
-------
Pillar Point
Otor lsoaphaeroma
o r r (ton en 9 e
lanirorsis
klncaidi
Idotea
•one erevensis
I.
«p. (1uv.)
stephensenli
tophlpoda
C- •
compact u
CnuridM
¦PP.
Dec n pod *
Oncer
oregonens is
Crypto1i thodea
sitchensls
Pagcrus
hifsutiusculus
Pagettia
gracllis
Insecta
Diptera
spp. (juv.)
Brvo*oa
®PP ¦
2 n
+ n
0.01m*
0. 2m1
0.4
±1.0
o.i
±0.6
1.0
*2.2
1.3
±2.8
0.1
±0.2
0.1
~0.2
5.1
±5.6
n
0.1
±0.2
0.1
±0.2
0.1
±o.i
wt
<0.1
<0.1
0.1
±0.1
±0.1
<0.1
<0.1
<0.1
<0.1
0.0
±0.0
0,4
+n, 3
£
0
0
33.5
±53.2
0
wt
8.0
±14.7
0.5
±0.6
0
0.3
±0.5
1.3
±l.n
2 , 3
±3.5
0.4
±0.7
0.3
±0.2
0.4
±0. R
2.6
±2.2
0.2
:0.4
<0.1
-------
Pillar Point
Specie* richness 124
Diversity - H' 2.CM
Total nu»*ier 3910.6
Total bloatas 2843.8
+ 0
-------
. a r i * i J , i * ; i r IV i r, • ' fine-".--..:. hi-nt'i
>. 1 Itfv. ten . . June Id1", ixprcSitd as n ich
*¦ s r 3 n J a r a ,I«*v i ri t : or. (N = -'«>.
Cn idjria
Ant horoa
-.pp.
Platyhel*lnthes
app,
NcatrCri
¦pp.
?ia
• p.
Avpn i ssa
coli»Mana
Calrptraea
f#.sc igiaea
Lacuna
varlegaca
¦' te it; i sus f rora the subtidal zone
,inJ«Vr S i .im.iss ( pj per O . "»5 rr.~
-5ra
-10m
wt
wt
l.fi
+ 1.7
0,1
±0.1
o. 3
to.1.
<0.1
12.0
±4.3
<0.1
6.0
:2 . 7
<0.1
19.3
10.1
<0.1
4.8
±2.1
<0.1
0.5
±1.0
<0.1
0,3
~0.5
0
<0,1
0.3
*0.5
n
-------
Pillar
Marparites
pupi1lus
Thais
las*llosa
Bivalvla
CI tnocard lm
nuttal111
Lu c 1 n oaa
tenulsculpta
Hacom
nasut a
w.
sp, (juv . )
Xya
aie^aria
Krsella
tualda
So len
sicirlus
Tellina
•PP.
TranMnnella
tantilia
Tie&ui
c*pax
Vi ne 11 d a
°] 1 eochaeta
son.
Subtidal
-5m
-1©B
0.3
'0.5
0
vt
<0,1
2.1
<1.7
0.3
~n.5
vt
o.r
<0.1
-------
Polvchjeta
Arabellidae
Arabella
tricolor
Capitell Ida#
Capltella
cap Ilit¦
Medionastus
Bp .
Motor^ast?is
1 .ncatu.*
n .
tenuis
Chaetooterldae
Splochaetopteru*
coatarua
CIrrat ulIda*
¦ p.
Clrratulus
ci rrat u«
Chaetotone
aetoaa
Dcrri1lefdue
ProtodorviIlea
uracil la
Clvcerldae
CIycera
ni«r lcsna
ron lad ldae
CItc inde
•rwigera
Subtldal
-5-1
-10m
wt
wt
0.3
±0,5
<0.1
43.5
±8.7
01.5
*57.3
n
cO.l
<0.1
3.3
±3.8
19.0
±10. 6
0.3
±0.5
0.3
±0.5
<0.1
-------
Pillar
(Von iada
Mculita
He«ionidae
. A
Gyp t i 3
brevlpalpa
Mieropodarkr
dubla
Luabrincrldae
Lurtrinerls
•P-
Maldanld
Fudywnt
Bp.
^•phtyidae
*«phtra
caecolde*
cal ifomtensis
H .
cl1 lata
S.
Bp .
Here Idae
Sere is
ap. (Juv.)
PIatvnereis
b? canalieulata
i"Viuph i dae
'Viuphla
elegant*
Subt idal
-5m
-10m
wt
wt
1.0
~''.8
0.5
M .0
1.3
~1.3
3.1
±3.6
17.3
-fl. 5
<0.1
n
0.3
~0.5
2.0
±1.6
5 .0
±4.2
0.5
~0.6
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
*0.1
0.3
i0.5
0.5
±0.6
1.3
±1.0
1.0
±0.8
0.3
±0.5
4.8
r2.5
0.5
~1.0
0.3
10.5
0.3
~0.5
0.8
0.8
±1,0
2.8
±3.1
1.3
±0.5
<0.1
<0.1
<0.1
<0.1
<0,1
-------
Fillar "oint
f^rhln1 ldae
Scoloploa
¦P-
0#*ni idae
Myriochele
o'-ulat a
f^enia
fualforal¦
Parson Ida#
Aricidea
ap.
P«ctinarlIdae
PectInarla
grcnulata
Phyllodoce
Fteone
lotiga
F,
tubcrculata
Eulalla
vlrld is
Ph
-------
H.
lunulata
S«*ellidae
Chrme
acaudata
Potmi lis
¦yrlopa
Sabellm
«#dia
Seal lorefEBldae
r>cal ibrtg*a
inflaltsiB
SIgallonIdHe
Pho!oe
¦Inuta
Splraldae
Halaroceroo
glutaeua
Polydora
sociallis
Prionojpio
clrrifera
P.
¦teenscrupi
Splo
f1licornis
Sfl1ldae
F.xofcme
•p.
Piotiosyilis
ur»M
S uhtIda
-5m
•1 t®
^,1
: 0. S
0
0.3
±0.5
A. 5
11.0
0.3
t 0. 5
150.0
~3*.3
0.8
* 1 .0
a
wt
0. 1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.i
n,:
n. 5
0.3
..5
0
0,3
±0.5
0.3
±0.5
0.3
±0.5
0.3
±0.5
0.8
±n.i
0
3T.5
±18,3
0.3
-0.5
0.3
~0.5
<•1.3
<0.1
<0.1
<0.1
<0.]
<0.1
¦0,1
-------
Pillar Point
Streptosvlx is
1 at 1 p.'l pa
«Jy 11 iH *
Ir- i-c ;lrrata
'•-ills
¦ DP .
fTnnimren
r" ' - ico si a
• pp.
.epccstraca
Neballa
puget t mis Is
Craatea
Dlaaty1 is
*p.
DiasE vlopti *=
sp.
Lawpropidae
Tanaldacea
Anatanals
norvanl
I.^PLcche. 1 a
dub i a
1 nopoii;
*.ep,a
sm«rl(.a
' 'otea
sp. ( j UV. 1
7.. r 1 ropi i «s
n n ,
SubtIda I
-5b
¦ lOm
0.
to.
0.
~0.
ft,
to.
wt
<0.1
'0.1
<0.1
#
o
0
2.5
12.7
wt
<0.1
SI .8
• A
0.1
* «">. 5
1 .0
t2. o
4.1
±2.9
ft. 8
110. i
i c 1
•17 2
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
11.5
14,7
0
<0.1
0.3
~0.5
<0.i
0.3
±0.5
0.3
tO.5
0.3
to 5
5.5
<3.1
0.3
~0. 5
0.3
±0.5
0
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
-------
Jaeropsis
d tibia
Munna
ap.
Par^nthura
cleft*""
^ynidotea
bicuspids
S.
sp. Ouv.)
A*phipoda
Can .idea
*P
roti > ¦
sp.
Caprrl1
Capre II i»
sp.
Trltella
pilinana
Decapods
Va1Ida*
ap. (1uv,)
Pinnothet idae
sp.
Pagurtdae
spp.
Puge ttla
Rracilis
PhoronIda
spp.
Suhtldal
-5m
-lira
0.5
±1.0
0,5
i] ,0
0.3
*0.5
0.5
~1.0
0.3
tO.5
114. 5
±34.3
2.0
±2 , 5
0
0.5
±1.0
1.3
±1.5
0
] .6
R.5
5.1
wt
<0.1
<0,1
<0.1
<0.1
<0.1
0.1
±0.1
<0.1
<0.1
-------
Pillar Point
Fchinoderwat*
Opbiuroidea
sp. (juv.)
Holothuroidea
J-eptoaynapta
clarki
Species Richness
nivcraley - H
Total nuaber
Plant blcwiss
Animal hloaass
Total Biomass
223
Subtidal
' ',31
-1 Hn
wt
vt
1.3
11.5
n. 3
¦¦1.5
«\1
77
2 .82
690.7
0.0
<7.9
<7.9
86
3.26
221.3
0.0
<8.4
<8.4
-------
«:>• Ja. North Beach Cobble (sand-gravel-. ori;,i:iisns from the high (•+•6') intertidal
zont collected spring i477, expressed ,is number and./or I? i .miss (g) per -.'IS (N"=20), 0,05 (N=4), and
'".2 m*- +¦ standard aeviation, ^ _
0.01 a 0.2 0.05 ¦ * 15 c«
# wt # wt # wt
srtM 0 0 1.5 <0.1
¦pp. ±1.3
.toda 0 0 1.0 <0.1
¦pp. ±1.2
Kollusca
'3«fitjT>poda
AoMMldM 0 0 0.3 <0.1
¦p. (Jut.) 10.5
Colli Mil* 0.1 < C.l 0 0
digitalis 10.2
LIttorina O.k 0.0 0 3^.5 1.7
scutulata ±1.6 ±0.0 ±37.3 ±1.8
L. 0.1 <0.1 0 7.8 1.1
•ltkaoa ±0.2 ±7.5 ±1.1*
0 0 0,5 < 0.1
p*r*cna ±1.0
Thais 0 0 0.3 0.1
lniUoia ±0.5 ±0.3
Annelida
OllgochMta 0 0 7.0 <0.1
¦pp. *5.7
Polyefaawti.
Reraidac
let* is 0 0 6.3 <0.1
rexllloaa ±6.2
I. 0 0 1.5 < 0.1
¦p. ±3.0
Syillda*
Syllis 0 0 0.8 < 0.1
ap. ±1.0
-------
Sorts Befedi Cobble +6
0.01 m 0.2 m 0.05 »3 x 15 cm
# wt t wt 0 vt
C-ustaeea
Clrrlpedia
Balasus
0
0
16.0
0.9
glaadula
~26.1
~1.8
Hith—' uj
0.2
< 0.1 0
6.8
0.1
dalli
±0.7
~12.8
~0.2
Isopoda
Enxphurou
0
0
18.3
< 0.1
¦e4ia
113.1
/¦¦phipoda
g&isaarldea
0
0
0.3
< 0.1
8pp.
?0.5
Specie* rlchMsc, 16
mnrrnitj -H' 1.99
Total ruber 537
Total Mcmii 19-5
J ? 5
-------
T^ble lb. North Beach Cobble (cobble over sand) benthie organ isms from the mid ( + 3') lntert idal zone
collected spring 1977, expressed as number ind/or biomass (g) per 0.01 (N-20), 0.05 (N»4), and
0,2 (N«4) + standard deviation.
0.01 u2 0.2 m2 0.05 m2 x 15 cm
§ wt # wt f wt
<0.1 0 0
poly» lptuclM
Polyslptaai* <0.1 0 0
Caiilarla
ABtboplrar* 0 0.5 0.2 0.3 0.1
•Jjvaatlsalm *1.0 ±0.k ±0.5 ±0.1
0 7.3 < 0.1
•pp. ±3.8
Mollti»ca
Aiphlanrt 0.1 < 0.1 0 0
¦p.(Jot.) ±0.k
Gastropoda
Acsaeldae 0.7 < 0.1 0 0
vp. (Jut.) ±1.3
OolllMlls 0.5 0.C 0 0
pelt* ±0.9 ±0.0
C. 0.5 0.0 0.3 0.1 0
ftrlgmtella ±0.8 ±0.0 ±0.5 ±0.2
Littorla* 2.2 0.1 0 0.1
•cntalit* ±2.9 ±0.2 ->¦ .3 ±0.1
LittoriM k.k 0.3 1.8 0.9 31.3 1.1
altkana ±l.9 ±0.6 ±2.k ±1.5 ±26.1 ±0.6
0.1 0.0 0 0
fMMtrmtft ±0.2 ±0.0
1. 0 0 1.3 0.1
pcrsaaa ±2.5 ±0.2
I. 0.3 0.0 0.3 0.J* 0
sevtw* ±0.8 ±0.0 ±0.5 ±0.8
-------
Notoaeaea
sp.
Searlcala
dira
Thais
enarglnata
T.
lanellosa
T.
ap. (Jmr.)
Eivalvia
Hysella
tunlda
Jfytilua
•p. (Jut.)
Protothaca
¦taalcwa
Annelida
Ollgocbaeta
•PP-
Poljchacta
Capltallidae
Capltella
capltata
Clyceridae
Heaipodua
bo re alls
Luabrlaeridae
Lumbrlneris
•P-
>»orch Beach Cobble -*-3
0.01 a* 0.
vt 1
0 0.8
~1.5
0 n. 3
±0.5
0 0.3
±1.5
0 0.3
±1.5
0 0.5
±1.0
0.1 <0.1 0
±0.4
0.1 <0.1 0
±0.3
0.1 <-0.1 0
IP. 2
0
227
2 2
n 0.05 n x 15 ca
wt # vt
0.2 0
±0.4
0.8 0.5 0.3
±1.5 ±1.0 ±0.6
0.3 0
±0.5
0.1 0.5 0.3
±0.2 ±0.6 ±0.4.
0.2 0.3 <0.1
±0.4 ±0.5
0
0
0
3.8 < 0.1
±1.7
±0.5
0.3 <0.1
i0.5
0.5 < 0.1
±1.0
-------
North Beach Cobble +3
3.01 n
# wt
Nereidae
Me re ii
vzxllloaa
M.
¦P-
Spionidae
Polydora
proboscidea
Pygoepio
elegana
Syllidae
Sjrllls
sp.
Cruntacea
Cirripedia
Balanus
cariosus
B.
glandula
Chthanlu*
dalll
Iwfodt
Exosphae rona
Media
Gooriaogpliaeroaa
oregoncn*c
Asp hi pods
ganaridea
•PP.
Decapod*
Hcaigrapaua
nudus
0
0
0
0
0.4
±1.3
0.3
±0.7
0.3
±0.9
0.1
±0.2
0
0.1
±0.3
<0.1
< 0.1
<0.1
0.1
to.2
0.2
±0.7
0.2 n"
# vt
0.05 m * 15 fa
P vt
0
0
2.8
±2.2
4.8
±5.*
<0.1
0
0
3.8
±4.3
0.3
±0.5
<0.1
< 0.1
6.3
±4.3
<0.1
0
0
o
0.5
±1.0
2.3
±4.5
0
0.1
±0.?
0.1
±0.2
0
0
48.5
±40.8
19.0
±22.0
0.0
±0.1
0.0
±0.1
0.5
±1.0
<0.1
1.3
a.Q
4.6
±5.6
0
228
-------
North Beach Cobble +3
0.01 as2 0.2 m2 0.05 m2 * 15 eta
Wt 1 Wt # wt
Insects
Dlptcra 0.2 <0,1
larva ±0.5
Species richness 38
Diversity -H* 2.48
Tocsl mariner 950.5
Total bloaass 32.1
229
-------
Table 3c- Worth Beach Cobble (cobble over sand) benthic orfcmi.sms from the low (+0') incertid.il zone
collected spring 1977, expressed ss number and/or biotnass (g) per 0.01 (N-20), 0.05 (N-4), and
0.2 (N-4) bi + standard deviation. -
0.0) m 0.2 m 0.05 m % 15 en.
t wt * wt # Wt
ThlorophytJi
Cladophoraceae <0,1 0 0
ap.
Enterooorpha <0.1 0 0
llnza
Mcioatrcma 0.0 0 0
fuactm *0.0
M. 0.2 4.3 0
•pp. t0.6 *8.5
Ulva 0.9 8.2 <0.1
¦pp. ±1.1 ±8.0
Bacillarlophyta
Patmales <0.1 0 <0.1
Phaeophyta
Alarla 5.5 142.6 0.0
app. *13.1 ±212.0 ±0.0
Anallpua 0.6 0 0
Japoalcua ±1.6
Deamraatia 0.0 0 0
aculeata ±0.0
Ectocarpua <0.1 0 0
ap.
Fucita 0,1 13.3 0
dlatxchua ±0.3 ±21.0
Laainarlalea 0.0 0 0
ap. ±0.0
Laathaala <0.1 0 0
dlfforala
Maraocyatls 0 31.3 0
luatkaana ±62.6
230
-------
North Beach Cobble +0
Petalonia
faacla
Kalf&iaeeae
Sphacclaiia
faconia
Rhodephyta
Antithanmlcm
defectum
Antlthaaolooalla
paciflea
Caillthaanlon
/Pleoaosporlun
Cerasiua
paclflcuB
Choreocolax
polyalphonlAe
CrytofMtssla
•p.
Clgartlna
paplllata
Glgartinaceae
spp.
BarpoaIphoala
pltmula
Laureocla
¦pectab11Is
Hoilenb«r gia
aublata
Hyrocnena
epp.
0.01 »
# %ft
0.2 nT
* wt
0.05 to *15 cm
« wt
<0.1
r
<0.1
~
0
< 0.1
<0.1
0
0
0
0
< 0.1
<0.1
<0.1
0.0
±0.0
0.0
±0.0
0
< 0.1
0
<0.1
0.0
±0.0
0
0
¦1
0
2.2
±3.8
0
0.0
±0.1
0
0.3
±0.5
0
0
< 0.1
0
1.7
~2.6
0.4
±0.6
0
0
0
0
!31
-------
North Beach Cobble +0
0.01 a"
# vt
0.2 a
# vt
0.05 n>~ x 15 era
# vt
Iridaej 2.0 80.7 0.3
cordata ±4.5 *79.2 ±0.6
I. 0.1 0 0
app. ±0.1
Odoethalla <0.1 0 0
floccoia
O. <0.1 0 0
¦p.
Patrocclla / /
¦iddendorfii
Pikaa 0.0 0 0
caiiforalc# ±0.1
Plocaraiura 0.1 0 0
cocclneua ±0.2
P. 0.1 0 0
app. ±0.1
PoIyncura 0 0.1 0
latiaclaa ±0.2
Polyalphoola 0.0 0 0
heodryl ±0.0
P. 0.1 0 0.2
pacific* ±0.3 ±0.2
P. 0.1 0 <0.1
paniculata ±0.4
Porpbyra 0.0 <".l 0
app ±0.0
Ptaroaiphxila 1.7 0 0,2
biplxmals ±2.3 ±0.2
P. <0.1 0 0
dandroldaa
! 32
-------
North Beach Cobble +>")
0.01 a
f wt
0.2 n'
# wt
0.05 w*" * 15 ca
9 wt
Scageit* <0.1 0 <0.1
occidentals
Sperwatophyta
Ph-rlliapadlx 0 0 0.0
acoulcrl ±0.0
Platyheialnthea
Turbellaria
•pp. 0.6 <0.1 0 0
±1.6
Ncaertu 0.3 <0.1 3 1.3 <0.1
¦pp. ±0.9 *2.9
Noaatoda 0.6 <0.1 3 5.8 <0.1
¦pp. ±1.4 ±4.0
Molluaca
Aaphlnaura 0 0 0.5 < 0.1
¦pp. ±0.6
Cyanoplax 0 0.3 0.0 0
dentlena ±0.5 ±0.0
Gastropoda
Acaacidae 0.2 <0.1 J 1.8 <0.1
spp. (Juv.) ±0.7
Aaphlaaa 0 0 0.3 <0.1
colunblaho, *0.5
Col)iaella 0 0.3 0.3 0
pclta ±0.5 ±0.6
Lacuna 7. 7 0.0 0.J <0.1 5.8 0.0
~arlegata ±7.1 ±0.1 ±0.5 ±2.8 ±0.0
Notoacau 0.1 <0.1 0.5 < 0.1 0.3 0.0
scutua ±0.2 ±1.0 ±0.5 +0.1
Onchldclla 0.1 <0.1 0 0
boraalla ±0.2
233
-------
Blvalvta
Macoma
¦P-
Mya
arenarla
Myaella
t ib» Ida
Mytilua
•pp. (Juv
Venerupls
japonica
Annelida
Ollgochaeca
•pp.
Polyctiaata
Aranlcolldae
Capltallidae
CaplCella
capicata
Medlomastua
¦p.
NotcHsaACus
Canula
H.
¦p.
Cirratulldae
app. (Juv.
Cirratulua
clrratua
North Beach Cobble +0
0.01 m
0.2
¥t
Wt
0.05 m x IS cm
9 wt
<0.1
0
0
0
0
0
<0.1
< 0.1
<0.1
0
0
0
0
0
0
0.
+ 0.
0.
±0.
0.
to.
0.
±1.
0.
to.:
15.8
±7.0
0.3
±0.5
o.a
±1.0
0.3
to.5
0.5
tl.o
0.3
±0.5
1.8
+ 3.5
0.3
±0.5
< 0.1
< 0.1
<0.1
< 0.1
1.9
±3.9
< 0.1
<0.1
<0.1
< 0.1
<0.1
<0.1
< 0.1
<0.1
-------
f
North
Beach Cobble +0
0.01 m2
i -> 2
0.2 m
0.05 m2
x 15 cm
f
wt
# wt
0
wt
Tharyx
0
0
1.0
<0.1
¦ultifilis
il.4
Dorrilleldae
Protodorvillea
0
0
0.3
< 0.1
gracilis
±0. 5
Eunlcidae
Eunice
0
0
1.4
< 0.1
valena
±2.0
Clycerlda*
Hcalpodus
0
0
6.0
<0.1
borealIs
±0.8
Lunbrlneridae
Liabrlnerla
0
1
<0.1
0
0.3
<0.1
±0
4
±0.5
Mereldae
0
5
<0.1
0
0
®PP-
il
4
Merels
0
1
< 0.1
0
0
vexlllosa
10
2
n.
0
5
0
0
¦pp.
±1
6
Platynerela
0
1
O
V
0.3 <0.1
< 0.1
bicanallculata
±0
3
to. 5
±1.0
Onuphldae
0
2
<0.1
0
0
ap.
to
7
Onuphia
0
0
5.0
A
o
H*
aclgaatla
±3.8
Ophallldae
Araandia
0
0
0.5
< 0.1
brevls
±0. 6
2 35
-------
North Peach Cobble +0
0.01 a
# vt
0.2 a
> wt
0.05 m x 15 cm
& wt
Phyllodocidac
Eteooe
long a
Hcalonura
coineaul
dlfflcilia
Phyllodoce
Maculata
Phyllodoce
»p.
Polyooidae
Hanaothoe
tabrlcata
S~ lidae
spp.
Fabrlcia
oregonica
Spioaldae
Malococeros
glutacua
Polydora
Columbians
P.
proboacidea
Pygoapio
clegans
Splo
f lllcomls
Spiophancs
bonbyx
Q
0
0
n.i
±0.3
0.1
±0.2
0.5
±2.0
0.2
±0.7
<4.2
±17.4
0.7
~2.2
0
<0.1
< 0.
< 0.
< 0.
< 0.
< 0.
0
0
0
0
0
0
0
0
3
0
0
0.3
±0.5
0.3
±0.5
0.3
±5
0.3
~0. 5
0.3
+ 0.5
0.3
±0.5
0.3
±0.5
12.5
±11.7
0.8
±1.5
4.0
±73
0.5
+0.6
0.5
±1.0
2.0
±2.2
<0.1
<0.1
<0.1
< 0.1
<0.1
< 0.1
<0.1
<0.1
<0.1
< 0.1
<0.1
<0.1
<0.1
236
-------
North Beach Cobble
Syllidae
Exogone
geraalfera
E.
lourei
Syllls
•PP-
Terebellidae
¦PP-
Crustacea
Ostracoda
•PP*
Clrripedla
Ealanua
carlosus
Ba.'.mua
glandula
B.
ap. (juv.)
Mysldacea
Archecnysis
grebnltzkli
Cuaacea
CumI la
vulgaris
Laapropldae
spp.
Tanaidacea
Leptochelia
dub la
0.01 m
wt
0.5
±1. 3
0.7
±2.2
0.2
t0. 7
1.2
±4.7
0.4
±0.8
0.1
±0.4
0.1
±0.4
0
<0.1
<0.1
<0.1
<0.1
<0.1
0.0
±0.1
< 0.1
0
0
0
0
0
0.1 <0.1 0
±0.3
0.1 <0.1 0
±0.3
0.1 <0.1 0
+ 0.2
0.1 <0.1 0
+ 0. 3
23?
+0
0 2
Wt
0.05 m" x 15 cm
# wt
0.3
±0.5
1.0
±1.4
6.0
±2.9
2.0
±2.3
<0.1
<0.1
< 0.1
<0.1
0
0
0
0.3 < 0.1
±0.5
0.5
±1.0
0
0.3 <0.1
±0.5
0
-------
laopoda
Exoaphaerona
aspl 1 cauda
Exosphaeroaa
¦edlfl
E.
onetone«¦
E.
•pp. (Juv.)
Gno r Iibo s p ha e r oaa
orgonense
Idotea
¦mtereyenala
i.
wogneaenskil
I.
spp. (Juv.)
Huana
ehronatocephala
Anphipoda
Gawsarldea
•P
Decapods
Cancer
oregonensiB
CrangOD
Bucltella
North Beach CobbK +0
n.01 m
wt
0.2 n"
¦? wt
tn x 15 cm
wt
0.2
±0.7
3.7
±8.2
6.
tl3.
1.
±2.8
n.5
±1.0
4,
±6.
fi.
113.1
0.2
±0.5
<0.1
<0.1
< 0.
< 0.
< 0.
< 0.
< 0.
< 0.
0
0
0
0
T
1.0
±0.8
0
0
0.1
±0.1
0.5
±1.0
3.n
±2.4
0.8
±1.5
0.5
±0.1
0
0
4. a
t3. 7
n
<0.1
<0.1
<0.1
< 0.1
<0.1
0
0.3
±0.5
0.0
to.I
15.8
±6.2
<0.1
0.1
tO.2
0.3
tO.5
< 0.1
0.2
±0.4
0
o
o
0
238
-------
North Beach Cobble +0
0.01 m2 0.2 n2 0.^5 m2 x 15 cm
vt ? wt * wt
Iiuccta
DlpUran 0.2 <0.1 0 0.8 <0.1
larvae ±0.5 *1.5
Bryozoa
Bryotoa 0 0 0.3 <0.1
¦pp. ±0.5
Species richness 125
Diversity -H' 3.24
Total traafeer 1681.1
Total blOMSS 465.6
2 39
-------
Table «,i. Morsu Creek (sand-gr.ive i -cobble) bent hie urg.ini sro.s from the hi-*h (+<'¦>') i nt ert i ¦
zone i'i < 1 1 e»" t ed spring 197? through winter 1978, expressed as number ,mcl/or niorass (•'
pi-r 0.01
f. 0 0 0 0 0 ~~0.3 < 0.1 0
ap. A *0. *¦
Oligocha«ta 0 0 0.3 <0.1 0 0 2.C < 0.1 0
app. *0.5 *2 .8
Crustacea
Cirrlpadia
Baianus 0 0 0 0 0 0 0
plandula
B. 0 0 0.1 «. 0.1 0 0 0 0
•pp. (luv.) +0.5
:'>o
-------
.)a]
I
Pall 77
Win 78
0.05 «2
0.01 »2
2
0.2 ¦
0.05 m2
0.01 m
2
0.2 ¦
0.05 e2
J? vt
# wt
# wt
# vt
# wt
# wt
#
wt
0.3 < 0.1
0
0
0
0
0
0.8
<0.1
51,5
+0.5
0.3 < 0.1
0
0
0
0
0
1.0
<0.1
*0.5
+1.4
0
0
0
0
0
0
0.3
< 0.1
+0.5
0.3 < 0.1
0
0
0
0
0
~0
+0.^
~0
0
0
0.3 < 0.1
0
0
0
+0,<
0
0
0
~0
0
0
4.0
0.4
+3.4
+0.2
0
0
0
0
0
0
~0.5
0.0
+1.0
+0.0
0.3 < jo.l
«
0
0
0
0
0.3
< 0.1
+0.5
K>.5
0
c
0
0
0
0
0.3
< 0.1
+0.5
0
{
0
0
0
0
~0.5
< 0.1
+1.0
0.8 0.1
0
0
0
0
0
0
+0.5 4o.i
"0.3 <. 0.1
0
0
0
0
0
0
+0.5
0.3 < 0.1
0
0
0
0
0
0
+0.5
~0.3 < 0.1
0
0
0
0
0
0
+0.5
2.0 < 0.1
0
0
0.5 < 0.1
0
0
1.0
< 0.1
+2.ft
+1.0
+1.2
0 I
0
0
5.5 0.3
0
0
20.3
o.e
+11,0 +0.5
+33.0 +0.b
0 1
0
0
"0
0
0
0
240
-------
Morae Creek +6
Spr 77
Sua 77
0.01 «7
0.2 m1
0.05 m7
0.01 m2
0.7 m2
0.05 HI2
# Mt
# «t
# vt
# wt
# wt
# wt
Crustacea (cont,)
Clrripedia
ChthasMlun
0
0
0
0
0
0
dalli
Myaldacea
0
0
0
0
0
0
¦p. (|uv.)
Tanaldacea
Leptichelia
0
0
0.3 * 0.1
0
0
0
duhla
laopoda
EitasphaercMia
0
0
0.3 < 0.1
0
0
0
sp,
+0.5
Tnorlwoaphaeros*
0
0
"0
0
0
16."* 0.2
oreponense
+32.5 +€,3
CuMCtl
0
0
0
0
0
0.3 < 0.1
m
TI
1c
<
V
*o.*
Aaphipoda
Caaaarldea
0
0
5.5 4 0.1
•r
-
-
up.
+1.9
Allorchestes
-
-
-
0
0
0.3 € 0.1
angustus
+0,5
Corophlua
0
0
0
0
0
0.3 <, 0.1
hrevls
+0.5
C.
0
0
0
0
0
~o
sp.
Orchestls
-
-
-
0
0
0.3 < 0.1
¦P.
+0.5
Orchestoldea
m
-
-
0
0
~1.5 4 0.1
pupettensl*
+ 1.7
Faraaoera
w
-
-
0
0
99.0 0.2
colimMana
+125.4 +0.
Inaecta
Oipteran
0
0
0.3 40.1
0
0
0
larvae +0.5
Species Richness B 16
Diversity, Hl 1.31 0.76
Tota L Musber 51.5 614.5
Total RIomii (y) 4 0.H <§.0
-------
Cr**k
JO.01 ¦
# wt
Fall 77
0.2 .2
# vt
0.05 ¦
# «t
0,01 ¦
# wt
Vif 78
0.2 b2
f vt
0.05 •
# wt
0
0.3 <. 0.1
+0.5
0.5 <0.1
+1.0
7
0.4*
313.5
<5.0
15
1.72
211.0
<11.5
0.3 « 0,1
+0.5
16.3
+32.5
0.3 t ft).I
+0.5
0
0
0.5 40.1
+1.0
0
0
0
0
5.8 < 0.1
+7.7
o.3 < §o,i
+0.5
"0.3 4i
+0.5
"0
0.1
0.3 < 0.1
+0.5
~1.5 4 0.1
+1.7
~99.0 0.2
+125.4 +0.1
0
0
0
0
55.3 0.2
+31.5 +0.2
0.3 <0,1
+0.5
0
0
0
0
0
0
0
0
6.3 < 0.1
-MS. 3
~0
0.3 <0.1
+0,5
4 1
-------
lible 4b. Morse Creek (c-obb le aver sand) bertthic organisms t rom tin- mid (+ J') inli-rt ida!
/one 1 ItU't t'd spring 1977 through winter 1978. expressed as number and/or biomaxs 1^)
jut O.fil (X=2'j), 0.05 (N«4) , and 0,2 (N=4) m ' + standard deviat ion. t
Chlorophyta
Cladophoracsea
sp.
Snteromorpha
1 Irtca
Honoa troma
sp.
r.ponfo«nrph«
coai a
S.
sp.
Ulothri*
¦ p.
Ulva
•p.
Uronpora
Bp.
Baclllarlophyca
Fennalee
• p.
Fhacophyta
Alaria
ip.
Dcamaraatla
aculcata
Fucui
Hiatlchus
Laminarial®*
•PP.
Pctalortla
fascia
Rhodophyta
Acrochaatluw
¦P-
Ahnfaltla
pllcata
Crytoalphonla
woodi I
Endocladla
mi rIcata
ClptartlnacM*
•p.
Gl^artloa
paplllata covplax
0,01 a>
# vt
Spr 77
0.2 m?
# wt
?
0.05 «
# wc
0.01 m?
# wt
Sum 77
0.2 m7
# vt
2
0.0! m
# wt
<0.1 0
<0.1 0
0 0
0 0
<0.1 0
<0.1 0
0 0
<0.1 0
0.0 0
*0.1
0 0
<0.1 0
<0.1 o
0 0
0 0
<0.1 0
0.0 0
+C.1
<0.1 0
<0.1 0
0 0
0.7 0
+1.3
0 0
0 0.2
+0.1
0 <0.1
0 40.1
0 <0.1
0 0
0 0.3
+0.5
0 <0.1
0 <0.1
0 0
0 0
0 0.4
+1.9
0 0,1
+0.5
0 0
0 0
0 0
0 0
0 0
0 0
0 0.?
+1.6
0 0
3.4 0
+3.4
"o 0
0 0
0 0
0 0
5.4 0
+4.S
0 0
0 0
0 o
0 0
38.1 0
+43.*
"0 0
(\3 0
+0.6
<0,1 0
0 o
0 0
0 0
0 0
o.e o
~0.6
?42
-------
g)
I
Fall 77
Win 78
0.05 m
# wt
0.01 m
# wt
0.2 ¦
# wt
0.01
# wt
0.01
# «t
0,1 ¦
# wt
0.05 •
# wt
0
0
0
0
0
0
0
0
0.1
0
0
0
0
0.?
+0.2
0
0
0
0
0
0
0
0.9
+1.3
"c
o
0
0
0
0
0
0
0
0
0
0
0
0
0
*0.1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
? 0
0.1
1 0
J o
I
i 0
0
0
<0.1
0
0
0
0
cO.l
0
0
J
0
0
0.9
+4.0
0
0
0
3.9
+B.9
0.7
+1.4
0
0
0
0
0
0
i0
1°
0
0
I 0
o
0
0
0
0
0,1
0.?
+1.8
0
0
0
0
0
5.0
+6.1
0
0
0
0
0
0
0
0
0
0
0
0.1
40.1
0
0
0
0
0
e.i
*0.1
0
0
0
0
0
0
-------
Rhodophyta (cont.)
Irldaea
cordata
I.
¦p.
Odcnthalla
fIccoasa
Polyalphonla
hendryi
P.
pacific*
P.
ap.
Perphyra
»,p.
Rhodochorton
¦ p.
RSodom*la
larl*
Rhodymenla
¦ p.
Cnldarla
Anthotoa
•p.
Anthopleura
alegantlnnlM
Pl*tryb«lailnth«B
Turbellarla
¦ pp.
K«Mrtw
¦pp.
KaplectonnM
praclla
NaaMtoda
¦P-
Holluaca
A«phlnaura
¦p.(Juv.)
Cyanoplax
dantlana
Gaatropoda
Aeaaaidaa
ap.
Spr 7 7
0,01 m? 0.? c? 0.05 in2
# wt # wt i vt
0 0 0
0 0 0
0 0 0
0 0 C
*0.1 0 0
<0.1 0 0
0 0 0
0 0 0
<0.1 0 0
<0.1 0 0
Marse Cre»k +3
Sua ? 7
2 2 ,
0.01 * 0.7 m 0.05 *z
# Wt # Wt # wt
0 0
0 0 0
0 0 0
0 0 0
CO.I 0 o
0 0 0
0.1 0.6 o
jO.l ±0.7
0 0 0
0 0 0
0 0 0
0.4 <.0.1
+1.1
0,1 <0.1
0.7
1.0
+2.0
~0
0.0
+0.1
0
0
0
0
0
0
0.3 *0.1
+ 0,5
0.1
+0, 5
0.1
+0.2
0.9 <0,1
+3.«
0.3 <0.1
+0.5
O.f* <0.1
-Hi .0
0.1 < 0.1
40.2
~o
0.3
+0.<
"*0.3
+0.5
0.0
+0.1
~0.0
+€.01
i6.o o.e
+4.5 +0.1
0
0.2 <0.1
+0.4
~0.1 <0.1
+0.2
0
0
32.5 <0.1
4-8.5
-o
0.1 <0.1
+0.2
6.5 < 0.1
+8.5
0.6 <0.1
+1.0
26.a
-------
t Creek +J
Fall 77
Win 78
O.CT- m2
4 wt
0.01 m2
* wt
0.2 ¦"
# wt
0.CK ¦
# wt
0.01 ¦
# wt
0.2 ml
# wt
0.05 ¦
# wt
0
0
0
0
0
0
0
0
0
0
0
0
<0.1
0
0
0
0
0
0
40.1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
*0.1
<0.1
0
0
4 0.1
iO.l
0
0
0.5
+1.0
~0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0.3 C0.1
+ 0.5
0
0
0
0
0
0
0.8 0.1
+0.5 +0.1
0
0.8 4
+1.0
32,3 <#.1
+8.5
~0
3.1
26.8 <
+26.6
0.3 4
+0.5
0
1.0 4
±1.4
241
.1
.1
6.6
+ 7.2
0.1
0.1 <0.1
±0.2
0
0.1 <0.1
+0.3
0.6 40.1
+1.0
0
0
0.3 0.0
+0.5 +0.1
6.8
-------
Mora* Creek +3
Spr
77
Sua
77
0.01 m2
0.
2 m2
0.05 m2
0.01 B2
0.
2 w2
0.0'
#
vt
#
wt
~ vt
#
Ml
#
wt
#
Hoilusca(cont.)
Ga« tropoda
Arehidorla
ap.
Colllaella
0
0
0
0
0
0
H{aitalin
C.
0.1
<•0.1
0.3
0.1
0
0.2
0.2
6.3
2.5
0
p®lta
+0.3
+O.S
+0.2
+0.4
+0.8
+11.2
+4.2
C.
0.?
< 0.1
0
0
~0.4
0.2
~0.5
0.1
0
strlpatcl la
+0.4
+0.9
+0.2
+1.0
ip.l
Lacuna
0
0
0
~b.l
<¦"0.1
~0
0
variegata
+0.2
Lit torin*
0
0.3
0.2
0
0
0
0
scutulata
+0.5
+0.1
L.
25.fl
3.7
2.5
"o.s
95.a 5.3
6.6
0.9
0.5
0.1
10.3
sltkana
+•11.9
+2 < 3
+2.6
+p.B
+41.6 +1.4
+ 12.5
• -1
+1.0
+0.2
+12 .3
Margarltes
0
"o
~0.3 <0.1
"o
0
"o
puplllui
+0.5
NotOICH«l
0
3.B
2.7
~~0
0.1
0.0
2.5
1.1
1.0
fenestrate
-1-2.9
+0.9
+0.2
+0.0
+2.1
+0.t
+2.0
N.
0
0
0
0.2
~0.0
"o
™o
peracna
+0.5
+0.1
H.
0
0
0
0.6
"o.l
5.3
1.6
0
acutua
+0.9
+0.2
+5.4
+l.f
Ocanebra
0
0
0
"*0
"o
0
lurlda
Onchidella
O.R
1.6
3.R
2.6
1.3 C 0.1
0
0
0
horaa11a
+1.1
+4.7
+2.2
+1.0
+1.5
Thai*
"o
"o
0
0
0
0
•¦•rpdriaca
T.
0
0.3
0.2
0
0
2.5
< 0.1
0
laaalloaa
+0.^
+i).5
+4 .4
T.
0
0
0
0
~0
J:?
sp.(1uv.)
Blvalvla
Cllnocardiuai
0
0
0
0
0
0
nuttsllll
Ma com
0
0
0
0
0
0
lnqulnata
M.
0
0
0
0
0
0
ap. CJuv.)
Huaeulus
0
0
0
0.1
< 0.1
0
0
pypMaus
+0.2
Myaalla
0
0
3.5 <0.1
"o
0
3.5
tualda
+2.4
+0.6
Mytllua
0
0
"o
0
0
"o
2
¦
%»t
0.5
0.1
+0.1
<0.1
call fornl*nuB
144
-------
f Creak +3
Pall 77
Win 78
0.05
0.01 m1
0.
- 2
2 m
0.05 m2
0.01 m2
0.
7 92
0.05 m2
#
wt
#
we
#
ttt
#
we
0
we
#
wt
#
we
OA
4 0.1
2.5
1.5
+0.2
+3.0
+1.8
0
0.1
0.0
0
0
"o
0
0
+€,3
+0.0
0
"0.2
"0.1
1.8
1.6
0.5
0.0
0.6
0.4
2.5
2.5
0
+0.5
+0,5
+2.2
+1.2
+1.0
+0,1
+0.9
+0,7
+1.9
+3.0
0
"0.4
0.0
~0
"o
~0.6
"0.0
"l .8
40.1
1.0
<. 0.1
+0.8
+0.1
+1.4
+0.0
+1.3
+1,4
0
7.2
0.0
0
0.3
<¦ 0,1
"0.3
4 0.1
0
~0.3
< 0.1
+6 ,4
+0.1
+0.5
+0.6
+0,5
0
*0.1
4 0.1
0
"1.5
0.0
0
0
"o
+0.3
+3.0
*0.1
10.3
0.5
1.9
0.1
0
"3.5
"0,1
5.4
1.2
0.3
0,1
5.5
0.0
+12.3
jjO.5
+2.5
+0.1
+3.9
+0,1
+10,0
+3.4
+0.5
~o. 1
+3.7
+0.1
"o
"0
0
"0
"o
~C
0
1.0
0.1
0.3
0.1
5.8
3,4
0
0.6
1.1
5.8
2.?
0
4-2.0
+0.1
+0,4
+0.3
+2.6
+1.7
+0.8
+3.1
+3.9
+1.7
"o
"o
0
0
0
_o
0
0
0.6
0.3
7.5
4.6
0
0.8
0.3
11,5
5.8
4.5
0.0
+0.9
+0.5
+5.0
+2.4
+1.0
+0.5
+5.5
+2.0
+3.8
+0.1
0
~0
0
0.3
40.1
"o
0
"o
+0.5
0
0
0
"0
0
0
0
0
0
0
0
0.1
*. 0.1
0
0
+0.2
0
0
0
0
"0
1.0
0.8
0
+2.0
+1,7
J.i
<0.1
0
0
0
0
0
0
0
0
0
0.5
40.1
0
0
0.3
0.3
+0.6
+0.5
40.5
0
0
0
~0
0
0
"0.3
"0.1
40.5
+0.2
0
0
0
0.3
<• 0.1
0
0
""*0.5
"o.l
+0.5
+0.®
+0.1
0
0.3
< 0,1
0
~0
0
0
~0
+1.1
3.5
4 0.1
"o
0
0
0
1.3
4 0.1
3.3
4 0.1
•*6,4
+2.5
+3.9
**0
0
0
2.0
4 0.1
0
"O.J
0.1
"o
+2.7
40.5
+0.1
244
-------
Morse Creek +1
Spr 77 Sub 77
0.01 0,2 m 0.05 m 0.01 0.2 m~ 0.05 m
» wt # wt » wt # wt # wt # wt
MolIubci (cont . )
Blvalvla
Hytllua 0 0 0 0 0 0
•dulis
H. 0.1 <¦ 0.1 0 0 0.3 0.0 0 0.3 « 0.1
ap. {.) +0,? +0.7 +0.1 +0.5
Frotothaca 0 0 0.5 <0.1 0 0 0
ataalr.^a 1.0
Trcaus 0 0 0 0 0 0
capax
Annelida
Archlannellda
Saccoclrrldae
Saccoclrrua 0 0 0 0 0 0
¦ p.
Polvehaata
Arenlcolidaa 0 0 1.8 0,1 0 0 0
ap. +2.7+0.1
Aharenicola COO 000
paclflea
A. 0 0 0,5 0.I 0 0 0.5 * 0.1
¦p. +0.6 +C. 1 +1.0
Gapltallldaa
Cap' talla 0.1 <0.1 0 16,fl 0.1 0 0 41.A <0.1
capltata +0.2 +15.4 +0.0 +6.2
H#dlo«Macy8 0 0 1.0 «. 0.1 0 0 0
ap. +2.0
Notoaaatua 0 0 0 0 0 0
tanula
ClrratulIda* 0 0 0 0 0 0
•P-
Clrratulua 0.1 < 0.1 0 6.3 0.1 0 0 8.3 < 0.1
cirratua +0.7 +4.0 +j0.0 +6.9
Dorvllleldae
Protodorvlllaa 0 0 0 0 0 1.0 <0.1
piracllla +1.4
Glycerlda*
Hmlpodua 0 0 0 0 0 0
borealla
245
-------
Creek +3
Fall 77
Win 78
0.05 ¦
# vt
0.01 m7
# wt
0.2 »
# wt
0.05 «
# wt
0.01 ¦
# wt
0.2 m2
# wt
0.05 ¦
# »;t
0
0
0
0
0
0
0.3
40.1
+0.5
0.3 « 0.1
0.
5
0.0
0
0
0.2
4.0.1
0
0.5
«.0.1
+0.5
+0.
8
+0.1
+0.4
+0.6
0
"o
0
0
"o
0
0.8
0.6
+1.0
+1.0
0
0
0.
3
0.1
0
0
0.3
0.0
~0.3
0.1
i°-
5
+0,2
+o, s
+0.1
+0.5
+0.1
0
0
0
0
0
0
1.8
4.0.1
±2 .4
0
0
0
0.5 <.0.1
0
0
0.8
4 0.1
+1 .0
+ 1.0
0
0
0
~0
0
0
0.3
40.1
+0,5
0.5 * 0.1
0
0
0.8 <0.1
0
0
0
±1-0
+1.5
41.B <0.1
0
0
21.5 «.0.1
0.3
4 0.1
0
16.8 40.1
+6.2
+21.0
+0.5
+ 17.8
"o
0
0
0.5 4 O.I
~0
0
"0
+1.0
0
0
0
0.3 <0.1
0
0
0
+0.5
0
0
0
0.3 4 0.)
0
0
0
0.5
8.3 < 0.1
0
0
1.8 < 0.1
0
0
2.0
< 0.1
+6.9
+3.5
+2.2
1.0 (0.1
0
0
0
0
0
0
+1.4
0.5 < 0.1
+ 1.0
245
-------
Morne Creek +3
Spr 77 Sun 77
0.01 m7 0.2 m? 0.05 »2 0.01 m2 0.2 m2 0.05 m2
# W t # Wt # Wt # Wt # Wt # Wt
Annelida (com . )
Polychaeta
Nereldae
Nereis 0.1 <0.1 0 0 0 0.3 <0.1 0
vexl Iloaa +0.7 +0.5
N. 0 0 0 0.1 < 0.1 0 0.3 < 0.1
np. +0.5
flatynerels 0 0 0 0 0 0.3 < 0.1
Alcana 1lculata +0.5
Ophelliilae
Arnatv! 1 a 0 0 0.5 40.1 0 0 0
hrevin +1.0
Ovenlloae
Owen la 0 0 0.1 0 0 0
fus;iormi*
PhyIlodocldae
Eteone 0 0 1.3 0.1 0 0 1 .? (0.1
lonva +1.9 j<). 1 ±1-7
Fula1 la n 0 0.3 0.0 0 0 0
viridls +0.5 +0.1
Sabs 11ldae
Fahrlcia 0.1 CO.I 0 0 0.3 <.0.1 0 0.3 <0.1
oreponlca +0.^ +0.7 +0.5
F. 0 0 0 ~0 0 ~0.3 <0.1
«p. +0.5
Splnnldae
Kalacoceroa 0 0 0 0 0 16.0 <0.1
plutaeua -tI4.4
Polvdora 0 0 0 0 0 0
colunhlana
P. 0 0 0 0.1 CO. 1 0 0
sp. +€.*
Pygoaplo 0 0 0 0 0 0 0
elegant
Scolalepla 0 0 0 0 0 1.0 cO.l
up. C +1.2
Splo 0 0 0 0 0 ~0
f1llcornla
Sy 11 ldae
nyllis 0 0 0 0.1 <0.1 0 0
adanantea +0.1
-------
Cr««k +3
Fall 77
Win 78
0.05 a2
» wt
0.01
# wt
0.2 wf
# wt
0.05 ¦*
# %»t
0.01 B2
# wt
0.2 a
# wt
0.05 b2
# wt
0.3 t 0.1
+0.5
0.3 ( 0.1
+0.5
0.1 <0.1
+0.2
0
0.1
±0.2
0
0.9
+ 1.9
1.3 < 0.1
+2.5
0
0
0
0
0
0
0.3 <. 0.1
+0.5
"0
0.1 <0.1 0
+0.1
0.5
+1.0
<0.1
0.5 < 0.1
+0.6
1.8 < 0.1
+1.7
"0
0
0
0
0
4.5 <.0.1
+2.6
0
3.3 < 0.1
+2,8
"0
0.3 t 0.1
+0.5
"0.3 i 0.1
+0.5
0.3 40.1 0
±1.7
0 0
2.5 <0.1
+3.1
~0
0.1 4 0.1
+0.3
~0
0
0
16.0 < 0.1
-t 14.4
~0
0
0
1.0 <0.1
+1.2
"o
0.5 <. 0.1
+0.8
0
0
0
0
0
0
0
0
0
0
91.5 <0.1
+56.3
2.3 <0.1
+3
0
+€
0
41
0
< 0.1
2
3
5
5 < 0.1
0
0.5 <0.1
+1.0
0.6 < 0.1
+ 1.5
"0
0
0
0
0
0
0
0
0
0
3.0 < 0.1
+2.5
~2.3 < 0.1
+3.9
~0
0.3 < 0.1
+0.5
0.3 40.1
+0.5
"0
-------
Horse Creek +3
Spr 77 Sum 77
0.01 m? 0.2 »2 ( .05 m2 0.01 «2 0.7. m2 0.0? «2
# Wt # Wt # Wt # Mt # %Jt # >t
Annelid* (cont.)
Polvchaeta
Svllldae
Svllis 0 0 0 0.1 < 0.1 0 0.5 < 0.1
•ipp. ^0. ? +1.0
01lKOchaet« 0.1 40.1 0 0.3 <0.1 0.2 <0.1 0 1.0 < 0.1
*pp. +0,2 ±0.r> +0.5 +1.4
Phoronida
Phorenopa is
harwwrl
Crustacea
0a tracnda
•PP.
Clrripedla
Ralanun
carlnaua
B.
glandula
B.
spp, < |nv.)
ChthamaIus
da 11 i
l.l 0.3
+3.4 +l.c
0
0.3 <0.1
"o
0
0
0
0
0
0
3.0 0.1
+6.0 +0.1
"*0
0.1 <0.1 0
+€.3
4.7
0
+1S.P +1
7.0 0
+22.7 +0
175.4 r
+¦> 14 .7 *•
0
I
+2
0
0,1
+0.2
25.0 0.4
M*.9 +0.7
24.0 "0.4
+33.0 +0.6
69.5 "0.3
439.H +-0.2
0
Cuaiacea
Cume11 a
vu 1 pa r i s
Ti aldarea
Anatanal*
norma" 1
Leptochella
duUa
0.1 CO. 1
+0.?
0
0
0
0
0
0
c
0
0
0
0.3 < 0.1
+0.5
T aopoda
("nor iaoaphaaroaia 0
oraRonenae
Ianlropals 0
kinraldl
Idotaa 0
srhailtt (
I. 0
woanese>sVi 1
0
0
0
7.*
+2 . <
t .4
+ 1.?
1.0 <0.1
+ 1.4
~0
0
0
n.i
-------
Cr««k +3
0.05 e
# wt
0.01 ID
# wt
Pall 77
0.2 ¦*
# wt
0.05 ¦
# wt
0.01 r
# wt
Win 7B
0.2 «2
# wt
0.05 a
# wt
0.5 < 0.1
~1.0
1.0 < 0.1
+1.4
0.3 <0.1
+0.7
0.2 < 0.1
+€.4
2.3 <0.1
+3.2
0.5
+1.0
<0.1
0.3 C 0.1
+0.5
O.P < 0.1
+0.5
0.3 <.0.1
+0.5
0.9
+2.1
<0.1
1.0 <-0.1
+ 1,4
11.5 <0.1
+23.0
25.8
0
4
0.4
0
3
21.0
12.5
6.3
0.3
O.B
0.1
0.5
4.2
+4P .9
+0
7
+1.3
+0
9
+42.0
+25.1
+11.2
+0.5
+1.9
+0.3
+1.0
+B.4
"*24.0
"0
4
46.9
6
2.3
0.2
37.0
2.B
26.5
3.5
0
+33.0
+0
6
+34.1
+4
3
+3.9
+0.2
+51.5
+4,6
+34,3
+4.9
"69.5
3
"0.5
To
1
~0
"0
0
0
+39. B
+0
2
±?
~0
0
0
0
0
0
0
0
0
0
0
0
129.« 15.5
+204.0+2 7.1
0
0.3 < 0.1
+0.5
0.3 <0.1
+0,*
0.3 <. 0.1
+0.5
0.1
+0.2
0
0.10 0
0 0
0
0
0
0
1.0 < 0.1
+2 .0
"o
0.3 < 0.1
+0.5
1.3
+2.6
~0.?
+0.7
~e
1 .0
+3.0
0.0
+0.1
<0.1
0.1
+0.2
0.5
+1.0
"o
0.0
+0.1
5.3 ?.0
+10.5 +3.9
"11.0 ~3.7
+1?.« +4.3
52.0 0.7
+51.4 +0.P
0
l.S
+2 .2
0.1
+0.1
0
0
0
0
0
0
4.3
+*.6
2.1
+3.K
0.5
+1.0
"o
0
0
<0.1
24/
-------
0.01 III
# wt
Sfr 77
0.? *
# wt
Morse Creek +3
Sum 77
0.0'
?
¦
wt
Crustacea (font.)
Iaopoda
Idotes 0.1 <0.1 0
spp. ( juv.) +0.7
Llmnorla 0 0
a 1 pa rum
Amphlpoda
Camwarldea 0 0
"PP-
Allsrchestes
anguntun
Anpithoe
ninulana
Corophlu* 0 0
brevta
C. 0.1 <0.1 0
up. 0.7
Hyale
frequens
Orchaatla
»P-
Parawoara
"pogatlea" n. np.
Fa raphoxu*
*pino«us Tvpe B
Pontoganela
cf. lvancvl
Derapoda
Cancer 0 0
¦agister
C. 0 0
oreponenal*
Cryptol1thode* 0 0
¦P. (Juv.)
Nealgrapaua 0 4,0 17.1
nudua +? . ? +7 .9
H. 0 ~0
oregonenala
H. 0 0
¦p. (juv.)
Pagurua h. 0 0
hlrautluaculua
t. 0.1 <0.1 0
app. (Juv.) *0.7
Pugettla 0 0
¦ p. (juv.)
0
0
0.3 <0.1
+0.5
0
0
0
0
0
0
0
0
0
1.3 0.9
+1.9 +1.4
~~0
0.01 «i
# wt
0.2 m2
# wt
0.05 m
# wt
0.3 <0.1
+1.1
~0
0.3 <0.1
+0.7
0.1 <0.1
+0.2
0.2 < 0.I
+0.7
~4.s < 0.1
+ 1?.4-
4.6 <0.1
+17.5
0.1 <0.1
+0.2
0
0
0
0
0
0.3 <0.1
+0.6
0.1
40.4
'0.7
~0.4
~0.1 *0.1
+0.?
0.1
0.0
+0.1
0
0
1.0 <0.1
+ 1.4
0.3 <0.1
+0.5
~0
0
0
0
0
0
0
2.0 < 0.1
+3.4
0.3 < 0.1
+0.*
l.B < 0.1
1.0
0.5 <0.1
+0.7
~9.5 < 0.1
*19.0
9.5 < 0.1
+ 19.0
0
0.3 < 0.1
0.3 < 0.1
+0.5
"0.3 < 0.1
+0.5
0.5 < 0.1
+1.0
0 7.3 * 0.1
+2.4
f.O 36. <5 ~0
+3.4 +32 .«•
"0 0
3.3 1.0
42.4 +1.0
0
3.3 < 0.1
+? .6
~2.0 1.1
+1.6 ~l.B
~i.3 ro.i
+1.3
~0.5 < 0.1
+1.0
? a %
-------
Creek +3
Fall 77
Win 78
0.05 m'
# wt
0.01 m
# wt
0.2 ¦
# wt
0 .Ofi »
# wt
0.01 m2
# wt
0.5 m2
# wt
0.0* m7
# wt
2.0 < 0.1
+3.4
~0 .3 < 0.1
+0.8
0.1
+0.3
0
<0.1
0
0
0.5 <0.1
+1.0
~0
0
0
0
0
0
0
1.8 < 0.1 3.7 <0.1 3.5 0.1 5.3 <0.1 0.4 <.0.1 0.3 <0.1 3.0 <0.1
+1.0 +3.9 +3.^ +0.1 +4.6 + 1.6 +0.5 +4.1
0.5 < 0.1
+0.7
~9.5 < 0.1
+ 19.0
9.5 < 0.1
+19.0
"0
0
167.6 0.1
+175.7 +0.1
0
0
41.? <0.1
+67.8
20.7 <0.1
+38,U
0
0
38.3 <,0.1
+73.2
0
3
< O.i
-
+c
5
~0
3
< 0.1
-
+0
5
"o
3
< 0.1
-
+0
5
0
5
< 0.1
-
+1
0
0
0
0
0
7
.3
* 0.1
0
+2
.4
"0
0
0
0
3
3
< 0.1
0
+2
6
2
0
1.1
0.1
+1
6
4-1.8
+0.?
1
3
<"0.1
0
+1
3
~0
5
< 0.1
0
+1
0
0.0
40.0
0.3
0.4
0
0
0
0
+0.5
+0.8
"0.5
. 1
0
0
0
0
+1.0
+4.2
0
0
0
0
0
4.0
7.9
3.0
0.0
0
5.8
18.4
0
3
0
4
+5.0
+6.5
+6.0
+0.1
+5.6
±19.7
+0
5
+0
8
0
5.0
0.2
0
0
2
3
"0
4
+6.2
+0.7
+3
9
+0
8
0
"o
0
2
0.1
0
~2
8
~0
I
+0
5
+5
5
+0
2
0.8
0.2
0.3
<,0.1
~2
4
0.0
1.0
0.2
~l
3
~0
8
+1.0
+0.2
+0.5
+5
5
+0.0
+1.4
+0.2
+1
5
+1
3
*0
"1.5
0.1
~0
2
4 0.1
"0
0
+2.4
+0.?
+0
4
0
0
"o
0
0
248
-------
Inaccer
Dlptaran
larvae
Species Richness
Dlveralty, H1
Total Number
Total Bloam as
Morse Creek +3
Spr 77 Sua 77
0.01 a7 0.2 m7 0.05 a7 0.01 a2 0.2 a2 0.05 a2
I Vt # Ht f «t # Wt # Vt # «ft
1 .2 <0.1 0 0 20.9 <0.1 0 2.3 <.0.1
*?.4 +40-0 +1 .9
59 76
2.34 2.04
1613.1 73S5.0
163.3 261.fi
269
-------
» Creek +3
Fall 77 Win 7F
0.05 a2 0.01 m2 0.2 m2 0.05 »2 0.01 «2 0.2 «2 0.05 m2
# «rt # wt # wt # wt # wt # wt # wt
2.3 <0.1 20.8 *0.1 0 2.0 <0.1 2.2 <0.1 0 9.0 <0.1
+1.9 ±^3.8 +4.0 +3.H +13.7
70 64
1.97 2.50
£319.4 3378.5
229.0 271.9
1
I
249
-------
r«Dla 4 r Norw r*^k < ,* p M> I» ^ver +*i>4) fwnchl? organtVM* # ro« the I .*w » > ! n' «»r' Mil f.>i+ <•
«»d/cr i ft > p^r r ^! <<• .>r>J , ">. '"?"¦< ' •#» 4 , , *n.J 'I ? • S- , * >,i .'• t *rrj 1r v: .u i ¦
77
Soft 77
O.Ol ¦
JL fflL
OJ •
0.05
0,01 ¦
J- XL,
p.j ix
Mil!H
f.
AUrti
- 0,1
d
0.9
*o.i
o
0.9
<9.9
9.9
? 1.4
<0.1
9,0
*0,1
. O.K
0,9
i I.*
9.1
'0,1
^0.1
0
0
©
0
9.9
19.S
9
9
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e
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i. ?
~o.§
1M.)
~ 194.4
. 0.1
0
O.i
* 0.1
0.1
0
0
0
0
0,7
' I §
0
*.4
* 7,3
- 0.1
to.)
* 2*.S
e
0
0
e
0.4
* 0.5
0
0
0
0
0
0
31,5
*2$.5
109.4
^177,2
FbU 77
3.0> m
i vt
0.31
OJ
0.05
0
0
0
0
0
0
0
0
0
0
0
0
0
0
D
0
0
0
0
0
0.1
0
0
0
0
0
0.1
'O.i
0
6
0
0
0
0
0
0
0
0
0
0
0
0,2
* 0,2
0
294,7
? 145.2
0
0
n
0
0
0
0
0
0
0
0
0
Q
0
-------
I
17
0,01 h2 0,1 m7 3,0) J
* * pt » *
<0.1 o o
<0,1 0 0
o Q o
<0.1 0 0
<0.1 0 0
11
-------
*mpnyi
pifltUit «9la
iHMmiimm
M)«n
lott—%,,»
P*lyatfhMl*
pttfta
s4ll fvrtla
S*r
e.ei J 9,t •1 o.o) a2
J »i ~ w < »
1«8
»,t
0
13.5
<•.1
8
8
<8.1
0
0
n,i
47*.3
0
t*4.3
1104,1
8.1
4.0
0
£®.J
*4.J
8.4
0
0
M.I
0
8
0
<0,1
8
0
8
8
0
0,8
0
0
18.1
8.8
0
0
*8*1
0
0
0
0.0
0
0
18a
0
8
0
<0.1
8
0
(0,1
0
0
Iter** Cr«*h *0
»«¦ 7 7
0.01 i2 0,1 a1 0.W a2
# wt # «ft # w*
1.1 18.9 0
-7.7 UK*
ooo
0 0 0
I).ft J93.0 0
ti.s tiie.#
1 .4 t) 0
r*.l
<0,1 0 0
0 0 0
0 0 0
0 0
0 0 0
0.1 0 0
t 0.1
<0.1 0 0
.0,1 0 0
*0,1 0 0
ooo
0 0 0
Fall n
0,01 ¦* 0.2 m7 0.0} m1
I m * * vt
1.3 23.6 0
£3.4 *21.0
0 0 0
0 0 0
1.8 43,6 0
-1,3 *75.1
<0,1 0 0
tO.l 0 0
<0.1 0 0
0 0 0
0 0 0
<0,1 0 0
*0.1 0 o
ooo
0 0 0
(0.1 0 0
ooo
0 0 0
-------
\
I
m. (*¦».)
Ht "
0,01 ¦' 0.1 •' 0.05 **
t 51 i H i 3
•0.1 0 0
O.J 0 0
tl.l
o.e o o
* 0,0
0 0 0
0 0 0
0 0 0
S.* <0.1 O.J 0.! 2.5 O.I
tl.t 10.5 to.* 12.1 10.1
0 0 0
taHtUrt* 1,0 '0.1 0 1,1 ^0.1
* 2.J tT.O
0
0
0
"ter»« Craak *0
0.01 •
Sa 77
0.1
Q.OJ ¦
I Vt
0,01
rail 1J
0.2 a3
0.05
0.7
* 1*4
0
1.2
4,2
0
0
: 0,1
0
0.*
0,6
0
0.1
t 0.2
<0,1
0.5
'0.3
0.1
tO.2
1.5
? 3*0
0*5
f-L0
0
0.1
*0,1
^0.1
0,1
'0.2
0
0.1
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0 0 0 0.1 0.1 0 0
• 0.1
0.* O.i 0 0.* . 0.1 0 0 1.1 0.0
•o.j • o. * • i.? 'o.i
0 0 0 0 0 0
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1.* e.l I.O 0.5 t.B 0.1 8.1 0.0 0 l.J 0.1
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tyniw iMmh lit
¦Hanrtcy - l .l*
total nrt ii * ,4**. I
DM*! Mam*m U) W* .8
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la " r«u n
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# wt # w i rt # yt # « < wt
o.a o.i o o.i -o.i
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l 11
J.55
4,341,5
m .o
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-------
Table -.d . Morse Creek (gravel-cohb] e
collected 7 June 1977, expressed
x* + standard deviation.
Chlorophyta
Dcrtnttia aarlna
Sallcyacla ovalla
Spongooserpha sp.
Bacillariophyta
Pton«l»> >pp.
Fhaaophyta
DwirMtli llgulata
D. virldia
Kctocarpacaaa ap.
Ectocarpua ap.
LMintriilti app.
Ralfsia pact flea
Sticcyoalphon tort ilia
Syrlngodcrea abyaslcola
Abodophyta
Acrochaatlua ap.
Ahnfcltla pllcata
Antithassmion defectum
Callophyllla flaballulata
Coralilnjcaaa ap.
Dalaacarlacaaa «p.
Gracilaria ap.
Haloaacclon glandiform*
By—nana ap.
Irldaa sp.
LfCafauchaa paci flea
Payaaonal11a paciflea
PlatythMmlon hataroraorphun
Polyneura laclaslma
Pulyslphonia paci flea
Ptaroaiphonla dandroidca
ihodochorton ap.
hen chic r, rcanisns from the subtidal ,'ont'
f number nnd f c-r b (p) r>er ".'i5 )
-10 m.
wt
< 0.1
< o, 1
<-0.1
< 0.1
<0.1
co.l
'0.;
<0.1
0.1± 0.1
-------
Horse Cr**k Suhtidal
-10 ¦.
# wt
Fori fara app. v/
Cnldarla
ltydrozoa app. -<0.1
TubularLa 3.814.5 <0.1
Anthoioa app. 2.3*3.9 <0.1
Hilcafa ap. 0.5 4 0.6 <0.1
Imrtia app. 5.5±1.7 <0,1
HMCoda iff. 21.5 14.8 <0.1
Molluaca
tephlnaura ap. (Juv.)
Mopalla ap. B
Toaicalla linaata
0.3 x 0.5
0.3 * 0.5
0.3 t0.5
<0.1
0.010.1
<0.1
Caatropoda
Molldacaa ap. (Juv.)
Anphlaaa coluablaoa
CallioaCoM llgatwa
Calypcraaa faatlglata
Colllaalla ochracaa
Crapldvla ap.
Margaritas pupillua
Margarltaa/Llru.aria
Naelca cla«aa
Trlchocropla cancallata
0.3 t0.5
12.5 ± 11.7
0.5 t 1.0
3.0 r 1.4
,3 ±0.5
.3 t0.5
.3 *0.5
,3 * 0.5
3 i 0.5
3 i0.5
0.
0.
0.
0.
0.
0.
<0.1
0.8 *1.5
0.4 i O.b
0.1 to.2
0.0 to.l
0.0 *0.1
< 0,1
<0.1
<0. 1
<0.1
Blvalvla
Cranalla dacuaaaCa
Huallarla kannarlayl
Ma coma aacCa
M. ap. (juv.)
0.3 t0.5
0.3 ±0.5
0.3 t 0.5
4.0 t 3.7
<0.1
<0.1
0.010.1
0.0 to. 1
-------
Horse Creek Subtldal
-10 m.
vt
BivalvLa (cone.)
IMlolui recent
tttaculua diaeora
Myaella tumida
Pro t o t Ka c a s t ne »
Tranaennella Cantllla
0.3t 0.5
0.3 i 0.5
19.8 t 5.0
1.8*1.7
0.5 10.6
<0.1
<0.1
<0.1
0.4 to.9
<0.1
Annelida
OllgoehMCa epp
5.8 ' 4.3
<0.1
Polychaeta
Aepharetldae epp.
Aaabellldea litturalla
0.32 0.5
0.3 t 0.5
'0.1
< 0.1
CaplcellIda*
Capitalla capltata
MadioaaaCua ap.
0.3?0.5
25.5 t10.4
<0.1
*0.1
Cirratulldae
Chaatoioae setoaa
Clrratulua cirratua
Tharyx auitifilie
1.3 *1.9
3.5 13.0
8.8 18.3
<0.1
<0.1
<0.1
Oorvilleldae
Protodorvlllea gracilis
1.5 13.0
<0.1
Flabelligerldaa
Pheruaa pluaoaa
0. 3 ±0.5
<•0.1
Clycerldae
Clycera ancrlcana
G. ap. (Juv.)
0.3 t 0.5
0.3 i0.5
<0.1
<0.1
Gonladldaa
Glyclnrfe picta
3.5 11.7
<-0.1
26-S
-------
Mors* Creek Subtldal
-10 n.
wt
Haaionlda*
Nicropod«rk* dubia
7.3t 4.0
*0.1
Maldaeidaa ap. (Juv.)
tUlochalla rwbrocincta
Vlcosacha lusbrlcalla
7.8i 4.6
0.3 f0.5
0.3t0.5
*0.1
*0.1
<0.1
HaraldM
Kerala ap. (juv.)
PlaCynerala blcaaaliculata
0.3 t 0.5
3.5 f 1.7
<0. 1
*0.1
Ophe 11Idee
Armandla br«vla
1.8 /1.3
*0.1
OrfclnlldM
fceoloploa ap.
1.3 ±0.5
*0.1
Owvitiidaa
OMnla fiulfonli
0.8 ±1.0
*0.1
Paraonldae
Arlcldea ap.
0.5 ±1.0
*0.1
Ptayllodocldae
Eteooe longa
¦alalia aangulnea
Phyllodoca vadalrcnala
P. ap. (jo\ )
0.3 t 0.5
0.5 ±1,0
0.3 *0.5
0.5 ±0.6
*0.1
*0.1
*0.1
*0.1
Polyneidae
ianethM labrlcata
1.0 to,8
*0.1
Sabellarlldee
Saballaria camtntariua
0.8 ±0.5
0.1
labellldee
Chone ecaudate
1.8 t1.0
*0.1
-------
terse Crack *ubtldal
-10 is.
# vt
Scalabrapaldaa
Scalabragaa In flatus
Sarpwlidaa
Spirorbla aoarchi
5. ap.
0.5 £0.6 <0,1
0.3 * 0.5 <-0.1
0.3 * 0.5 <0.1
Splonidwi
Polydora inuti 0,3 *0,5 <0.1
P. social 11 0.3 * 0.5 <0.1
P. ap. 0.3 £0.5 <0.1
Prlonoaplo cirrifara 1.0^0.8 <0.1
P. ifMliHl 5.3 £3.4 <10.1
lylllda*
Bxogona g«aa»ifcra 0.511.0 <0.1
K. loarai 0.8 ±1.0 <-0.1
I. app. 19.5 t 2.1 <-0.1
Syllia app. 0. j ±1.5 <0. 1
Yaraballldaa ap. 0.310.5 <0.1
fipuncula
Colfingia vulgaris 0.31 0.5 <0.1
CruaCacaa
Oatracoda app. 0,b f1,5 ^ 0.1
Ctuucaa
Cuawlla app. 0.8 10.5 <0.1
Maatylia app. 6,8 f 6,4 ^0.1
Laptoctu&a app. 2.0 ±2.7 <-0.1
Tanaldacaa
UpMehilli dubla 240.3t4o.B O.ltO.
1?* S
-------
Iiopodi
laniropa.• analoga
Jacrcfitii :.bla
Asphlpt *
Crart nandax
C. ati< -.cx
MmA.idu app.
Dcc«|}&4«
Cancer roductus
Carldaa ap. (juv. )
Diacoraopagurua achnltti
Eualua ap.
Pagurldaa ap. (juv.)
Pagurua bcringanus
Pir.nlxa ap. (juv.)
Pugattla gracilis
Schinoderaata
Ophiuroidaa app.
HoloChuroldca
Mopalla intaraadla
Spaciea richness
Diversity - H'
Total nuabar
Total bicMis (g)
; L d,1 1
-10
vt
10.
5.
5 t b. /
0i6.7
<0.1
<0.1
0.5 £ 1.0
0.3 t 0.5
101.5 £41.8
<0.1
<0.1
0.2 tO. 1
1.0 i 1.4
0.5 ± 0.6
0.5 tl.O
0.5 ±1.0
5.5 £2.4
0. 3 i0.5
1.01 1.4
3,0 ± 2.2
0.6 * 0
<-0.1
^0.1
<0.1
0.0
0.4
<0.1
<0.1
to.
to.
8.316.2
<0.1
0.5 t0.6
<0. I
127
2.72
581.1
<14.4
-------
Table ia. P*:.-ket t Point r rr.r r - in i-nr.ive> br-nthi •
tidal zon.? . I Iv.-tf.' ?• ¦>? in?. I f.r- iuch mntt-r 197*
it: pt-r ¦% '"'j ;'i=2:
f!<* i * ¦5»r?"K.rs tin- ii nr. \ / ir.tt r-
t'XPft'Sst-J is nunhi-r -tii»! • s'r
tor o- *• standard Avv I > t i; >n .
Fall 77
# vt
0.8 <-0.1
t0. 5
6.8 *0.1
±4.1
0.3 -0. I
*0,5
0.3 <0.1
*0.5
0
0
2.0 <-0.1
*1.2
0
1.8 ^0.1
•* 1.0
20.& <0. 1
*8.9
0
Win 7 8
# wt
0.8 *0,1
1 0.5
8.5 d).i
16.2
0
0
0
0
0.5 tO.l
ri .0
0.8 tO.l
*1.0
0
19.0 < 0.I
2-9.5
0
-------
Beckett Peine +6
Spr 77 Sun 77
I vt # vt
itereidae
Ranli 2.0 '¦O.i 0.3 ^
vexilloea ^ l.t ±0 %
V. 0 0
•p.
Flatyaerels 0 0,3 v/
blcaualleulata -0.5
Platonic 0.5 <0.1 0
• pp. ^ 0,7
Syllida*
Kxogone 0 0
lourel
Syllit 17,0 <0.1 3.5 <-0. I
• pp. <11.3 ±4. 5
Crustacea
Cirrip*dia
Balanua 0 2.8 0.7
glandala ±4.3 *0.6
1. 0 0.5 <0.1
ap. (Juv. ) -1.0
laopoda
¦xoaphaaroata 0.5 <0.1 0
mdla ^ 0. 7
Jfesphipoda
Gaaarldta 1.0 <0.1 _
•pp. ±1,4
On II ¦ mi ¦¦¦Hi in 11 j111 iii _ 0,3 <0.1
confervlcolua 1 0.5
OrchMtoldea _ 0.3 <0.1
pwgsttenals 1 0.5
Pall 7 7
# vt
Win 78
# vt
0
¦ 0.1
0.3
± 0.5
0.8 <0.1
-0.5
0
0.
± o,
0.
±0.
0.
± 0.
0
<0. 1
< 0. 1
< 0. 1
o. 3 <0.1
-'0.5
1.5 <0.1
±1.7
0
3.5 <0.1
± 3.7
12.0 4.2
ill.2 ±4.2
0,3 <0.1
±0.5
10.8 4.2
H1.9 — 6.6
0
61.3 0.4
£ 30. t ±0.2
2.0 <0.1
±2.5
0.8 <-0.1
±1.0
0.8 <0. 1
±1. 0
-------
Beckett Point
Spr 77 Susa 77
# at # rt
Insacta
Dlpc«rin 0 0.3 ^0.1
larvaa f0.5
Sftclti richnaas 10 13
Diversity - B* 1.43 2.04
Total Bv~tb«r 81.5 15.8
Total bloMia (g) 4 1.0 <1.7
Fall 7/
# Vt
Win
#
0
15
1.44
110. I
< 5.8
0
12
1.69
47.6
<5.3
-------
>b, Beckett Point .'protected «>anm the ml ti (+ 1* ) i n t f r i i d'i I
zone '?o ] 1 (?*'ted sprint 19 7? through wirter 1478, t'xnressed as number irni-'or bionass (<».) per
O.Oj {"=2; spring) and n. 025 (»*A; summer-winter) a- + standard deviation.
Spr 7 7
# wt
Sue 77
# wt
Full 77
# wt
Win 78
# wt
Cblorophyta
Kacaroaorph*
coapraasa
S.
lruratirvalls
E.
1 ins*
Uiva
• p.
0.
taeniae*
Uroapora
•PP-
taci1lcrlophyta
Pannalaa
¦PP-
nMophyti
Pal
-------
Beckett Point +
Spr 77
# vt
Sua 77
# vt
taMrtaa
'CP-
•PP-
Moileaea
AaphlaMra
•pp. (Jo*.)
Gastropoda
•PP- (J"v•)
41om41«
Alvlnia
¦pp.
CotllMlla
itrlgtUUi
Parts!®*
occl4antala
ImkLqom
vesimla
Lacwna
varlagaca
Utterlu
antalata
L.
altkana
Hargar1tea
pupillua
Octnabra
lurlda
Thai*
¦PP-
1.5 <0.1
1 0. 7
0.5 < 0.1
±0.7
1.0
±1.4
0
0.5
1 0.7
0.5
*0. 7
0
*0.1
41.0 <0.1
±58.0
0
0.1
± 0.1
0.1
±0.1
2.5 *0.1
13.0
0.3 <0.1
t 0.5
0.6 <0.1
~ 1.0
1.0 - 0.1
11.1
0
0.8
±1.0
0
* 0.1
C.? <0.1
±0.5
0
0.0
f 0.1
0.0
±0.1
<• 0.1
13.2
0.3
10.5
0.3
±0.5
0
0
0.8 <0.1
±0.5
Pall 77
# «t
Win 78
# vt
5.3 <0.1
±4.0
6.8 <0.1
±7.0
0
6.3 0.1
±3.0 ± 0.1
1.3 <0.1
1 0.5
21.5 0.J
20.6 ±0.1
1.0 0.1
± 2.0 ±0.2
0.8 <0.1
* 1.0
10.0 0.2
i 9.9 i0.2
77.3 0.3
12.4 10.1
0
0
0.3 <0.1
±0 '
r <0.1
io.s
0
2.5 <0. 1
+ 4.4
0
0
0.3 <0,1
t 0.5
0
0
0
0.3 <0.1
±0.5
0
0
0.3 <0.1
to.5
0
0
0
0
-------
Backatt Point +3
Bivalvia
Ads I a
ca1Iforn1ana1¦
ClinocardiwM
attttalill
Mjrttlla
twl(U
Kytilua
•Pf. (J«v - )
Procothaea
ataaiMa
TraBaaoaalla
cantllla
Traaua
tapa*
Annallda
Oltgochaata
•PF-
Polyctvaata
CapltallliM
Capitalla
capitata
H*diaMat«a
MP-
HotaaMCm
taaaia
Ohaateptaridaa
Spiechaatoptarw«
coatariw
Spr 77
# vt
0.5 <0.1
t 0.7
0
1.0 *0.1
" 0.0
5 c 0.1
t 7.6
4.5
t 6.4
3.0 *0.1
±4.2
0
1.0 <-0.1
t 1.4
2.0 <0.1
*1.4
0
1.0 <0.1
t 1.4
0
Sub 7 7
# vt
0
0
17.3 0.1
±8.5 10.1
4.3 0.1
i 1.3 ±0.1
1.3 *0.1
±1.5
3.3 0.0
¦*2.6 10.1
14.6
t 0.„ '9.2
2.5 -0. I
12.1
0.5 <0.1
11.0
1.0 *0.1
I 2.0
8.5 <-0.1
*9.3
0
Chryaopatalidaa
Palaoootua
ballia
0
0
Fall 7 7
# wt
Win 78
# wt
0
8.3 0.4
i 10. 1 ±0.5
17.3 0.1
±11.4 ±0.1
225.3 0.5
1 65. 7 ±0.2
43.0 0.4
± 33.5 t 0.4
9.5 0.2
17.2 10.1
4.5 1.3
13.1 11.6
3.0 -0.1
2 2.2
0.5 '0.1
i 1.0
0.3 <0.1
t 0.5
3.3 <.0.1
±3.3
0.3 <-0.1
• 0.5
0.5 <0.1
i 0.6
0
0
3.8 c 0.1
i 1.5
11.5 <0 1
1 2,1
0.8 <0.1
t 1.0
0.5 *0.1
* 0.6
0.5 2.0
£ 0.6 ±-1.7
3.8 <0.1
±5.6
0.3 <0.1
t0.5
0
7.8 <.0.1
i 7.8
0
0
-------
Beckett Point +3
Spr 77 Sua 7 7
P vt # vt
Clrratalidaa 0 0.3 <0.1
•pp. Gov.) -0.5
0 0
•F9-
froto4orvl llu 0 0,3 <0.1
gracilis to. 5
Glycarlda*
Baalpodua 68.5 / 43.5 y
borMlia ±2.1 ±2.5
MacionIda*
Mlcnptdarkt C 0.3 <-0.1
dabla 1 0.5
Ophlodroni 0 0
teral4M
hnli
•p.
riitjiMnii
bicaulioiliCi
OnaphlidM
Otwphla 0 0
•P-
OplMiildM
Anndia 0 0
br«vla
0ph«lLa 0 0
liaaclna
0
0
0
0
Fall 77
t vt
Win 78
# vt
0.5 <0.1
t 0. fa
0,3 <0.1
t 0.5
0
0.3 <0,1
f 0.5
78,8
±4.6
s
30.3 s/
±11.1
11.3 <0.1
±3.3
1,8 <0.1
±1.7
0
0
1.3 <-0.1
±1.0
9.8 <0.1
±10,2
0
0
0.5 <0.1
i 1.0
2.6 <0.1
±1.7
0.5 <0.1
* 0.6
0
0
-------
Beckett Point +3
Spr 77 Sum 77
# «t # trt
lidaa
Ovnlf f. 5 <0.1 7.0 y
fvalforals 17.S *6.5
Phyllsrfocldaa
Itaooa 0 0
1®U£JI
•alalia 0 0
faadrilebata
I. 0 0
a«ogt»l.i*aa
Pbyllodoci 0 0
maculate
r. o o
•pp-
Plalootdaa
Pitloo* 0 0
•p.
Polynoidaa
BarMtbea 0 0
Ivwlata
Saba11Ida*
¦PP.
Slgall«ni4aa
Pholoa
¦iottta
iploaidaa
Poljr4ora
eolanblana
0
0
Fall 77
# vt
Win 78
# »c
207.3 y
i 66. 2
30.5 ^0.1
f 24.0
1.3 *0. I
11.0
0.8 *0.1
11.5
2.6 '0.1
*1.7
1.3
-------
ficckcct Point +3
Spr 77 Sua 77
# vt # vt
SpLonidaa (coot. )
Polydora 0.5 <0.1 0
proboacLdaa ±0.7
P. 0 0
•P-
Prloooaplo 0.5 *0.1 0
cirri fara *- 0.7
P. 0 0.5 <0.1
•taanatrupl 1 0.6
Py|o*plo 0 2.5 <0.1
• lagan* *- 2.9
Scalalapi* 0 0
•P-
Splo 0 1.3 *0.1
fllleornla i 1.0
SplefhatiM 0 0
bewfay*
Sylllda*
Irnii 0 0
bravipharyngla
Kxegooa 1.0 < 0.1 0
ItUtl 1 1.4
•phaaroayllla 0 0
plrlfara
SyllU 1.0 *0.1 1.0 <0.1
•pp. 11.4 11.4
fmbtllldt* 0 0
•If-
PolTCirrua 0 0
•p. A
Croat mm
Ostracoda 0 0
¦PP.
Fall 7 7
# «fC
Win 78
# vt
0
0.5 -0.1
-0.6
1.0 <-0.1
* 2.0
0.5 <0.1
*1.0
47.0 <0.1
±47.3
0.5 <0.1
* 0.6
14.3 <0.1
¦* b. 6
1.5 <0.1
t 1.7
0
0
0
0
0
0
0
0
0.3 <0.1
± 0.5
1.3 <0.1
t 1.0
0.5 <0.1
f 1.0
0.3 <0.1
±0.5
0
0
0
0
0.5 <0,1
±1.0
0.3 <0.1
±0.5
0
1.0 iO.l
t 1.2
-------
Clrrtp«41«
glaadela
B.
• p. lica«4a
B.
mdla
&sorl»o®phacrom
enpctsit
Aapbipoda
AllorchMCCB
sngmtu*
A&lKogmmmrmm
coafcrvlcolaa
Capralla
laavl«jctila
Corophioa
•pp.
Qmm»ri4*«
*PP-
Qrehaataldaa
l*ag«tcwili
Beckstt Point +3
Spr 77 Sua 7 7
# wt # wt
3.0
± 4.2
0
0,8
*1.1
117.8
*83.7
0
2.4
£ 2.0
0
23.0
t 3.7
0
< 0.1
3,8 * 0.1
13.3
7.0
*».7
4.3
J 6.0
i0.1
r 1
±0,2
0
0
2.0 <-0.1
t 1.4
0.5
10.9
28.5
±15.2
0
«. 0.1
0.1
to.2
0.5 <-0.1
±1.0
0.3 *0.1
±0.5
1*11 77
# %#t
Win 78
# wt
19.8
± 26.2
0.5
i C.6
3.4
±3.5
<0.1
1.0
±1.4
0
0.0
£0.1
1.3 *0.1
J 1.5
15.0 <0.1
t 12.9
0.3 <0.1
-0.5
40,0 0.3
±14.9 ±0.1
0
0
0
0.3
± 0.5
0
<0.1
0.8
t l.o
5.3
i 3.8
10.0
* 6. 7
^0.1
<0.1
0.1
0
0
0.8 <0.1
*1.0
-------
Beckett Point +3
Spr 7 7 Sues 7 7
# wt # wt
Dicapoda
Caacar 0 0.3 0.0
Hgiittr *0.5 *0.1
ItaitrapM* 0 0.3 0.2
orcgonensis 10.5 -0.4
Pagunu 0 0.3 0.0
hiraatiuaeulua *0.5 *0.1
F. 0 2.8 <-0.1
¦pp. (Jov.) -2.1
Plnnlxldaa 0 0.3 0.1
a pp. 10.5 10.2
lutcta
D.pt*ran 0 0.3 <0.1
lanrM 10.5
Phorooida
Pborcxiopaia 0 0.3 *- 0.1
hara*ri ±0.5
Echlno4«nMta
(ehlooidM
Bao^rutar 0 1.0 0.1
•xcMtrlcui 1 0. B *0.1
HolottaiTold««
Laptoayuapta 0 0.3 <0.1
clarkl i 0.5
Sp«cl*a rirhnan 23 55
Mv«r*lty - H1 1.89 2.20
~ziml nwbar 1 72.5 276.7
fecal blomasa (g) £ 2.7 <-20.9
Fall 7 7 Win 78
# wt # wt
0 0
2.8 0.1 0
-1.5 0.1
1.3 0.1 0
*1.0 *0.1
1.3 0.0 0
*1.9 *0.1
0 0.3 *0.1
t 0.5
0 0
6.0 cO.l 1.3 tO.l
*4.1 *1.3
16.8 0.0 0.8 0.0
tn.9 ±0.1 *1.0 io.l
2.0 0.1 0
±4.0 tO.l
74 29
2.73 2. 15
965.7 99. 1
<13.3 <4.1
-------
•pp.
^ /
Table k . Beckett Point ^protected san (S»*»' suramer-winter) m- + 'itandaru .<«.•%•».-it ion.
Spr 77 Sum 77 Fall 77 Win 78
# vt # wt # kC # vt
Chlorophyta ,
Intcroaorpha 0 v Q 0
Lntaatlnalls
E. C
llMI
uivc o y J 0
•p.
Uroapora 0 0 J 0
•PP-
Rhodophyta
Caraniuai 0 J •/ 0
pad flew
Cnldarla
tothotM 0.5 <0.1 0.8 <0.1 0.8 <0.1 1.8 <0.1
¦p. ±0.7 + 1.0 +1.5 +2.9
PlatybalvlEithaa
TurbtlUrli 0.5 <0.1 0 0.5 «¦ 0.1 0
¦pp. +0.7 +1.0
26.5 <0.1 21.5 / 8.3 *C. 1 12.0 <0.1
•PP- +12.0 + 9.3 14.0 jl2.4
Nanatoda 45.0 y 7.3 <0.1 220.8 <0.1 41.3 <0.1
•pp. ±12.7 ±3.9 ± 188.8 ±15.3
HdIluica
A*ptaln«0.1 0
•PP. (J«v. ) ±0.5
Aflaja 0 0 0 0.3 <0.1
dlo«ttd«« ±0.5
281
-------
leckctt Point +0
Gastropoda (cone.)
Alvlnla
• p.
Cyllehna
¦P-
Farculw
occidental*
BunlnoM
vaalcula
Lacuna
varlagaca
Litcorlna
sickana
Marg«rlc«a/
Llntlarla
Micralla
Cabtroi*
McaMrltts
¦•ndlcua
Odoateatia
•p.
Qc*n*bra
lurida
Blval -"U
Cllnocardlun
nuttallli
Ct*m11i
dacuaaata
Hiatal la
arctica
Lueinoaa
tcnalscu1pta
Hmcams
inquinata
Spr
7 /
Sua
77
#
wt
#
wt
2.5
<0.1
0,8
<0.1
+ 3.5
+ 1.5
*" 6.0
0.1
0.3
0. 0
* 8.5
+ 0.1
~ 0.5
+ 0. 1
0
0.3
i 0.1
<- 0,5
0
0.3
^0.1
10.5
4.U
< 0.1
16.3
0. 1
+¦ 0.0
13.4
t 0. 1
0
0.3
< 0.1
*¦ 0.5
0
0
1.0
< 0.1
0
1.4
1.5
0.9
0.5
0. 3
2.1
11.2
10-6
10.4
9.0
< 0. 1
0
.12.7
0
0
9.5
0.9
0.3
0,1
13.5
ll-i
10.5
10.3
0.5
< 0. 1
0
10.7
_ 1.5
<. 0.1
0
+ 2.1
1.5
<0.1
0
12-1
0
0.3
0.1
10.5
10. 1
Fall 7 7
# wt
Win 7£
# wt
7.3 <0.1
I 3.6
0.5 0.I
10. 0 10. 1
0
1.8 <0.1
±1.5
3.8 - 0.1
l2.t>
0
0.3 <0.1
* o. s
0.8 <0.1
II . 0
0.5 <0.1
±0, h
0.5 -
-------
Becktte Point +0
Blv«Ivla(coot.)
MacoM
nctuci
M.
»*Cti
n.
• pp. (Juv. )
Hyatt la
tnl la
Mytllaa
adulli
H.
•P. (j«v.)
Fanop#
ganaroaa
Protathaca
aeaainaa
Tal llna
•odasea
T.
¦ pp. (Jttv. )
Tran«*nnalla
tantil* -
Traaaa
capax
£nn«lldi
Oligochaata
•PP-
Polychaata
Capital 1 ids*
¦ p.
Capital la
capital*
Spr 7 7
# vt
0
0
3.0 <0.1
11.4
212.S 0.9
1181.7 10.9
0
2.0 <0.1
12.8
0.5 0.3
10,7 10.4
9.5 0.1
19.2 10.1
0
6.0 <0.1
11.4
24.5 O.i
1 5.0 10.1
0
11.0 <0.1
110.2
0
4.0 <0.1
Is.?
Sum 7 7
# vt
0.3
1.1
+ 0.5
12.2
0
0.3
< 0.1
10.5
244.3
0.4
1205.3
10.2
0.3
0.1
10.5
10. 1
1.0
<0. 1
10.8
0
0.3
0.3
10.5
10.5
0
5.fa
o.:
28.0
0.L
120.4
_0.1
1^,3
' 6
115.8
11.7
1.3
<0.1
10.5
0
0
Fall ??
# vt
Win 78
# VC
0.3 0.1
10.5 10.1
o
0
4*7,5 0.8
1 b9.0 10.5
0
0.3 <0.1
10.5
0
7.3 o.y
15.4 11.7
0
1.5 0.0
13.0 10.1
5. fa 0.1
14.9 10.1
4.3 0.8
13.0 11.1
31.0 <0. i
110.2
1.9 < 0,1
12.0
3.5 <0.1
12.4
0,8 0.3
11.0 10.5
1.5 <0.1
-3.0
2.3 <0.1
13. 3
220,3 0.4
lo4. 7 10. i
0
0
0
7.0 0.1
12.6 10.2
1.5 0.2
12.4 10.2
4. "! 0.1
13.9 10.2
12.0 <0.1
13.7
i.8 1.1
12.1 12.2
3.5 <0.1
13,3
0.8 <.0.1
11.5
4.5 <0.1
le.4
-------
Backett Point +0
Spr 7 7 Suit 7 7
# vt # vt
Caplcalllda* (cont. )
Madlonaatua
15.5
< 0.1
6.3
<0.1
¦p.
* 12.0
±4.7
NotOMCtua
0
2.0
s/
lioeatua
±1.8
St.
11.0
< 0.1
10.3
< 0.1
tanwlt
±2.8
±18.5
V.
3.5
< 0.1
0
•p. (Juv.)
±5.0
Chaafceptarrlda*
ttaaechaatoptarua
5.0
<0.1
13.3
y
taylorl
±5.0
±12.6
SpiectMfc* m^ptmrum
0.5
<0.1
4.0
/
cottirw
1 *
o
•
±1.6
Cirratnlidaa
0
0
(J«*.>
Tharjnt
1.5
< o.l
0.3
< 0. 1
sulci fills
12.1
±0.5
Dorvillaldaa
Dtorviliaa
0
0
svdelphi
Fr»to4crrvlllaa
4.5
< o.l
1.0
<0.1
gracilis
±6.4
±0.0
61 year 14m
0
0
•pp. 0"v-)
Clycara
0
0
anrictn«
6.
0
0
taaaalata
Hmmtpodua
47.0
< 0.1
17.3
y
boraalla
±19.8
±4.9
Pall 7 7
# wt
Uln 78
# wt
31.0 <0.1
3. b
0
7.3 <0.1
13.8
0
11.3
111.7
3.3
±3.3
3.H
±2.6
0
< 0.1
< 0. 1
<0.1
2.5
±1.3
0.5
±0.6
<0.1
<0.1
2.5
±2.9
2.3
±1.7
< 0.1
<0.1
1.5 <0.1
±1.0
0.5 <0.1
±0.6
6.0 <0.1
±1.6
0.3 <0.1
±0.5
1.3 <0.1
±1.0
0.3
±0.5
0.3
±0.5
0.3
±0.5
12.0
±1.4
< 0.1
v-'
< 0.1
< 0.1
0
0
0
12.0 <0.1
±2.9
-------
Beckett Point +0
Goniadlda*
Clycinda
plct«
lasloaldM
•P- CJ"V. )
Baalonld
¦ p. A
Mlcropo4arkc
daMa
©phiodromt**
pttgattamala
Hagaloaidaa
Magalc »
Japeokci
H.
pitalkai
feldanldaa
• p. (J«v- >
Axiotfcal la
nkroclncti
Haphtyldaa
RaphCy*
CMCOi d*t
¦ .
•p. (j«v.)
¦•raidae
•p. (Jhv.)
¦•rata
•p.
Platynaraia
blcaoalLculat*
Spr 7 7
# wt
9.5 <0.1
±6.4
0
0
5.0 < 0.1
12.8
4.3 0.0
18.5 t 0.1
0
0
1.5 < 0.1
±2.1
0
0
0
0
14.0 <.0.1
±2.8
12.5 < 0.1
±0.7
Sua 7 7
# «t
1.0 < 0.1
±0.8
0.5 <0.1
±1.0
0
4.6 <0.1
±1.0
1.0 cO.l
±0.0
0.3 <0.1
±0.5
6.3 <0.1
±7.6
0.8 <0.1
±1.0
0.3 < 0.1
±0.5
0
0
0
4.8 <0.1
±3.3
7.5 y
±5.1
: k'i
Pall 77
# wt
Win 78
# wt
< 0.1
v. <0.1
_,.o
4.3 < 0.1
i*-6
20.3 <0.1
±10.8
12.3 <0.1
±4.0
0
2.0 <0.1
±1. 2
4.0 <0.1
±2.2
0.8 <0.1
±1.0
0.3 0.1
±0.5 ±0.3
0.3 <0.1
±0.5
35.3 y
±*1.9
9.5 y
±6.5
42.3 y
±16.0
5.0 <0.1
±0.0
0
2.3 <0.1
±2.6
To.5 <0.1
±5.5
6.8 <0.1
±5.9
3.3 < 0. 1
±1.3
0
1.3 < 0.1
+1.5
o.8 y
±1.0
0
0.5 <0.1
±0.6
21.5 <0.1
±9.7
8.5 <0.1
±3.1
37.8 <0.1
±9.0
-------
Beckatt Point +0
Spr 11 Saa 77
# *ft # vt
Couphi daa
Mopatra 0 0.5 <0.1
ornata *0.6
OphcilldM
Araeuila 1.0 <0.1 l.S <0.1
brtvla +1.4 +1.3
OrblnlidM
Scoloploa 3.5 <0.1 1.3 <0.1
•p. i 2.1 ± 1.0
OMQlldM
Omnia 15.5 <0.1 25. & /
fmalfonda ±20.5 ±4.8
f*ctlurlidM
Pcctlnarla 0 0
gram lata
Pbyllo4oclda«
1.0 <0.1 1.8 < 0.1
ieaga ± 0.0 ± 1.5
C. 0 0
tubarcvlata
K. 0 0
¦P. (J*v-)
¦ulalla 3.5 < 0.1 0.5 <0.1
^ngulou ±3.5 ±0.6
Phylledoc* 0.5 <0.1 0
iroclandlca t 0. 7
P. S.5 < 0.1 7.0 y
ucvlati ±10.6 ±4.6
P. 0 2.5 <0.1
ap. A ± 1.3
Pall 77
# vt
Irtn 78
# wc
0.3 <0.1
0.5
43.0 <0.1
20.0
18.5 <0.1
>- 6.8
2.5 <0.1
11.3
5.0 <0.1
4,6
34.8 <0.1
12.3
21. 8 < 0.1
10.5
0.5 <0.1
+ 0.6
0.5 < 0.1
I 0.6
0.3 <0.1
t 0.5
0.3 <.0.1
. 0.5
5.b <0.1
; 1.9
0.3 <0.1
± 0.5
0
0
3. G <-0.1
± 0.6
0
21.8
; 9.6
y
0. 3 <0.1
0.5
5.3 < 0.1
±3.5
0.8 <0.1
± 0.5
-------
Beckett Point +0
Spr 7 7 Sub 7 7
# vt # irt:
Phyllodoci Jm (cont.)
Phyl lotfcce 0.5 <0.1 0.3 <0.1
•p. +0.7 i 0.5
Polynoldm 0,5 < 0.1 0
•p. +0.7
Arctoooa 0 0
vlttata
Uarwrtho* 1.5 <0.1 0.5 <0.1
labrlcaca ±2.1 +0.6
H. 0 ~0.6 <0.1
lwwlaca ^0.5
L*pl4aath«ala 0.5 <0.1 0
b#rk#l«ya« +0.7
Slgaleald««
Fhel*« 1.0 <0.1 0
¦Imita +1.4
SploaldM
Halacocaroa 0 1.5 <0.1
gl«tMua +1.9
Pslyfera 37.5 < 0.1 259.8 y
social!¦ +53.0 +118.5
P. 0 68.0 y
ap. A +52.6
P. 0
• p.
Prioooaplo 4. 5 <0.1 2.6 <0.1
itMnitrvpl +2. 1 +2.2
Pygoaplo 2.0 < 0.1 1.3 <0.1
•lagana +0.0 ±1.2
Scolalapla 0.5 < 0.1 G
ap. +0.7
spio ~o 25,o y
fl llcomls +6.2
Fall 7 7
# wt
Win 78
# wt
0
0.P
< 0. 1
-1.0
0
0
0.8
< 0.1
"1.0
0
1.3
<0.1
±1-3
5.8
< 0.1
±5.6
T9.8
y
±22.5
1.8
<0.1
+ 2.1
5.8
<0.1
T //. 5
11.0
<0.1
±3.4
l.fc
<0.1
+2.2
"o
0.8 <0.1
+1.0
0.5 <0.1
+ 1.0
0
0.3 <0.1
±0.5
0.3 C0.1
±0.5
1.0 <0.1
±0.6
0
0
0.8 < 0.1
+ 1.0
~3.5 <0.1
±5.2
0
0
4.S ^0.1
+ 2.4
"C 5 <0.1
+ 0.6
~0
1.0 <0.1
+ 1.4
-------
Beckett Point +0
Sptool&M (coat.)
SpiOfhetM*
ba«i>TX
SylliJmm
trmi*
bwmtphmrjngim
foogoa*
gMlftri
S.
Ivwrmi
SphMrosyllla
plrlf«r«
S,
•p.
Syllit
¦ pp.
Cmttcat
D«cr*co
-------
Backett Point -Kl
Spr 7 7 Sub 7 7
# Wt # Wt
Cruatacaa (cone.)
I&opoda
tftoaphaaroaa 1.0 <0.1 0
aapltcanda *1,4
lanlropala 0 0
klncalal
taphlpoda
Capralildaa
Capralla 0 0
iMvitSCull
r inniii 1 Im 70.5 <0.1
ap>p. JQ
Fall
#
77
wt
Win 78
# wt
0.8 <0.1
1.5
"o
22.0 <0.1
*15. b
0.3 < 0.1
+ 0.5
0.3 <0.1
+ 0.5
20.5 0.1
+ 13-7 +0.1
0.3 <0.1
+ 0.5
0
-------
(cone.)
fcmeogancia
cI. ro*tr«t«
Sy^cfeiUdltm
r*cCip«lp«i
9»c«po4a
Cancar
C.
w.
C«cldM
•p. (J«V.)
Craagoaida#
¦p. (Jb*.)
htaotlj
•P-
Paganu
»Pf»* (J«v- )
PImUi
¦p.
PlBOOtlMrldM
»pp.
NpCt&i
grac'li.
Ptioroeiia
Pharanopaia
hmrmmrt
khiao4»rMti
fteblasldaa
Daxsdraatar
«xc«ntricws
Backctt Point +Q
Spr 7 7 Sum 77
# wt # vt
0.3
+ 0.5
< 0.1
—
3.3
+ 0.5
<0.1
0
0.8
+ 1.5
<0.1
0
2.0
12.7
<0.1
0
0.3
+0.5
<0.1
0
0.3
10.5
<0.1
0
0.3
+0.5
<0.1
J
1.3
10.5
<0.1
1.0
< 0.1
0.8
<0. i
n.4
11-5
0
0.5
10.6
< 0.1
0
6.8
15.4
<0.1
1.5
< 0.1
0.3
< 0.1
12.1
10.5
4.7
0.1
22.6
1.9
12.5
10. 1
14.9
12.8
2 ^ !
Fall
#
77
vt
Win 78
# vt
0
0.3
I0-5
0.3
10-5
0
0
< 0. 1
<0. i
0
0
0
0
0
0
2. b
15.5
0.3
_0.5
0
0.1
to. 1
< o. l
0.3 <0.1
±0-5
C.3 <0.1
10.5
0
0.5 <0.1
+ 1.0
24.8
+4.6
0.4
k0.1
12.0 0.3
+6.2 +0.1
-------
Bcckctc Point +0
Bolotk*reld«a
LcpCeaynapCa
clarfci
Spr 77
# vt
5.5 0.1
+ 6.4 +0.1
Sua 7 7
# vt
2.6 0.1
+ 1.0 +0.2
Sp«ci«s rLchnM*
Dlvcrilcy - ¦'
Total rt mb«r
cal bioaui (g)
68
2.52
1241.5
<61.3
S3
2.51
1464.6
<69.5
291
Fall 7 7
# vt
Win 78
# vt
3.5 0.0
1.9 +0.1
93
2.13
242 7.8
<77.0
2.o y
: i. 2
79
2. 30
1144.6
^69.6
-------
Table Beckett Point (sand.i bentn 1 c orga^isas fror thr subtida 1 zone collected 6 June 1977 , expressed
as number and/or bioaass (g) per 0.U5 m + standard deviation i N=-» J ,
-5 m.
# wt
-10 m,
# wt
Chlorophyta
Ulva «p.
Ihodophyta
Orraaalue nibnta
Cnidaria
Anthosoa
Actlncrla ip.
FLatyhclainthaa
Turballarla a pp.
rtaa app,
coda »pp.
Hoiluaca
Gaatropoda
Ac—Id— an. (Juv. )
Aglaja dioaadaa
Ai viola ap,
iapblaaa coluablana
CrrithiopaLa ap.
Goltihaliidag
Crapidula app. (juv.)
CyiIchna ap.
iMloowvtiiculi
Lacuna variagaCa
Margarlcaa pwplllua
Hargarltaa/Llrularla
Mitral la carlnara
Kitrail a tubaroaa
Maaaarlua Mndicua
Odoato*ia ap.
Turbonl1 la ap.
0.5 t 1.0
< 0.1
<0.1
0.11 0.2
0,3 J 0.5 <0,1
13.0 ±5.8 cO.l
8.0 — 4.2 -.1
0.3 10.5 <0.1
0
15.8 18.1 <0.1
0.3 1 0.5 <0.1
0.3 i 0.5 ^0.1
0.5 11.0 <0.1
0.5*0.6 <0.1
0.5 £0.6 < 0.1
0
9.515.9 <0. I
1.312.5 0.110.2
2.0± 1.4 <0.1
0. 3 ± 0. 5 <0.1
2.512.7 0.110.1
0
0
1.3 +0.1 <0,1
20.0 ±2. 5
18.0 ± 7.9
0
0.3 10.5
17.0 112.5
0
0
0
0.5 11.0
1.3U.5
0.3 10.5
3.314.3
0
0
0
It.3 ±18.6
1.0 10.8
2.8 *4.9
0.8 1.0
<0.1
<0.1
0.1 10. 2
0. 3 10.6
^0,1
<0. 1
--0.1
-0.1
0.1 ±1.2
0. 0 1 0. 1
<0.1
<0.1
-------
Backatt Point Subtlda1
livalvia
Acll« cMtrauli
ClLaocardlv nsttal111
Cranalla tectiitta
CryptoiBy* californica
Uiclneu txwlmlpti
Lyaoila californica
MacsM tp. (>v- )
Kya trmrla
Myaalla Ciwlda
NytllM »f. 0"v- )
Macula Catmia
Fode4«MM capio
Tallloa ap.
AooalLda
OllgochMCa *pf.
PolychMta
AHpharatidaa ap.
hir>tt arctic*
Capitallliaa
Caplttlla capieaca
IMlimtu ap.
¦oCosaataa llnaatua
ChMtoptarldaa
MaaociMMtoptania caylorl
Pbyl lochMtoptarua p roll flea
tflMhMtopttnii coat an*
Oorvll laid**
Frototforvlllaa gracilis
Glycaridaa
Ciycara ap. (Juv. )
wt
¦10
wt
0
0
D
o.s ; l.o
2.» t1.9
0.3 t 0.5
10.0 17.1
0
64.5 t 16.5
0
0
0.3.' 0.5
18.8 til .3
<0.1
<0.1
<0.1
0.2 10,3
0.1t 0.1
<0. 1
0.5*0.5
4.6 t 3.6
<0.1
0.3 t 0.5
0.3 i 0.5
1.8 i 1. 7
0
J. 8 ± 2. 1
0
1C.3 - 10.3
0.3 £ 0.5
153.b t 44.4
0.3 t 0.5
0.3 £0.5
0
7.3 ±6.0
<0. 1
<0.1
0.3 10.5
<0.1
0.1 t 0.1
<0.1
0.2 10. 1
<0.1
<0.1
0.6 t 0.6
1.3 ±1.5
<0.1
0
0.3 t0.5
< 0.1
0.3 10.5
0
<0,1
0
0.3 r 0.5 <0.1
0
1.0 t 2.0 <0.1
1.8*2.1 <0.1
1,5.' 1.7 <0. 1
1.8 t 2.9 <0.1
0
5.5 ±3.1 <0.1
14.0 ±13.9 < 0.1
41.3 ^49.3 <0.1
8.0 14.9 <0.1
0.3 >0.5
< 0. 1
0.8 t 1. j
<0.1
0.3 ±0.5
- 0,1
-------
Beckett Point Subtidal
ClyearlAav (cerat. }
Hnr9*dB* boreal 1*
Goatmild—
Cld«rk« tfabia
Qphiedromu pagettan ita
Lawbrloartdaa
Lnahrinarls tp.
Magaloaidata
N*|«loiui plUUal
UltaliM
tal*tb«lli rvbroclncta
¦eclyeene ap.
ilcaauclM Laabrlcolus
B. pmmtMmt*
R*phtytA»*
Iteptitya farru&inaa
M. ap.
I. ap. /
0.5 11.0
68.0+51.7
<0.1
<0.1
0,3 + 0.5 <0,1
0.3 ~0.5
<0.1
-------
Beckett Point Subtidal
Ophallidaa
Anundla bravla
OrhlnlldM
Scolopoloa
OwaniLdaa
Ovania faiUonli
FactlnarfIds*
PKtiDirla graoulata
my 11 ode da*
Staeoa looga
tulalia aanguiaaa
Phyllodoca grocnlandica
f. Mculaca
F. ap. A.
P. ap. (Jwv.)
FolyvolttM
larmothoa imbrlcata
H. 1unalata
Tanoola kltaapanala
Safeallldaa
Chona leal da*
Polydora llalcola
-S a. -10 m.
# nt ' ____ wt
14.5 13.9 <0.1 3.011.6 <0. 1
2.010,8 <0.1 1.311.3 <0.1
t.S £ 7.2 <0.1 0.3 tO.5 <0.1
0.3 1 0.5 <0.1 0.3 10.5 <0.1
0 0.3 1 0.5 <0.1
7,8*5.1 <0.1 12.5*4,2 <0.1
0 0.3 ±0.5 <0.1
0.5 * 1.0 <0.1 4.015.7 "0.1
2.3 14.5 < 0.1 6.8* 5.2 < 0.1
3.3 ± _. 1 < 0.1 32.5 *5.7 <0.1
2.8 1 1.0 <0.1 3.8+3.0 <0.1
0 1.3*1.0 <0.1
0 0.3+0.5 <0.1
0 0.3 10.5 <0.1
0 0.3 10.5 <0.1
0 0.310.5 <0.1
0 4.012.6 <0.1
0 0.5 11.0 <.0.1
-------
Splouldaa (coat.)
Poison pygldlalla
P. aoclalla
P. *p.
FrioMspio itMMtrupl
Spio flllcornla
Splophaoaa bombyx
SyLlfdM
Exogora* ap.
Sphatroajl11a brandhoracl
Syl11a app.
T«r«b«l 114m app.
Slpuncvla app.
Golflngla psjacccnala
Cniatac*a
Oatracoda app.
L«pCoatr«ca
Itaballa pwgaetcnala
Cwacei
Cwiilii vulgarla
Olutjlli ap.
Lap&octau ap.
TaoaldacM
L*ptoch#lla dubIa
laopoda
Llancria algarw
Manna itblfuata
Faranthura *l«gans
lackcct Pcint Subtldal
-5 a.
» «rt
1.3 ±2.5 <0.1
J.8 i 3.0 <0.1
1.0 *0.8 <0.1
81.5 £ 5.8 <0.1
O.bi i .0 <0.1
0.5*0.6 < 0.1
0.5 * 0.6 <0.1
Q.b 4 1.5 <0.1
0
0
0
0
2.8 i 2.1 <0.1
2.011.8 < 0.1
0.3* 0.5 <0.1
0,3 i 0.5 <0.1
0.3 1 0.5 <.0,1
63.0114,5 <0.1
0
0.3 * 0.5 <0.i
0. 3 ±0.5 <0,1
-10 IE.
# vc
0
54.0 *39.1
0.811.5
3b.5 t 10.b
0.3^0.5
0
0.5 10.6
0
/.*¦ 4.1-i
0. 3 ±0.5
0.3 i0,5
0.3 10.5
42.3 ±16.3
5.6 ±4.9
0.3 ± 0.5
0.3 ± 0.5
0
15.8*5.7
0,3 1 0.5
0
1.3 t 1.5
< 0.)
<¦0.1
<0.1
- 0.1
iO. 1
*0.1
<0 1
<0.1
<0.1
-0. i
CO. 1
<- 0.1
<0, 1
<-0. 1
<0. 1
<-0.1
-------
Bccke t Point Subcldal
-3 a.
vt
• 10 03.
# wt
Avphipoda
Ciwirldti app.
Caprcila ap,
Hetacaprc1la k«nn«rlyi
Decapoda
Calllanaaaldae ap. (juv. )
Cancer oragoncnais
Crangon ap.
Hippolydda* ap, (Juv . )
Pa£urida# ap. (juv.)
Pagurua ba**lngatma
Finnothartda* ;jj. (juv .
Pugettij gracllia
phoronlda
Phoronopaia hamcrl
Echinodarmata
Ophlaroldaa app.
Holothuroldaa
Lvptoaynapla clarki
148.b i17,6
0. 3 t 0.5
1.3 t 0.1
0.3 t 0.5
0.3 tQ.5
2.5 * 2.1
i.3 t 1,9
0.8 ± 1.0
1.0 t 2.0
1.0 i 1.2
2.K t 2.1
0.1 ±0,1
'0.1
4 0.1
*0.1
0.2 t0.4
O.Ot 0.1
*0.1
< 0.1
<•0.1
<0.1
0.0
: 0. t
151.3 ±42.0
0
0
0
0
0
1.3 ±1.0
ii.ot;.o
o
3.3 ±1.7
4.8 ±3.8
0.5 ±0.6
C. 3 £0.3
< u, 1
<¦0.1
*0. 1
^0,1
¦0.1
0.5 <0,1
0.3 t 0.5 i 0.1
0
Spcclaa richness
Divarally - H'
Total nuabar
k 3
2.77
70.8
90
3. 05
8 / / . 1
ToCal blomass (g)
-------
3^. t « i-xp-jiru sanc!-f»ravvl ) benchic organisms t ruir trie n igft < *o ' ) inCerl j
sprinK tnri.-i j/u winft-r ly>5, expressed as number .»nu •" :>r biomds.s {p, / per u.
n r J ii^v l «i; j , •
5 or
Sun
A-.ne 1 Ida
01 igocha#t»
spp •
Cri. *i ec«i
Aoritpodfl
»#7CTwridea
¦ pp.
Orchestoidea
Duget .rniU
S»-*eci£S R I chrwaa
1
1 v- r < 11 v, K
Tki', rtiiJser
Tr a 1 Blomaaa
wt
0.2 <0.1
¦* o. s
0.2 <0.1
±0.5
2
0. b*5
U.4
<•0.2
wt
le.f-.
i]4.3
1
0.00
1*. h
<0.1
<0.1
Fs 1 ! 7?
* wt
Win 78
# wt
0.7 <0.1
± r,. ¦>
1
0.00
0.2
<0.1
-------
; i l I rs'j . Uungr-r,»*ss. bp:t • t- x:: j s e tt sand-gravel; oentbic ufK^nistas trom cat nuc i+J,) int> r t 1 d a 1
l ,r.i- .oiiev'.eu s p r i * . i trirough winter 1978, txpressed as rural* i- r and/or bi jibjss (gi
-j , u ) t,' * standard Jeviation '1 N = .. .
Spr 77
• Mt
Sum 77
# Mt
Fall 77
# wt
Win 78
# wt
b m. : I e m
•op-
> -me ltd*
Oilcech««ta
¦pp.
Crustacfa
Ai^jhipoda
Cwir idea
«PP-
Orcb«f ,-oide»
p>iK«' tens 1*
0.2 <0.1
* 0.5
13.h 0.0
± 2n.t> iO.l
0.6 <,0.1 0
A 1.3
0.2 <0/1
*0. S
0.8 <0.1
* 0.8
28.6 tO.l
4.15.9
I; n lfi Richneaa
Diversity, H*
Tot» 1 Ni,*b*r
Total Bloaats
2
0.08
13,8
C 0.1
2
0.56
0.8
<. 0.2
1
0.00
0.8
< 0.1
1
0.00
23.6
< 0.1
-------
¦ 1 <• 'jc *un f! s s v Kpcsed ^and-gravei) benthic organisms from tfu- low ( »G 1 ) intcrtidal
uv.r ^ " I i e» ' eo tprin^ i 7 7 trirougn winter H 78 , expressed as number and/or Didnass g )
pr*r j. A -jf + star.oar: deviation lN=5j,
4o\laaea
Gait ropoda
A«phl«««
co lua*>larvc
L*CUR4
variant ta
Spr 77
# wt
0
n
Sum 7 7
~ wt
0
0
Fall 77
# vt
0.9 <0.1
+ 1.9
0.2 <0.1
+ 0. 5
Win 78
# wt
Btvalvla
Cllnocardlu*
•PP-
CreneI la
dacuaaata
0.2 <0.1
+ 0. s
0
c
0.2 <-0.1
+ 0,5
0
0
\nnrilda
Jllgocliacta
Dorvlllaldaa
frot edorvl I lea
graeilia
Sptooldc#
•pp.
0 ruatacea
Anphipoda
Ciwr d#a
•pp.
Faraantra
"Sarrata" n. sp.
0.4 <0.1
+ 0.9
0
0
2.0 <0.1
+ 2.2
n
0.2 to.l
+ 0.5
7.0 <-0.1
+ 5.2
0.2 iC.l
+ 0.5
"o
24.6 iO.l
+ 24.1
3.4 iO.l
+ 4.6
Rlchnraa
^Tfllltv, H1
' oea 1
Total 81c
3
0.69
2.6
< 0.3
2
0.13
7.2
CO.2
:>
0.27
26.0
<0.5
1
0.00
3.4
c o.i
] y
-------
• r *>-i. i'unit-ni s-i .Spit i .^,ncJ-s»ravc' I> bt-nt hi.
zone ¦' 1 ! t*.;-: L-d #t June 1 4 7 7 , r-xpressea as nu
—- *¦ st.ir.d.ird ;k-'. i it tr" ',", = ...1.
-5
Chlorophyta
CHattoac/rpha call fornlca
ladllirlophTti
Prmultn
Tbmm~_ ©phvta
Alar la app.
CoaCarta coatata
DtawmtU aculeata
D. llguUtt var. ligulata
Ictocarpacca*
ft IdatnU app.
Gtffordia ovaea
Baploglata andcraonll
Hxtophjllua a«aafle (luv.)
Laainartal** app.
L«*dnarla app.
Kaifaiacca«
ftalfala paclfica
StlctToalphon tort Hit
Syrlugodcraa abyaalcola
Usoddphyta
Aci jchattim app.
intlthtnlor defectum
A. app.
lotryoclad is ^aeudodichocooa
C« 1111 Ha«s. Ion / Pleaoa j>or lu«
Choreocolax polyalphonla*
Cruorlopala wltuirii
Crypeonaala borcallr
C. ovallfolia
luthora frutlci-loaa
Gl&arinaeeae
Conlotrlchua alaldli
I'M
¦rg.inisns fr.T. the ^ubtidal
• t r an'i'or (;u p<-r
m. -10 ra.
vt. * vt •
O.O-tO. I
0.1*0.1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0.2*0.U
0.2*0.2
0.1*0.1
O.HO.O
0.1«0.
0.0M).
0.0*0.
0.1-0.
0.0*0.1
0. W0.1
1.U2.0
0.0*0.1
0.1*0.1
0.1*0.1
o.lto.o
0
0
O.O-tO.l
0
0
0
0,010.1
0
0
0
0
0
0.1*0.0
0.0*0.1
0
0.CK0.1
0.0*0.
0.0*0.1
0.0*0.1
O.OtO.i
0.0*0, 1
o.oto. 1
0.0*0.1
0.110.1
-------
Dungep#ss Spit
I
*be4«phrra (cone.)
Gracl Urla ipp.
Cyncgongrui leptophyllua
Hiifwnifi app.
Bollanferrgl* aubulaea
Irldaaa ipp.
laptofaucNra paclflca
Ma^traooptera app.
Rsyaaowtlia pacifica
Pttycodryo laab«lll*e
Flacythawlon pec datum
P.
P. «pp.
Polys iphonl* hendr vi
P. paclflca
?orphyra app.
Pteroaipbeola d«ndrold«a
Khedyveata app.
SctMgrMM lnterrupca
Sfuaaariactcc
Tharttf1lnpalr pcggiaaa
Caldarla
Rydroaoa
Coryot Cnbuloaa 0
Antbosoa
0
Plat jbm lalnthfa
TurbelLarla app. 0
Nrwrrtaa app. 1.3*
Hoiluaca
Aaphlntura
Cryptobranchla concrncrlca q
SubtIda 1
-5 m. - 10 m.
wc. # wt.
0 0.2 *0.2
0 0.0 *0.1
o 0,1*0.1
0 0.1*0.1
c 0.1 to.l
o 0.1 *0.1
0 0.0 *0.1
o 0.1*0.0
0 0.0*0.1
0 0.0 * 0.1
o 0.1 10.1
o 0.1 to.2
o 0.11.0.0
o 0.1 *0.1
o 0.1*0.2
o 1.01-1.0
o 0.0 10.1
o 0.1 *0.0
O.OtO.l 0.0*0.1
0 0.1*0.0
0.3 * 0.5 40»1
0.5*1.0 <0.1
6.8 ± 2.6 <0.1
43.81 32.7 <0.1
0.3 ± 0.5
-------
Dung*n*aa Spit Sub^ldal
Melloaca
Gastropoda
Achilla coludblana
CtlyptrMi faatlgata
lae«M varltgata
ftoriarleea/LirularLa
Narprltci pupillua
Ml trail* tab*roaa
lUtlej cLauaa
Trlchecrepia canccllrtta
tivilvil
CllDOcardlu* nuttalll
CrtMlU d*tuaaata
fenlUrU k»nr*rleyi
Hacmn ap.
M. if, juv.
Mrs# ar«aar is
NtmIU tuaidi
Protothacn itialwi
rriBMRM 1 li cant 11 1a
Trcaua capax
Aim* llda
011gacha«ta app.
Myc ha* t a
Acroclrrldac app.
Capita 1114a*
Nfdlowicua app.
lotMatut 1 lnratua
Clrratalida* app.
Chaatotona aatoaa
Caul*ri*lla app.
Clrratulua clvracua
Yhary* aultlf111*
DttrvllLldtda*
Procodorvlll*a gracllla
Clycvrlda*
Glyclnd* plcta
¦10 m.
wt.
-5
wt.
1.8*3.5 <0.1
0.3*0.5 <-0.1
1.0*1.4 <0.1
0
0.3*0.5 <0.1
0.3*0.5 <0.1
C
0
0.8 11.0 <0.1
4.3*1.9 <0.1
2.0*2.7 <0.1
3.8*2.2 <-0.1
0
0
0.3*0.5 <0.1
0.3 *0.5 <0.1
0.5.0.
1.8*2.
0
0
0
0
1.
0
2.
0
5 '• 1.0
0*2.2
<0.1
<.0.1
<0.1
<0.1
0
3.3*4.6
0.3*0.5
2.3*1.7
0.3 *0.5
C.5*0.6
3.5 *4-4
0.5 ± 0
0.5*0.'
0.3*0.5
<0.1
0.4*
<0.1
<0.1
0.2*0
<0.1
<0.1
<0.1
<0.1
.9
5±1.7
3.
0
0
0
0
0
0
0
0.5 tO.6
<0.1
<0.1
13.5*8.8
0.5*1.0
53.0 *9.3
1.3*1.5
1.8 ±3.5
3.8 ±1.7
1.3*2.5
1.3 * 1.5
2.5 ±3.0
22.8*29.3
0.3 *0.5
<0.1
<0.1
0.0*0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
rn
-------
Dungeness Spit
-5 a
#
A:.wlidi (cent.)
FelrcKatti
Heaieaidac
GypCla btfvlpalpa 0
HIcropodarka dubla 0
Haldanlda* (juv.) 0
Axlotto 11a rubroclncta 0
Baphtylda*
Naphtya cillaca 0
H. f#rrugln*a 0
H. a?. 0
HereIda#
Ikrtii ap. 0
Platjrwrtli blcanallculata 0.3*0.5
Oplke lilda«
ArwodU bravla 0.3 ±0.5
TravlaLa for ill 0
Orblalldae
Scoloploa ap. 0.5*1.0
Ow*niid«*
Owvnia fualforaia 0
Phyllodeclda#
ItteM louga 0
KuLalla aanguloea 0
Fhyllodoca ap. (juv.) 0
Polynolda*
hraethw labrlrata 0.5 ±1.0
Sabcllldac
Chow ap. G
Sabelia aadla 0
Sea I Ihr* gvtidac
Scallbrcfpaa infla'tra 0
Sarpullda* app. 0
Splrorbla ap. 0
Ifi
SubtIda 1
wt .
¦10 ¦.
wc.
<0.1
<0.1
<0.1
<0.1
0.3 ±0.j
11.0 ±3.4
3.8 ±3.3
1.3 ±2.5
0.3*0.5
0.5 ±1.0
0.3*0.5
0.3± 0.5
3.5 *2.7
0.5*0.6
0
6.0 *3.4
0.5*0.6
0.3*0.5
0.5 *0.6
0.8 *1.0
0.5 *0.6
0.3 *0.5
5.3 *4.7
0.3*0.5
1.3*2.5
<0.1
-------
Di^ngrnes* Spit
-5 n.
#
AoneliAa (cotic .)
PolychMta
SlgallonldM
Ffwle» alau'.a 0
SplOAldM
MalacOcarsa f.lutaeua 2.814.9
Prionoaplo cirrlfera 0
P. aCetnatrupl 0
Sple filicornl* 0
Splophanea boobyx 0
Syllidae
Izogon* app. 0
I. feaaifcra 0
I. lourel 0
SphArroayllla pirrifera 0
Syllld#i loogocirrjta 0
Syllit app. 0
Tcrebcllldae
Fbl^clrnia ap. A u
f. ap. 1 0
P. ap. 0
Archlaaovllda
?elyg®r«iiid*e
Polygordlua ap. 0
Cruacacaa
Miitylia ap. 0.8*0.5
La^ropidae ap. 0.5*1.0
LtptoeiM ap. 0
TaaalJacca
UptocSflli dubla 0.310.5
V
Subc icJfc 1
wt.
• 10
wt.
0.5 tO.6
-0. 1
<0.1
0.3*0.5
7.3 * 3.8
1.3*0.2
0.3*0.5
<¦0.1
<0.1
<0.1
<0.1
20.6 * 24.5
0.3*0.5
4.3*4.0
1.3*1.5
1.0*1.2
5.0*1.4
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
9.6*11.3
1.0*2.C
7.0 *8.1
<0.1
<0.1
<0.1
5.0*2.5
<0.1
<0.1
<0.1
1.0* 1.4
0
0.3*0.5
<0.1
<0.1
<0.1
12.3*14.8
<0.1
-------
Dungeneas Spit
-5 a.
#
Cruacacca (cont.>
laopoda
Exoaptoarooa aaplicanda 0
I, rheariMiruai 0
E. ap, (Juv.) 0
G«®**iwerpha#re«B 2.0*2.7
Xrfoeaa (p. (Juv.) 0
MMptkip&ta
Caaarld«i 19.0* 23.6
D*c«p«i«
Ca 11 IimmldM so. (juv.) 0.3 ±0.5
Hrmigrapaua or* jconenala 0
fUppolyt ldae 0
!*jitfa« (juv. ) 0
Pagurtdae ap. (1uv.) 0.3*0.5
fliuwtHrldat (juv.) 0
Pugattla gracilis 0
Pft®r®«lda
Fhoronopiii h*r«ri 0
I yOXO#
Cchlnod#rwta
Qp^luroldea app. 0
Hel^churold«a
L«pto«faapta c larkl 0
Sp*cl*t Rlchnaaa 28
Diwrilty, 2.43
Total nuafetr 42.9
Total tlowsas (r) <2.8
Subtldal
-10 m.
wt. # *t.
0.3 t0.5 <0.1
5.8 t 7.0 ^0.1
1.8*3.5 <0.1
<0.1 0
0.3 tO.5 -( '
<0.1 28.0 *8.5 <0. 1
<0.1 0.3*0.5 <0.1
0.3*0.5 <0.1
0.3*0.5 <0.1
2.8 * 3.0 <0.1
<0.1 0.5*0.6 <0.1
0.3 * 0.5 <0.1
0.8 * 1.5 <0.1
0.5*0.6 <0.1
0 0.0*0.1 <0.1
1.3 t 1.3 <0.1
0.3 ± 0.5 <0 1
136
3.82
338.2
< 16. 1
-------
Ar>nc ' ( j
igochaefa tpp.
Pol ¦ *>Mtl
S v i i i d
f. xodorv* ioure i
Cru»
A(«phipo^«
naswMirid^a >pp.
8p#ei«?s Mtcht«e«B
m v*r»i c V, - 1
Tor m1 Member
T <> r d 1 81 1TMI s s
> r*• * " ir '' ¦ - ¦ : i -n i' \ - w .
i
^ h
» wt
1 .' *1.1 <0.1
U.2 * 0,/ < 0.1
' i-.< <0.1
3
0.7 b
2,2
0.3
-------
' , . " * - .".u rt it1,:! . -'t-
r .«?¦>! ', -r i, im,! s ¦ n i r
Ann*1Ida
01lgochavta spp.
Cruttacea
Iaopoda
Cnorimmhaeroma oret^onenae
Atafhi poda
Gamarldea «pp.
Dipt*ra lirvir spp.
Species Ric^rteis
Dlwnitv, Hl
Tota 1 Nunber
Tot a I ''
^r.iv«. ; i v-nthu1 ¦ • r < m i ->!r.s t roTi t hv
l i* 17 ~ . . vprrPh.-i! .i •>
- - •»[ .!p'; ? r«.i : • , . 11 i ¦ i V- S i .
11
1.4 + 1.5
< 0.1
1.6 + 2.1 <0.1
il8. 8 * 30.^ <0.1
7.8 - r-.?
U
0.63
34. ^
<0.4
c 0. 1
-------
.win ?*, ; • ¦v v '-xk/d : ' c ~
(-0- > in* t rt ! .1 ;i 1 ?."n« r
¦ r t. • r « •» < ;;t. r
Plat lair.thes
Turbc1 larla app.
Anr* 1Ida
Pol vc'ia^ta
Arenlcolld-e app.
CaplteIlldae
Mertlwaaatua acp.
Capit#lia capltata
Notomaat us ten«ls
Gtmladldae
Glvcinde pices
Lunfcr inerIdae
Lo-nbrlner 1 a sonata
Lu^nbrlner ia »pp.
N*r*Ida*
Hrrsla app.
Orblnidae
ScolopLoa app.
Paraoclriae
Par«oneils plat vhranchia
Splonidae
Ma lacpcfroa c lutarua
Prlonoaplo ateen; erupt
Cruatacea
Aiphleoda
Ca-naridea app.
Eotaua torfua waahtnrtonlanuB
Spec Ira R1chn#aa
Div*rait, h'
Total Nu<»b«T
Total Blomaas
! h«*nti>: • riUi-ii ii.-ri
¦ M'i'ted .spri:;,; i v / ^ x:i f. J
'.'J" + st rri j j :.
-K)
* Wt -
fi. 'i /o. 1
1.4 1 . 7 < 0, '
0.2 i 0.4 < ^ . 1
^~ * Q .' - . I
o.fi ± 1.~ 2
72.4
<1.5
-------
Neiwr?*a apu.
H^nstcsda spp.
Mo Ilusca
B1va1via
Mvse xla tuusida
Protothaca itaminea
i> lllna ap.
Anrw>! Ida
Oli#ochaeta app.
Pslvcharta
Cap 1te 1 lldae
M»dio(aa«tua sp.
Clrratu1idae
TV.jr,x imtlrifills
^>onladldae
CIvcinde picta
Nephtvldae
fwpphtv* cae col C" »
K. »p. Muv.)
Ke reidae
Nerei* procera
N. ap. fjuv.)
Platynerela blcanalieu lata
Orb In Ida#
Gainer la unclnara
S^olwplos ap.
Owenlldae
Qwenls fualfor-lB
Pi.vSlodocidae
Ereonr lonea
PS llodoc# "wc>. 1»* i
#
™ ^ rn.
wt.
i. 31 e.r
. 8 t 1.0
0.3 4 0.5
0.: t, 0. ?>
1.0*0.3
0.3 t 0."
20.5*13.0
2 s, s * 3 ' • ^. *>
3. S ± ?. 4
0.3*0."
0.3*0.-
0.3*0.^
OJtO.S
0. 8 ± 1.
0. 5 ± 1.0
3.5 ± 1.3
33.5*38.2
0.3 tfi.s
0.8 i1.0
CO. 1
<0.1
-------
Twin Rivers linbtirfal
Annelida (eont.)
Pol venae1 ta
Spionidae
Polydora socialls
Prionospio iteenstrupi
Spiophanes boob *
C" itti*. a
atracoda spp.
Cut«c?«
Olactvlgpals sp.
Isspoda
Edotoa sub littoralla
GnorIraoaphaerona or< goivnie
Svnidotea bicuspids
AwphIpoda
C4tmar idea s~p.
Specief Rirhrsess
Divert if'., H1
Total Number
Total 91onaaa
22 June 1^
wt,
0.3*0.5 <0.1
7.r, i3.fc <0.1
0.3 ±C. 5 <0.1
0.3*0.5
8R.3*2S.5
n.Stfl/
I * .r' *21.8
0.3 i0.r
74.3 »2i.9
2
1.-.
2.
<0.1
0.1 t 0.1
<¦0.1
0.1t 0.1
cO. 1
0.1t C.l
<2.8
-------
Table 8a. North Beach Sand (exposed sand) benthir •-rganisms from thi- high Oft*) inttrt id.il zone
collected spring 1977 through winter 1978, expressed as number and/or biomass
-------
North Beach Sand + 6
Spr 77 Sam 77
# w # wt
Cra*tae*a
Oatracoda 0.2 <0.1 0
•pp. + 0,5
Clrr&pcdla
Calanua 0 0.2 <0.1
carlo* im + 0.5
B. glandula 0 0.4 <0.1
+ 0.6
Mystdaeaa
ArehMoeyaia 0 4.0 <0.1
grebrutakll + 5.4
?anaidae«a
Lcptochalla 0 0
Isopoda
KuMiplMcroaa 0 0
aedia
Laalropala 0 Q.l
kiacaldl + 0.4
Amphipoda
CnMiridaa 0.2 <0.1 0
¦pp. *0.5
Dvcapeda
Pagurua 0.4 < 0.1 0
¦pp. (Juv.) 2 °-6
Spccica Richnrua 10 8
Diversity, Bl 2.02 1.48
Total Wuafeer 4.6 7.8
Total lieaua (g) <1.0 <0.?
zi:
Fall 77 Win 78
;> J#t ~ Wt
0 0.2 <0.1
± 0.5
0.2 <0.1 0
± 0.5
0.2 <0.1 0
+ 0.5
0 0
0.2 <0.1 0
i 0><;
0 0.2 <0.1
+ 0.5
0 ~ 0
0 0.4 0.1
± °*6
0 0
6 5
1.15 1.55
2.4 1.4
<0.6 <0.5
-------
Table 8b. North Beach Sand (exposed sand)
collected spring 197? rougn winter 1
on IN*'
Spt 77
~y
O.Oj m- stantla'd Ion iS=ri).
wt
art-tea 0,2 <0.1
•op. ~ 0.5
XimC(Hi 0.2 <0.1
app. +0.5
Melius ca
Gaatrepeda
Lacuna 0
variagata
Littorioa 0
scutulata
Bivalvia
Myaalla 0.2 <.0.1
tuasida 4- 0 5
Tel Una ~0
•p.(jUT.)
Annelida
Folycha#ta
Aranicolida*
Abaraoicola 0.2 <0.1
ap. +0.5
Capita It Ida*. ~
Capltalla 0
capitata
J&g«lc»id»e
Magalima 0
picalkal
Naraldac
Hercia 0
mllloia
faraonida*
Paraonc I la 7.8 <0.1
platybranchia +12.5
1c organisms frc-m tie "ud (+2*) inr ?r t id.i i zone
express^<¦ <.! as number .ini'or blomass (g) per
Sua 77 Fall 7 7 Win 78
# wt # wt § wt
0.6 <0.1 0.2
+ 0.9 +0.5
0 0
0 0.4
+ 0.6
0 0.2
+ 0.5
0 0
0 0.2
+ 0.5
0.4 <0.1 0
+ 0.6
0.4 <0.1 1.4
+ 0.9 ++ 1.7
0.2 <0.1 0
+ 0.5
0.2 <0.1 0
t °*5
6.6 <0.1 0.6
+ 5.<> +0.9
<0 I 0
0
<0.1 0
<0.1 0.2 <0.1
+ 0.5
0
<0.1 0
0
<0.1 0
0
0
<0.1 1.8 <0.1
+ 1.8
-------
North Beach Sand +2
Annelida (coat.)
Polycit*tC«
Mag* loalda*
Magtlona
pltalkal
Herald#*
Htull
vczilloaa
FiraoaldM
Paraonalla
plaCTbranchla
Sptonlda*
Ma iac«c#roa
glattot
Pygoaplo
• lagtaa
SyllldM
Scrtptoayllla
latlpalpa
Cruacacaa
Clr*lp*dia
ftalaiwa
carlo*u*
I. flandola
Hyaldacaa
AretUMOayal*
grabltakll
Cuaaec*
Diaatjlopais
¦ cp»
Taiuldacca
Leptochalia
dub la
I«Op«4a
Eaocphacroat
¦edia
SPT 77
# trt
Sua 77
# vt
0
0
7.8 '0.1
¦ 1 12.5
0.4 <0.1
±0.9
0
0
0
0
0
0.2 <0.1
to. 5
0.2 '.0.1
± 0.5
0
0.2 <0.1
+ 0.5
0.2 <0.1
+ 0.5
§.6 £0.1
t 5-6
0
5.2 <0.1
t 2-2
0.2 <0.1
+ 0.5
0
0
0 4 <0.1
t 0.6
0
0.2 <0.1
± 0.4
0
3! 5
Win 78
# vt
Fall 77
# Mt
0.6
+ 0.9
3.2
f 1.6
<0.1
<0.1
1.8
+ 1.8
1.0
+ 2.2
1.8
+ 2.1
0.4
+ 0.6
<0.1
<0.1
<0.1
-------
th Beach Sand
Spr
77
Uvm
77
#
wt
#
m
Crustacea (coot.)
Aaphlpoda
Sohaustorlus
120.4
0.3
303.6
0.4
•pp.
+ 28.1
+ 0.2
+
129.4
+ 0.1
Paraphozus
- Q
0.2
<0,1
c.f.abronlua
+
0.5
f. splnoaua
0
0.2
<0.1
Typt i
¦r
0.5
F.app.
0
0
Dacapoda
Crangoa
0
0
franaciscoroa
Spaclaa tlchnasa
9
13
Diversity, I1 0.33 0.26
focal Muster 130.0 318.6
Total Slows a (g) <1.2 <1.4
3 i h
F.1I 77
# we
Win 78
# irt
76.2 C.2
+ 60.6 + 0.2
99.4 0.1
«• 39.1 +0.1
7.8 <-0.1
+ 74
10
0.70
91.2
*1,4
10.0 <0.1
+ 7.8
0.2 0.2
ir 0.5 +0.4
10
0.66
117.6
<1.1
-------
Tabls 8c. North Beach Sand (exposed sand) benthic orpanisms fron the low (+0") intertidal zone
collected spring 19 7 7 through winter 197H, express* d as numbt'i and/or bioraass (r) per
t. 05 m2 + standard deviation (S=5),
Spr 77 Sua 77 Aut 77 Win 78
fr jrt # vt # wt # wt
Cnidarla
AochocM
Anthoplaura 0 0.2 <0.1 0 0
flfgiDtliiiai + 0.5
hart*! *pp> 0.3 <0.1 1.0 <0.1 0 0.6 <0.1
+ 1.3 + 0.7 + 0.6
lteaatoda app. 0 0 0.2 <0.1 0.2 <0.1
+ 0.5 +0.5
Nollidci
GastrojMMC
Uttorina 0 0.2 <0.1 0.2 <0.1 0.2 <.0.1
illkiM 2. 0,5 + 0,5 + 0.5
Bivalvla
Myaalla 0.4 <0.1 0 0 0
Cualda + 0.6
Annelid*
Ollgochaata 0 0 0 0.2 <0,1
•»< °-5
PolycKasta
Araolcollda*
Aturaclcola 3.2 <0.1 0.2 <0.1 1.2 *0.1 0.2 <0.1
(clapr*dl)ocaanica + 2.0 + 0.5 + 1.0 + 0.5
Capltcllida* 0 0.2 <0.1 2.4 <0.1 2.0 <0.1
Capita 11a +0,5 +2.2 + 1.9
eaplcata
ConladIdm
Glyecrld* 0 0 0.2 <0.1 0.8 <0.1
plcta +0.5 +0.5
Nag*lonlda*
Hagalona 0.? 0.2 0.1 0 0.6 <0.1
plctlkal +0.5 +0.5 + 0.9
317
-------
Worth Beich Saiul +0
Spr 77
Spr 77 Sua 77
# Wt * vt
Annelida (cone)
Polych»»ea
Bapht y Idae
Maphtys 0 0
•pp.(Jo».)
ii. eallfornlcnaia 0 0
•rbtnlidaa
Seeiopolot 5.6 v 6.6
-------
Morth Beach Sand +0
Spr 77 Sua 77
# vt # vt
Cr>iatac*A (coot.)
Archaeovyal* 0.6 ^0.1 59.4 0.2
grebnltskll + 1.3 — 52.6 +0.1
CVMCHI
DUitylopili 0.2 <0.1 2.0 <0.1
spp. +0.5 + 2.6
Laapropidae 0 0.6 <0.1
•pp. + 1.3
Iiopoda
3xMphMrOM 0.6 <0.1 0
wdia + 1.3
GnorlaoaphaeroaM 0.2 <0.1 0
oregonense ±. 0.5
Tectlcepa 0.2 0.0 0
pugettenala + 0.5 t. 0.0
Asphipoda ~
CaMarldea 0.8
•pp. — 1.3
Sohaustorlua 147.0 0.3 49.6 0.0
•pp. ±128.6 10.4 1 61.7+ 0.1
Paraphoxus - 0.6
c.f.abronlua t. 0.6
F. SplMMUS - 0.2
Type B +0.5
t. spp. 1.6 <0.1 0
t 1.5
Species Richness 16 21
Dlwrsltyi H* 1.11 1.56
Total Muaber 260.4 184.6
Total Bloaass <1.4 <1.7
319
Fall 77
* vt
Win 78
* vt
0.2 <0.1
+ 0.5
0.2 <0.1
± 0.5
0.6 <0.1
t 0.6
0
0.2 *0.1
+ 0.5
1.0 <0.1
- 0.7
0
0
10.8 <0.1
± 3.9
139.6 0.0
+ 62.1 +0.0
8.0 <0.1
t l-9
139.8 0.1
+ 92.9 +0.1
0.2 <0.1
5
18
± 0.5
1.02
189.6
<1.8
0
19
1.36
264.8
< 1.9
-------
Table 8d. North Beach Sand (sand-gravel-cobble) bench!c organisms fro^i the suhtidal zon#>
collected 24 June 1977, expressed as number and/or bxomass (g) per 0.05 m- + standard
deviation (S»4) at -10m, per 0.1 r«2 4. standard deviation (N*=2) at -5m.
-5 m. -10 a.
# vt. # wt.
•P-
Chlorophyta
Iryopilt 0 <0.1
coreleulaas
B. ip, 0 <0.1
ChMCCMprpha <0.1 <0.1
californlca
Dirbfili <0.1 <0.1
¦arlaa
•acilarlophyta
PiaaaUi <0.1 0
¦PP-
tbmophyta
OiiMnitU 0 <0.1
•eultaU
D. ligulata 0 < 0.1
war. Uguliti
Ictocarpalts <0.1 0
•p.
Laadaaria 165.5 0
aaccharina + 234.1
L. ap. 0.5 0
+ 0.7
Lawlaariaecac 0.1 0
t 0-1
Pterygophor* 916.3 0
californlca — 1295.8
talfaia
pacific#
Sphacalarla 0 <0.1
•p.
Svringodaraa < 0.1 < 0.1
abyaalcola
120
-------
(torch Beach Subtldal
Ihodophyta
Aerochaatiua
•p.
AaCltlmilon
d*f«CtUM
A. daodroldaua
A. kyltnii
AnclchMoiOMlL* pacific*
var. pacific*
lowacaa lscmia
lotrycladla
pwwi od 1 c hoc OH
&otryoglo«aua
farlovtaaliiai
Calltthaanloa/
Ptaoooapor luw
Callophyllla
f laballulata
C. hMMphjrlla
C. plnnata
C, ap.
Caraalw
eallfmmicini
C. gardniri
C. atrlccua
C • ap *
-5 au
# we,
-------
North Beach Subtldai
thodophyta (cone.)
ChorwocoLax
p«ly»IpbooLa*
ConaeaatlMa
il^Ui
Coralllnaacca*
•p.
Cruorta
pre fur •'<%
Crypto. i
bora .»
C. obovaca
C. ovallfclla
C. ap.
0» Leasarl«cea«
•p.
Dtlasaarla
d*clpl*n«
Kuthora
{rutlculosa
Cigartine
•p.
Goctlaophyllua
akottabcrgll
Cracilarla
•p.
Crlfflthala
pacifiea
¦slOMCciOa
g Land 1 forme
Halyaanla
cocclMa
Htrpoaiphonia
grandla
H. pluaula
-5
wt.
-10
wt,
<0.1
4.6
1 6.5
0
s/
<•0.1
0
0
< 0.1
— J.J
< 0.1
0.5
+ 0.6
<0.1
<0.1
0.3
+ 0.1
0
0
1.6
1 2.3
0
0.1
t o.l
0
0
s/
0
0.1
1 0.1
0.4
f 0.8
— u . j
0.1
i o.i
0.0
i o.i
<£0.1
0.1
t 0*1
"o.i
t o.i
o
0.8
± 0.7
<0.1
1.7
1 3.0
0
O.i
t 0.1
0
-------
North Beach Subcldal
-5 m. -10 a.
# wt. # wt.
fthodophyta (coat.)
H* r®roa tphomla
<0.1
0
JtpMlU
Hollcobcrgta
0
<0.1
¦ubulata
Hywnena
2.0
0
aaithll
f 2.8
H. ap.
0.8
+ 1.1
0
Iridaea
<0.1
1.8
ap.
t 3.6
Laurcncla
5.2
0
apoctabllia
+ 7.3
Ht^riaopCtr«
0
£0.1
pUtyphylU
IfeoagardhlelU
<0.1
0
balltyl
NltnburglJ
0
<0.1
¦odtriMlina
Nlcophyllua
3.8
0
alribllt
t 5.3
Odcmchalli
<0.1
0.1
f Ioccm*
± 0.2
0. vaah'nftonle..«li;
0
0.1
t o.l
OpunClclLa
70.0
0
califarnica
1 98.9
hcrocclli
V
0
aiddendor £11
PcyiiOMllli
s/
y
paclflea
Phycodry®
0
0.0
laabcliUt
1 0.1
Platyaiphooia
CO. 1
0
clev*Land11
Placythaaanlon
0
<0.1
twtcroaorphua
323
-------
North Beach Subtidal
-5 ea.
# wt.
fthodophyta (coat.)
PlaCythMsaloa <0.1
pcetloatua
F. vllloaua <0.1
Plocmlaa 0.4
tirtila^lMoi ++0.5
F. cocelaem 0
\ ap. 0
Folymira 0.3
latisaljH + 0.3
*-0.1
hcadryl
P. pacific* <0,1
P. icopalwlua <0.1
Ptvrochaodrla <0.1
woodll
Ptaroaiphcmla 1.1
blplmiata i 1.3
P. der«irold«s 1.3
t 1-6
P. gardn*ri
t. gracllia <0.1
PCIlota 0
flllclna
Rhodoptllum 0.3
plaaotin I 0.2
Ahedyaanla 0
pcrCuaa
ft. aclpitata 0
ft. ap. 0.7
+ 0.0
324
-10 «.
# wt.
0.2
t 0.2
<0.1
0
0.0
+ 0.1
u. 0
+ 0.1
0.2
+ 0.1
0.1
± 0.2
<0.1
0
0
<0.1
0.7
+ 0.6
C0.1
0
0.1
1 0.1
0.1
± 0.1
6.7
1 13.4
<0.1
1.2
+ 0.9
-------
North B*ach Subtldai
Rhodophyta (cont.)
Sarcodiotheea
furcaca
Scagclia
occidental*
SquaaarlaccM
•PP-
Stcnograw*
lntarrupta
Thurvtcllopaia
pcggLana
Tralllialla
lncricata
Cnidaria
Kydrozoa
¦PP-
Anthozoa
Salcaapa
d«c*a£entaculata
PlaCyhelalncha*
TurbalLarla
•PP-
Neaartca
¦PP*
Neaateda
•pp.
Melluaca
Aaphlnaura
•p.(Juv.)
-5 w.
vt.
-10
wt.
0
<0.1
<0.1
<0.1
0
<0.1
0.4
• J. 0.7
~~ 0.5
- 0.8
0.2
± 0.2
-£0.1
J
0.5
+ 0.7
<0.'
0.3
+ 0.5
<0.1
1.5
±2.1
7.0
± U. 2
33.0
+ 24.0
<0.1
< 0.1
< 0.1
4.8
1 3.4
13.0
+ 6.6
< 0.1
<"0.1
0.5
+ 0.6
0.1
+ 0.1
32 5
-------
North Beach Subtldal
Holluici (cont.)
Aaphlneura
Wpldosona
¦ertcnsll
Toniccll*
llncata
Gaatropoda
Amti
aitrt
AlvlnLa
•p.
Aaphlaaa
coluablana
CalllMtOM
llgatua
C»lyptra«a
faselglata
C«rlchiopala
¦P-
Colltmllm
Kkraw*
Lacuna
~ariegata
Lai la
cocfcrrellll
Hargarlre*/
Llrularla
M. poplllua
Raelca
clauaa
Odoatenda
• p.
S*ari#«la
dira
-5
wt.
-10 m.
# Wt.
3.0
±4.2
0.1
0.2
0.3
t 0.5
0
0.4
0.9
2.5
t 3.5
3.0
t 1.4
2.0
t 2.8
1.0
i 1.4
+ 0-5
- 0.7
1.0
± 1.4
94.0
•t 125.
-------
No'' - Subtldal
- > is.
# wt.
tolluaea (coot.)
Blvalvla
Aatarte 0
CB^IICtl
Clinocardiuai 137.5 0.1
mittailll i 154.9 I 0.2
Cr*o»lls 1.5 <0.1
d#cu*aata "L 2,1
Cryptowya 0.5 <0.1
callfornlca — 0.7
Intodvm 0
aaxicola
Clfeywris 0.3 2.5
iubobtd«ta t 0.5 1 5.0
Bualtarla 0
ktnmrrlyl
Uimi 0.5 <0.1
clatula jl 0. 7
Maeoaa 0
•p. (Juv.)
My** 11a 10.0 < 0.1
tualda i 4.2
Scavl* 0
rubroplcta
Tflllna 3.0 <0.1
•p. — 4.2
TranMonalla 10.5 <0.1
taut 111a 1 4.9
Traaua 0
canax
Aria* 1 Ida
Ollgochaata 17.5 <0.1
app. + 9.?
PolycKacta
Acroclrrlda* 0.5 <0.1
•pp. — 0.7
-10 a.
wt.
3.8
3.6
1.5
1.3
32.3
14.9
0
2.3
+ 0.3
0,0
1 0.1
0.1
± 0.1
0.3
t 0.5
< 0.1
0.3
i 0.5
0
43.2
t 86.5
0.5
1 1.0
1.0
t 1.*
0.3
i 0.5
< 0.1
< 0.1
2.7
±5.3
2.3
± 1.7
0.3
+ 0.5
0.0
± o.l
<0.1
62.5
+ 23.2
<0.1
0
-------
North Beaeh Subtldal
Annelida (cone.)
Folychaeta
Aerocirrldae
Marrochaeea
•p.
Capltellidae
Capi tella
capltata
Hedimaatua
•p.
Rot (Meatus
lioeatua
M. eeauia
Clrratulidae
•pp.
Caulleri* 11a
gracilis
Cirratulla
cirratua
Dor*1lieIda*
Ffotodorvi Ilea
gracilis
Eualcida*
Eunice
valeoa
Clycerldae
Glycera
capicaca
Head pod ua
borealla
Bsalonldae
ap. A
Micropodarke
dubia
Ophlodrowus
pugettenets
-S n.
wt.
• 10 w.
wt
4.5
+ 3.5
3.5
1 2A
0
1.0
1 1.4
0
3.5
1 0.7
29.0
— 25.5
0.5
10.7
7.0
19.9
4.5
t 6.4
54 0
t 12.7
0.5
t 0.7
<0.1
<0,1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
< 0.1
2.5
i 1-7
2.0
+ 2.3
0.3
t 0.5
2.3
t 2.6
0.3
± 0.5
8.5
1 6.0
0.5
1 1.0
5.0
± 3.2
1.5
+ 0.6
0.8
1 1.0
8.5
±4. *
0
20. j
1 15.3
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
< 0.1
<0.1
<0.1
328
-------
Month Beach Subtldal
-5 at. -10 m.
» wt» # wt.
llda (coat.)
MaUanldac 7.5 0.1 1.3 0.1
¦pp. (Ju».} +7.8 * 1.0
Axiochalla 0.5 0.1 0
rubroclncta + 0.7
HI come he 2.5 0.1 0
W»rlealla t 4.5
I. pcriouti 4.3 0.1 0
t 4.9
¦aphcyida*
¦aphtjr* 0 0.8 0,1
callfornicnala t. 1*0
1. ap, (Juv.) 0 0.3 0.1
i 0.5
¦braida*
krali 5.5 0.1 0
p*UBlci ± 7.8
H. «p. 1.0 0.1 0
± 1.4
Plac jmarela 6.0 0.1 10.8 0.1
blcaoallcu lata + 2.8 + 5.6
©naphid*#
Oouphla 28.5 0.1 0.5 0.1
atlgmtla 37.5 ± 1.0
Opbaiildji#
Ophelia 0 0.3 °.l
llaacloa ±0.5
Owrnlldac
Omnia 1.0 0.1 1.8 0.1
fualfonia ± 1.4 ± 1.5
Phyllodocldac
KuLalia 1.5 0.1 0
¦aeroeare# i. 2.1
K. algrlmculata 0.5 0.1 0
- 0.7
B. quadrloculata 1.0 0.1 0
t 1-4
I. vlrldl* 1.0 0.1 0
i 1.4
129
-------
Berth R#ach Sub tidal
-5 a. -10 m.
# wt. # wt.
Annelida (cone.)
Folychaata
Phyllodocl4ae
Eulalla
0.5
<0.1
0
•p.
++ 0.7
Braiomtra
~ t*.0
<0.1
8.8
<0.1
colmauldifflci.il*
± 0.0
± 2.1
Phy ll«*!oca
1.0
<0.1
0
(rotalaadlei
± 0.0
P. Bf. (jU¥.)
0.5
<0.1
0.5
<0.1
I 0.?
± 0.6
Pllarglda*
Slgariira
8.5
<0.1
0
cantaculata
± 12.0
PliiMldM
rialooc
0
0.3
<0.1
¦p.
±0.5
Polywoldae
1.5
<0.1
0,3
<0.1
¦p.
+ 2.1
± 0.5
lir«Bthot
2.5
<0.1
0.3
<0.1
l^rleata
± 0.7
± 0.5
Mar*©th®«
0
0.3
C0.1
luaslaCa
±0.5
2.5
<0.1
0
•?-
±3.5
3.5
<0.1
2.3
<0.1
fuodati
14.9
1 1.2
Oriopal*
45.5
<0.1
0
minuca
i m.3
Potaallla
0.5
•""1
a
O
V
0
InK+raadla
+ 0.7
Scalabragaida#
flcalcl>regaa
0.5
<0.1
0
lnf Latua
± 0.7
St rptilida*
Cruclgera
0.5
<0.1
0
zygophor*
i 0.7
110
-------
North Beach Subtidal
Annelida (ccraC.)
Pelychatta
9«rpy*
-5
wt,
-10 m.
vt.
1.0
~ 1.4
~ 2.5
± 3.5
0.5
± 0.7
0.5
± 0.7
42.0
± 5 9.4
27.5
t 38.9
0.5
± 0.7
3.5
±4.9
13.0
± 2.8
0.5
i °»7
"" 2.0
± 1-4
9.0
t 0.0
0
<0.1
co.i
<0.1
<0.1
<0.1
<0,1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
0.3
±0.5
0
0
1.8
± 1.3
0
1.5
fl.J
0
2.3
± 1.5
13.5
± 6.6
3.3
+ 1.5
<0.1
<40.1
<0.1
<0.1
<0.1
<0.1
3 31
-------
North Btach Subtldal
Annelida (cmt.)
PolychMta
SyUidw
Braela
brvvlpharyngea
Ezegooe
g#«sal f#ra
I. louni
I. app.
Plonoayllla
uraga
Sphaaroayllla
pArlftra
Syllia
araiilaria
SyllU
•p.
Trypaaoayllls
TtrtbtllidMf
Folyclrrla
ap. A
P. ap. B
Archlanotllda
Poljrfordlldaf
Pelygerdlua
¦P-
SaeeoclrrldM
Saccoclrru*
•rotlcua
SlpuncuLa
Phaacoloaoaa
agualiil
-5 a.
-t.
-10 m.
1.5
f+ 2.1
" 7.5
t l0*6
4.0
1 4.2
28.0
t 26.9
0
4.0
1
2.0
± 2.8
5.5
+ 4.9
2.0
± 2.8
0.5
1 0.7
0.5
L 0.7
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
40.1
<0.1
<0.1
1-3
1 1 9
7.0
Z 2.7
0.3
1 0.5
12.8
t 6-2
0
2.8
+ 1.'
<1.8
19.2
2 3
+ 1.9
•'0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
52.0
1 38.2
<0.1
28.0
1 22.3
1.5
t 1.7
<0.1
<0.1
0.5
+ 0.7
<0.1
-------
North Beach Subtldal
-5 !¦.
-10 at.
#
we.
#
wt
Cruataeaa
Oatraeoda
6.
<0.1
0.5
<0,1
•PF-
+ 6.4
+ 1.0
Cirriptdla
la lamia
1.5
0.5
0
•P-
+ 2.1
+ 1.0
Ltptoatraca
Seballa
0.5
<0.1
0
pogatcenala
i °-7
Nyaidacaa
Archa«omy»t«
1.0
<0.1
0
grabnlcskll
+ 1.4
Cuaaeaa
Cttawlla
0.5
0.1
0.5
<0.1
•P»
0.7
+ 1.0
+ 1.0
Eiaatylopala
J. 5
<0.1
0
ap.
+ 0,7
Tattaldacaa
Laptoctwlla
17. S
<0.1
8.8
<0.1
dub la
+ 13.4
+ 6.9
laepdda
Gaaciaitiropaia
0.5
<0.1
0
pa—mophlla
+ 0.7
Cirolana
1.0
<0.1
0
harfwrdl
+ 0.0
C. vanc«uv*ranala
0
0.5
0.0
+ 1.0
+ 0.1
Exeaptfeaarom
0.5
<0.1
- 0
amelicauda
+ 0.7
K. aadla
6.5
<0.1
0
+ 9.2
E. tbonburum
58.5
0.0
<0,1
+ 32.7
3-0.1
+ 3.4
lanlropala
1.5
<0.1
" 0.5
<0.1
analog*
+ 2.1
+ 6.9
I. trldana
0.5
<0.1
0
+ 0.7
Ti3
-------
North Beach Subtidal
Cruaeac** (e«*nt.)
laopoda
Ldotaa
rufvsccna
I. ip. (juv.)
Jserapsis
dubi*
J. bb«u
J. MtMl
JanirLlata
occldaotalaa
Llaaorta
alsama
Humw
ataphciUMnl
M. ublqulca
H. ip.
Amphtpod*
Gaamar idea
*PP*
terophiufs
•p.
Gapr«llld*a
Capreila
callfornlca
C. ap.
C. ap. (Jut.)
Mr Cacaprclla
kcntwrlyi
.10
we.
wt.
2.5
± 2A
0,5
t 0,7
5.5
t 7*8
2.0
+ 2.8
0.5
± 0*7
0.5
+ 0.7
13.5
+ 19.1
0
0.5
1 0.7
0.5
± 0.7
427.5
+ 328.8
0
19.0
+ 26.9
4.0
± 5.7
1.0
+ 1.4
0.5
1- 0.7
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
<0.1
0.6
+ 0.9
<0.1
<0.1
<0.1
<0.1
0
0
0
0
0
0
0
0.3
± 0.5
0
31.5
t 3.7
8.3
± 3.4
0.3
+ 0.5
0.3
+ 0.5
~~ 0
<0.1
0.1
~ 0.1
<0.1
<0.1
<0.1
-------
North Beach Subcidol
-5 a.
# vt.
Cruatacaa (coot.)
AapttiMcia
Caprcllidca
Trltalla 0
pillaaiu
Decapod a
Ca&ccr
1.5
0.3
ore gone na la
+ 0.7
±0.3
Cancar
0.5
0.6
productua
+ 0.7
±0.8
C. ap.
+ 5.7
± 0.*
Pfayllollchodaa
*~0.5
0.1
papllloaua (Juv.)
+0.7
Ftnalxa
~0.5
0.1
ap. (Juv.)
+0.7
Pui«etia
3,5
0.1
gracllia
+ 3.5
3 35
-10 a.
# irt.
0.3
t0.5
<0.1
0
0
0.3
±0.5
S
1.0
±2.0
1.0
1.2
0.3
0.5
1.0
1.4
1.8
0.5
0
0.0
±0.1
<0.1
0.0
±0.1
<0.1
4.5
± 3.1
0.1
-------
North Beach Subctdal
-¦> IB.
* wt,
Bryoioa J
•pp.
Echlnoderaata
Ophluroidea 0
•pp. (Juv.)
Kolothuroldaa
Lrptoaynapta 4. 5 0.1
clarki +0.7 +0.1
Aaeldlacea
Pyura 0*5 6.7
haustor + 0.'/ + 9.5
Specie* Xlchnea* 201
Diversity, i1 1.44
Total amber 1410.3
Total bie«ii 1202.1
) JO
-10 m.
# wt.
0.3
+ 0.5
1.5
+ 1.3
0
151
3.86
372.1
73.7
0.1
0.1
+ 0.1
-------
Table 9a. Kydaka aeacn (e.xposec sand; bench ic organisms from the nigh (+6*) intertiaai zone
collected spring 1977 through winter 1976, expressed at. rmiaber and/or bioraass (g) per
0,05 bi ~ sr jndard deviation < N*5) .
Annelida
Ollgochaeta
• pp.
Polychaeta
CapitaIlldae
•PP'
Splonlda*
Pygoaplo
rlegana
Syllldae
SyIlls
• pp.
Crustacea
Itopoda
Clrotana
vancouverer.sl*
Amphlpoda
Gamaridea
•pp.
Itchyrocerus
angulpea
Orchettoldea
pugcctens1*
Insect*
Dipterin
Larvai spp.
Specie* Rtchnesa
Diversity,
Total Number
Total Blcxnaca (f.)
Spr 77
# Wt.
0
0,2 <0.1
+ 0.4
2
0,64
0 . .-5
<0.2
Sun 77
# wt.
0.4 -cO. 1
+ 0.5
3.6
r 2.1
0.2
+ 0.4
0
0.2
t 0
2.0
1.7
1.4
2.h
*0.1
< 0.1
<0.1
<0,1
<0.1
fc
1.23
7.2
<0.6
Fait 77
# wt,
O.n <0.1
1 0.9
0.2 <0.1
i 0.4
0.? <0.1
i 0.4
0
3.8 <0.1
+ 3.3
4
0.71
4.0
<0.4
Win 78
# wt.
0.4 <0.1
+ 0. s
0
0.2 <0.1
+ 0.4
0
2
0.64
0. 6
< 0.2
-------
lable Hh. Kydaka Aeacn '.exposed sand) Stnthic organisms
collected spring If 71 through winter 1978, expressed
U .03 b1, + standard deviation (. N — 5 .
Nemertea
•pp.
Annelida
Polychaeta
Ncphtyidae
Nephtya
califernlensla
Phylloducidae
Eteone
long*
Splonldae
Pygoaplo
elegana
Crustacea
Myaldacca
ArchaeMryala
grebnltzkll
Aaphipoda
Gaonarldea »p.
Kohauatorlua
washlngtonlanua
Species Klchneaa
I
Diversity, H
Total Kuaber
Total Biomaaa
Spr 77
# wt.
0
0
0
0.2 <0,1
+ 0,4
0,2 <0.1
t 0.4
0
0
2
0.69
0.4
<•.2
Si m 77
# vt.
0.6 <0.1
+ 0.9
0.2 <0.1
+ 0.4
0,2 <0.1
± 0.4
0
10.6 <0.1
t 7.1
0
4.8 <0.1
t 2.6
5
0.87
16.4
<0.5
i the raid l»3'i mtertidal zone
number and/or biomass igl per
Fall 77 Win 78
# wt. # wt
0 0
0.4 <0.1
0.5
rs
2.0 <0.1
+1.2
0
0
2
0.4 5
2.4
0
0.6 <0.
1 1.3
1
0.00
0.6
<0,2
<0.1
-------
Table 9c. Kydaka Beach (exposed sand) benthic organisms from trie law (<-0') inr e r 11 da 1 zone
collected spring 1977 through winter 1973, expressed as number and/or biomass (g) per
0.05 a- + staiHJard deviation C»'«5).
Annelida
Oligochaeta
•pp.
Polychaeta
Mage l
-------
Kvdak# Beach -H)
Spr 77 Sum 77
# «, # we.
Cruatacaa
Aaphipoda
Gaiwarldea
Sohauitorlua 10,8 <0.1 0
•P • — 1^.6
Paraphoxua 0.2 <0.1 0
¦p. - 0.6
Spectra tlchtwaa f 5
Dlweralty, H 0,61 1,25
Total nu«b«r 13.8 5,6
Total biataa«« (g) <0.6 <0.5
Fall 77
# wt.
Win 78
~ wt.
U
0
7
1.77
2.8
<0.7
0
0
1
0.00
0.4
<0.1
-------
Table Id. Kydaka Beacn >. .sand ; Dentine organisms trora
expressed as number and/or biomass (g) per 0.05 ra
-*» tn
#
Rbodophyta
Ant ithaimlon defectun
ChoreocolAx polyaiphemlae
Leptofauchea pactfica
Polysiphonla hendrvi
Pteroalphonla klplnnata
P. gracllla
Cnldaria
Hydrozoa app.
Neawrtea app. 8. 5 ±7.0
IWnatoda app. 3.3 IS.3
Mo!, luaca
Gastropoda
Ajaphlaaa coluablana C
Callloaeoa* ap. 0.3*0.5
Lacuna vartegata 1.8*2.8
Naaaarlua mndlcua 0
Qdoatoad* ap. 0
Trlchocropala cancellata 0
Blva1*1*
Cltnocardlum nuttallli Q. 3 ± 0. 5
Hlatc1la arc lea 0
Luelnoma tenuleculp^.i 0.3*0.5
Macoaa ap. 0
Myaella Cumlda 30.5*22.
Solen alcarlua 0.3 ±0,5
Tclllna ap. 1.3 ±0.5
Tranaennella tant111a 0
Anne 1Ida
Ollj^ochaeta app.
0.3± 0.5
subtidal son'.' collected 2 i .June 1^77.
standard deviation (N=4j.
-10 m.
we. # wt.
0 CO. 1
0 <0.1
0 <0.1
0 <0.1
0 <0.1
0 <0.1
0 J
<0.1 12.3 * 5.8 0.0±C.l
<0.1 23.8 ±12.6 0.0 ±0.1
0.3 *0.5 0.1*0.2
0.0 ±0.1 0
0.1*0.1 0
0.3*0.5 0.2*0.3
0.8 ±1.0 0.1*0.1
1.3 *2.^ 0.1 *0.2
0.0x0.1
0.0*0.1
0.1 * 0.0
0.0 ±0,1
0.1 * 0.1
0.8 t 1.0
0.8 t 1.5
8.8 *5,0
0.5 *1.0
82.8*79.5
0.3*0.5
2.8*1.5
*5.8*23.7
0.1*0.1
0.0 * 0.1
0.6 *0.3
0.0*0.1
0.2*0.1
0.0 *0.1
0. 1 t 0.0
O.w *0.2
<0.1
0.R ±.1.0
0.0*0.1
-------
Annelida (cont.)
PolyehMta
AnpHarceldae .p. (juv.)
Caplte 1lldae
Copier lla eapltaca
Nadlcmaatua ap.
Hfttonitui llneatua
N. ap.
- soptellida# app.
Cirraculldae ap.
Chaetoxona attoa*
Flabelllgeridae
Pharuaa pluooaa
GlycarIda*
Give*ra capltata
Healpodua boreall*
CVnladldae
Clycinde plcta
He* loti Ida#
Hlcropodarkr dubia
Lusster Irvertdae
LtMrf>rlnerla ap.
Magelonldae
Hagclena pltelkal
Maldanlda* app.
fflcoaaehe p«raonata
IWpht ytda#
IWphtya catci
N. ca*coldea
H. ap. (juv.)
Nartldae
Harala proccra
M. ap. (Jin .)
Platvnrrela btcanaliculata
Kvdaka Beach SubtIda 1
-¦> ti,
# Mt .
0
13.0 % 8,8 <0,1
12.8* 11.9 <0.1
0.5*1.0 <0.1
0
0
0
0.3 ± 0.5 <0.1
0
0
0
1.010.8 £0.1
0
0
0.5 i 0.6 40.1
1.8 12.6 <,0.1
0
0
1.5tl.3 <-0.1
0.3*0.5 <0.1
0.H0.6 <0.1
0
2.3 * 3.9 <0.1
-10 f..
# vr.
0.3 tC/ <0.1
IS.3*14.5
83.5*52.7
0.3 ±0.1
0.5 i1.0
0.3 10.5
1.H 1.0
0
0.3*0.5
0.3*0.5
0.5 *1.0
2.5*2.1
1.0 * 1.4
1.5± 0.6
0
17.
3.5 *-3.3
0.8 * 1.5
1.8 * 1.3
1.0 * 1.4
0
0.5 4 0.6
2.3 *4.5
0.0 i 0.1
0.0 i 0.1
-------
Kvdaka Beach Subtiaal
-5 m.
* wt.
Ann*114a («ont.)
Pol vch®ef,a
Onuph Ida*
Onuphia ?l*Rana
0.8 * 1.0
<0.1
0. atlfcinatla
0.3 t0.5
<0.1
O. ftp. (Juv.)
0.3 i0.5
<0.1
Ophellldat
Arwn
-------
Annelid® (coot.)
PolyeHarta
Spionidac
Polydora aoclalla
P. ap. A
Prionoapio cirrlf*ra
P. ateenatrupi
Pygoaplo elagana
Spto fillcornia
Splophanea btwsfeyx
Svllida#
Ex©*One R*anlf*ra
E, app.
Sph«*roay I Ua plrlfera
StrcptoaylHa latlpaloa
Syllla ap.
Terebellldae
Thelepua ap.
Slpuncula app.
Cruacaeca
Oatracoda app.
lcp teat: race
Nrballa pugectenais
Mvaldacca
Arehaatwyaia grebnltxkil
Cuaacaa
Cuaalla vulgaris
Diaatylla ap.
Dlaatylopaia ap.
Laapropldar app.
TMMidacaa
Anatanala ssertaanl
L*ptocbella dubia
Kvdaka Reach Subtidal
-5 m, -10 m.
* wt. # wt.
3.0*2.5 <0.1 2.« O.OtO.l
0 0.3 1 0.' < C. I
0 8.0*3.5 G. 0 » 0. I
37.0*19. L <0. 1 192.5 *70. 5 O.OtO.l
0 32.0 *3^.2 O.OtO.l
0.3 *0.5 <0.1 0.340.5 0.0*0.1
1.8*1.5 <0.1 3.8 i 1.0 O.OtO.l
0 0.8*1.5 O.OtO.l
0 1.8*7.8 <0.1
0 0.31 0.S 0.0 *0.1
0.3* 0.5 <0.1 0
0 0.3*0.5 O.OtO.l
0 0.5 t 1.0 0.0*0.1
0.3*0.5 <0.1 0
9.8*4.7 0.1 t 0.0 78.0*29.1 O.ltO.O
0.3 £0.5 0.0 * 0.1 0.3 * 0.5 0.0*0.1
0.3*0.5 O.OtO.l 0
0 0.3 * 0.5 0.0 ± 0.1
0 2.3a 1.5 0.1*0.0
23. 8*8.8 O.ltO.O 0.3*0.5 0.0±0,1
0.3 t 0.5 0.01 0.1 0
0 0.5*1.0 C.OiO.l
0 8.0*2.9 0.1*0.0
-------
Crmtacaa (cent*)
Iaopoda
Cyjfthura iwinda
Edotea iublltcoral it
LlanorLa algaru*
Hunna atephertacnl
Faranehura elegana
Synldotea blT^cuapida
Amphipoda
Ca pre 11 idea
Trite 11a plliaonla
Gamaridea app.
Decapods
Cancer or*gonenala
Plnnotherlda* ap. (Juv.)
Pugettla gracllla
Phorontda
Phoronooal? harmrri
Bryoioa app.
Cchlnodcrwita
Ophiureidca ap.Cjuv.)
Echliwldca
Dcndraater axcentrlcit
Helothuroldea
brptoaynapta clarkl
Speclea Rlchneaa
I
Dlvcralty, H
Total Ruofcer
Total Blooaaa
Kydaka Reach Subtidal
-5 <*. -10 ni.
* wt. # wr.
0 0.5±0.6 0.1*0,1
1.0 ± 1.4 O.ltO.I
0 3.8 1 7.5 0.0*0.1
0 O.Sil.O G.OiO.l
0 2.3 t 1.0 O.liO.O
2. (i 12.5 O.ltO.I 0
3.0 t3.b O.ltO.I
103.8.60.7 0.2 t 0.2
0,3 t 0.5 0.0±0.1
5.8 i 11.5 C.Oi 0.1
0
0
128.8*51.4 0.3 t 0.2
0
0.3 t 1.0 0.5 ± 0.6
0.3 t0.5 0.0 t 0.1
1.0*0.8 O.OtO.t
v/
0.8 i 1.5
0.0 t 0.1
y
0.3t0.5 O.OtO.l
16.8*18.6 0.3*0.2
54
2.55
311.5
<.5.7
¦! j 5
0.3 t 0, 5 O.DiO.l
0
0
96
2.83
870.5
Cll. i
-------
I'.i. I jHt- ^ t .'-.n
z. • .¦> 1 l»^ led -,;r.;,ini .in- ? smt
• r i ¦ u v. n v i ,i t r I '4 . H , fXp ri's-u-,; i nur
"1
» = - ; ^jcnt-r-wm:, r> * st. ;n .•.? r.!
r *.
: i + s %
vi.it I • n .
i n r >¦ r; :.! : i
! ) ss < i: I :>.• r
Spr 77
# vc
Sum 7 7
# wt
Fall 7 7
# wt
Win 78
# wt
Ncncrtea
• PP.
ParantMrtai
peregrins
0
8,0 otho*
iabricaCa
0.5 <0.1
t0.6
j..
-------
Cmatac#a
Cumacca
CumIIi
vulgaris
Tanaldacaa
Laptochalla
dubla
Aaphlpoda
Gaprallldaa
• pp.
G*»aridaa
•PP.
Fiimiiut
¦Pf- (j«v. )
Pugactla
gracllii
Inaacta
Dipt«t«o
larvae
Dolichopodidaa
•pp.
Sperlaa richness
D1 varsity - H"
Total maabar
Total bl(MM«« (g)
• town +ti
Spr 7 7 Sua 7/
# wt # wt
4,5 «• 0.1
t 6,4
18.5 <~0.1
t 26.2
0.3 4 0.L
* 0.5
32.5 <-0.1
±46.0
0
0
2.0 *0.1 0
1 2.8
U 0
b
0.07
7511.0
<0, 1
4
0.02
1110.7
<0.4
Fall 7 7
# wt
Win 78
# wt
3.3 -G.l
* 6.5
3.5 <0.1
±4,4
0.5 *.0. I
*0.6
0
2.0 <0.1
±1.4
0 0.3 *0,1
±0.5
0 0.3 <-0.1
0.5
0 0
0 0.3 -0.1
±0.5
4 14
0.07 0.73
J7A.3 46.9
<'¦ i <1.3
-------
Table 10b . Janestsrvn <, prot vc ted sand) ht-ntli ir organisms
collected spring 19 77 through winter 1978, expressed
0. 03 spring) and 0,i>2 > (N-4 » s ¦ irane r-winter} rs~
Spr 77 Sum 77
# vt 0 wt
Chlorophyta
Diva C 0
CHniiti
U. 0 0
•p.
thodophyt*
0 0
•atorlanua
Cnldarla
Anthotoa 0 0
•pp.
Hydrosoa
Corfu# 0 0
• p.
iMHirtc* 1.0 <0.1 1.5 <0.1
»n?. -1,4 M . 7
Pirmtacrtu 0 0
parcgrlna
Mraacoda 2.0 *0.1 10.3 <0. 1
¦ pp. 12.8 ±12.3
Molluica
Gaacropoda
Acmaaldaa 0 C
ipp.
Aglaja 0 0
dloMdla
Lacuna 0 0
varlagata
Oplachobranchla 0 0
34 S
in- mid (+1.4'} Inters idal ?.ou«.*
Rbcr and/.T l>i.->mdss (g) s.u-r
ndard vf.*t
Fall 77 Win 7 b
t vt # Wt
/
V
' y
/ 0
0.5 0.1
£0.6 £0.1
0.3 <-0.1 0.3 <0,1
*0.5 «0.5
0.3 <0.1 0
-0.5
6.3 <0.1 11.8 ^0.1
44.7 ±6.4
0
n
3
<•0.1
±0
5
0.5 M).1
0
3
^0.1
10.6
±0
5
o
o
0
3
<¦0.1
±0.5
to
5
0
0
3
<-0. 1
±0
5
-------
J*«a£ovr +1.4
Blvalvla
Macou
lnqulr.ata
M.
Dllltl
M.
¦ pp.
My t i 1««
¦P-
Tranacaaalla
cancllla
Annelid*
OligochMta
• pp.
PolychMCa
ArenicolldM
• M>- (jw.)
Capitallld*«
Capitalla
capitata
H*4tc»M«tus
•P.
¦otoaaataa
ap.
Clrratulldaa
• pp.
Gonladidaa
Clyclnda
plcCa
Lunhrinarlda*
Lusbrlntrla
• pp.
Spr 77
# wt
Sum 77
§ vt
0
0
3.0
*1,4
0.5
*-0. 7
0.5
±0.7
63.5
t 9.2
7.8
10.6
0.2
*0.2
<0,1
0.4
*0.2
0.5 3.?
10.6 ±4.7
0
0
19.0 0,1
±17.3
38.0 <0.1
*24.0
40.3 <0.1
± 24,4
2.5 <0.1
±3.5
17.5 *0.1
±28.7
43.5
* 24.8
±l'i
i . "f
0
<0.1
cQ. 1
90.8 <0.1
*69.4
1.5 -0. 1
4 / 7
0.5
t0. 7
40,1
j
to.5
<0.1
3-'. 9
Fall 7 7
# vt
Win 78
# vt
0.5 3.6
*1.0 *7.6
1.8 8.5
-1.7 f 9.7
0
0.3 0.1
* 0.5
180.8 0.8
116.4 11.0
542.0 0.2
*1046.0 t0.4
176.3 <0.1
*21.6
75.3 0.1
*36,2 *0.1
0.5 <0.1
L 1-0
0
0
0
0.3 <-0. 1
to.5
0
0.3 2.0
*0.5 *3.9
0.3 <0. 1
* 0.5
0
132.0 <-0.1
1 23.0
12.5 <-0.1
±10.0
53.3 <0.1
*25.1
23.5 <-0.1
* 8,9
0
o.e co.i
*1.0
0.3 <.0.1
10.5
0,5 <0.1
±1.0
0
-------
Janaatovn +1.4
Maidanldaa
¦pp. (jwv.)
N*phtyldaa
Maphtya
tuci
H,
caccoidaa
H.
call fomlanals
M.
•PP. (J«v.)
Herald**
•PP. (ad*It)
I.
¦pp. (Juv.)
Fl«tyn*r*la
blcanallcalata
Ophalilda*
Araandia
bravla
Orbinllda*
Halaarla
¦ pp.
Scoloploa
¦pp.
Paraoolda*
Paraoaalla
platybranchia
Spr 77
# vt
1,0 <0.1
i 1.4
2.5
-------
JaaaaCovn +1.4
Spr
77
Sua
77
#
wt
#
vt
Phyllodocldaa
0
0.5
<0.1
• pp.
f 0.6
If—in
19.0
<0.1
43.8
<0,1
ionga
* 5.7
f 13.7
E.
0
0
tmbmstmtwta
Eula 1 ia
0.5
<•0.1
0
aanguia«a
* 0. 7
Pfeyllodoc.
1.0
< 0.1
0. 3
<0.1
¦MulaCa
± 1.4
i 0.5
P.
0
0.5
<0.1
•»P*
* 1.0
Polynelda**
0
±°*1
i 0.1
•pp.
45
Spionld#«
0.5
<0.1
40.0
* 0.1
•PP. (j«v. )
±0.7
*- 72.2
Mai*c#€#*o»
287.5
<0.1
20.5
<0.1
gl«CMWS
*369.6
*14.3
Polr*ora
4,0
<0.1
47,0
<0.1
k«pl japonic*
* 1.4
t 27.4
P»
0
,0,3
<0. 1
qwadrllobaCa
i 0 ,5
P.
0.5
<0.1
0
• p.
± 0.7
Prloneaplo
0.5
<0.1
0
cirri fin
± 0.7
Pygospio
41.0
< 0.1
1287.8
< 0.1
•lagans
*??.6
± 1495.7
Seolalapla
0
0
•p. C
SpLophanaa
0
0
boabyx
33!
Fall 7 7
# wt
Win 78
# vt
0
17.8 iO.I
i 5.7
0.5 <0.1
11.0
0.3 <0.1
±0.5
8.0 <0.1
t 4.2
0
0
0.5 <0.1
i o.i
297.3 v/
120.8
73.5 v'
t 14.1
0
0
0
68.5 <0.1
t 1.5
0.3 <0.1
1 0.5
0
0
33.8 <0.1
-16.8
0
0
8.5 <0.1
±8.1
0
0
0
326.0 <0.1
i 38.5
6.S <0.1
1' %
u
0
0
49 5 <-0.1
±35.2
0
0.3 cO.l
to.5
-------
JttMstovm +1,4
Spr 77 Sun 77
# « # wt
SylltdM
txogon* 2.0 <-0,1 0
loural *1.4
Str*if»te#ylll» 2.5 < 0. t 1.8 tO.l
latlpalpa f 2.1 1 3.5
Sy1 lis 0.5 0.1 1.1 <0.1
¦PP.
Crustacea
1 0.7 ±1.5
CuMlla 2.5 <-0.1 0.8 *0.1
vulgaris *-3.5 10,5
Tanaldac*a
LaptochalU 0.5 <0.1 1.0 *0.1
uubl* ±0.7 f-
Isopoda
bMphMrsai 0 0
m*4lm
Aaphipoda
taf>«lice« _ 2.b <-0.1
asasslill 11.7
tepltho« _ 0.3 -0.1
1ac«rtosa * 0.5
Aaiasgarawnta _ 0.3 tO.l
confarvlcolus i 0.5
Capralla 0 0
la«vlusculs
Capr«lll4*« 0 0
•pp.
Corophlun 42.3 ^ 0.1 24.3 /_ 0.1
schema lew ^ 2 2, J 1 6.6
C. 0 28.5 -lO. 1
•alaonia * 27.2
Pall 77
# wt
Win 78
# wt
1.0 <-0.1
~0.8
1.0 40.1
1 0 b
0
4.0 cO.l
* 5.0
1.6 *.0.1
i 1.5
2.5 zO.l
±1.0
3.3 <-0. 1
12.0
1V0 <0.1
±
-------
Aaphlpoda (cont.)
Corophiiw
•PP-
Caawarldaa
• PP.
Paraphilia
aplnoaua typ« A
>e >(v
cf roatr.ta
Baeapoda
Carldca
•pp.
Plnnlva
•p. (j«v.)
Plnnotharldaa
¦ pp.
Dpotabli
pug«tt«nsla
Xcblnod«rMC«
S0leeh*troi
-------
Tjble iUc, J h t iwi < pr^tei'mi nan.:; benihic iire.ini^ns fr.ira tU,- low (h>' ) im <.• rt : d:» I ;nt-
.'Oilcctt'-j spr j ne If •• t h rin«r» »intcr 1 '¦? ,* K. citiirt >=*!•.i js r.-jni'v-r and ''«*r H I •>«•, ».«.«» <-.* • r
"\ 1
sprina I and 'i. fN«j; s:inni: r-w i-i t »• r i *n~ * st.-v i ./
blpirmata
P. 0 0 0 ^
•p-
CMdarla
Ar.tho«oa 0.5 <0.1 6.0 ^0.1 2.5 0.0 0.3 <0.1
•p. -0.7 15,6 11.9 *0.1 10.5
Ha)camp* 31.0 0,1 2.5 <0.1 0 2.0 <0.1
dac«Ht«ncacul aca ±11.3 *0.0 *-5.0 14,0
Kowrtti 2.5 <0.1 0.8 <0.1 3.0 <0.1 2.5 <-0. 1
8pp. ±2.1 -0.5 il.e 11.3
Wmmmtodm 0 0.3 <0.1 1.5 <- J. i 8.5 <0.1
•pp. i 0.5 i 1.3 ±9.5
Molluaca
Gastropoda
Alvlnla 0 0 0.3 <0,1 0.3 <0.1
•p. f 0.5 -10.5
Lacuna 0 0 0.3 <0. 1 0
varlagata -0,5
Acolidldaa 0 0 0.3 <-0.1 0
ap. (Juv. ) £0.5
-------
Bivalvla
CTjptamym
call fomlci
Hacom
Irqulnata
Macoaa
naawta
M.
¦ pp.
Protothaca
•taslnaa
Tranaaci'valla
tantllla
Traaaa
capax
A/in«llda
Ollgochacta
"PP-
Polychaata
ArvnicolidM
•P- (J*v.)
BranchioK* dan*
vlnc*nt1
C«pltallIda*
Capital la
capltata
HfrdicNMftttia
¦ p.
Clrrat«llda«
• P.
Spr 7?
# Ml
0
0
0
16.0 0.1
1 27 . 7 10.2
0
39.0 0.2
144.2 *0.2
0
584. 5 <-0.1
t 135.0
1.0 <0.1
£ 1.4
12.0 <0.1
-* 2.6
53.0 '0.1
* 6.5
3a.5 <0.1
1 2.1
.4.5 <0.1
l 20.5
JaractCown -H)
Sun 7 7
# vt
0.3 <0.1
?0.5
0.3 0.3
-0.5 * 0.5
0
0.5 0.3
*1.0 -0.7
0.3 0.9
i 0.5 * 1.8
13.5 <0.1
*1.7
0
238. 3 <0.1
t ft4.7
1.5 '0.1
*1.9
4.5 < 0.1
t 4.4
15.fi <0. 1
1 4.9
37.5 <0.1
* 14.8
1.5 <0.1
13.0
Fall 7 7
# vt
Win 77
# vc
0.5 0.5
10.6 - 0.6
0.5 0,1
10.6 -0.2
0
0
0
64.0 0,3
130.7 -0.3
0
495. 5 < 0. I
140.5
2.0 <0.1
t 1.4
2.8 <0.1
13.0
: 4. 0 <0.1
•».0
.4. 3 <0.1
i 9.4
1.0 <0.1
l 1.4
0
0
0.3 0.1
? 0.5 10.3
0
0
39.3 <-0.1
-16.6
1.8 0.4
1 0.5 10. 7
676.3 ~'
i.189.6
4.0 <0. 1
-'2.9
2.3 <0.1
13.9
58.3 <0. I
1 23.5
32.8 <0.1
i 13.9
3.8 <0.1
13.5
-------
Jaaeatowo +0
Spr 7 7 Sum 7 7
# wt # wt
CirraCwlida* (coot. )
Clnitilu 1.5 <-0.1 0,8 <0.1
cirratua *2.1 *1,0
tbmtjn 4.5 < 0.1 2.5 <0.1
¦alCifllU 3.5 i 2.4
Bcrvillaidaa
¦orvillaa lfc.5 *0.1 4.5 *0.1
rvdolphl * 23.3 ^ 2.7
Protodorvl1 lea 2,0 *0.1 0
gracilis t 2.8
tunicoidaa 0 0
•P-
Goniadid*#
Clycinda 0 0
plcta
Baatenlda*
Gypcia 0.5 <0.1 0
fcravipalpa *0.7
Ophiodroaftia 3.0 <-0.1 2.3 <-0,1
p«g*tt«nala 1 1.4 *1.5
0. 0 0
•p. k
L«*brii»«ri
-------
MaidanldM (coot.)
Axlothalla
mbroclncCA
Buclfwun*
d«lln«ata
l«phtyiAa«
itephtr*
CMC*
M.
ea«cold«a
N.
cali fornlanala
ft.
• p. (Jmv. )
tartla
vaxllloaa
M.
•p. (J«v.)
Platyn«rili
blcanalleulata
OnupMia*
Onuphis
lrldaictm
0.
ItigMtwC
oph«nid««
Armandla
bravla
OTblnlldaa
Italnarla
dcndrltlea
acown +0
Spr 77 Sua 77
» wt ~ lit
49.5 <0.1 26.3 <0.1
•40.3 -til,6
0 0
0
0
0
0
0
0
0
0
0
0
0.5
i 0. 7
< 0.1
0,3
10.5
1.0
i 1.4
0
<•0.1
<0.1
0.5 <0.1 0
t 0. 7
0.5 <0,1 0
±0.7
4.5
12.1
<0.1
0.5
i 0.6
< 0. I
0.5 < 0.1 0
10.7
Pall 7 7
# vt
Win 78
# Ml
20.8 0.2
£ 7.4 -0.2
0.3 <0.1
£ 0.5
0.5 <0.1
*1.0
0.3 <.0.1
-0.5
0
0
0
0.3 <0.1
1 0.5
20.5 0.0
± 4.4 *0.1
0
0
5,3 <0.1
± 1.9
0
b. 0 ^
1 5.4
0
0
0
0.3 <0.1
i 0.5
0.3 <0.1
— 0. 5
0.8 <.0.1
<1.0
5.5 <0.1
* 11.0
8.3 <0.1
17.9
0
0
7.0 <0.1
±3.2
0.3 <0.1
i 0.5
-------
JaMBtwm -H3
Spr 77 Sum 7 7
# trC # vt
Orblolida* (coot.)
tULoarli 27.5 <-0.1 11.3 <0.1
unclnata ±17.7 i 2,2
0 0
•p.
Phyi. lodocldaa
IC*on« 2.0 <0.1 0.5 <0.1
lcm ft a i 0,0 * 0.6
1. 0 0
tab«rculaCa
ftalalia 1,0 <0.1 0
• angwlriaa *1.4
I. 0 0.J 0.1
•p. {Jmv.) 0.5
Fhyllo4«c« 2.0 <0.1 1.3 <0.1
¦aculata *1.4 ±0.5
P. 0 0
•pp.
Polyaoidaa
Eanaotho* 2.5 <0.1 9.5 <0.1
labrlcaca 13.5 13.0
H. 1.0 tO.l 0
•p. {Jmv.) t , +
Sab«lllda«
Ukxm 0 0.3 ^0.1
•p. ±0.5
S p hm to4o r i d mm
SphMrodorvpala 0 0
•twit*
Sploal4««
Halacocaroa 32.5 <0,1 15.0 <0.1
glutaa..* i 19.1 t 10.8
Fall 77
# vt
Win 7B
# wt
3.5 *0,1
1 7,0
13,8 <0.1
10.8 V
* 7.5
6.8 <0.1
±3.9
0,3 <0. 1
±0.5
0.3 <-0.1
± 0,5
0.8 <0.1
±1,0
1.0 ^.0,1
± 0.8
0
0.8
±1.0
-0.1
3.0 <0.1
±3.1
2.5 <0.1
±2.4
6.3 *0.1
± 3.4
12.3 0.2
± 2.1 tO.2
1.3
i 1.0
0
v/
0.3 <0,1
£ 0.5
67,3 <0.1
- 2. 1
101.3
i 37,5
y
-------
Jaewatown +0
Spr
77
Bum
77
Fall
? 7
Win
78
#
wt
*
wt
#
wt
#
wt
Splooldaa (cont. )
Folydora
0
0.3
tO.l
0
0
•P-
£0.5
Pygoipto
0
0
0
0.3
<-0. 1
clagana
- 0.5
Spio
0
0.3
<- 0.1
0
0
filicornia
* 0.5
SyllldM
•ranla
0
0.3
'0.1
0
0
brrvlpharyngaa
*0.5
txogoo®
0
0
1
5
-------
Jaasestown +0
Spr 7? %mm 7 7
# vt # wt
Tarawa 11 14m (cant. )
Falycirrua b.5 <-0.1 3.3 *0.1
•p. i 0. 7 13.6
Sipuncwla
Calling!* 0 0
pvb«tt«nai»
Cr«•£•«•*
UptMCtaci
MaNtlia 0 0
p«g*t »
Oacracoda 0 0
•pp.
Mysidftcaa
Arche#eey«l« 0 0
grabnlttkll
ilia 0 0
vulgaris
Uftocvai 0.5 410.1 0
•p. -0.7
Tanaldac**
Laptochalla 40.5 *0.1 37.3 *0.1
*ibla 120.5 ±22,9
laopoda
tzQ*ph&*trma 0 0
a^lLcawd*
U«t*a ~ 0
¦p. (J«v.)
> f\ (
Fall 7 7
# wt
Win 7 8
# vt
•.0 *0.1
t 6.7
2.0 *0. 1
- 1.8
0.3 <-0.1
* 0.5
0.3 *0.1
? o.s
1.0 <-0,1
i 0.8
0.8 '0.I
*1.0
/. .5 <0.1
*9.0
0.3 - 0.1
* o.:
o
0
0
44.0 <0. 1
110, 2
52.0 <0.1
*19.3
0.3 <.0.1
I 0.5
0.5 *0.1
*1.0
l.tt <0.1
1 ,
0
f 1
A.. -J
-------
Ja«e»tovn +€
Spr 77
Sum
77
# vt
#
wt
laopeda (cone.)
Phyllodorma
0
0.3
^ 0 1
abdoaiaalls
* 0.5
Synldotaa
0
0
blcuaplda
Aaphlpeda
r«ilidc«
Gapralla
0
0.3
<0.1
•P.
t0.5
riM>rld«a
29.5 <0.1
-
¦ pp.
19.2
JnpithcMi
2.8
c0. 1
lacartoaa
±5.5
CallLopialla
0.3
* 0. 1
praCtl
*0.5
Corefhiua
0.5 0.1
0
»9P.
0. 7
Mailt*
0.5
*0.1
dentata
±1.0
Paraphomia
0.3
< 0.1
alalli a
i 0.5
•
1.3
«*0. 1
aplnoacs Cypa A
f 2.5
PoaCsganala
0,3
cO. 1
cf. Ivanovl
1-0.5
r.
5.5
1 0.1
ef. roatraCa
t 10. j
Tritalla
0
0.3
*0.1
pillaftna
1 0.5
Hacapeda
Calllanaaaldaa
0
0.3
« 0. 1
ao. (j«v.)
±0.5
CarIda*
0
0
¦ PP.
361
Fall 7 7
# wC
Win 78
# ¥t
0.5 *C. 1
i 1.0
0
0
0,3 <0.1
i|),5
54.5 0.4
* 9.3 -0.3
16.5 <0.1
! 5.5
0.3 0.1
0.5
3.5
1.7
0.1
0.3 iO.l
±0.5
0
0
0
0.8 <0.1
±1.0
-------
Decapod* (cone.)
Hapcacarpus
¦P-
Planix*
littoral is
P.
Bp. (jUV. )
Upogabia
pwgattanals
Inaacta
Dlptara
larva
Phoroa&da
Phoronopala
haracri
Schlnod«raaCa
Opharoldaa
¦PP-
Holocburoidaa
Laptoaynapta
clarkl
Spaclaa richnaaa
Dlvaralty - N1
Total nuabar
local blo*aaa (g)
Spr J 7
# vt
0
0
2.5 0.2
tO.7 ±0.1
6.0 2.0
*0.0 *1.1
Jaaxatovn +0
Sub 7 7
# vt
0
0.3 0.0
t 0.5 * 0.1
0.3 <- 0. 1
* n '
5
0
1.0 <0.1
± 0.0
1.0
± 0.8
<0.1
1.5 <0.1
i 0,7
0
0.5 <-0.1
i 1.0
0
11.0 ^ 0.1
- 4.2
47
2.36
1295.5
15.1
2.S < 0.1
* 1.5
57
2.42
605.4
6.3
FaU 77
# vt
Win 76
# vt
0.5 0.0
t 1.0 ± 0. 1
0
3.5 <0,1
5.5 7.2
10.8 £8.0
0
0
1.3 <-0.1
*1.3
8.3 cO.1
*5.6
0.3
± 0.5
<0.1
0.3 <0.1 0
± 0.5
8.8 <0.1
i 8.3
6.3 0,1
12.5 0.2
4.8
-------
Table lOd. Jamestown (sand) btnthic organisms from the subtidal zone collected 7 June 197?,
expressed as number and/or hiotr.aS:-> (.£) per 0.05 tn*- + standard deviation
•-¦r
"hinropVyta
Chaetcmorpha
ra!1for a lea
r,i d^phora
pygma ea
"ntoc ladia
viridl*
< P.'
< f . ]
r
r i
Fhaeophvts
Ec t r.ct r pus
sp .
:,a »-1 * 1 p«
P ' ' f S i a
sp.
"t 1 c t j "«s 1 ph'-n
t orr111<
*vr i ngoderma
ahy^slcrla
i .
<.r , i
r
r£ mi 'in
call f orr ! c sm
< r ^ j
r,. i
±C A
ip.i
t.r .1
r.r
4 / - I
r 1
±cr-
r
-------
Jaxps t run
•Ihr-rfophyta S; i^r. t , )
n,o» feer> 10x
pol ysipbc>niac
"thoriaceae
Eijthnra
?r if ic<; 1 osa
rauchpa
!acInlet*
r fvnlofri chu»
alstdtl {» f.. elegans)
"rict lar la
verrucosa
Hvomogortgus
1rptcpSv!1 us
f«\i 1 ynenia
callfrrnica
HerposIphonla
prandls
Hyiwnen*
ap.
Iridaea
sp.
Mewhranoptera
placvphylla
Pcyssmel T la
pa-lflca
Platytlvemniri
h e C e r omor pV.; irr
P ,
pec t Ina t . ir
V.
reversuw
r.
vt11os um
f 1 da'
V f ,
r .
< P. 1
1. e>
± ' J>
r
11 .n
± i , f
< r.1
*-< s
< e.i
-t 1 P . s
< <..!
c
r,
<0.1
0
< p, 1
r
<0.1
p
r
n
n
< f1.1
-------
Jarf-1". ti-.vr
ilfip''r fihvta f ccr. t , )
Plocaaiua
coccLmu*
Polyncvra
UtllllH
Polyilpbonla
handryl
P.
pacifiea
Ptaroalphonl*
blplOMtl
V.
d«ndrold«a
Rbodoptiltta
Khed^ul*
¦P-
Scagclla
occidtfltil*
fC«OOgr«HM
Intarrupta
Porlfara
¦PP-
Cnldarta
AaChosoa 2.8
•pp. 12.5
HalcMpa 0
dceaaCaataculata
Platyh*lalnchaa - Turballarta 0
¦Pp.
Rnirui 4.8
¦ Pp. 2.4-8
0. 1
±0.2
7.2
±12.9
0.2
i0. 1
*0.1
0
0.6
10.2
0
0.1
±0.1
<0.1
0.1
10.0
v'
0,1
to. 1
<-0.1
1.5
-*-1.9
0.3
±0.5
0.3
10.5
8.5
ri.7
o
0.2
±0.4
<0.1
0
<0.1
0
^0.1
<0.1
<0.1
<0.1
<-0. 1
-'0. I
<0. i
<¦0. 1
-------
¦ateda
•pp.
Mollusc*
^hlnwti
Nepalta
lignoaa
Gastropod*
Alvlata
• p.
Aaq»hl«aa
coltmfciaaa
BLtclaa
aacbrlchcil
Caljrptraaa
faatigata
Ccrlthioftals
•P-
Colliaalla
ochracaa
Lacims
varl«|«t<
Hargaritaa/
Uxslarla
Margarltaa
pupilIns
VaaaarlM
•M^icwa
¦atlea
clatua
Odoaeoalc
•F-
0®tsop©ta
tabvlata
•toiro Subtldal
66.8
137.5
-5 m.
wt.
<0.1
¦10 n.
21.3
11.3
wt.
<0.
0,3
10.5
0,3
10.6
20.5
t 7.2
0
3.a
±4.4
2.0
±2.2
2.3
±2.9
0.3
±0.5
0.3
±0.5
9.8
±6.2
0
0
0
0
<0.1
0.0
±0.1
0.1
±0.1
<0. I
c0. 1
«.o. i
<0.1
I. J
11.5
0.3
±0.5
0
1.1
±1,3
0
0.8
±1.5
11.3
± 6. 7
.'.8
±4.3
O.j
±0.5
0,3
±0.5
1.5
±3.0
0.3
±0.5
<0.1
<0.1
<0.1
<0. 1
<0. I
<-0.1
<0.1
<0.1
<0.1
< 0.1
w_
-------
Jamatovn Subtidal
-5 w. -10 a.
i jjt. # wt.
Cutropoda (coat.)
Thau o« 3
-------
llvalvla (coat.>
taaellU
twmmm
capax
Ana*IId*
AreklmMllia
PolfgOTdlM
¦p-
Tolfchmtm
tepbar«tlda«
Awpkarat*
arctlca
¦PP • CJ** •)
ifiitobrwchilM
AplitobrtBchu
¦p.
CaplttllidM
Capital!#
capita*a
HlilWMltM
• p.
C&aaeoptaridat
nif I lochaatoptaws
prolific*
SplochaaCopeartt*
ceataraa
CIrrata114a*
•PP.
Subtidal
-5 -10 m.
wt. # vt,
<0.1
<0.1
7,8
ii.y
1.3
±1.3
<0.1
0.1
10. 2
<0.1
12.0
±3.2
1.5
i 2.4
0.3
10.5
<0.1
tO.l
<0. 1
<0.1
1.0
10.8
39.8
z 15.4
<0.1
<0.1
<0. 1
<0.1
<0.1
4.0
14.1
1.8
±1.0
32.3
±38.6
< 0.1
<0.1
<0. 1
-------
Clrratwlldaa (cont . )
Cha«t:0xon#
lltOII
ClrratnlHS
clrratus
Th*ry*
¦uUlfillB
Oorvillaid##
frotodorvlilaa
gt»clllt
Dor* .a*
rvdolphl
Clycarldaa
Clycara
• p. (Juv. )
s*
¦wrlcim
C.
capi tal a
Con ladIdaa
Clyclnda
plctc
Gonlada
MCUlltl
Haslonlda*
Syptii
bravlpalpa
Hicrop+darka
dvbla
Lu*fcrlnartdaa
Lumbrlnarla
*p.
JlDMUovn Subtida
•5 m, -10 m.
wt , # vt.
3.0
* 1.4
0
3.8
17.5
11.8
i 7.3
I . 5
±1.3
0
1.5
11.0
1.0
t 1,2
2.0
i 0.8
0
c
20, 3
t 8. 7
4.5
*1.0
<0,1
<0.1
c 0.1
< 0. 1
<0. 1
t 0.1
* 0.1
«• 0. i
<0.1
5.5
i 3.9
0.3
20.5
b. 6
tb.b
2, b
Jf 2. 2
0
0.5
±0.6
0.5
— 0.6
0
2,8
t 2.8
0.8
t 1.0
0.5
•tO.6
31.8
t LI. 1
4.3
*2.2
<0.1
<0,1
<0. 1
4, 0.1
<0.1
<-0. 1
<- o. 1
«-0..
<¦0, 1
tO. 1
tO.i
-------
Raldatiidaa
•Pf. (J«v-)
Cuclywna
• P-
SiCOMCh*
f*ara©nata
Maphtyldaa
Napnty*
¦ p.
M.
clllata
Paraliaa
MaraLa
•P- (Juv- )
N.
procara
PlaCynarala
bLcanallcvlaCa
OnuphidM
Omtphi#
4rt tfttMni
0.
0.
• p. (J«V.)
Ophallldaa
Aaaotrypana
•alogaatar
Amanita
bravla
Travlala
forbaall
¦ town Subtldal
•5 Qi.
•10 m.
14.3
*10.4
5,3
t 10.5
20.0
t 7.4
wt.
< 0. 1
*o. i
< o, i
10.0
±8.5
4,5
*7.7
2.0
±2.5
wt.
«cQ. i
<.0. 1
4.0. 1
0.3
* 0.5
<•0.1
0
1.0
<0.1
11.4
0.3
<0.1
1.0
4.0, 1
i 0.5
10.6
0. 3
4 0.1
0.5
40. 1
1 0.5
-*0.6
16,3
<-0. 1
8.8
tO, 1
17.7
t 8. 7
0
1.0
^0. 1
tO.8
0.5
'•0.1
0
ti.O
0
2.3
<¦0. t
41.7
0
0.3
<0.1
10.5
3.3
<0. 1
0
1 2.1
0.5
11.0
< 0. i
-------
Jnwatovrt Subtidal
# _ wt.
Orbinlldae
Scoloploa 2,8 '0,1
• p. 11.7
OimliiM
OmoIi 6.5 <0.1
fMlloraia 9
Paraoaida*
Arlcld«« 1.6 < 0.1
• p. 2-1.0
NctUirlidM
PactInaria 0
grata* lata
Phyl LodocldM
Iiaoat 0.5 <0. 1
lOOfa t I.0
Bulalis 0.3 cQ.l
••againaa - 0.5
Phyllo4oca 0
• p. A
Fhyllodoca 1.3 ^0.1
fro«olandica ±1.3
Phjrllodoc* 0
¦acwlaCa
Fhyllo4oca 1.3 < 0. 1
¦p.
-------
Janastcvn Subtldal
* vt.
Chooc 1.5 < 0, i
•caudata i 1.3
C. 0
• pp.
Btacho&a 0
aoalla
laKKBOM 0
• p.
PoCMilla 0
lntar—ifla
f. 0
¦frlopt
Sabclla 0
¦*4L«
Icalabragaldaa
Scalabvagaa 0
lnflaCBM
Sarpvl Ida*
SpLrorbls 0
¦P.
Slgaileaiia*
Fholoa 1.5 <0,1
vicnita *0.6
Splooida*
Laaalea 0
cirraCa
N*lic»ctrn 0.5 0. 1
glwtaama 0.6
Polydora 0
• p. A.
-10 m.
# wt.
3.8
i 1.5
5.0
i 5. h
4.0
12.2
2.0
+ 4.0
0.3
±0.5
0. 5
i 1.0
13.5
2 10.2
< 0. I
< O.i
<0,1
-------
Jancacowr. Subtidal
-5 a. -10®.
# vt. vt,
Splenica#
-------
J*a««tovn Subtldal
Tar*b«llida« (cont. )
Plata
criKiti
Polyclrrua
apt A.
troclaa
graffi
-5 m.
# lft.
0
1.3 <0.L
*1.0
0
OIlsochMCa 13.5 <0.1
• pp. *14.9
Slp«nc*llda 1.8 <0.1
«rp. *2.1
Ph«acol»a«w 0,3 < 0.1
•gMBlali 1 0.5
Prlapvll**
Frlapalai
etetialua 0
CruiticM
Oatracoda lfc.b <0.1
a pp. ±6.7
Laptoatraca
Itebalia 0.3 < 0.1
pwgattanai. t *0.5
c«.«
Ha 0.3 <0.1
vwigarla i 0.5
Maatylla 0.3 <. 0.1
ap. t 0.5
Laapropl rfa« 0 5 <0.1
app. 1 1.0
-10 is.
# vt
0.5
i 1.0
0,3
* 0.5
1.0
± 1.4
?l.O
r 19.9
0.3
t- 0.5
0.3
±0,5
< 0. j.
<0. 1
< 0.1
<0.1
<0.1
<0. 1
0.3 < 0.1
*0. 5
lfc.8 <0.1
— 11.9
0.3 <0.1
i 0. 5
3.0
t 1 ,2
2.5
* 1.9
0.5
t 1.0
< 0.1
<. 0. 1
<.0.1
-------
Jamestown Subtltial
Taneldaca*
L#S>toctal1*
(tub la
laopoda
Bplcarldaa
•P-
Kkeaptatroan
¦»dt*
B.
rhowiwra*
CborlBioap.ia*ro»a
©r*goo«n«a
Ja«ropala
4«bla
Farantb«ra
•lagans
Aapttipo4«
Coropblius
•PP.
gmrldta
•PP.
Sttcapoda
Carlda*
•p. (Juv.)
Ciangen
Montcalla
Crangon
•P-
¦aptacarpua
aclaftoni
Hlppolytlda*
•P-
- 10 it. .
vt.
-5 ®.
wt.
132.3
0,0
68. 5
<¦0.1
t 41.9
t 0.1
t 43.6
0.3
< 0.1
0
-* 0.5
0.8
<0.1
0
* J .0
i.C
* 0.1
0.5
<•0, 1
t 1 .2
t 1.0
0
0.3
<•0.1
*0.5
0
0.3
4 0.1
10.5
0
1.3
< 0.1
tl.5
0.3
i 0.1
0
i 0.5
B3.0
0.2
76. b
0.1
1 26.8
t 0.2
±48. 1
± o.l
1.3
<0.1
1.3
<.0.1
t 1.9
±1.5
0.5
0.1
0.5
0.1
11.0
- Li. 1
11 . 0
10.2
0. r
0.1
0
i 1.0
*0.1
0.3
0.1
0
i 0.5
10.2
2.3
< u. 1
0
i 4.5
-------
Jawatovn SubtIda 1
-5 m.
vt,
-10 a.
# vt.
Dacapoda (cont.)
07«geni«
gracllia
Pagyru#
barlnganua
P8gHTld««
¦p. (j»v.)
Planlxa
•p.
PlnnattwrLda*
• p. (j*v. )
gracilis
Pugcttla
producta
PugaCtla
«p. (J«v.)
Spirontocarla
prltmota
0.3
*0.5
0
2.3
±2. 1
0
±Q
.3
0.5
t 1.0
7.3
£ h , i
0,
t 0,
0
t o
3
5
0.3
5
0. 1
t 0,5
0.1
Jto. 1
<0.1
<0.1
<0,1
0.0
±-0. 1
0.0
to.I
£ 0. 1
0.1
*0.1
0.3
-0.5
4.8
* 5.0
0
1.6
-1.3
1.5
±1.7
0
0.5
— 0.6
0
Q. L
1 0.3
<0.1
< 0.1
<0.1
iO.l
Phoronida
PhoronopaLa
harvari
2.3
t 0.5
<.0.1
kryosoa
¦PP.
Echlnodar*ata
Ophluroldaa
(Juv.>
tiolothuroldaa
Cupencacca
• p.
Lapcoaynapti
dark!
7
± U
0. 1
0.1
t- 0.
0.3
0.5
0.3
A 0.5
0. 3
1 0.5
0,1
t 0.1
0. 1
10.2
< 0. 1
-------
6p«cl«t rlchr.caa
DivcraLty - M*
Total niMb«r
Total blomaaa (g)
atovr. Subcldal
-5 m, -10 m.
# VC , # Wt .
136 15b
3.49 3.81
bbb.2 543.2
39.i 3.8
-------
378
-------
Table i. Tongue Point (rock). Complete list of ail benthie organism?
collected from spring 1976 through winter 1978. Checks indicate
tide heights where organism was. found.
+6'
+ 3*
-5m
-1 Otn
Chlorophyta
Blidmgia sp.
Cha.'tcsraorpha sp.
/
r"
/
Cladophora Hp.
*
/
V
/
Codi.im sp.
r'
Derbesia marina
/
~'
Ent>;roraorpha compressa
/
E. crinita
/
E. incest i rial es
/
¥
/
r
E. liriza
~
~
/
i
Mcnostroma fuscuns
~
/
/
Rhii.oc Ion turn r i par i Lm
/
/
t
f
Spongoroorpha coalita
/
/
¥
V
S. spineseeno
~
/
/
/
Ulothrix sp.
/
Uiva spp.
/
/
/
/
Ulvaria ap.
r'
/
Uroapura sp.
~
Baci11ariophvta
Pennales spp.
V
/
r
/
/
Phaeophyta
Alaria spp.
~
»•'
i
r
/
r
Analipus janonicus
/
/
/
CoatHria costata
/
/
/
Desnarealie aculeata
/
D. Hgulata
/
/
*
D. v i r id is
379
-------
TONGUE K)INT
+6'
~ r +0 *
¦5m -10m
Ectocarpus parvus
Fucus dis11cus
Giffordia ovata
Hedophyllum sessile
Larainaria groenlandica
L, saccharina
L, set. che 11 i l
¥
/
/
Leathrsia diffurraia
/
/
Nereocystis leutkeana
Petalonia fascia
Phaeostroph ion i rrej'.u Lare
I'terogophora califoriiiea
Py1aiel1 a teiu 1 1 a
Ralfsia pacifica
Saunderse11 a simplex
Sphacelaria norrissn
S. racemosa
Syringoderraa abyss i cola
Rhodophyia
At \ othat'C i um sp, / / / / /
Ahnfeltia piicut a /
Amp 11 s i phon l a pacifica
Ar.t i t hamn ion defectum / / r
A. dendroideum / / /
A. kylinii / / / /
Ant i thatnri lone 11 a pacifica var,
pacifica r ~'
ionneinai»onia nootkana /
losaiella orbigiana / /
B. pbiraosa / / / /
Calliarthron tubeiculoium / / /
Cm11ithaanion biserlatum /
C. pikeanum / / / /
3 HO
-------
ONGUE POINT
Ca11ophvIlls edentata
C. flabellulata
C. f lrma
C. haenophylla
Cetami lut californicum
C , pac i f i cutp
C. washingtontense
Choreocolax polys 1phoniae
C lathroraorphum parcum
Corallina Vancouveriensis
Constantinea simplex
Cruoria profunda
Crypconemia oLtovat.a
0. ova 1 i folia
Cryplofiiphonia wood is
Del esseria decipiens
Endocladia muricata
Erythrotrichia parksii
Far 1owia mo 11i s
Fauchea laciniata
Gigartina papillata
G1 oiopt* 11 i « lur • hta
Gon iraophy 11 um sknt t sbt?rg 11
Gon i oc r i chun elegant)
C.rac 11 ar i a verrucosa
Gratwloupia dorvphora
Gnitithsia pac i f ica
ilalosaec ion glandi tormt*
Halymenia coecinea
Harveye11 a rairabilis
Herposiphonia grandi«
Her pos i phon i a plutaula
-------
TONGUE POINT
Htiterosiphoma sp.
HiLdenbrandia occiden tale
HoLtenbergia nigricans
Hollenbergia subuiaia
Hvmenena flabelligera
H. multiloba
H, setche11ii
1r;daea cordats
I, cornucoptap
I. heterocarpa
I, Iinear e
I.aurencia spectabilis
Leptofauchea pacifica
'.ithophy 1 luni ap.
Lithothamnian califormcura
L. murieatuo
Me lobe si a tnediocria
Membranoptera ditnorpha
M. mulc i ranoaa
H . p 1 atyphy11 a
Mesophy11um conchatura
M, 1 amf 11 a t um
MicroclaJia borealls
M. couIteri
Neopt i Lota cal.ifomiea
N . hypno i dee
Odonthalia flocoaia
U. kamtacftatica
Opuntle11 a cal ifornica
Petracelia oiddendorfii
PeyiiuneIlia pacific*
Phycodry• sp.
Platythamnion pectinatum
P. vil lo«um
Pleonoaporium Vancouverianun
-------
TONGUE POINT
Plocamium coccLneum
P. paciiicu ra
I', tenue
Polyaiphonia her.dryi
P. pacifica
P. paniculata
P. acopuloru m var. villosu m
Forphyra paeuiolanceolata
P. ami th i v
P, sp.
Pterosiphonia bipinnata
P. dendroidea
P. gardneri
P. grac i1i.h
Prionit i« lanceolata
Ptilota filicina
Ptilota pectinate
Pti loth&mnionopsiB lejolisea
Rhodochorton purpureum
Rhodoglossum call t orm cum
Rhodomela larix
Rhodopt i1 urn plumojum
Rhodymen1 a californica
pac i f i c
R, p«lmat a
R. pertusa
Rhodymenioco1 ax bot ryoidea
Scagelia occidentale
Spermatophyta
Phy11ospadix scouleri
Potitwra spp,
Halichondria panicea
Hallclona permol1 lb
Oph1itaapcngia p«nnata
-------
TONGUE POINT
Cnidar i a
Anthozoa
AnthopLeura elegantissima
Bnlanophy11ia elegane
Tealia sp.
Hydrozoa spp.
Aglaophenia sp,
Plumularia sp.
Sertular«lia sp,
Sertularia sp,
P1 atyhelminthes
Turbellaria spp.
Nemertea spp.
Eraplectonema gracile
Nematoda spp.
MoLlusca
Aroph ineura
Cyanoplax dentlen»
Kathar i na tanicat a
Lepidoioria cooperi
L, raertenaii
Mopalia ciliata
M. hindaii
H. lignoaa
M. muacoaa
M. ip .
-------
TONGUE POINT
Ton 1 eel La 1 meat a
Gast ropoda
Acteocina sp.
Acinaea mitra
Aeolididae Bp.
Alvinia sp.
Ampli1hbu columbiana
Balcts montei eyene is
Bar 1pela haliotiphila
Bittium eschiichtii
CalllOHtana iigatum
Cal/ptraea last igiata
Cerithiopsis sp.
Ceratostoma tollatum
Collieella digitalis
C. ochracea
C. pelt*
C. strigatella
Crepidula aculeata
C. iduncai
C. formcata
Ciepipatella 1ingulata
Cryptobranchia concent rica
Cy1lchna ap.
Diaphana 3p.
Diodora aspera
Fu«i t r i Con oregwnenaia
Cranulina margarilu1 a
Hernialopona baculum
Lacuna variegate
Lirularia lirulata
L. succincta
-------
TONGUE POINT
Lit tor iua scutulata
L, sitkana
Hargarites pupillus
Mi t re L1 a car inata
M, gouldi
M. tuberusa
Moelleria sp.
Naasarius mendicus
Notoacmea fenestrate
N. persona
N. scutum
Ocenebra lurida
OdostotBLfl sp.
Oenopota tabulata
Onchidella boreal is
Retuoa »p,
Searleaia dira
S iphunakia thersites
Thais canaliculata
T, emarginata
T. 1 ame 1 l oa a
Tricolia »p.
Tr ichotropib cance Llata
Velutina laevigata
Vermetua comp^ctua
Bivalvia
Adu I a call, f orn i ens i a
Chi amys hastata
hntodeama uaxicolf.
-------
TONGUE POINT
Hiatella arctica
Lasaea cintula
Lyonsia call form ca
Modiolus rectus
huseuLi.a diacors
M, pyg naeus
My sella t'jmida
Mytilus ca1itorniamis
M. edulis
Nftastoma sp.
Fer« i te 11 a pen i ta
Prutothaca staminea
Traitaennella tanc.il la
Treius capax
Zirfaea pilsburyi
Anne Ixda
Arch ianne1ida
Folygordiidae
Polygordius up,
Oligochaeta bpp.
Polychaeta
Ampharetidae spp.
Arfin co1idae up.
Branchiomaldane vincenti
Capi te11idae
Capilella caplc at a
Chry Htipet a I idae
Paleanotua belli#
-------
TONGUE POINT
Cirratul ida<-
Ciiratulus cirracus
Docecoceria concha r um
Tharyx multitiLis
Eun i cidae
Eunice va1 ens
He s i onidae
Micropodarke dubia
Oph iadroraus pugettensis
L.umb r i ner i dae
J.umbr inervs inflata
Mai danidae
Nicoraache persunata
Nere idae
Nereis pelagiea
N. vexillosa
Platynereis bicanaliculsta
Onuphidae
Onuphis stigmatua
Ophe L i idae
Arraand i a brevis
-------
IONGUE POINT
Orbimi da;?
Nsuieris dendr * f.ica
N, quadriciispida
Prot oar icie11 a oligob*anchia
Sculoplos spp.
Hhy L i odoe idae
hCeone longa
tula! ia bi I im>aca
!•,, nignmacul at a
K. quaarluculata
L. sa.T^u Hie a
t, vindio
Phy1!odoce sp.
Pi 1flrgidao
Sigsmbra lenfaculata
Polv"ioirl«t
Atctcnoe vittata
Mai oaynda Drevi«eU)t,a
Harmothoe imbi icata
Polygon i idae
Pt'iai.Hn! aspera
Sabei1idae
Chime ecaudata
Eudistylii p.)' ynoi ph*
K. Vancouver!
Fabricia curtegoniea
-------
TON (J UK POINT
Oriopsis miiiuta
Sabella media
S-ibe I la 51 at te sp.
Schi Z'"branch La insignis
Sabe 11 ,ir 1 i da«*
Iciatithy t and armatus
Sabellari.a ci'tr.>'n!ar ium
Scrpu!idne
Spirorbis nak.imi.ra i
S . spit 1 1 1 UTl
Si ga?on;due
I'ho 1 in1 mi nut a
jphaerodor i dat
Sphar rudoropsis uunuLa
jpionidae
Ma L 1 umb i *na
P. pygidvalis
P. spongi c
-------
TONGUE, POIN T
xogone getmiiterti
K. 1oure i
Odontosyllis sp.
Sphaet'osy 11 i s brandhors
S. pirifera
Syllis adamantea
S . a l ten sal a
S . hv a \ v na
S, stewaiLL
lernbellldae
iUa^iutrUt1 ciirat a
N i <: o 1 •; a /. o s c e r i c o L a
PisCa b ri: v i b raticii i a t a
Polyc1rr us spp .
3 i fwncu 1 i da
Phascolosoma agassizii
Crustacea
! is t r a coda spp.
Lirripedia
Balanus cariosus
a. glandula
H, nubilus
Chthansalus dalli
FoLlicipes polymerue
Amphipoda
Al 1 or cheste* (. ... -j 11a
Araphi 1 uchus sp. A.
Ampithoe ginulins
A. pp. C
A. me a
-------
TONC-UK HO INT
Aoroides culnmbiae
Atylus LnviUenai5
CaprelLa ^.slifornica
C. drepanechir
C. greenlyl
L. 1aevi uscu1 a
('.. swndax
C . n^t.i lens ; s
C. vv r r ucos a
Ceradocus .-ipinicauJus
Ct;: cop? compactuj
.'oruph iiuii brtv 1 s
'¦Jyin.wiuu:iciiiata
lieutt'l Is califormea
totsau stoi1 us sp.
Er1ctUonlup braailiensis
Gniiuiir op&i b sp, B .
0 . thouipsoa:
Hy&le anceps
(i. trequens
H. grand 1 corn i s cal it'ornica
H. pugett crisis
Ischyrooerus anguipea
J«nsa fa 1 ca t a
Kermystheus ociosa
Lepidepecreura sp, A.
L. gurjanovae
Lev cot hoe ep. A,
Maera aimile
Mf:lita californica
Metacapre.1 In anomala
M. kennerlyi
Metopa ristella
Najna ct. cone 11lorura
-------
TONGUE POINT
U11goch:•nur lighti
Orchomene sp, A,
0, sp- 8
i'anoploea hedgepsthi
Par-i 11 or ciie:-: tes ochocensis
Paramoera serrats
P, sp. E,
Paraphoxus spp .
i-arapleustes nautilus
P. pugettensis
P. sp, fi.
Photis b11 areata
P brevipes
P. sp. k.
Pleusirus secorrus
Pleustos depress..
Podocerus crista'.us
Folycheria osburni
Ponrogonij : a sp. D
P, intermedia
P. rostrata
Stenothoides sp
Synchelidium rectipalmura
Tr i l e 11 a pi 1 itnana
Cumaced
Bath/cuma sp.
Cornelia vulgaris
Diastylopsio sp.
1.arapropi dae sp.
Lepcocuraa sp.
Vaurtthorapsonia or.
-------
ToNGUi, POINT
Oecapoda
Cal1iannssidae sp. I juv , )
fani-i.-r or t'gon^ns 1 s
C. product us
Cry ptoIi thodes aitchensis
L, C y p i c u s
1) 1 scorsopaguruh uiimi ct l
11assuchinus term imanus
Fab i a su by uadr ata
Hfpta -Krpus stimpsoni
Lt'bbinjs sp. ( juv . }
M im.i 1 us t ii 1 i a t w
Ot'd i g(,u t bu J ir ermis
Oivgoma gracilis
Pachycho103 rudis
bor inganus
P, caurinus
P. da 11 i
P. h. hirautiusruluH
P, (tenner lyi
Pi" I ro 11 f, t hi.- * f r i orn«*rus
i'hy i I ol l r hodes papillosum
Hinno tlu*r es pu k e 11 »'iims
P<4: f I t i :i ^ r.i> ill*
S;u ron( ocai is prumota
1 nopatla
Hupy r i da«' sp.
Ciroian* hurtordi
i.o UnCliura iquaiulitv tiM
Dynmnifn«* 1 1«
D glabra
1). slit'a r»*r i
-------
TONGUE POINT
+ tl
~J'
+0
-5m
I Um
bxosphaf lom.i amp Li cauJa
K . me d i a
E, rhomburum
Gnoi Lit.oh pha-Toma oregor.eusf
Inniropus anal ">ga
1 . V i no» i d ?
I, minuLm
I, iridt'ns
I'Jo lea a c u 1 e a l a
i , njontf feyeiis i
f . reset* t«
I , schmt 11i
I at on«p:i
I . WtiblV.'S'-DSk i 1
JatropHis dub * a
J, lobata
J , set ObA
Ligia pal l \ i i
Limnoi-ia alga mm
Mt:nna chriMJUttocepha I a
H, Ht^phrnsenl
M . ub i i|ii l t a
Paranthura o 1 -gai;n
SynidaiM pet t lboneae
S . s p , A
'1 una i iJj'e i
Anai an* 11 uormarn
!,epl octie I i a dub l a
Pancnlui c«" 1 i tornion*lb
Insect*
ChironoEBidle »p,
CoLeoptera ip.
Coll oi*bo 1 a »p.
Dipteran larvae
l>ol i rlujpouidae sp
tphy .!r t dui- it...
~
/
-------
TONGUE POINT
~ u »J *0' - bm --1 Urn
Pycrsogonida
Ache 1la chr1 af a
A spineecta
Arnmot ht: I L a tuborculats
Halosoma corapar f nn
H . v l r l d i rU I i na ; e
Phoxi ch i L i d l urn t t-nicu at utr:
Fychogonum st « n: ns i
Bryozoa sop.
'6 i c r i s i a s p , ~
C he 11 o s 11 »Ria t n s d . /
LllSia sp. r
rilicrisia sp. ~'
F L u s t r e i 1 a s p . »•
HeLc^rnpora magna »
H. Sp. r
H i ppi/tsio/i by a I l nn /
M< tub rail i pr. r H sp, /
tth i nodel tnat r
Afi 11 roiJca
Hi'iirivia leviustula /
b'ptaster i a» he r sr t in r /
thtinjierias k i.-e!11 •»r i
Pis ««*•«¦£ ckhriei enus
Kiii i no i dt- a
S t r on #y I ot fiu iolui dr oebic h l ens i .1
S. irani iicinua
S, purpurati
-------
TONGUE POINT
1othuroidea
uuiuiciir l a rainut'i
C. pseudocL-ratH
Kiipent «c t « qu i nqm'semi t a
Lt?pf os;, napta c ' ark i
Parapt i chopus c a111 orn i crsiK
Thyone benti
Uph k ;.ro i d« a
\mph l phoL is sq'Odisijta
U i amph i ou i a oci'uientaho
ttin r da I a
A.Hi idiucca sp.
-------
Table-
r
Pillar Point (nH'k id il ; '"ino-rvJIun subt !«!nl J
mpleit- list of 1 1 bpnthic organ itsras collected from ••piing 1^7^
rou^li spring 1^7 7. Ch« -its tn-iu-.itv tide height whi« rt* organism w;i>t
f: iHind
h I ¦¦ '•! p h vf a
Hrvopsis sp.
• Urn u>ru>» pha sp.
('iadophorn.-t?:.*.' sp.
Enter phi i rinil a
K . j i! I e-it i ;l.l 1 J ;>
li, Lin /a
Mo no s t rorra t us cum
I'M ztu-1 cai hp r i pa r t um
Spon;»,«'Rnirplia ci;,i 1 < ta
S,
Tide Ht
-0*
• 5m
¦ 1< )ni
U K>t fit i x kj>
lilva sp;>.
Uro-spot-i hp
Hac i ila r i ophv La
Pi'nn,:;les spp
Phiii-aphv t .i
A1r i
Asia 1 isuis
Alciria spp
MS leys
Cvmat tir ri' t r i i f cat a
IVsn nest 1actil eat a
I), it »,n 1 at a
a "pus s ; ¦11 a;u;
f y rtv, i,« nn.'i'.i or.^ i 1
K<* idmaisnla s imp 1 ex
Finns d I sf I rh'ss
t 1 f t s>rd 1 a Hp,
Hedopiiv L luff. Be:inllf
i.ami ita r t jstm a 1 and i ra
i. , h.ii i !u r i -sa
L. si l.-h.' nil
i.rat hc.«. la •'. ! 1 t orni s
Ni' r »»o. v i r in tuetkrana
' V1 .1 11 a 1 a*. ! a
"tl. i ~*. ' . ,>1 . »l i r r < "m ' • •
-------
PILLAR POINT
+ t;
I'hcH'oph;, i a (tout . )
K i i < - i.i i111,i t' ^ P .
^phace1 a r la racruio-.
M tv-1 V1..1»phon tat 11 i s
Tide Ht
+ J' -0*
¦ >m
10m
Khodophvta
Ac rut*hat-1 ium sp, >¦ »
Aim t o I H a giga<"t inoid.-s •
A. pi u-at,j » > ~
*s: s t it h.itnn 11 - (s dp t i'c t um
A, dend ro i Jeum
A, kvllnii
Ant i i b itrn initi' ! 1 ;l >;! .in
A. pact? H: a
11 . f -
bos It-1 l,i planvsa
L:U 1. f t luirenion artiLum
C. p} ki-atuKTi
1 lophv] ! Is i i rna
Li"-;m(uni t'.UOP. 1 (llliim
('horedco lax po I vh iphnn i a<*
Curallirii) vancouver J p»sIk
I'rvntus J i .i wo mi t I
f)c* It* k t* i i a dre 1 p 1 fii* .
!)i I h.-.i eal f fornJ ca
Endue!adia nairicaua .
Krvt hropliv > i um do lcs-,at i' drs
I : r 1 ''hi a parks i S
{«i ga r t i n.» pap 1 H At i
i In 1 ope 1! i M ! i! r cat a
• ra• ! 'art a audi-r soil 1 I
{ - h j - - i i. -'.u : i
'.rat i 1 ar iupti11 a orv/oMt s
'giHigriJH Icpi.iphv 11 a*
-------
{'[MAR POINT
Rhiuiophvr a (11.itr , /
Ha Li.p.
Itiiiiit'.i .'¦> : ¦).:! a
I. heteroiarpa
i thut h.inin 1 mi c.i 1 1 t orn j cum
i.nphos iptu.-ni a i i 1 urn
Men;!' ranopt *• ra d lrr.^rpha
M. f"s] I f i ramo.-ia
M. latvphy ! ia
Mt iphvSlun i on,-hat urn
M11" r ¦ n• | ;iil tbt,r e,i 1 i s
Nti'ma i i hi v 1 m i rs i ho i dt?s
Neoaga i ait If I la ba i ley i
Neop L 1 ini,i ! en I(i i drs
U. r.'il 1 t.)! uica
Oi!r S 11
rule Ht,
+fj' + \' 4-0' - 5m - i Ott.
Phyt t<-! v s i sain111 f»e
F i kea . a I i I. u ii 11 .<
P ! at vt h.inri Icm hi-! e rumei phum
i'. |H 1 11,.I! ;im
w(ll...^ura
Kl'H ,:n i n::; l j tneu^j
P . t CITIi'
i'<- | vs I |»h.*n » i i.iMitrv :
r . j> j, M I. ,i
i ¦ * !' ' ' V
(1 i t-1 1 ;>h -ii i a h i | irtna'a
I .It nil I i 1 ilea
PC, H tit a tilt e 1 na
Ft Met ii.imn iui«<>p>< t «. 1 e i >> ! i
-------
PILLAR I'OIV?
+6'
H::; 11 • !sy f a (run? , )
KtrxuH'hurton purpurea®
Rhodug lossurn < a I i I o rn I c um
RUOiJotiiei .» l.irix
Htuidymen la ju» I but t«i
Sca^c i ia u-T Idt-nt a K'
Sr:ithura naiadum
"1 I H a't 11> 11 a h y nd»< r at
Sj>».-r3,.itophyta
F''hv I 1 »;p:u! i >. h (on ! <¦ r i >
t'oste .j n.arlna
Pu f t!cia spp.
Hal i >. Uoridrla pauicra
lia li< tonia p'rD' Hi,-,
t'pn 111 asp.:-m;i a pennaia
l-n Id i1" i a
:tv d ro/.'.a «;pp .
At iiufnaria :ip ,
Oln> I I a sp.
Phi a lid La ,sp.
An rln.i.'oa
Antlmplftira e !. j t i;, J a
S.' vpho/oa
Ha I Icl vst us aur I. ii 1 a
Plat vtu> I m i n t hen
Tin he 1 i ar I a •}<;•.
Nrm-- r 11-a spp ,
Aiipt. » IH'i'llK sp.
> f i «>h! a! i:; us sp ,
Nrf! : 11 IH; t -ipp .
*ti •! i j- \ a
Amph ! ni -,i i
I !i,l» f lip i i'U 1 i
s v .mop i a * ¦ Ii*.! t Stun
-------
r
.t itr« t u«" 1 t .•«
¦ -5>d f fa i- iHji .i
r. i ru-. l
. 1 i
i'fl t I'l' i J .1 I 1T t i' » t
•-11 upt» i I
\e%> 1 J iTuLh- >p;>.
A 1 v ! i>! ,i sr..
Amph ( s1 urrh L,in.i
.\reh! ii r i ch, t i 1
t'.'a i v<-. t as t f.ij:'t t <>
\ .i" ' ' ! 'i U'p its ' • ,x -
1 ' ' s i i »t n; i t i U s -t
v'. , '.'l' 1 I i
<". t; t r i g:1' e 1 1 a
(¦ rep i ,i< • i .i .
til ns.i t |1,U silum
1 ¦ ' 1:1 1 V.J ; 1 l' »-i l .1
I I t ¦ <>r i na s, ul.it.i •
' . v ! ; k.i-i.i
Ma vy.i r i i pu|> 1 1 1 us
Mi t rt'l i.'i i ai in.H i
Mttf i U't ! a sp .
N.il o.ii'ini'.'i {• t*r i¦ n.i >
N. hi s:Lutr. *
iMii' fiiml,, «»(i .
0i>- h i i- i I;! hi>i !», i i ••
' l.i ! i m-'-l ,i I ! »
I'h.t i •, • -ma It' .!.•«! i
: . c-i.H't! t n.il a
5. I ,iirn;-J 1-.
! url-on S ' I .t ¦<[> .
»V i >it i n.i i tii'v i g«.if ,i
VlTBt'i '.IS , OltiilllillS
-------
!' t S,:,AK POINT
>6' +3* +H' - 'm -1
H i va I v i a i t tun . )
Auul.i a IS.. rni crisis
i' I i tun¦ .i :"ii i um mi 11 a 111 i
H < a t I i-i arctica
Laraea <- i stuid
I uc I n;-roa ienm *J 1 n t a
M-n or.i.i uasata
Muscisl uk Py knt-vte11s
My a artnar la
Mys<-1 I. tumid.-;
Mv til wh iM L i t a rr» i
M, edit 1 1 s
t'anopea KA'ni" rusa
!Vn i i e 1 La i ••!', rad !
P r= >1 DI hac.i t.aii; i iiim
So l<-n sir >r i i;-
I V 1 1 i ihl spp .
1'rar, anr1 I a t r:t i .a
i ri'Hus capax
Zi rI a pishrvi
Aiii'.i" ; iii 1
01 , /arhaet I spp .
I'tv 1 vchnr ta
Apli rod I t i d.U' >rpp .
Ar;:tjc I 1 i 'iar
Ar abt' 1 1 a Iriroim
¦\ t n'.U > >) 1 Id-li'
ismalila se vlnri-nt i.
1 1 1 d. u
i i
,i > ,io I t at a
* .>? 5 •< t as
Iki J 1 i l V <1 { U '-t
I. «T1:, I s
il.U'! o|5{ !»r I I ¦ r
• i111 h.i. i \ . t - •; • .»st.uuii'
\
-------
PI* \R POINT
t-6'
*• i' +0 "
5 m
li)tp
'ulv h.n'ta tcont . )
( iirld.it
Pa 1eanot us be Li is
(It r ratu 1 idae
thaetuzorie setosa
i' 1 rr.uijlus c 1 rralut
1 t,.i rvx n;u 11 i f i 1 is
i)o rv i 1 I e ! d.it
r rot iu!crvi I lea grac I i ts
i: lab',-1 1 l r id ae
"!ii>rbs;i p I amusa
<;iy<'er Idae
*:iyt-or;i nmt" t 4 cana
',-n * .tu idao
s.Ivi'UhIi' arnitgi-ra
C . p 1 r t .1
hruHiii'..
maculata
lies lt>iii.J,ii>
t>yp L t hi i-vlpalpa
Henien Ida-- Hp. A
MlcrciputJarkt' dub la
Hphi idromus pugetLrnsIs
!,uffibr tnei i ciae
t. imbr Iner Is In! lata
i.onata
Ma j «ian 1 da v
KucLvmem* sp.
Nepht vidai»
Nephtyn raeca
N, fTH'CO i J(\i
S . f.i 1 i torniensls
ft. cltlata
N» i v I il.n*
Ner»'iN brandti
N,
N . vi' * 1 . losii
Plar vn«»r«-1 •» t icanal iculata
-------
PII LAR POINT
+6'
+ 3'
+0'
¦5m
1 Otr
Po 1 v chai'ta (cant . i
Onupniciai*
Onuphis e leg,ins
«). 1 r lclt'si-ens
0, stigma I is
wphe L i ! li.ie
Armartdia brevIs
Orblniidae
Maineri* dendritica
N. uncinata
• i op I or sp,
l 'v
! S diU
My!
Owe-
:it; ueulata
u:; L form Is
ParaonLdae
Arici di'.'i sp.
Pec t i iur 1 Idrit'
PectInaria granulata
1'hy i 1 odoc ldae
Eleant
E. tubercu lata
Eu1a 11a btlineata
!•'. quadr ioculata
E, »anguinea
E. viridls
Hhy1ludoce groenlandica
P. maculata
P, mu111serat a
Polynoidae
Ha Iobv dna brevisetoaa «¦ *
Marmot hoe irobrlcata
ti lunulata ~
Sahu liar' * da**
Sabellaria cementart urn »
Sabe 11 Ida."
Chont? ecaudata
Eudistylia Vancouver!
Fabr ic i.i oregun lea
-------
PILLAR POINT
+6' +-}' J-U' -bm ~10m
Pa 1 y<'h;n>ta tcont. )
S.'ihc 1 i 1 dae (font. i
Manayunkla aestuarine r
(i r i o p s 1 s rn 1 n u t a '
Pot ami 1 l.i my r lops r >
So be 1 1.1 media '
Srnlzobranchia ins v i .< ' 1
Seal Ibr.-^miclde
S c a 11 b r e a,raa In: I a t un 1 *
So rpti 1 idae
hpirorblrt 'lakamurai '
S. spIr i11 urn *
S i I Lon i dai"
PhnLue rntnuta T ' '
Sp lor1! dae
Mcil.u i)> eroi' glutaeus ' • •
Polydora culurr.biana ' '
P, litnlcola
P. S.!C 1 j1 Is
Is. sp. A *
Pr 1 oil!)npio cirrifera ' *
P. .stcenstrupi * »
Pvkosdlo o1y
Splo fLllcorairf ' '
Sp lopHants bomb v x '
Sv 1 I (dae
Hrrtnla brevi phn rvngea *
Kxiijionn rtemni f i r a * * '
K . 11)• i r «> 1
i'doiHosv Ills sp. '
I'lonosv Ills uraga '
Spii.it' > os v 1 1 is p 1 r i ! era *
St reptosivl 11'« lat Ipalpa '
Sv 1 1 1 d»m loiiRiu't r r ata '
Sv 1 L i -i adamant t»«s ' '
S. hyalina 1
S , pu 1 v*t.r a '
. s i i-wart I ' '
-------
PILLAR POINT
+6'
I'ol/chaeta (cent.)
T erebu11 idae
Thelepus crispus
S ipur.cul i 111 »'
Leptostraca
Nebulla pugettensls
Cumarea
Cumclla vulgaris
Ola,® .vllti sp.
Dlastylopsls sp.
L&mpr -lIdae sp,
Lupti jma sp,
ana tdacea
Anatanals nonaanl
Leptot hella dub la
Pani ol us cal1fornlensla *
Pspudotana is orn1 a tus
40 ;
-------
PILLAR POINT
+6'
Isopoda
Aega symmetrica
Cirolana harfordl
Cryptothir balani »'
Dynamenella glabra
D. sheareri »'
Edotea subllttoralIs
Exosphaeroma amplicauda »
E. media
E. rhomburum
Gnorlmosphaeroma oregonense /
Ianiropsis analoga
1. kincaidi
Idotea aculeata
I. fewkesi
I. montereyensis >
I. rufeseens
I. wosnesenski i »
Jaeropsis dubia
J. setosa
Limnorla algarum
Munna chromatocephala
M. stephenseni
M. ublqulta
Munnagonlum waldronense
Paranthura elegans
Synidotea bicuspida
S. pettiboneae
Amphlpoda
Accecomoera vagor
Alioichestes angustus
Ampelif ^ agassiz L
A hancocki
Amplthoe huaerails
A. me a
A, slmulans
Anlsogamarus pugettensis
Aoroldes coluabiae
Atylus levidenaif)
A. tridens
408
-------
PILLAR POINT
+6'
Amphipoda (cone.)
Calliopiella prat11
Calllopius sp. A
Caprella incisa
C. irregularis
C, Iaeviuscuia
C. riatalensls
C. pustulata
Caprtila verrucosa
Cercops coopactua
Corophlura brev Is
Deutella californica
Erlcthonius brasiliersls
Gammaropsls chompsoni
Hyale anceps
H, frequent
Ischyrocerus angulpea
Ischyraceridae sp, 2
Jaasa falcata
Megal uropua Inngimerus
Mellta californica
M. deadlchada
Monoculodea zernovi
Najna conslliovum
01lgochinus llghti
Orchotnene sp, A
0. sp. B
Parallorchestes ochotensls
Paraphoxus cognatus/simulana
P. nautilus
P, splnjsus
F. varlatua
Parapleuates pugettensis
P. nautilus
Photla sp. A
P. bifurcata
Phocis brevipee
Pleusyaetes sp. A.
Pontogenela Ivanovi
P. roacrata
409
-------
PILLAR POINT
+6' +3' +0' -5m -10m
Amphipoda (conc.)
Protooedela po. A r f
Synchelidium rectipalmum ' »
S, shoemaker! »' '
Tritella pllimana *'
Westwoodilla caecuia ~' »
Decapoda
Cailianajsidae sp.
Cancer gracilis
C. oregonensis
Crangon nmnitella
Cryptolithodes sitchensis
Hemigrapsus nudua
Oedignathus lriennls
Pagurus granoaimanus
i'. h. hirsutiuaculus
/
Pandalus montagui tridens
Pirinocheridae - p.
Pugettia Rraclils
Telrae&sus cheiragonua
Insecta
Coleopteran larvae
Dipteran larvae
Phoronida
Phoronis ijimai
Bryozoa spp.
Crista sp.
Kilicrista sp.
Flustrella sp.
Heteropora magna
Hippothoa sp.
Membranipora membranacea
Microporella sp.
EchinodPimata
Opiii .rnidea spp.
Aateiwidca
Leptasterias hexact is
410
/
-------
PILLAR POINT
+&'
Echlnodcnnata (cont.)
Holothuroldea
Eupentacta quinqueseiiiita
Leptosynapta clarki
Chordata - Ascldlacea spp.
411
-------
Table 3. North Beach Cobble (cobble over sand). Complete list of all benthic
organisms collected from spring 1976 through spring 1977. For subtid.il
see North Beach Sand, Table 8. Checks indicate tide heights where
organism was found.
„ +6' +3' +0'
Chlorophyta
Cladophora sp. - »'
Enteromorpha intest in;j lis /
E. 1lnza / /
Monostroma fuseurn /
Rhlzoclonium sp. r
Spongomorpha coalita / »
3. spinescens f'
Ulva spp, / /
Bacillliiriophyta
Pennales spp. /
Phaeophyta
Alalia spp. > /
Anal1 pus Japonicus /
Desmarestia aculeata / ~'
Ectocarpua sp. »
Fucus dlstichus /
Hedophyllum sessile /
Lftminaria saccharlna /
Leathesia dlfformis /
Nereocyst is luetkeana /
Petalonia fascia / ~'
Phaeostroohion trregulare / ~'
Ralfsiaceae spp. r
Scytostphon lomentaria /
Spharclarla norris si 1 /
S. racewosa »
Rhodophyta
Ant I thamn ion defectum J
A, dendroideum /
Antithamnlonella glandulifera .
A, pac if lea ~
Bosaiella plumosa ~ /
Calliarthron tuberculosum /
Geranium pacificuo /
Choreocolax polyeiphaniae / /
Cryptoaiphonia woodli *
Crytonemla sp. r
Dilsea califomica »'
Cigartinaceae spp. / »
412
-------
NORTH BEACH COBBLE
Herposiphonia grandis
H. plumula
Hollenbergia nigricans
H. sublata
Hymenena flabelligera
Iridaea cordata
I. heterocarpa
Lauren :ia speccatiiis
Loaentar ia balleyara
Membranoptcra weeksiae
Ml croc lad la boreal is
Odonthalia floccosa
Petrocelis midd^ndorf i i
Pikea californica
Plat vthamnion v11 losiur.
Pleonosporium sd.
Ploeamtum coccineum
Polyneura latissima
Polysiphonia hendryl
P. pacifica
P. panicuiata
Porphyra spp.
Pterorhondria woodii
Pterosiphonia bipinnata
P. dendroldea
Rhodoirela larix
Rhodopcilum plumosum
Scagelia occidentale
Spenaatophyta
Phyllospad Lx scouleri
Cnidar ia
Hydrczoa
gpp.
Anthozoa
Anclicpleura elegantissima
Platyhelmlnthes
Turbellaria spp.
Nenertea
spp.
-------
NORTH BEACH C03WLE
+6'
+3*
+0'
Nematode
3pp.
Moliusca
Amphineura
Cyanoplax dent lens
Katharina tunicata
Lepldozona mertensii
Mopalia lignosa
M. nuscosa
Gastropoda
Alvinia sp.
Amphlssa columblana
Archidoris montereyensis
Collisella digitalis
C. pelta
C. strigatella
Lacuna varlegata
Li tf.orlna scutulata
L. aitkana
Margarltes puplllus
Notoacmae.i f eniatrar.
N, persona
N. scutum
/
/
/
V
/
/
/
Qnc'ni del la borealia
Searlesla diva
Thais emarginata
T. lamellosa
/
/
Sivalvi a
Adula californiensia
Crenella decussata
Macoma sp.
Museulus pygmaeus
Mya arenaria
Mysella tumida
Mvrilus tdulia
Fretoehaca atamiuea
Solen sicariuB
Transennell* tantilia
Venerupie japonica
Oligochaet.
app.
414
-------
NORTH BEACH COBBLE
Polychaeta
Arenicclidae
Abarenicola sp. (juv)
Capitellidae
Capltella capitata
Decamastus gracilis
Mediomastus sp.
Notomastus tenuis
CIrratuiidae
Cirratulus cirratus
Tharyx multifills
Ik?rvilie Idae
Protodorvilisa gracilis
Kunicldae
Eunice valens
Clyceridae
Hemipodus boreal is
Lumb.iner idae
Lumbrinerls lnflata
Nephty idae
Nephtys caeca
Nere idae
Nereis vexillosa
Platynerei8 bicanaliculata
Onuphidae
Onuphis atigmatis
Ophellldae
Anaandla fcrevis
Ophelia limacina
Otbinliiia«
^roloplos app,
Paraonldae
Paiaonella platybranchia
-------
NORTH BEACH COBBLE
Phyllodocidae
Eteone longa
Eulalia quadrloculata
E. sangulnea
Hesionura colneaui difflcllis
Phyllodoce maculaca
Pisionidae
Pisione sp.
Polynoldae
Halosydna brevisetosa
Harmothoe imbricata
Sabel1idae
Fabricia oregonica
Schizobranchia insignia
Splonidae
Malacocerus glutaeus
Polydora columbiana
P. proboscidei
P. sp. A
Prionoaplo ateenstrupi
Pygospio elegana
Spio clrrifera
S. fllicornis
Spiophanes bombyx
Syllidae
Exogone gemmifera
E. lourei
Syllis adamantea
S. hyallna
Terebellldae
Eupolymnla heterobranchia
Crustacea
Ostracoda
spp.
Cirripedia
Balanus carloaua
B. glandula
Chthaaalus dalli
-------
NORTH BEACH COBBLE
Mysldace.i
Archaeomyais grelmltzkii
Cumacea
Cumella vulgaris
Lampropidae spp.
Tanaidacea
Anatanais notmani
Leptochelia dubia
Pancolus californiensis
Isopoda
Dynamenella shearer!
Exosphaeroma atppi 1 cauda
E. media
E. onctoncum
Gnortmosphaerotta oregonenae
Idotea aculeata
I. nonterevenais
I, resecata
I. vasnesenskii
Jati'opsis lobata
Munna chromatocephala
M. scephenseni
Aaphipoda
Accedotaoera sp.
Ampithoe Bimulans
Anonyx sp, B
At ylus tridena
Calliopiella prattl
Caprella inclsa
Corophlua brevis
Eohautorlue washingconianus
Guernea sp. A
Hyale frequena
Iachyrocerua anguipea
Megaluropus longitnerua
Melita callfornlcd
Ollgochinus lighti
Orcheatla ap.
Parailorcheatea ochotenals
-------
NORTH BEACH COBBLE
Paramoera mohrl
Paraphoxus spinosus
Parapleustes pugettensis
Phot is brevipes
Pontogeneia cf. ivanovi
P. sp. D
Synchelidiua shoemaker1
Decapoda
Cancer gracilis
C. oregunensis
Crangon munltelli
C. nigricauda
Hemigrapaus midus
H. oregonensis
Pagurus granosimanus
P. hlrsutiusculus
Ptigeteia gracilis
Insecta
Dipterarv larvae
Bryozoa
spp,
Enchinodcrraata
Holothuroldea
Eupentacta qulnquesenlta
-------
Table 4. Morse Creek (cobble over sand-gravel intertidal; gravel-cobble sub-
tidal). Complete list of all benthic organisms collected from spring
1976 through winter 1978. Checks indicate tide heights where organism
uas found. +6' +3' +0' -5m -10m
Chlorophyta
Eliding la minima / /
Cladophora sp. / /
Derbesia carina »'
Enteroaiorpha flexasoa /
E, intestinal is / /
E. Unza /
Halieystls oval is *
Honoatroma sp, * /
Rhi^-ocloniuifl sp. /
Spongomorpha coallta / /
S. spinescens »' /
Ulothrli sp. / /
Ul"a spp. J J
Uroiipora sp. »' J
Bac i1lari ophy ta
Pennales spp.
Phaeophyta
Alaria spp.
Analipua japonicus
De9irc.ri.st i a acuJsata
D. liguuata
D, vlridis
Ectocarpua simulant
Fucus dlsticbup
Hedophyl]um sessile
Iaminarla groenlandlea
L. saccharin?
L . aetchel1i i
Nereocyatis luetkeana
Fetalonia fascia
Ralfila pacifies
Sphacrlaria racesosa
S. subfuaca
StictyoBlphon toitills
Syrtn&oderoa abyisslcula
f f /
» v *' v
¥ ¥
¥
t
t r
/ / /
J
/
v
/
V ¥
r
i
¥
i
/
/ /
¥
/
/
r'
t
Rhodophyta
Acrochaetiu® sp.
Ahnfaltla plicata
Antlchaanlon defectum
A. dendroideua
/
/
V
£10
-------
MORSE CREEK
Rhuduphyta (cont.)
Bang la fuscopurpu eum
Boss leila plumosa
Bof rye plus sum farlowianum
Callltharanlon plkeanum
Callophy11 Is flabellulaca
Ceramluo pacificum
Choreocolax polysiphon!ae
Coralllna Vancouver lens is
Cryptosiphonia wnodii
Jelesserla declpiens
Endocladia muriraia
Erythrot richia parksil
Clgart ina paplllata complex
Gonimophy1lum skottsbergii
Cracilarla verrucosa
Tide Ht.
+6' +31 +0' - rjm
•10m
Gymnogongruj Itspcophyllis
Halosacclon gla.idiforme
Halymenia cocclnea
Hildenbrandia sp.
Hyraenena sp.
Iridaea cordata
I. heterocarpa
Leptof aur.hea pacifies
Membranoptera sp.
Microcladia borealle
Odonthalia floccosa
0. waahlngtoniensis
Petrocells middeadorftl
Peyssonella pacifica
Platythanmion heteromorphma
Pleononpor iua sp.
Plocaaium coccineum
Polyneura laclsslma
Polyaiphonia hendryl
P. pacifica
Porphyra niniata
/
/
J
*
~
Prionitis lyallli
Pteroaiphonla biplnnata
P. dendroidea
¥
/
420
-------
MORSK C.REEK
Rhodophyta (cont.)
Ptilota filicina
P. pectinata
Rhodochorton purpure»jm
Rhodomela larix
Rhodymenla californica
R. pacifica
P.. palmata
R. pertusa
Sarcodiotheca furcata
Scagelia occidentale
+6'
+ 3*
Tide Ht
+0'
Forifera spp.
Halichondria panicea
Haliclona permollts
Ophlitaspongta pennata
C'nidaria
Hydrozoa spp. /
Tubular la sp, *'
Arithozoa
Anthopleura elegantissima / »'
Halcampa sp.
Tealia sp,
Platyhelointhes
Turbeilaria spp. / r'
Nemertea s,jp, / » /
Emplectonema gracile *' »'
Nematoda spp. / »' i'
Mollusca
Amphineura
Cyanopla* dent lens / /
Katharlna tunicata /
Mapalla sp. B
Tonlcella 1ineata
421
-------
MORSE CREEK
+6'
lychaeta (cont.)
Conladidae
Glycinde pi eta
Heslonidae
Mlcropodarke dub i a
Ophiodromus puget t ens is
Lurabrinei idae
Lumbfiner is inflata
Mage Ion i dae
Magelona pitelkai
Maldanldae
Ax lot he 1 la rubrocincta
Eue 1 ymene sp .
Nicomache personata
N. lubricalls
Nephtyidae
Ne^htya caL;a
N. caiifcrniensis
No re'dae
Mlcronereis nanaiaoenuis
Nereis pelagica
N. vexilloaa >'
Platynereis blcanallcjlata
Ophel1idae
Armandla brevis
Orb in ii dae
Scoloploa sp.
Owen J jaa
Oweni.'i f us Ifonni s
Parann idae
Aricidea sp¦
Paraonella platybranchia
Pectinariidae
Pectinarla gr&nulata
Phyllodocidae
Eteone longa
E, tuberculata
-------
HORSE CREEK
46'
Cast ropoda
Tide Ht.
+3' +0' -5m
¦ lQrn
Acmea mitra
Afcolididae Sp.
Alvinia sp.
Araphlssa columbiana
Archidoris montereyensis
Barleeia haliotlphila
Calliostoma ItgaCum
Calyptraea fastigiata
Colllsella digitalis
C. ochracea
C. pelta
C. strigatella
Crep idula sp.
/
/
Lacuna varlegata
Lirularia lirulata
Littorina scutulata
L, sitkana
Margarites pup i1lus
Nassarius tneridicus
Nat lea clausH
Notoacroea fenestrate
N. persona
N. scut'.'ai
Ocenebra lurida
Orichldella boreal is
Seariesia dira
Thais emarginata
T. lamellosa
Trichotropis caru-el lata
Divalvia
Adula californlenais
Clinocardium nuttallii
rrenella decussata
Cryptonya califomica
/
V
J
/
Hiatella arctlca
Huailaria kennerlevi
Luclnoma tenuiscuipta
Lyonsia califomica
Hacooa Inqulnata
M. nasuta
H. secta
/
J
422
-------
MORSE CREEK
Bivalvia (Cont.)
Tide Ht,
+6' +3' -K)'
-5m
' .'ID
Modiolus rectus
Musculus Jisnora
M. pygmaeus
Mya arenaria
My s e 11 a tumida
Mytilus califomianus
M. edulis
Procothaca staminea
Tellina nuculoides
Transennalla tantilia
Tresua cap ax
Annelida
Oligochaeta spp.
Polychaeta
Arapharet idae
Ampharete arctica
Asahellides 1i t coral 1s
Arenicolidae
Abarer.icola pac if lea
Branchiomaldane vincenti
Capitellidae
Capitella capitata
Mediouastus sp.
Notomastus tenuis
ChrysopetRlidae
Paleonotus bellis
Ci rratulidae
Chaetozone setosa
Ci rratulus cirratus
Tharyx mult if ilis
Do rvilie idae
Protodorvl1 lea gracilis
Flabelligeridae
Pherusa plumosa
Glyceridae
Glycera asericana
Hemipodus boreal is
423
-------
MORSE CREEK
Tide Ht,
+6' +3' +0' -5m -10m
Pbyllodocldae (cont.)
/
Eulalla quadrioculata /
E. aanguinea /
E.virldis / /
Heaionura coineaui difficills /
Phyllodoce maculata / / *
P, madeirensis J
Polynoidae
Harmothoe imbrtcata / /
Sabellarlidae
Idanthyrsus armatus /
Sabel larla cem^ntar1 urn / v
Sabe11idae
("hone ec.audata /
Fabric la oregonlea / »•'
Jasniineira sp.
Manayunkia aestuar ine /
Schlzobranchla inslgnis /
Sabe11a sp, /
Sabel1astarte sp. /
SealIbregmidae
Seal ibregtna inflatum
Serpulidae
Spi rorbis moerchl
L. nakamural
S. spirillum
Slgalionidae
Pholoe minuta
Sphaerodorldae
Sphaerodorun pap 111 1fera /
Spionldae
Malaeoceros glutaeus J /
Polydora armnta /
P. Columbians / / / *
P. commensal Is /
P. proboseidea / /
P, socialIs / / /
P. spotiglcola r
P. »p. A / /
Prionospio cirrifera / /
P. steenscrup 1 »" / /
425
-------
MORSE CREEK
Tide Ht.
+6' +3* +0'
Spionldae (cont.)
Pygospio elegans »' /
Scolelepis sp, C r /
S, squaniata *'
Spio filicornis »' >
Spiophanes bombyx
Syllidae
Autolytus sp.
Exogone gemmifera
E. laurel
Eusyllis sp.
Spaerosyllls pirifera
Syllis adamantea
S. heterochaeta
S. longoclrrata
Trypanosyllis gemmipara
Terebe1lldae spp.
Arehlannellda
Polygordidae
Polygordlus sp,
Saccoc1rridae
Saccocirrus sp. ~'
Hirudinea sp.
S1punrui a
Uolfingla vulgaris
Phascoloscma agassizii
Pycnogonida
Halosoma coapactuns
H. viridintestinales
Nymphon grossipes
Phoxlchilid'.um femoratum ^ ^
Crustacea
Ost raroda spp. " •'
Clrripedia
Balanue cariosus ^ *
B. glandule »'* *' >*'
B. nub Ilia /
Chthaaalua dalli ~*' *'
/
426
-------
MORSE CREEK
+•6"
Crustacea (cont.)
Leptostraca
Ncballa pugptcensis
Cumacea
Cume11 a vulgaris
Dlastylis sp,
Lampropidae spp.
Leptoeuma sp,
Tanaidacea
Anatanalu normani
Leptochelia dubia /
PancoLus callfornlensis
Iaopoda
Cacclanlropsis psararaophila
Dyrtamenella shearerl
Exosphaeroma ampllcauda
E. media
R. rhomburuo
(incir lmosphaeroma oregonense »'
lanlropsls analogs
I. kincaidi
I. magnocula
Idotea aculeata
I. aontereyensis
I. achmittl
I» wosnesenski1
Jaeropsls dub la
J. lobata
Llmnorld algarum
Munnfl chromatoeepha la
M. stephensenl
Paranthura elegans
Amphlpoda
Accedomoera vagor
A. ap. C
Allorchestes angustua /
Ampellsi-a crlstata
A. pugetlca
AmpIt hoe lacertosa
A, me a
A. simulans
Anorsyx lat 1 coxae
Aoroldes colutabiae
AtyluB levldenal9
-------
MORSE CREEK
Tide 'r.
+6' 4-3' -5m
Amphipoda (cont .)
Caprella laeviuscula
C. mendax
C, natalensl1?
C. striata
Ceicops compactus
Corophiud brevis
C. cra.-.s lcorne
-10m
Eohaustorius wash ing tan Ianus
Gammaropsis chompsoni
Guernea sp. A
Hyale frequens
Ischyrocerus sp. I
I. jnguipes
Jassa fa 1 cata
Kerrayscheus ociosa
Maera simile
Me 1ita calilornica
M dentata
M, desdlchaila
Najna const 1 iorutn
Orchestia sp.
Crchestoidea pugettensis
Parallorehestes ochotensis
Paramoera columbiana
P. 'pugetica" n. sp,
P. serrata
Paraphoxus cognatus
P spinosus / obtusidens
F, spinosus Type B
/
Parapleustes pugectenuis
Phot is hrevipes
F. sp, D
P1 eijs 1 run secorrus
Pontoganeia cf. ivanovi
Stenothoides beringiensi§
Synchelidium rectipalmun
Westwoodilla eaecula
Decapoda
CallianAssidae ap. (juv, >
Cancer naglJter
C. oiegonensia
C, product us
428
-------
MORSE CREEK
+6
Decapoda (com.)
CranRon dalli
C. muni tella
^ryptolJthodes sp. (juv.)
Di.scorsopaguru3 schmitti
Eulaius sp. (juv.)
Heml gr ip.jus nudus
H. otegovu-nsls
Heplacarpus ap. (juv.)
Hippolyce clat A
Qedlgnathus inermis
Pagurun h. hirsut*usculus
P. berlnganus
Pinnixa fabz
Pugettia gracilis
P. product us
Telmessus chelragonua
Upogebia pugettensis
Inserta
Dipterart larvae /
Phorcnida
Phoronopsis harmeri
Bryozoa spp.
Flustrella sp.
Hlppothoa hyalina
Membranipora sp.
Echinodermat^
Ophluro idea
AmphlpholIs squacata
Holothuru idea
Cucumarla sp.
Eupentacta pseudoquinquesemtta
Leptosynapta clarki
Hopalia intermedia
429
-------
Table 5. Beckett Point (protected sand-gravel intertidal; sand subtid.il).
Complete list of all benthic organisms collected from spring 1976 through
winter 1978. Checks indicate tide heights where organism was found.
Chlorophyta
Enteromorpha
c^mpressa
E. Intest inalis
E. linza
+6'
+3'
/
/
K)'
-5m
• 1 Om
Ulva taeniata
Urospora spp.
/
Bacillariophyta
Pennale9 spp.
Phaeophyta
Feidmania
SPP-
Rhodophyta
Ceraraitim
nibrum
C. paciiicum
Polysiphonia pacifica
Folysiphonia paniculata
Porpnyra 3(>.
Spennatophyta
Zostera marina
Cnidaria
Anthozoa
Actinaria spp.
Platyhelminthea
Turbellaria ipp
Nemertea
spp.
Nematoda
spp.
Mollusca
Gastropoda
Aeolldidae
sp. (juv.)
Aglaja
dlomedia
Alvinia spp.
Araphissa
columbiana
Barlecla
haliotiphila
Cerithiopsis bp.
*
/
¥
/
/
/
y
4 30
-------
BECKETT POINT
Collisella
ocrachei
C. pelta
C. strigatella
Crepidula adunca
Crepipetella llngulata
Cyliclma sp.
Fartuluo occ identale
Hamlnoea vesicula
Lacuna var iegaca
Lirularia lirulata
Littorlna scutulata
L. aitkana
Margarites pupillus
Mitrella carinata
M, tuberosa
Nassarlus mendicus
Natica clausa
Notoacmea perpona
N. scutum
Ocenebra lurida
Odostomla sp.
Pollnlcea lewlsli
Solarlella obscura
Thais spp.
Turbonilla sp.
Bivalvia
Acila castrensis
Adula callforniensls
Clinocardium nuttallli
Crenella decussata
Cryptomya callfornlca
Hlatella arctica
Luclnoma tenulaculpta
Lyonsla callfornlca
Macoma lnqulnata
H. nasuta
M. sects
Modiolus sp.
Muaculus pygaaeua
Hya arenarla
Myaella tualda
Mytilua edulls
Nucula tenuis
-------
BECKETT POINT
Panope generoca
Pododestaus cepio
Protothaca stamlnea
Paephidia lordi
Saxidomus giganteus
Tellina carpenter!
T. modesta
Transennella tantilla
Tresus capax
T, nuttallii
Annelida
Oligochaeta
spp.
Polychaeta
Anpharet idae
Araphrete
arctlca
Arenleolidae
sp. (juv.)
Capitellidae
Capltella
capltata
Medlomaatua sp.
Notomastus 1lneaius
N. tenuis
Chaetopteridae
Mesochaetopterus taylori
Fhyllochaetopterus prolifica
Splochaetopterus costarua
Chrysopetalidae
Paleonotus bellia
Clrratulldae
Chaetozone aetosus
Clrratulus cirratus
Tharyx aultlfllia
Dorvilleidae
Dovillea annulate
D. rudolphl
Protodorvillea gracilis
Eunicoidea
sp.
Clyceridae
Glycera aaericana
G, capitata
G. tasaelata
HcaipcHua boreal is
U 32
-------
BECKETT POINT
Grtniadidae
Glycinde annigera
G. p
-------
BECKETT POINT
+6*
Phvllodoc idae
Lteone Longa
E. tubercuiata
Eulalia quadrioculata
E. sanguinea
K. viridis
+3'
+0'
-5m
-10m
Hesionura coineaui difflcilis
Phyllodoce castanea
P. groenlandlca
P. maculata
P. £l ^ . A
Ptiargidse
Pilargis bcrklc-vae
Slgambra tentaculata
Pisionidae
Plsione sp.
Arctonoe vlttata
Harmothoe imbricata
H, lunulata
Lepidasthenia berkleyae
Tenonia kitsapensis
Sabel i idae
Chone ecaudata
Sabella media
S«rpuJIdae
Splrobla nakamurai
Sigalonidae
Pho1 on minula
Splonldae
Malacocerus gl utaous
Polydrra caulleryi
P, columbiana
P, Iimicola
P. proboseidca
P, pygidialis
P. quadrilobata
P. nor tall a
P. sp, A
Prionospio c 1 r r 11c ra
P, steenstrupi
Pygo splo elegana
Scolelepia sp.
Splo f11 lcornls
Splophanes berkeyorum
S. bombyx
Svllidae
Bran la brevlpharvngla
Exogone gemnifera
E. louie1
Sphaerosy11 is brandhorsti
S. plrlfer^
L"U,
-------
BECKETT POINT
Terebellidae
Polycirrus sp. A
Ar chi annelida
Polygordlus sp.
Protodrilidae sp.
Stpuncula
Golfingia vulgaris
C. puget tensls
Crustacea
Leptostraca
Nebalia pugettertsis
Ostracoda
spp.
Cirripedia
Balanus glandula
Cumacea
Cumella vulgaris
Diascylis sp.
Difistylopsls sp.
Eudorella sp.
Lampropidae sp.
Leptocuma sp.
Tanaldacra
Leptochelia dubia
Isopoda
Clrolana vancouverlensis
Dynamenella shearer 1
Exosphaeroma ampllcauda
E. meoia
Gnoriidosphaeroma oregonense
lanlropsia kincaidl
Idotea reaecata
Limnoria algarum
Munna ublquata
Paranthura elegans
Mys idacea
Archaeomys i a grebn it zkt1
Amoh1 poda
Aliorchestes augufltus
Ampeltsca agaesizl
A. pugetlea
Amp It hoe lactrtosa
/Vnisi'ganmaruH iijnfi'rv i.olus
Anroides columb1ae
-------
BECKETT POINT
+ft'
+3*
+0'
-5m
-10m
Amphipoda
Byblts serrata
C.iprella laeviscula
Corophium ascheruslcum
C. crass i corne
Dullc.hla sp. A
Gamnaropsis thompsonl
Cuernea sp. A
Hlppomedon sp. B
Tschyrorense anguipes
Metacaprella kenner1yi
Melita dentata
M. desdichada
/
/
J
0'-v,;omene sp. A
Orchestoidei pugettensls
J'achynus
cf. barnard 1
Piiramoera "senata" n, sp
Paraphoxus sp. A
P. sp. C
P. spinonus
Phot Is b. L-vlpes
Pleuslrus secorrus
Podoceropsis
inaequistvlls
Puntogenpia
rf. rostrata
Protonsedeia sp. A
Syrii he! idlum rectipalmum
S. .shoemaker j
Trltella ptlSmana
Westvoodllla caetula
Oecapoda
Cal1lanassidae Bp. (juv.)
Cancer gracilis
C. maglater
C. oregonensls
C. productus
Crarigon nigrarauda
Hemigrapsus nudus
H. oregoneneis
436
-------
BF.CKf.TT HOIST
+6
+ 3'
Hippolytidae
sp,
Labi doch1 rur. Kplendcb rrns
OrtgonJa gr id 1 Is
Pagurus btrluganus
P, h. hirsutiusculus /
Plnntxa llrtornlis /
P. Occident a!1m
Pugettia gracllis
Telmc-ssus i he i ragjnus
Upogcb 1.1 pugot t ens i s /
1 lificr tn
Dlpteras. larvae
/
Phoronida
I'horonupals harmcri
Phc-r t 1 s i j iin.i i
T
Ech inudi' rma t a
Erli ino 1 u'j
Dendraster i xconr rlc us
r
Holot huroi dea
Lept oaynapt .1 r 1 ark 1
Hfilpndia Intermedia
/
Ophiyoldea
Anphipholis squanuta
437
-------
Table 6. Dungeness Spit (exposed gravel-sand). Complete list of all benthle
organisms collected spring 1976 through winter 1978, Checks Indicate
tide heights where organism was found.
Genus Species Tide Ht.
*6" +3' +0' -5m -10a
Chlorophyta
Chaetomorpha californica /
Bac illariophyta
Pennales spp, J
Phaeophyta
Alaria spp, /
Coitaria costata /
Desmarestia aculeata /
D. liguldta var. ligulata /
Ectocarpaceae /
Peldmania 8pp. /
Giffordia ovata /
Haplogloia anderaonii /
Hedophy11um aeasile /
Laminaria »pp. /
Ralfsia pacifica /
Stictyoaiphon tortilis /
Syr ingodenna aby«a icola /
Rhodophyta
Acrochact ium app. /
Ant ithannion defectum /
Botryocladia pacudod i chotoaa /
Call ithannion/Pleonoiporiua /
Choreocola* polyaiphoniac
J
418
-------
Dungeness Spit
Cruoriopaia aestuarii
Cryptonemia boreal is
C. oval1 folia
Euthora frut iculoaa
Gigartinaceae spp.
Goniocrichum alsidii
Gracilaria spp.
Gymnogongrus leptophyllus
Halymenia spp.
Hollenbergia subulata
Iridaea spp.
Leptofauchea pacifica
Meubranoptera spp.
Odonthalia ap.
Peyasonelia pacifica
Phycodrys laabel1i ae
Platythamnion pectiatum
P. v i 11 oa urn
Polysiphonia hendryi
P. pacifica
Porphyra spp-
Pteroaiphonia dendroidea
P. gracilis
Ptilotdt filicina
Rhodoptilua pluaoaun
Rhodytaenia app.
Tide Ht.
+6' +3* ~0' -5m -10m
/
/
/
y
*
/
/
/
/
/
/
/
/
/
/
/ /
~ /
/
/
/
/
~
r
i
/
l* 39
i
-------
Dungeneas Spit
Scagelia occidentals
Stenogramne interrupta
Thurete11 ops is peggiana
Tide Ht.
+6' +3' >0' -5m
/
•10®
/
/
/
Cnidaria
Hydrozoa
Coryne tubulosa
Hydruidea app.
Tubularia sp.
Anthozoa
Halcampa sp.
/
/
V
Platyhelminthes
Turbel1 aria spp.
Nenertea spp.
Neraatoda app.
/
/
/
/
/
/
/
Mollusca
Aaphineura
Cryptobranchia concentrica
Leptochiton internexus
Gastropods
Aeolididae app.
Aaphiaaa coluabiana
/
/
/
/
/
.43
-------
Dungenesa Spit
Tide Ht.
+6' +-3' ~0' -5m - 10m
Calliostoma 1i gaturn /
Calyptraea fastigata / /
Lacuna varlegata / , /
Margaritea pupillus /
Mitrella tuberosa /
Natica clausa /
Thais canal Leulata /
Trichotropia cancellata /
Biva1via
CIinocardiujs nuttal 1 ii / / /
Crene L L a decasaata / / /
Humilaria kennerleyi /
Macoma sp. /
Modiolus sp. /
Mya arenaria /
Mysella tumida / /
Protothaca ataminea /
P, tenerr lui /
Tellinasp. /
Transent«lla tantilia / /
Tresus capax /
Ann#Iida
Oligochaeta app. / / / / /
441
-------
Dungeneas Spit
Tide Ht.
+b' +3" +0'
¦5m
¦10m
Polychatta
Acrocirridae 8pp.
Arenicolidae spp.
Capite11 idee
Cipiceila capitata
hedionwatua op.
Notomastus 1ineatus
N. tenuis
/
~
/
i
/
Cirratulidae
Chactozona setosa
Cauleriella Bp.
Cirratulus cirratus
Tharyx multifilis
Uorv ille idae
Protodorvil1ea gracilis
/
/
/
Gor. iadidae
Glycinde picta /
Heaionidae
Gyptii. brevip tlf •> /
Micropodarke dubia /
Mald&nidae
Axicthella rubroc incra /
lie lymene sp. /
Nephtyidae
Nepbtye ca?ca /
N. ciliata /
N. ferruginea /
Nere idac
Nereia sp. /
Platyneie i» bi canal iculata / /
Opheliidae
Armandia brevia / /
Trav'in forbeaii /
442
-------
Dungeness Spit
Orbiniidae
Scoloplos 8p.
Oveniidae
Owenia fusifonnis
Par&onidae
Paraonella platybranch ia
Phyllodoc idae
Eteone longa
Eulalia sanguinea
Polynoidae
Harmothoe imbricata
+6'
Tide Ht.
+3' +0' -5m -10m
Sabellidae
Chone sp.
Sabella media
/
Sealibregmidae
Scalibregma inflatum
Serpulidae
Spirorbis Bp.
Sigalionidae
Pholo« ninuta
Spionidae
Malacoceroa glutaeua
Polydnra -]uadr i lobata
P. Nocia1.is
4-'.i
/ / /
/
-------
Dungeness Spit
Tide Ht.
+6' +3' +0' -5a -10m
Prionoapio cirrifera /
P. steenscrupi /
Spio filicornis *
Spiophanes botnbyx *'
Sy11idae
Eusyllis sp. »
Exogone gensnifera /
E. lourei /
Sphaerosyllis pirrifera /
Sy 11 is heterochaeta /
S. longocirrata /
Terebellidae
Polycirrus sp. A /
P. sp. B /
Archiannelida
Polygordiidae
Polygordius sp, /
Sipuncula
Phasco losoaia agassieii /
Crustacea
Cirripedia
Balanua rostratua alaakensia /
Ciuaacea
Diastylis sp. / /
LaapropWUta / /
Leptocuma sp. /
Leptostylis sp. /
444
-------
Dungeness Spit
Leptostraca
Nebalia pugettenais
Mysi dacea
Archaeomysis grebnitzkii
Tana idacea
Leptochelia dubia
Isopoda
Exosphaeroma amplicauda
E. rhoctburum
Cnor imosphaeroma oregonense
Idotea sp. (juv. )
Tide Ht.
+6' +3' +01 -5m
/
/ / /
-10m
¥
/
/
/
Aruph lpoda
Accedomoera vagor
Ampelisca criatata
A. macrocephala
Anoryx lacicoxae
Aoroides columbiee
Atylus levidensis
Caprella laeviuscula
Corophium crass icorne
Eohauatorius Washington!anus
Guernea rp. A.
Ischyrocerus anguipes
Melita californica
M. dentata
M. desdichada
Orchonene sp. A.
Orchestoidea pugeCtensis
Parameera serrata n. sp.
Paraphoxus spp.
Paraphoxus spinosua
Parapleustcs sp.
Photis sp.
Synchelidiua shocaakeri
Tiron bioccllata
/
/
/
/
/
/
/
/
/
/
/
/
/
/
/
/
/
/
/
~
/
/
/
/
4 «15
-------
Dungeriesa Spit
Tide Ht.
~61 +3' +0 -5m -10m
Decapoda
Call ianassidae (juv.)
-------
Table /. Twin Rivers (exposed grave 1 -rwinrl) . Complete list of all benthi
organisms collected spring 19/0 through spring 1977. Checks indicate
fide heights where organism was fount',
,, . , ^' -t-'V - Stl
i.hlorophy ta
Er t eromorpha 1 i n *. a »
Ulva spp. ~
Baciliarlophyta
Penna'es spp. >
Phaeophyta
Deamares t ia viridis >'
Raifsia sp. »
Syr ingodenna abyssicola >'
Rhodophyta
Acrochaetium sp. »
Antlthamnion dencroideum *
Ceraroium vanh ingtoniense ~
Cracilaria verrucosa >
Craci1 aria sjoestedtii >'
Gvmnogorigru^ leptcphyJlus »
Hollenbergia auiuilata ~
Irldaea sp. »
Neopt ilutd aspleniodes *
Platythamnion reversurc >
Platythmanicn pectinatum *
P1 atythamnion villos-im /
Pleonosporlum sp. »'
Polysiphonia hendryi <
Polys1phonia pacifica >
Polysiphonia paniculata >
Porphyra sp. »'
Pterosiphoma bipirmata »
P. dendroidea '
Pt i lota Cilicina >-
Khodyraenla palmata >
Sarcodiotheca furcata >
Scagelia occidentale »
Stenograrame 'ntt-rrupta »
Cnidaria
Hydro^oa
spp.
Scyphozoa
Hallclystufa auricula »
Ant'-'nzoa
Halcarapa decent en t .icu lata »
Platyhelminthes
Turbeilaria
spp.
Nemertea
spp. »' »'
Amphlporus sp. »
Nematode
spp. '• /
-------
TWIN RIVERS
+6'
Xolluaca
Gastropoda
Aglaja diomedea
Calvptraea fastiglata
Crepidula perforates
Lacuna variegata
Li rtil aria I lrulata
Hargarities pup 111us
Thais .analieulata
Thais lamcllosa
Bivalvia
Clinocardium sp.
Crenel la decussata
Luc trio ma tenuisculpta
Macoma spp.
Modiolus sp
Musculjs sp
Mya sp.
My sella tjmida
Prctothac* scaralntM
Psephldla lordl
Te11 Ina sp.
Ti ansenne1 la tant i L la
Annelida
01Igcchaeta
spp,
Polychaeta
Arenicolidae
Abare, :ola sp, (J uv.)
Ampharetidae
Ampharete arctlca
Capltel1idae
Barantol la ainericana
Capitella capitata
Medlomastus spp.
Notomastus tenuis
Cirratulldae
Chaetozone aetusa
Tharyx multlfilis
(!1 yeer idae
Glycera sp.
Hemipodus boreal is
Gonladidae
Cllycinde pictu
Gonia.la sp. (Juv.)
Hesionidae
Gyptia brevipalpa
Mi'-ropodarke dubia
Luiabr ineridae
Lurnbriner i8 luti
L, conata
-------
TWIN RIVERS
+6
C ruiitncea
Oscracoda
spp. >'
Cirripedia
Bal.inus jp
Cumacea
Cumella sp.
Dlastv1 Is sp.
I) i ast y 1 ops I s sp .
Leptocuaa sp.
Tana 1 da> ea
Leptochelia dubia
1 sopoda
Edotea sub 1i 11oralis
f.norimosphaurona oregonen.se »
Idotea sp.
Svn idotea bicuspids
S. pettiboneae
Amphtpoda
Cap re >1 idea
Can r«i 11a grac i 1 ior
Tr I te L la pill m.ina
Camraar1 doa
Arcedorooera vagor
Ampelinca agassiz i
A, macrocephala
A. pugetica
Anonyx laticoxae
Aoroides coluacblae
Argisaa hanatipes
Atylus lnvtdensus
Eohaustorlus washing!: onianus
Gammaropsis thompsoni
Onernea sp, A
Hyale pugettensia
Ischyrocerus anguipes
Jassa falcata
I.epldepecreum sp. A
Mtlita desdichada
Honoculodi-s ?.ernovi
Orcheatia georgiana
Orchomene sp. A
ParaphoxuE spp.
Paramoera "aerrata" n.s.p. >
PnotIs brevlpee
Phot is sp. D
Pleusiru.® secorrus
Pontogeneia rostrata
Protomedia sp. A
SynchelIdium rectipalmum
S. shoemakei1
TrIon biocellata
Westwoodi1la caecula
450
-------
Mage Ion Idae
Magelona pitelkai
Ma I danidae
Euclymene dilinear a
"ephty idae
Nephtvb caecoides
N. eallforniensis
Nt»r«? idae
Micronereis nanainioensih
Nereis pelagica
N. proceri
N. vexillosa
P latynereis !>1 -am i 1 cul
Onuphidae
Onuphis iridescens
Opheliidae
Artnandia hri*vis
Orbiniidae
Nainerts uncinate
Scolop]us sp.
Oweniidae
Oven in f.-siformis
Paraonidae
Arieidea sp.
Paraonella platybranchls
Phyllodocidae
Cteone longa
E. tuberculata
Fhyllodoce maculata
Polynoidae
Arc rone vittata
Harmothoe imbricata
Sigal J on idae
PhoLoe minuta
Sphuerodoridae
Sphaerodoropsis minuta
Spionidae
Mai coceros glutaeus
Polyd ora hoc ialis
Prionuspio steenst nip 1
P. etrrifera
Spin f11 Icornla
Spiophanea buobyx
Syi1idae
Autolytus sp.
Exogone genraifera
E. loueri
Terebel lidae
sp.
Pycnogonida
Nyaphon pine 11 at-
-------
TWIN RIVERS
+6'
Deeapoda
Calllanassid.se sp. (juv.)
Cancer gracilis
C. oregonensls
Hippolv* IJ-e sp. (juv.)
Pagaurus beringanus
P. kenni'r ly i
Pinnixa sp.
Pugettia gracilis
I n a e c t a
Dipt era larva spp,
Phoron ida
Phoronib ij imai
Bryozoa
spp.
451
-------
Table 8. North Beach Sand (exposed sand intertid&l; sand-gravel-cobble
subtidal). Complete list of all benthic organisms collected spring 1976
through winter 1978. Checks indicate tide heights where organism wag
found. +6' +2' +0' -5m -10m
Lhlorophyta
Bryopsis corticulans *
Chat1 tomorpha cal ifornica / r
Derbesia marina / »'
IJpc i 11 ar loph v ta
Pennales spp. / /
Phaeophyta
Alaria sp. /
Agarum cribrosum /
Cosiaria costata /
Desmarestla aruleata »'
0. ligulota >'
Ectocarpalea np. »
Laminaria saccharina ~'
Pterygophora californica /
Spbacelaria sp. r'
Syiingoderma abyssicola ~ »'
Rhodophyta
Aerochaetium sp. • /
Alnf elt la plicata »'
Antithamnion defectum ~' <•'
A. dendroideum »
A. V y 1 in i i /
Antlthsmnionella pacifica /
A. glandulifera >'
Bonnemaisonia nootkana »
Botrycladia pseudodichotoma / /
Botryoglossum farlowlanium ~
Calliarthron tubercplosum /
Cal1ithamnion sp. / /
CallophyIlls firma /
C, flabelluljta / /
C. haenophylld >
C. pinnata /
Ceramium californicum / /
C. gardneri /
C. 8t r i c t um /
C. washin&to.iLense /
Choreocola* poly9iphoniae *'
Con st an tine a simplex ~
Crjorle profunda <•'
Cryptonemia boreal la ' /
Delesscr La decipiens / /
Euthora fruticulosa / *
Gigartin* sp. / /
Conlouphy Llum ekuttsberp.i 1 »'
45:
-------
NORTH BEACH SAND
+6' +2^ +21 -5m -10m
Gcniotrichum elegans >
Gracilarla verrucosa / /
G, sjoestedtll /
Griffithsia pacifica /
Hdlosacc ion glandiforme r
Halymenia coccineae /
Herposiphonia grand is / /
H. plunula /
Heteroaiphonia densiusrula J
H. japonica r'
Hildcnbrandia sp, >' /
Hollenbergia subulata /
Hymenena amithii v
Iridaea ap. / /
Laurencia spectabilis /
Lophosiphonia villum /
Membranoptera platyphylla /
Neoagardhiella baileyi /
Nienburgia anderaoniana / /'
Ni tophyllura mirabile »*'
Odonthalia flocossa / /
0. kamtschatlea ~' /
0. washingtonlens is / r
Opuntiella californica /
Petrocelis middendorfii /
Pey9sonellia pacifica / /
Phycodrys isabelllae /
Pikea robusta /
Platysiphonia clevelandil /
Platythamnion heteromorphum ,
P. pectinatum » /
P. villosum / /
Plocamium cartilagineura /
P. coccineum / /
Polyneura latlssima / /
Polyaiphonla hendrvi / /
P. pacifica »' /
P. acopulorum /
i'orphyra spp. / /
Pterochondria wood 11 ~'
Pteroaiphonia blpinnata / /
P. dendroidea /
P, gardnerl /
P, gracilis /
P. pacificum /
Ptilota filiclna /
Rhodoglossua callfornicum /
Rhodoptllum pluoosum /
Rhodymcnla palmata / /
R. pertusa /
R. atlpitata /
Sarcodlotheca 'urrata / »'
453
-------
NORTH BEACH
Scagelia occidentale
Stenogramme interrupta
Thuretel1opsis pegglana
Trailliella intrlcata
Spermatophyta
Zostera marina
Cnldaria
Hydrozoa
spp.
Anthozoa
Anchopleura elegantisalma
Halcampa decemtentaculata
Platyhelminthes
Turbellaria
spp.
Itaspiella armata
Nemertea
spp.
Nematoda
spp.
Mollusci
Amphineur a
Lepldo.-.ona mfcit" „j.
Mopalla 1ignoaa
M. Hp. v
Tonicella linuata
Gastropoda
Acraaea mltra
Aeolidldae sp.
Alvlnta sp.
Amphlsaa columblana
Calliostoma ligature
Calyptraea fast Ig'iatA
Cerithiopsls sp.
Collise11a ochracea
Lacuna var1egata
Laila cockerel11i
Lirularla llrulata
Llttorlna scutulata
L. sltkana
Hargaritiea pupl11 us
Notoacmea pernona
Nasaarius mendicua
Natlca clauaa
Odoatoola sp.
Searlesla dlra
Blvalvia
Astarte compacta
Cardlt ventrlcosa
Cllnocardiun nuttallll
Oeneila decussata
454
-------
NORTH BEACH SAND
+6'
Cryptomya californlca
Entodesna saxicola
Glycymeris subobsoleta
Humllarla kennerlyi
Lasaea cistula
Lucinoma tenuisculpta
Macoiua aecta
Haeoma spp.
Mya sp.
Mysella tumida
My111 us spp. »'
Protothaca staminea
Psephidla lordt
Semele rubropicta
Tellina spp.
Transennella tant i1la
Treaus capax
Zirfaea pllsbryi
AnnelIda
01igochaeta
spp. /
Polychaete
Acrocirridae
spp,
Macrochaeta sp.
Arenicolidae
Abarenicola sp,
A. claparedl ocean lea
Capltellidae
Capltella capltata
Medioaastus sp.
Notomastus llneatus
Notonnastus tenuis
Clrratulidae
Caulleriella gracilis
Chaetozone setosa
Clrratulua clrratus
Cirri forala sp.
Tharyx oultifilis
Dorvilleidar
Protodorvlllea gracilis
Eunlcidae
Eunice valens
Fiabelllgeridae
Pherusa pluiaosa
Glycerldae
Glycera capitata
Hemipadua boreal is /
Goniadldae
Glyclnde armlgera
G plcta
A 55
-------
NORTH BEACH SAND
+6'
Hes ion L^ai
Hesicnidae sp. A
Micropodarke dubla
Ophiudroaus pugettensis
Magelonidae
Kagelona picelkai
Maldanid aa
Axiothella mbrocincta
Euclymene sp.
N'comache lumbrlcai.ls
N. personate
Nepht yidae
Nephtys calIfovniensis /
Nereidae
Micron-jreis nanairaoensik
Nereis pelagica
H. vexillosa r
Platyriereis blcanaliculata
Onuphidae
Onuphis Iridescens
Onuphls stigmatis
Ophel1idae
Armand ia brevis
Ophelia 1imarina
Orbiniidae
Naineris uncinata
Scoloplcs sp.
Owenlidae
Owenia fusiforrals
Paraonidae
Paraonella platybranc'nia J
Phyliodocidae
Eteone sp.
Eulalia b11 ineata
Eulalia macroceros
E. nigrimaculata
E. quadrioculata
E. vlridis
Hesionura co ineaui dlfflcilis /
Phyllodoce groenlandica
P. oaculata
Pllargidae
Sigambra tentaculata
Pisionidae
Pisione sp.
Polynoldae
Harmoth *5 iobrlcata
H. lunu.ata
Sabellldae
Chon# ecaudata
OriopBIs nlnuta
Fotamilla intermedia
456
-------
NORTH BEACH SAND
+6'
Leptro^t r&^a
Nebalia puRpf.en: is
Mysldacea
Archaeomysis grebnltzkil
HolmesieIla anomala
Cwu-cea
C nine 11a a p.
Dlastylls sp.
Dlastylopais spp.
Lampropidae spp.
Tana idacea
Leptochella dubla
Isopoda
Caeclaniropsls psaumoplii la
Cli olaiu tiarfordi
C, vanc^uvirer.sis
^xosphaeroroa amplicauda
E. i -sdia
E. rhcoburum
Cnorimcsphaeroma uregonense
laniropais analoga
1. ktncaldI
I, trldens
ldotea ."ufescens
Ja^opais dubla
J. lobata
J. setosa
Janirllata occldentales
Llmnorla algaruro
Munna stephenseni
M. ubiqulta
Munnagonium waldronense
Pariinthura elegans
Tecclceps pugettensls
rtmphlpcda
Gammaridea
Accedomoera vagor
Ampeli sea cristata
A. pugctlea
Ampithoe slmulans
Anonyx latlcoxae
AorIdes columblae
Atylus levldensls
A. tridens
Corophlum crasaicorne
Eohaustoriuq washingtonlanua
Ericthonius brasiliensls
Gantmropsls Chonpsoni
Guernea sp. A
Ischyrocerus angulpes
Maera simile
Megaluropus longlmerus
458
-------
NORTH BEACH SAND
+6'
Scalebregmidae
Scalebregma inflatum
Serpulidae
Crueigera zygophora
Serpula vermlcularis
SpLrorbis nakamurai
Sigallonidae
Pholoe minuta
Sphaerodoridae
Spnaerodoropais minuta
Spionidae
Laonice cirrata
Malocnceios. glutaeus /
Polydora haaata
P. py^idialls
P. soclalis
P. spongicola
Prionospio cirrlfera
Prioncsplo steenstrupi
elegans /
Scoielepis sp. A
Spio cirrlfera
Spio fill cornis
Splophancs bomb'--*
Syllldae
Branla brc
Exogone gc
E, lourei /
Pianosylit
Ephaerosyll, a
StreptosyIlls i .11 . palpa
Syllis armillaris
S, heterochaeta
S. hyalina /
Trypanosy111b goumipara
Terebell.ldae
Polycirris sp. A
P, sp. B
Archiannellda
Polygordiidae
Polygordiua spp. /
Protodrillidae spp.
Sac core 1rrldae
Saccocirrus eroticus
Sipuncula
Golfingia pugetcensis
Phaacalosoma agassizii
Crustacea
Ostracoda
app. /
Cirripedla
Balanua carioaus /
B. giandula /
457
-------
NORTH BEACH SAND
±§1
Mellta californica
Melita desdlchada
Orchestia sp, (juv.) /
Orehomene sp. A
Paramoera "aerrata" n. sp. /
Paraphoxus spp.
P. c.f. abronius
P. spinosus type B
Parapleustes pugettensia
Phot Is brevipes
Pleusirus aecorrus
Pontogenela ivanovi
P. rostrata
Protomedeia 8p. A
Synchelidium rectipalmum
Caprellidea
Capre1la augasta
Caprella graclLior
Caprella californica
Metacaprella kenr.erly 1
Trltella pllimana
Decapoda
Cancer oregonensis
C. productus
Crangon alaskensis
C. franscincorum
C. raunitella
Hlppolyte clark1
Lebbeus sp,
Lophopanopeus bellus
Oregonia gracilis
Paguridae sp. (juv.) /
Phyllolithodes papllloeua
Pinntxa ap.
Pugettia gracilis
Inaecta
Diptera larva /
Bryozoa
spp.
Brachopoda
Teiebratalla transversa
Echinoderaata
Ophiuroidea spp. (juv.)
Holcthuroidea
Leptojynapta clarki
Mopadia intermedia
Aacidiacea
Pyura haustor
459
-------
Table 9. Kydaka Beach (exposed sand). Complete list of all benthic
organisms collected spring 1976 through winter 1978. Checks indicate
tide heights where organism was found.
Rhodophyta
+6' +3' K)' - 5m -10m
Anthitnamnion defectum /
Choreocolax poly9iphoniae /
Leptofauchea pacifica >
Polysiphonla hendrvi
Pterosiphonia bipinnata
P. gracilis >'
Cnidaria
Hydrozoa spp. *
Anthozoa spp, "
Nemertea spp. / / / / /
Nematoda spp. / *'
Mollusca
Gastropoda
Aglaja diomedea /
Araphissa col unbiana /
Cal1iostoma sp. /
Lacuna variegata *' »'
Nassarius r endicus
-------
KYDAKA BEACH
+6*
Gem iadidae
Giveinde picta
Hesionidae
Mlcropodarke dub ia
LurabrtnerIdae
Lumbrineris sp.
Magelonidae
Magelona plteikai
Ma? danid.ie
AxiotheIla rubrccincta
EucIyraene sp.
Nicomache personata
Nephtyidae
Nephtys caeca
N. raecoides
N, californiensi s
Nereldae
Nerei s procora
Platyneieis bicanaliculata
Omiphidae
Onuphis elegans
0. irldescer.s
0. stigmatis
Ophelildae
Annand1 a brevis
Orbinlidae
Naineris uncinata
Sfoloploa sp,
Owenlidue
Owenia fusiformis
Paraonldae
Paraonella platybranchia
Pectlnaritdae
Pectinatla granulata
Phyllodocidae
Eteonc longa
Eulalla virldis
Phyllodoce easterner.
P. groenlandica
P. maculata
P. multiserlata
P. sp. A
Polynoidae
Eunoe senta
Hanaochoe Itnbricata
Sabel1Idae
Eudistylia Vancouver1
Schizobranchia insignia
Sigalionidae
Pholoe mlnuta
Thalent'ssa spiaosa
461
-------
kydaka
Sp ion idae
Laonice cirratus
HalacocetOS glutaeus
Polydora eaulleryi
P. social is
P. sp. A
Prionos plo ctrrifera
P. steenst ri.
Pygosplo eJegans
Scolelepls squamata
Spio filicomis
3piophanes botnbyx
Syl1idae
Kxogone gemmifera
E. lourel
Sphaerosy 11is pirifera
Streptosy!lis latipalpa
Svl1 is sp.
Terebe11idae
Thelepus sp,
Fipuncula spp.
Irlapu1ida
Priapulus ^.audatcs
Crustacea
Qstraeoda spp,
Leptostraca
Nebalia puget t ens is
Mys 1 dicea
Archieomysis grebnitzkii
Cumacea
Cumella vu lgaris
Diascylis ap.
Diastyiopsis sp,
Lampropidae spp.
Leptocuma ap
Tanaidacea
Anatana is normani
Loptoche1 la dubia
Isopoda
Cirolana vancouverensis
Cyanthura munda
Edotea aublittoralis
Gnor imoaphaeroma oregonerise
Idotea wosnesenski i
Llmnoria algarum
Munna atephenaeni
Paranchura elegans
Synldotea bicuapida
A62
-------
KYDAKA BEACH
4-6'
Amphlpoda
Ampellsca agassizi
A, macrocephala
Ampithoe humeral.is
Aoroldes columbiae
Atyl us levidensus
A. tridens
Caprella laeviusc"la
Dullchia sp. A
Eohausrorlus sp. b
F, washiagtotilanus
Erlcthoniua brasiliensis
Hippomedon sp, A
Ischyroceri-s angulpes /
Lepidepecreum sp. A
Megalurnpus longimerus
Helita desdiehada
Monoculodes splnlpea
Najna consiliorum
Orchestoldea puget tens i s »'
Orchonene sp. A
Paraphoxus epistomus
P. milleri
P. vlgitegus
P. spp.
Parapleuatea spp.
Phot is brevlpes
P. sp. A
Pleustea depressa
Pontogeneia rostrala
Protomedeta sp, A
P. ap. B
Synchelidlutn shoemakeri
Tiron bior.el lata
Tritella plllmana
Westwoodilln catcula
Decapoda
Cancer oregonensls
Calllanassidae ep. (juv.)
Paguridae ap. (juv.)
rinnotheridac ap. (juv.)
Pugettla gracilis
Insecta
Diptere larvae J
Phoronida
Phoronopsls hannerl
Bryoroa spp.
Echinodermata
OphluroIdea ap. (juv.)
Echinoidea
Dendraster excentricus
Holothuroldea
L«ptosvnapta clarki
461
-------
Table 10. Jamestown (protected sand). Complete list of all benthic
organisms collected spring 1976 through winter 1978. Checks indicate
tide heights where organism was found.
+6' +1.4' +0' -5m -10m
Chlorophyta
Chaetomorpha californlca » '
CIadophora pygmaea ' '
Endocladia vi ridis *
Enteromorpha intestlnalis "
E. 1 inza »¦ *
E. prollfera ' *
Monostroraa sp. i
Ulva taeniata '
U, spp. r ' *
Urospora sp. r
Bar 111ariophyta
Pennales spp. *'
Pha eophyta
Cymathere trlplicata ~
Desmarestia sp. *
Krto carpus sp. ~
Laminaria sac char ina ~
Ralfsia sp. »' ~"
St 1ctyosiphon tort Ills *
Syringoderma abyssicola •
Rhodophyta
Antlthamnlon glandulifera >
Asterocolax gardner1 »
Bangla tenuis »
Calllthamnlon/Pleonosporlum ' ^
Callophyllis f label lulata r *
C. haenophylla »
C. marglnata •
/
Geranium callformcum
C. eatorlanum
C, waahlngtonlenst k' "
Choreocolax polysiphonlae '
Delessericeae spp. '
Euthora frutlculosa r
Fauchea lac Inlat a '
GonlotrIchure alsldli
Gracilarla s loestedli 1 *
f
G. verrucosa ' '
Gymnogongrus leptophyllus "
Halymenla callfornica r
HerposIphunla grandis *
Hymeiena sp.
Neopt .lota asplenoldes /
Nitophyllun sp. ''
Pcy mone111a pac1fica "
Plat ytbamnion heteromorphuo
P. pectlnatua
464
-------
JAMESTOWN
+6'
+1.4'
+0'
-5m
-10m
Platvthamnion reversum
P. vlllosum
Plocamium coccineum
P. tenue
Polyneura latissima
Polysiphonia hendryl
P. pacifica
Porphyra sp.
Pteroslphonia bipinnata
P, dendroidea
Rhodogiossum roseum
Rhodoptilum plumosutn
Rhodymenia palmata
Seagelia occidentale
Stenogranme interrupta
Spermatophyta
Zostera marina
Pori f era
spp.
Cn i d a r 1 a
Hyd rozoa
spp.
Coryue sp.
Antliozoa
spp.
Halcampa decententaculata
Platyhelmlnthes
Turbellaria spp.
Nemertea
spp.
Paranemert.es peregrine
Nematoda
spp.
Mo 1lusca
Amph ineura
Mopalia lignosa
Tonire11a sp.
Gastropoda
Aeo1ididae
sp.
Aglaja diomedia
Alvinia sp.
Amphissa columb Lana
Bittiura eachrichti i
Boreotrophon or' us
Caiyptraee fasti^ata
Cerlthiopaie sp,
Collisella ochracea
Lacuna varlegata
Margarlte9 puplllus
Naesarlua mendlcus
Nat lea clausa
/
/
»-
/
/
/
/
V
!
1
r
-------
JAMESTOWN
+6
Odoatomia ap.
Oenopota tabulata
Pollnices pallidas
Thais ap,
Trlchotropis cancel lata
Turbonilla sp.
Bivalvia
Card ita ventricosa
Chlamys hastata
Clirtocardium nuttal 1 i i
Crenel la decussata
Cryptomya californica
Luc 1noma tenuisculpta
Lysonia californica
Macoma balthica
M. crassula
H. Inquinata
M. nasuta »'
Modiolus rectus
Museulus diseors
Mya arenaria
Myie11 a tumida
Mvtilus edulis
Nuculana hamata
Nuculana minuta
Nucula tenuis
Protothaca staminea
Psephidia lordi
Solen sicar ins
Telllna sp.
Trachycardium sp.
Transennella tant111a »
Treaue capax
Yoldia sp.
Annelida
Oligochaeta Bpp. »
At chiannel 1 '.i
PolygordIidae
Polygordius sp.
Polychaeta
Ampharet idje
Ampharete arctica
Apiscobranchidae
Apietobranchua sp.
Arenicolidae
Branchionaldane vlncenti
Capiteilidse
Capital la eai itata *
McdlomaBtus sp.
Notomaatua lineatus
N. tenuis
466
-------
JAMESTOWN
+6* +1.4' +0' -5m -10m
Chaetopt i?r idae
Phylloch.'.-t :»pterus prolif ica * »'
Splochaetopterus costariun »' /
Cirratulidae
Chaetozone aetosa / «
Cirratulus cirratus / / / /
Tharyx mult If ilia v" / /
Dorvi1lcldae
Dorville » rudolphi >' r"
Protodorvi 1 lea gracilis »' / /
Eunicoidea
9p . V
Glyce ridae
Glycera americana / ,
G. cap 1 tat a alpa r .
Micropodarkt- dubia / /
Ophiodroir • pugettensis '
Nepht v' .1-
,»htys caeca J /
i. caecoides ~
M. cal1fornipneis / r .
N. clllata /
N- u i dae
Nereis pauc ldentata ~'
N. pelagica /
N. procera / /
N. vexi1losa J /
Platynereis biranaliculata / » / / /
Onuph idae
Onuphia 1ri descens / / »'
0, stigmatis r / /
Ophel1idae
Amaot rypane aulogaster »
Arnandta brev is / / J ,*
Travisla forbesi i / ,
467
-------
JAMESTOWN
+6' +1.4' +0' -5m -10m
Orb in i idae
N'aineris dendrit iea /
N. quadricuspida j
N, uncinata /
Scoloplos 9pp. i / / r
Oweni idae
Owenla fus iforals / /
Paraonldae
Arlc idea sp. »' / r*
Paraonella platybranchia /
Tectlnarlidae
Pectlnarla granulata /
Phyllodoc idae
Etecne longa / J / »'
E. tuberculata / /
Eulal1 a billneata /
E. nigramaculata /
E. sanguinea ~ / / /
Phy1lodocc groenlandica / /
P. maculata / / ~' / /
P. a p. A /
Folynoldae
Eunoe sp. / v
Harmothoe imbrIcata > / »' , *'
H. lunulata / > J
Sabel1idae
Chone ecaudata / »
C. gracilis /
Euchone anal is /
Eudifltylla Vancouver i / /
Laonome sp. /
Potamilla intermedia / ~
P. myrlops »
Sabella media / /
Sabellastarte sp. /
Sealebregmidae
Scalebregma In flat. / /
Serpulidae
Splrorbis sp. /
Sigalionidae
Pholoe mlnuta / /
Sphaerodoridae
Sphaerodoropsis mlnuta / /
Sp lonidac
Laonlce clrrata / /
Malacocerus glutaeus / / / /
Polydora sp. A /
P. kempl Japonica /
P. quadr 1 lobaca »' / /
P. social la /
Prion.»»pio c 1 rri t »ra / J /
P, t»n«trupi / / »
P v. • legana / /
Af.8
-------
JAMESTOWN
+6*
+1.4*
Scolelepis sp. C
Spio fillcornis
Splophanes berkeleyorum
S. bombyx
Sv11idae
Branla brevipharyngea
Exogone gemmi fera
E. lourei
P ionosy11 is uraga
SphaerosyHis brandhorot i
S. pirifera
SLreptosy 11 i:> latipalpa
Sy11 ides longocirrata
Sy11is alternata
Terebellidae
Eupolymnla heterobranchia
Neoamphitrite sp,
N ! co lea zoster 1 col a
Plata brevibranchlata
P, cristate
Folycirrus sp. A
Proc lea graffl
Theiepus crispus
HIrudinea sp,
S1 puncuI Ida
Colfingia pugettenslr,
Phascolosoma agasslzl 1
Pr lapulIda
Priapulus caudatus
Crustacea
Ostracoda
spp.
LepLostraca
Nebalia pugettensls
Mya idacea
AcanthomysIs sculpta
Archaeomysis grebnitzk.il
Cum.'icea
Cumella vu lgaris
Dyastylis 8p.
Dyastylopsie sp.
Eudorella ap.
Eudorellopais sp.
Lampropidae spp.
Lamprops Bp.
Leptocuma sp.
LeptoBtylis sp.
TanaIdacea
Leptochelia dubia
+0'
-5m
-10m
469
-------
JAMESTOWN
+6' +1.4' +0' -5m -10m
Isopoda
Dynamenella sheareri /
Exosphaeroma ampllcauda / /
E, media ~' / /
E. rhoaiburum / /
Gnorimogphaeroma oregonenese /
Idotea 3p. (juv ) /
Jaeropais dubia / /
Munna sp. /
Munnogoaium waldronense /
Paranthura e!egans >' /
Phyliodurus abdominal is /
Synidotea b i cusp i da /
Amphipoda
Ampe!isra agassizi /
A. cristjca / J
A. pugetica ~ / /
Ampithoe lacertosa >' /
Anisogammarus conlervicolus /
A. pugettensis> /
Anonyx laticoxae /
Aoroides columbiae / / /
Argissa hamatipes / »'
Atylus collingi /
A. cridens /
Byblis veleronis /
Cal1ioplella pratti /
Caprella laeviuscula / /
Corophium acherusicum
C. crassicorne / r
C. saloon is / »'
Guernea sp. A r /
HIppomedon sp. A /
Iachyroceru9 anguipes *' /
Lepidepecreum sp. A / /
Melita dentata / »' /
Orchomene sp. A / /
Paraphoxus cognarua r'
P. robuatus /
P. similia /
P. spinoeus . r
P. spinosua type A /
P. spinoaus/obtusidens /
P. tridentatus
Parapleuates sp. / /
Phot is brevipes / /
P. ap, D /
Pleuairun secorrus r
c£. Podoceropsis inaequiatylla /
Pontogeneia cf. ivanovi / / ~'
Pontogeneia cf. rostrata / / /
P. sp. D /
A 70
-------
JAMESTOWN
+6' +1.4' +0' -5m -10m
Protooedeia sp, A J
Stenothoides beringlensis \/
Sync.helidlum rectipalmum / / *
S. shoemaker!
Phoronls ijtmai J
Phoronopsis banner! /
Bryozoa
3pp. /
Echinodermata
Holothuroidea
Cucumarla sp. /
Eupentacta pseudoquinquesemita /
Leptosynapta clarki / r' / /
Molpadia Intermedia »' /
Ophiuroidea
Spp. r' / /
Amphiodla urtiea /
Amphipholus Bquaoata /
Ci amphioda perlercta r
471
-------
APPENDIX III
PHYSICAL PARAMETERS
472
-------
I Second w«»ar < lr d*r * -»r, *a
Bertli Btffvii
I«k(i ta*cN Beckrtt
S««4 CdMI* ^oi#t
H»U»T f/ ,19
tfrtm »~" 11.* v.2 n.o
IM l»H M>.) - ^ 1 1
P.U l»77 JO.I • 30.5
Water t»7« Ii.5 - 30.1
mt«* T—wrtBft T
Ifri^ 1*7) 12.0 10.3 10,6
m 1MB II l ft 7 12.4 * li .2
Nil 1*1? ft.5 - 7.ft
Ittnt«r 1I7S iJ - 6.7
Air TwimftX
Sprlac If 77 11.0 10.7 17.0
f II If 77 Ift,« - U.n
felt 1*77 §,^ - 4.4
trl»««r l*7§ ft.2 - *.*
gs,»sjynigiiff p.-1 »f.*^±gaA
V^rl^ If77 0 6 0
t !¦ II 1*77 0 -0
Pail If77 0 -0
*td«w 1 f 71 j[y - o
d I r ? ?? jf r 'Ivor, in; •* • • • p
Itorit Tonpu» Tvlr Plliir fyd«fe«
. <¦ ?fc.1
29.* 2V* 13.2 - - S0..*
30,* |t,* ?i,7 - - 7fl.1
10.t 8.« A.? no din 10.<. P.I
III 11.2 A,' - * t.O
1.3 ? .* M - - 8.0
6.1 T.5 V< - - ? .fi
12.1 S.* 10.3 !MJ d4r< fl.0 *.G
10. * 12 .0 12 ,2 - - i I .<•
7«* V<» *. \ - • *> ,0
^. 6 U .* *.0 - - 6 *
0 0 0 no diti 0 0
0 J 0 - - o
J ^
J Jj
c
J
-------
Vertli
ftifcarta
Port*
«**«*
Cc^hit fatal
Bf# tag)
¦pviii
rail IfH
Itft
1 my
3? Mil
n
10 J**.
7 AfH'ii
6 **rii
1 Jot?
I * Oe I .
> i Jan.
$ Aptl\
28 J\lT!t
I7 Oct.
/ Jtan.
D»jMfcn
12 *cf - - . * fv '
n ton. 6 frb. ft **'•, - * fmh
-------
Table 2. Sediment Type by Study Area - Tide Height
Tide
Study Area Height +6* +3'
Tongue Point
Pillar Point
North Beach Cobble
Morse Creek
Beckett Point
Dungenesa Spit
Twin Rivers
North Beach Sand
SCydaka Beach
rock
rock
cobble/sand
gravel
cobble/sand-
grave 1
sand-gravel
sand-gravel
sand-gravel
fine-med "and
fine-coarse
sand
rock
rock
cobb]e/sand
cob 'sand
sand-gravel
sand-gravel
sand-gravel
fine-med sand
fint-coarse
sand
jamestown
sand-gravel
fine sand
+0'
rock
rock
cobble/band
cobble/sand
f t ,i-med sand
sand-grave 1
and-grave1
(+2')
fine-mod sand
f ine-coars j
sand
( +-1. 4 ")
Fied sand
rock
fine-med sand
cobble/gravel
fine-med sand
sand-gravel
gravel
cobble/sand-
gravel
f ine-nu'd sand
f ine-coarse
sand
rock
fine-med sand
cobble/gravel
f ine-CEed sand
sand-gravel
fine sand-silt
cobble/sand-
gravel
f ine-med sand
f ine-coarse
sand
-------
KYCWA BEACH
STUDY ADEh
TfOi
S.PfiW
>*-
. W&M r*
SO M
SAN JUAN
V
1^11
Ri^fc
-------
60M
PILLAR POINT
STUDY AREA
-------
TW/N RIVERS
STUDY ABE A
<#£JST TWiN
l Rives
-------
tOH'
6i)£'
pG
iTUf'
A
1M
f Vf
,ol<
-------
MOffSE CREEK |
study AQEA i
ac3
-------
Qi.cncmoN
APPA
*. WOOOJ
DUNGFNESS SPiT
STOOY AREA
Figure i>
481
-------
\
30
u
JAfi/£6T0WN STUDY AREA
-------
3c
c^^/>
.^Q/j
4^r
-------
*x
------- |