50%

!!RH SEDIMENT
                           FIELD VERIFICATION PROGRAM
                               (AQUATIC DISPOSAL)

                               TECHNICAL REPORT D-85-7

               USE OF BIOENERGETICS TO INVESTIGATE THE
               IMPACT OF DREDGED MATERIAL  ON  BENTHIC
                  SPECIES:   A  LABORATORY STUDY  WITH
                      POLYCHAETES AND  BLACK  ROCK
                              HARBOR MATERIAL
                                       by
                         D. Michael Johns, Ruth Gutjahr-Gobell
                            Edgerton Research Laboratory
                               New England Aquarium
                            Boston, Massachusetts  02110
                                       and
                                   Paul Schauer
                          Environmental Research Laboratory
                          US Environmental Protection Agency
                          Narragansett,  Rhode Island 02882
September 1985
 Final Report
                          Approved For Public Release; Distribution Unlimited
                        Prepared for DEPARTMENT OF THE ARMY
                             US Army Corps of Engineers
                             Washington, DC  20314-1000
                        and US Environmental Protection Agency
                               Washington, DC  20460
                          Monitored by Environmental Laboratory
                     US Army Engineer Waterways Experiment Station
                     PO Box 631, Vicksburg, Mississippi 39180-0631

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   Destroy this report  when no longer needed.  Do not return
                      it to the originator.
The findings  in this report are not  to be construed  as an  official
     Department of the Army position unless so designated
                by other authorized documents.
       The contents of this report  are not to be used for
        advertising, publication,  or  promotional  purposes.
         Citation of trade names does not constitute an
         official endorsement or approval of the use of
                  such commercial products.
       The D-series  of reports includes publications of the
          Environmental Effects of Dredging  Programs:
             Dredging Operations Technical Support
           Long-Term Effects of Dredging Operations
       Interagency Field Verification of  Methodologies for
        Evaluating Dredged Material  Disposal Alternatives
                  (Field Verification Program)

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SUBJECT:  Transmittal of Field Verification Program Technical Report Entitled
          "Use of Bioenergetics to Investigate the Impact of Dredged Material
          on Benthic Species:  A Laboratory Study with Polychaetes and Black
          Rock Harbor Material"

TO:  All Report Recipients
1.  This is one in a series of scientific reports documenting the findings  of
studies conducted under the Interagency Field Verification of Testing and
Predictive Methodologies for Dredged Material Disposal Alternatives (referred
to as the Field Verification Program or FVP).  This program is a comprehensive
evaluation of environmental effects of dredged material disposal under condi-
tions of upland and aquatic disposal and wetland creation.

2.  The FVP originated out of the mutual need of both the Corps of Engineers
(Corps) and the Environmental Protection Agency (EPA) to continually improve
the technical basis for carrying out their shared regulatory missions.  The
program is an expansion of studies proposed by EPA to the US Army Engineer
Division, New England (NED), in support of its regulatory and dredging mis-
sions related to dredged material disposal into Long Island Sound.  Discus-
sions among the Corps' Waterways Experiment Station (WES), NED, and the EPA
Environmental Research Laboratory (ERLN) in Narragansett, RI, made it clear
that a dredging project at Black Rock Harbor in Bridgeport, CT, presented a
unique opportunity for simultaneous evaluation of aquatic disposal, upland
disposal, and wetland creation using the same dredged material.  Evaluations
were to be based on technology existing within the two agencies or developed
during the six-year life of the program.

3.  The program is generic in nature and will provide techniques and inter-
pretive approaches applicable to evaluation of many dredging and disposal
operations.  Consequently, while the studies will provide detailed site-
specific information on disposal of material dredged from Black Rock Harbor,
they will also have great national significance for the Corps and EPA.

4.  The FVP is designed to meet both Agencies' needs to document the effects
of disposal under various conditions, provide verification of the predictive
accuracy of evaluative techniques now in use, and provide a basis for deter-
mining the degree to which biological response is correlated with bioaccumula-
tion of key contaminants in the species under study.  The latter is an
important aid in interpreting potential biological consequences of bioaccumu-
lation.  The program also meets EPA mission needs by providing an opportunity
to document the application of a generic predictive hazard-assessment research
strategy applicable to all wastes disposed in the aquatic environment.  There-
fore, the ERLN initiated exposure-assessment studies at the aquatic disposal
site.  The Corps-sponsored studies on environmental consequences of aquatic
disposal will provide the effects assessment necessary to complement the EPA-
sponsored exposure assessment, thereby allowing ERLN to develop and apply a
hazard-assessment strategy.  While not part  of the Corps-funded FVP, the EPA
exposure assessment studies will complement  the Corps' work, and  together the
Corps and the EPA studies will satisfy the needs of both  agencies.

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 SUBJECT:   Transmittal of Field Verification Program Technical Report Entitled
           "Use  of  Bioenergetics  to  Investigate  the Impact  of Dredged Material
           on  Benthic Species:  A Laboratory Study with Polychaetes and Black
           Rock  Harbor Material"

 5.   In recognition of the potential national significance, the Office, Chief
 of  Engineers, approved and  funded the studies in January 1982.  The work is
 managed through the Environmental Laboratory's  Environmental Effects of
 Dredging Programs  at WES.   Studies  of the effects of upland disposal and
 wetland creation are being  conducted by WES and studies of aquatic disposal
 are being  carried  out by the ERLN,  applying techniques worked out at the
 laboratory for  evaluating sublethal effects of  contaminants on aquatic organ-
 isms.  These  studies are funded  by  the Corps while salary, support facilities,
 etc., are  provided by EPA.  The  EPA funding to  support the exposure-assessment
 studies followed in 1983; the exposure-assessment studies  are managed and
 conducted  by  ERLN.

 6.   The Corps and  EPA are pleased at the opportunity to conduct cooperative
 research and  believe that the value in practical implementation and improve-
 ment of environmental regulations of dredged material disposal will be con-
 siderable.  The studies conducted under this program are scientific in nature
 and will be published in the scientific literature as appropriate and in a
 series of  Corps technical reports.  The EPA will publish findings of the
 exposure-assessment studies in the  scientific literature and in EPA report
 series.  The  FVP will provide the scientific basis upon which regulatory
 recommendations will be made and upon which changes in regulatory implementa-
 tion, and  perhaps  regulations themselves, will  be based.   However, the docu-
 ments produced  by  the program do not in themselves constitute regulatory
 guidance from either agency.  Regulatory guidance will be  provided under
 separate authority after appropriate technical  and administrative assessment
 of  the overall  findings of  the entire program.
      ChoromokosT Jr., Pti.D.,  P.E.
Director, Research and Development
U. S. Army Corps of Engineers
Bernard D. Goldstein, M.D.
Assistant Administrator  for
Research and Development
U. S. Environmental Protection
Agency

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         Unclassified
SECURITY CLASSIFICATION OF THIS PAGE (When OMB Entered)
            REPORT DOCUMENTATION PAGE
                                                          READ INSTRUCTIONS
                                                        BEFORE COMPLETING FORM
1. REPORT NUMBER
                                     2. GOVT ACCESSION NO
 Technical Report  D-85-7
                                                       3. RECIPIENT'S CATALOG NUMBER
4. TITLE (and Subtitle)
USE OF BIOENERGETICS  TO INVESTIGATE THE IMPACT OF
DREDGED MATERIAL ON BENTHIC SPECIES:  A LABORATORY
STUDY WITH POLYCHAETES AND BLACK ROCK HARBOR
MATERIAL
                                                   S. TYPE OF REPORT 4 PERIOD COVERED

                                                     Final Report
                                                   S. PERFORMING ORG. REPORT NUMBER
7.  AJTHORf.J
 D. Michael Johns,  Ruth Gutjahr-Gobell,
 Paul Schauer	
                                                       8. CONTRACT OR GRANT NUMBERf.)
9.  PERFORMING ORGANIZATION NAME AND ADDRESS
New England Aquarium Edgerton Research  Laboratory,
Central Wharf, Boston,  Massachusetts  02110 and
US Environmental  Protection Agency, Environmental
Research Laboratory,  South Ferry Road,
Narragansett. Rhode  Island  02882	
                                                   \0. PROGRAM ELEMENT, PROJECT, TASK
                                                      AREA i WORK UNIT NUMBERS

                                                     Field Verification Program
                                                     (Aquatic Disposal)
U. CONTROLLING OFFICE NAME AND ADDRESS
DEPARTMENT OF THE ARMY,  US Army Corps  of Engineers
Washington, DC  20314-1000 and US Environmental
Protection Agency, Washington. DC   20460
                                                   t2. REPORT DATE
                                                     September 1985
                                                    13. NUMBER OF PAGES
 14. MONITORING AGENCY NAME & ADDRESS^/ different /ran Controlling Otllce)

 US Army Engineer Waterways Experiment Station
 Environmental Laboratory
 PO Box 631, Vicksburg, Mississippi  39180-0631
                                                    IS. SECURITY CLASS, (at thlt report)

                                                     Unclassified
                                                    IS*. O ECL ASSI FICATtON/DOWNGRADING
                                                       SCHEDULE
IS. DISTRIBUTION STATEMENT (ol title Report)
 Approved for public  release; distribution unlimited.
 17. DISTRIBUTION STATEMENT (at the ebeueet entered In Stock 30, II different Inrrt Report)
 IS. SUPPLEMENTARY NOTES
Available from National Technical  Information Service,  5285 Port Royal Road,
Springfield, Virginia  22161.
19. KEY WORDS (Confirm* on rtreret elde II neceeeery end Identity by block number)
 Bioenergetics—Technique   (LC)
 Dredging—Environmental aspects  (LC)
 Dredged material  (WES)
 Polychaeta  (LC)
 Benthos   (LC)	
                                           Marine pollution  (LC)
                                           Dredging—Connecticut—Black Rock
                                             Harbor  (LC)
20. ABSTRACT (Continue ea rente* elite ft mcMMqr end Identify by Mock number)
      Both solid  phase and particulate phase assays were conducted with two
species of polychaetes to determine the accuracy  and reproducibility of con-
ducting bioenergetic studies on  poLychaetes exposed  to highly contaminated
dredged sediment.   The two species  tested were Nephtys incisa, an errant
burrowing sediment  ingestor, and Neanthes arenaceodentata, a tube-building sur-
face feeder.  Exposure to various treatments was  for 10 days.
                                                                        (Continued)
DO
 FORM
t JAN 73
1473
EDITION OF I NOV 65 IS OBSOLETE
                                                      Unclassified
                                             SECURITY CLASSIFICATION1 OF THIS PAriC fWi.fi Dele Entered)

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            Unclassified
SECURITY CLASSIFICATION OF THIS PA.GEfWh«B D«<« Enl.r.rf)
20.   ABSTRACT  (Continued).

     Results with both species  of  polychaetes indicate that, with few exceptions,
all of the physiological parameters measured  (rates of feeding, growth, repro-
duction, and ammonia excretion)  can be  made with accuracy.  Changes in growth
(determined as dry weight) between treatments,  for example, can be measured
following a 10-day exposure period providing  that care is taken to adequately
size the individual polychaetes  prior to initiation of the experiment.

     The bioenergetic endpoints  measured in this study were found to be
repeatable.  In addition,  physiological responses were found to be dose-
dependent.  Dosage was based  on the relative  proportion of reference and Black
Rock Harbor sediment in a  particular  treatment.

     This investigation is the  first  phase in developing field verified
bioassessment evaluations  for the  Corps of Engineers and EPA regulatory program
for dredged material disposal.   This  report is not suitable for regulatory pur-
poses; however, appropriate assessment  protocols that are field verified will be
available at the conclusion of  this program.
                                                 Unclassified
                                        SECURITY CLASSIFICATION OF THIS PAGEfHTiwt D«(» Entered)

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                                   PREFACE






     This report describes work performed by the US  Environmental  Protection




Agency (EPA) Environmental Research Laboratory,  Narragansett,  R. I.  (ERLN),




as part of the Interagency Field Verification of Testing and Predictive




Methodologies for Dredged Material Disposal Alternatives Program (Field




Verification Program (FVP)).  This program is sponsored by the Office, Chief




of Engineers, and assigned to the US Army Engineer Waterways Experiment




Station (WES), under the purview of the Environmental Laboratory's (EL)




Environmental Effects of Dredging Programs (EEDP).  The OCE Technical Monitors




were Drs. John Hall and William L. Klesch.  The objective of this  program




agreement Is to verify existing predictive techniques for evaluating the




environmental consequences of dredged material disposal under aquatic, wet-




land, and upland conditions.  The aquatic portion of the FVP study is being




conducted by ERLN, with the wetland and upland portion done by WES.




     Although not totally inclusive, we would like to thank the following




researchers:  Drs. John Gentile, Gerald Pesch, John Scott, Mr. William Nelson,




and Ms. Carole Pesch for their many discussions and criticisms;  Mr. Michael




Balboni for  the use of and help with their benthic exposure systems;




Ms. Cornelia Meuller for supplying the _N. arenaceodentata juveniles;




Dr. Wayne Davis for use of  the  'ant farm'; and Dr. James Heltshe for advice




on statistical procedures.  We would especially like to thank Dr.  Anthony




Calabrese and Capt. Robert Alix of the National Marine Fisheries Service in




Milford, Conn., for boat time on  the R/V  Shang Wheeler.  This research was




supported by Cooperative Agreement CR809956  between  the US  Environmental




Protection Agency and  the New England Aquarium  to Dr. D. Michael Johns.




     The EPA Technical Director for the FVP  was Dr.  John H. Gentile;  Technical

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Coordinator was Mr. Walter Galloway; and the Project Manager was Mr. Allan

Beck.

     The study was conducted under the direct management of Drs. Thomas M.

Dillon and Richard K. Peddicord and under the general management of Dr. C.

Richard Lee, Chief, Contaminant Mobility and Criteria Group; Mr. Donald L.

Robey, Chief, Ecosystem Research and Simulation Division; and Dr. John

Harrison, Chief, EL.  The FVP Coordinator was Mr. Robert L. Lazor, and the

EEDP Managers were Mr. Charles C. Calhoun, Jr., and Dr. Robert M. Engler.

     COL Tilford C. Creel, CE, and COL Robert C. Lee, CE, were Commanders

and Directors of WES during the conduct of the study.  COL Allen F. Grum,  USA,

was Director of WES during the preparation and publication of this report.

Mr. Fred R. Brown and Dr. Robert W. Whalin were Technical Directors.

     This report should be cited as follows:

     Johns, D.  M.,  Gutjahr-Gobell,  R.,  and Schauer, P.  1985.
     "Use of Bioenergetics to Investigate the Impact of
     Dredged Material on Benthic Species:  A Laboratory
     Study with Polychaetes and Black Rock Harbor Material,"
     Technical  Report D-85-7,  prepared by US Environmental
     Protection Agency,  Environmental Research Laboratory,
     Narragansett,  R.  I., for  the US Army Engineer Waterways
     Experiment Station,  Vicksburg, Miss.

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                             CONTENTS


                                                               Page

PREFACE	    1

LIST OF FIGURES	    4

LIST OF TABLES	    5

PART I: INTRODUCTION	    7

       Background	    7
       Purpose	    8
       Scope	    8

PART II: MATERIALS AND METHODS	   10

       Sediment Sources and Exposure Systems	   10
       Experimental Organisms	   17
       Physiological Measurements	   18
       Burrowing Activity	   25
       Statistical Analysis	   27

PART III: RESULTS	   30

       Sources of Variation	   30
       Treatment of Data	   31
       Consumption	   32
       Production	   34
       Maintenance Costs	   38
       Partitioning of Energy Resources	   44
       Scope for Growth	   52
       Burrowing Activity	  54

PART IV: DISCUSSION	  60

PART V: CONCLUSIONS	  71

REFERENCES	  72

APPENDIX A: SOLID PHASE AND  PARTICULATE PHASE DATA SHEETS	  Al

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                          LIST  OF FIGURES
 No.

 1       Central Long  Island Sound disposal site and South
         reference site	  11

 2       Black Rock Harbor, Connecticut, source of dredged
         material	  11

 3       Sediment dosing system with chilled water bath and
         argon gas supply	  14

 4       Schematic of  the suspended distribution and dosing
         system used to expose juvenile Nj^ incisa to reference
         and BRH sediment	77	  16

 5       Schematic of respirometers (syringe and stender dish)
         used to determine oxygen consumption rates of
         N. incisa juveniles	  24

 6       Schematic of  the narrow, glass-walled aquarium used
         to determine burrowing activity and maximum burrow
         depth	  27

 7       Net growth efficiency of N_._ incisa juveniles exposed
         to various mixtures of reference and BRH sediment for
         10 days	  45

 8       Net growth efficiency of N. incisa juveniles exposed
        to various combinations of sediment and suspended
        particles for 10 days	  48

 9       Drawing of burrows created by IJ^ incisa juveniles
        exposed to reference and BRH sediments for 10 days....  54

10       Drawing of burrows created by N. incisa juveniles
        exposed to various combinations of reference and
        BRH sediment for 10 days	  56

11       Drawing of burrows created by N._ arenaceodentata
        juveniles  exposed  to  various  combinations  of
        reference  and BRH sediment for 10 days	  58

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                         LIST OF TABLES
 No.                                                           Page

 1   Sampling protocol  for  physiological measurements at the
    end  of  the  10-day  exposure period	   20

 2   Sources of  variation in respiration and  ammonia
    excretion rate  of  Nj^ incisa  juveniles determined during
    initial experiments	   31

 3   Food consumption rates of 11.  arenaceodentata  juveniles
    exposed to  various sediment  conditions	   33

 4   Changes in  dry  weight  of N_._ incisa juveniles  exposed
    to various  sediment  conditions	   35

 5   Changes in  dry  weight  of !*._ arenaceodentata juveniles
    exposed to  various sediment  conditions	   37

 6   Weight-specific respiration  rate of N^ incisa juveniles
    exposed to  various sediment  conditions	   39

 7   Weight-specific respiration  rate of N.  arenaceodentata
    juveniles exposed  to various sediment  conditions.	   40

 8   Weight-specific ammonia excretion rate  and 0:N ratios of
    N. incisa juveniles  exposed to various  sediment
    conditions	   42

 9   Weight-specific ammonia excretion rate  and 0:N ratios
    of N. arenaceodentata  juveniles exposed to various
    sediment conditions	   43

10   Cummulative energy values  for production and
    maintenance costs of 11. incisa juveniles	    45

11   Net growth  efficiency  of N. incisa juveniles exposed to
    various sediment  conditions	    47

12   Cummulative energy values  for production and
    maintenance costs of IS. arenaceodentata juveniles
    exposed to  various sediment conditions	    50

13   Net growth efficiency  of N. arenaceodentata juveniles
    exposed to  various sediment conditions	    51

14   Scope for growth  values for Nj^ arenaceodentata
    juveniles exposed to various sediments	    53

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No.                                                            Page

15  Burrow length and maximum burrow depth of N. incisa
    juveniles exposed to various sediment conditions	     55

16  Burrow length and maximum burrow depth of
    N. arenaceodentata juveniles exposed to various
    sediment conditions	     59

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           USE OF BIOENERGETICS TO INVESTIGATE THE IMPACT
       OF DREDGED MATERIAL ON BENTHIC SPECIES;   A LABORATORY
       STUDY WITH POLYCHAETES AND BLACK ROCK HARBOR MATERIAL
                        PART I: INTRODUCTION

                            Background

       1.  Effective short-term biological response measurements
which can adequately detect the effects of environmental concentra-
tions of contaminants have been called for by several international
commissions (International Council for the Exploration of the Sea
1978; Mclntyre and Pearce 1980).  In order to be of value the measure-
ments must have some relevance to ecological fitness.  In addition,
these relatively short-term laboratory effects tests (usually less
than a month) must have a predictive capability which allows estima-
tion of the degree of ecological change which will take place.
       2.  An effects measurement technique which may satisfy the
preceding criteria is the determination of biological energy balances
(Edwards 1978;  Capuzzo and Lancaster 1981;  Johns and Pechenik
1980;  Johns and Miller 1982;  McKinney 1982) along with its corol-
laries, including scope for growth (Warren and Davis 1967; Bayne
1975).  Previous studies using these principles have found a reason-
able correlation between changes in energy balances or scope for
growth and changes in population fitness  (Bayne et al. 1979;
Gilfillan 1980).  In a series of detailed field studies, for example,
Gilfillan and his co-workers (Gilfillan and Hanson 1975; Gilfillan

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 et  al.  1976;  Gilfillan and  Vandermeulen  1978)  found  a  reduced scope




 for growth  in the bivalve   Mya  arenaria  collected  from oil-impacted




 sites when  compared  to individuals  from  nearby, relatively clean




 reference populations.  These data  were  related to and predictive




 of  eventual changes  in population structure  observed in  the  impacted




 sites.   Changes in population structure  that Gilfillan could relate




 to  the  reduced scope for growth included reductions  in yearly growth




 rate and population  density.






                              Purpose






        3.   The biological effects portion of the Field Verification




 Program (FVP)  is being implemented  in two phases.  The first phase




 is to identify, biological test procedures that are responsive to




highly  contaminated  dredged material.  In this phase,  the applica-




bility,  reproducibility, and repeatability of the biological measure-




ment in  the laboratory are to be demonstrated.   The  second phase will




be to field verify the biological responses observed in the labora-




tory to  determine the predictability of  the laboratory-derived data.




The purpose of this  report is to describe the results of the first




phase in our efforts to apply bioenergetics techniques to two species




of polychaetes.






                                Scope






        4.   This paper describes efforts  to evaluate  the utility of




bioenergetics techniques with the polychaetes Nephtys  incisa and




Neanthes arenaceodentata and to determine what effects highly

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contaminated dredged Black Rock Harbor sediment have on these energy




budgets.  Nephtys incisa is a dominant infaunal macroinvertebrate




in Long Island Sound benthic communities (Sanders 1956, 1958; Carey




1962).   Impact from contaminated dredged material disposal on this




species may change the present community structure.   Neanthes




arenaceodentata,  on the other hand, is not indigenous to the Long




Island Sound study area.  Rather it is being tested as a possible




surrogate for those infaunal polychaete species that may occur at




any disposal site.  In disposal sites where indigenous species may be




difficult to collect, or maintain,  or study in the laboratory, a




surrogate species offers the opportunity to still determine potential




impacts on that general group of organisms.  In proposed disposal




sites,  where little scientific information exists on the indigenous




species, the use  of a suitable surrogate allows for the assessment




of potential impacts without the time-consuming learning curve




(i.e.  methods development, etc.) that would be needed with the




indigenous species.

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                   PART II:  MATERIALS AND METHODS




               Sediment Sources  and  Exposure  Systems




        5.   Reference sediment  for these  studies  was  collected from


 the FVP South Reference site  (40°7.95"N  and  72°52.7"W), which is


 approximately 700 m south of  the southern perimeter  of the Central


 Long Island Sound disposal  site (Figure  1).  Reference sediment was


 collected with a  Smith-Mclntyre grab sampler (0.1 m^) in August and


 December 1982 and May 1983  (collections  I, II, and III, respectively).


 Sediment from each collection was returned to the laboratory, press


 sieved  (wet)  through a 2-mm mesh stainless steel screen, homogenized,


 and stored  in polypropylene (collection  I) or glass  (collections II


 and III) containers  at  4°C  until used in  experiments.  Each container


 of  material was coded with  collection number, date,and jar number.


 (See Lake et  al.   1984  for  complete  details.)


       6.  BRH sediment was collected from 25 locations within the highly


 indistrialized Black Rock Harbor (Bridgeport, Conn.;  Figure 2)  study area

            2
with a 0.1-m  gravity box corer to a depth of 1.21 m.  The contaminated


sediment was homogenized, distributed to barrels, and stored at  4°C.   The


contents of each barrel were homogenized, wet sieved  through a 1-mm sieve,


distributed to glass jars,  and stored at  4°C until used in experi-


ments.  Samples of sediment were taken at various points in the


collections, mixing, and distribution procedure for moisture content


and chemical analysis.  (See Lake  et al.  1984 for details.)
                                10

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                                             SOUTH REFERENCE
                                                  • SITE
Figure 1.  Central Long Island Sound disposal site and
           South reference site
 Figure 2.  Black Rock Harbor, Connecticut,  source  of
            dredged material
                          11

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       7.  Two exposure regimes were used in this study.  One was a




solid phase assay in which the polychaetes were exposed to various




combinations of reference and BRH sediment for 10 days.  A 10-day




exposure period was chosen based on data provided in Pesch and




Hoffman (1983).  The other exposure regime used was a particulate




phase assay in which worms were exposed to either reference or BRH




particles after being placed in 100 percent reference and 100 percent




BRH, respectively.




       8.  For the experiments in which the worms were to be exposed




to only the solid phase of the sediments, approximately 400 ml of




the appropriate substrate mixture was placed in a 150- by 75-mm




dish, and then put in a water bath of 20°C (Figure 3).  Seawater at




20°C was allowed to flow through the treatment bowls at a rate of




approximately 50 ml/min.  Where both reference and BRH sediments




were used in a treatment, the two sediments were combined in a




volume-to-volume ratio and throughly mixed.




       9.  Implementation of the particulate phase assays required




the consruction of two identical sediment dosing systems to simul-




taneously provide either BRH or reference material as suspended




sediment.  The dosing systems (Figure 3) consisted of conical-shaped




slurry reservoirs placed in a chilled fiberglass chamber, a diaphragm




pump, a 4-jj. separatory funnel, and several return loops that




directed the particulate slurry through dosing valves.  The slurry




reservoirs (40 cm diam.  by 55 cm high) contained 40 I of slurry




composed of 37.7 & of filtered seawater and 2.3 Jl of either BRH
                                12

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or reference sediment.  The fiberglass chamber (94 cm by  61  cm by




79 cm high) was maintained between 4°and 10°C using an externally




chilled water source.  (The slurry was chilled to minimize microbial




degradation during the test.)   Polypropylene pipes (3.8 diam) placed




at the bottom of the reservoir cones were connected to the diaphragm




pumps (16 to 40 fc/min capacity) that had Teflon diaphragms.   These




pumps were used to circulate the slurry but minimize abrasion so




that the physical properties and particle sizes of the material




remained as unchanged as possible.




       10.  The separatory funnel was connected to the pump and




returned to the reservoir by polypropylene pipes.  The separatory




funnel served two functions:   (a) to  ensure  that  a constant head




pressure was provided by the overflow, and (b) to serve as a  connec-




tion for the manifold located  4 cm  below the  constant  head level.




The manifold served  to distribute the slurry  by directing a portion




of  the flow from the funnel  (through  6 mm inside  diameter polypro-




pylene tubes) through the  Teflon dosing  valves and back  to  the




reservoir.  At  the  dosing  valves,  the slurry was  mixed with seawater




to  provide the  desired  concentrations for the toxicity tests.




Argon  gas  was  provided  at  the rate  of 200 ml/min  to  the  reservoir




and separatory  funnel to  minimize oxidation  of the  sediment/seawater




slurry.  The dosing valves were controlled  by a microprocessor.




The microprocessor  can  be  programmed  to deliver  a pulse  with a




duration of  0.1 sec up  to continuous pulse delivery and at  intervals




from  once  every second  to once every hour.
                                 13

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           ARGON
          INJECTION
       SEPARATORY
         FUNNEL
                                 DELIVERY
                                 MANIFOLD
                                             DOSING
                                             VALVE
                                        TO EXPOSURE
                                          SYSTEM
                                   RETURN
                                   MANIFOLD
                                    SLURRY
                                    RESERVOIR
         CHILLED
         WATER BATH
Figure 3.   Sediment dosing system with chilled water  bath and
           argon gas supply
                           14

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       11.  Within the exposure system distribution jar,  seawater  at

20°C with the appropriate concentration of  suspended particles  was

allowed to flow through the treatment bowls at a. rate of  approximately

35 ml/min (Figure 4).   In order to maintain the particles in  suspen-

sion as long as possible, a small crystallization dish with a stir

bar was placed in the  middle of the exposure dish.  For continuity

throughout this report, the scheme used to describe the particulate

phase assay treatments in which there was both suspended  and  bedded

sediments was:  suspended particulate phase/solid phase.

       12.  In both the solid phase and particulate phase assays 15

worms of approximately the same size were placed in each treatment.

After the 10-day exposure, the worms were sieved out of the mud,

counted, and saved for physiological measurements.

       13.  During the experiment the worms were offered  prawn flakes

(Aquatic Diet Technology, Inc., Brooklyn, N.Y.) as a food source.

Previous research* has indicated that laboratory holdings of K._

incisa grow better when offered prawn flakes than when left without.

It is unclear, however, whether the prawn flakes are used directly

as a food source or whether the flakes act as a substrate for bacte-

rial growth, the bacteria then being utilized as the food source.

Prawn flakes is also the food source used in maintaining laboratory

populations of N. arenaceodentata  (Schauer and Pesch, In Preparation).

With this species, the prawn flakes are utilized directly as food.
    * Personal communication, Paul Schauer, July  1983, U.S.
 Environmental Protection Agency.
                                 15

-------
                   DOSING SYSTEM
                                      •Suspended Particles
    Spigot-
                            u
                   5OOOC
                                      -Distribution Jar
                                    —Stir  Bar
                EXPOSURE  SYSTEM
    Stir Bar
                          \\\
                                             Exposure
                                            "Container
        \\\ \\\\\\\\V\\\\\\ \\\\\\\\\\\\ \\\\\vs
Figure 4.  Schematic of the suspended distribution and dosing system
          used to expose juvenile N^ incisa to reference and BRH
          sediment
                             16

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                       Experimental  Organisms






       14.  Nephtys incisa were collected from Long  Island  Sound in




the vicinity of the South Reference  site.  Bottom samples were




collected using a Smith-Maclntyre grab and were initially sieved on




board the research vessel.  Worms which passed through a 2-mm sieve




but were retained on a 0.37-mm sieve were kept for laboratory exper-




imentation and were all juveniles (Carey 1962).  The worms  were




then placed in a single, unsieved Smith-Maclntyre grab sample for




transport back to the laboratory.  In the laboratory, the worms




were resieved with a 0.37-mm mesh using  seawater at temperatures




close to collection temperature (+ 2°C).




       15.  Following this, individuals were visually separated into




relative size classes for experiments.   Although somewhat subjective,




careful visual separation leads to a coefficient of variation in




dry weight at the start of an experiment of only 24 percent.  Other,




more accurate sizing techniques, such as wet weight determinations




employing a wet-weight-to-dry-weight regression curve,  are time-




consuming and, more importantly, may cause physical damage to the




worms from the handling required.




        16.  All  experiments were conducted at  20°C.   Laboratory




acclimation for  Nephtys incisa  collected at 20° + 3°C was a  minimum




of  3 days  to  allow  for  adjustments  to  laboratory holding conditions.




For worms collected at  temperatures below 16°C, acclimation  included




a  1° to  2°C increase  in temperature per day until  20°C  was  attained.
                                 17

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        17.  Neanthes arenaceodentata juveniles used in this study

were from laboratory cultures (original stock from D.J. Reish,

California State University, Long Beach, Calif.)•  Worms were cul-

tured in a flow-through system at 20° + 1°C and fed prawn flakes

according to techniques described in Schauer and Pesch (In Prepara-

tion).  Worms used in an experiment were from the same hatch and

were of approximately the same age.  Refer to summary data sheets

in Appendix A for complete information on the ages of the worms

used in each experiment.


                     Physiological Measurements


        18.  Although the fundamental integration of biochemical and

physiological mechanisms is complex and difficult to measure, the

net response can be measured at the whole organism level.  Bioener-

getic analysis compares the major anabolic and catabolic processes

that occur and allows for an evaluation of the relative partitioning

of available energy amongst growth and maintenance requirements.

        19.  The energy budget of the juvenile stage of an organism

can be described by the following formula:

                      C-P + R + E + F                       (1)
where

    C = total energy consumed

    P = amount of energy converted to tissue

    R = amount of energy used for maintenance measured via aerobic
        respiration

    E » amount of energy lost through ammonia excretion

    F • amount of energy lost through feces
                                18

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       20.  Scope for growth (SFG),  which is a derivation of  the




balanced energy budget formula, is an approach which provides for




an estimation of the potential for growth (Warren and Davis 1967).




In this approach, the amount of energy available for growth (and




reproduction in sexually mature stages) is estimated by the following




relationship:






                         P = C - F - (R+E)                   (2)






Since physiological measurements in SFG studies are typically made




over a short period of time (< 24 hr), P in this case is an instan-




taneous measure of growth potential.  Scope for growth is an index




by which the current physiological condition of an organism can be




evaluated.




       21.  The approach taken in this study was to attempt to (a)




measure all parameters in the balanced energy budget equation to




determine reliability of the measurements and ease at which the




measurements can be made and (b) estimate variability in measurement




so that sample size for the experiments can be determined.  Following




this, scope for growth values were determined where possible while




at the same time actual growth was being monitored.  Table 1 presents




the sampling protocol and the approximate times needed to make the




various physiolgical measurements.




       22.  For N^ arenaceodentata, values  of  consumption, produc-




tion, respiration, and excretion were measured directly.   Therefore,




scope for growth values as well as balanced energy  budgets could be




determined for this species.  With N.  incisa,  however, only  values
                                19

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                               Table 1


    Sampling protocol for physiological measurements at the end of

                        10-day exposure period
                   Physiological
                   Measurement
                     Feeding

                     Respiration

                     Excretion

                     Total Time Required
                     Feeding

                     Respiration

                     Excretion

                     Total Time Required
      Time
    Required
       hr
  Nephtys incisa



         2

         3

         5


Neanthes arenaceodentata

        24

         4

         a*

        28
* a » Pespiration and excretion rates determined at same time in syringe
      respirometer in a time of 4 hr
                                 20

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of production, respiration, and excretion could be measured directly.




Feeding rates, on the other hand, could not be quantitatively meas-




ured with any relability.  In failing to quantitatively estimate




food consumption rates,  neither scope for growth nor  a balanced




energy budget for II.  incisa could be derived,  although an estimate  of




the relative partitioning of energy between production and maintenance




could be made.  The measured parameters allowed for the calculation




of a net growth efficiency using the following formula:






              Net growth Efficiency = P/(P+R+E) x 100%        (3)






       23.  Net growth efficiency values offer insight into the




degree of integration among physiological processes.   It offers a




time course estimate of the cumulative effects a particular




condition has had on an organism.  To allow for comparison to N_._




incisa, net growth efficiencies were also calculated for N.




arenaceodentata.







Food consumption and assimilation efficiency




       24.  Food consumption rates of individual N^ arenaceodentata




were determined over a period of 24 hr using a preweighed amount of




prawn flake as the food source.  Following this period, remaining




food was taken from the bowl, rinsed in deionized water, dried to a




constant weight at 60°C, and weighed to the nearest 1 yg on a




Perkin-Elmer AD-2Z Autobalance (Perkin-Elmer Corp., Norwalk, Conn.).




Food consumption rates were taken as the difference in dry weight




of the food initially offered and the amount remaining at  the end
                                21

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of 24 hr.  In addition to collecting any remaining food particles,




fecal material was also collected, rinsed, dried, and weighed.




       25.  The efficiency of food assimilation could then be deter-




mined as the difference in energy content between food and fecal




material (energy content of fecal material - energy content of food




source[J/mg]/energy content of food source[J/mg] x 100).  The energy




content of both the food and fecal material was determined using a




wet oxidation technique in the presence of an acid-dichrotnate mixture




(Maciolek 1962).






Production




       26.  In all experiments, juvenile worms were used so that




changes in dry weight reflected only changes in growth (production).




At the beginning of an experiment, a subsample of individuals (10




to 15) was taken to estimate initial worm weight.  The worms were




quickly rinsed in deionized water, dried at 60°C for 24 hr, then




weighed on a Perkin-Elmer AD-2Z Autobalance to the nearest ug.




Following the 10-day exposure, 10 to 15 worms were taken from the




experimental conditions to assess their physiological state.




Individual respiration (see paragraphs 27-28) and excretion rates




(see paragraph 31) were determined prior to rinsing, drying,and




weighing.  Table 1 presents the time course needed to make the




physiological measurements for both Ni^ incisa and K[._ arenaceodentata.




Changes in growth were computed as being the difference in dry




weight from the beginning to the end of the 10-day exposure period.




After the worms were dried and weighed, energy values for  the tissue
                                22

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were determined using a wet oxidation technique in the presence of




an acid-dichromate mixture (Maciolek 1962).






Respiration




       27.  For N. incisa, routine rates of  oxygen consumption were




measured using a 3-cc syringe containing a small amount of surficial




sediment (0.5 to 0.75 cc) as a respirometer  (Figure 5).  A worm was




placed in each syringe with 1.5 cc of air-equilibrated seawater.




After 1 hr, the oxygen concentration of 1 cc of the seawater was




determined by injecting the sample onto the  face of a Radiometer




oxygen probe (Radiometer, Copenhagen, Denmark) fitted with a Radio-




meter thermostated sampling cell.  The syringe was then refilled to




1.5 cc, and the new oxygen concentration recalculated.  This proce-




dure was repeated once per worm.  Three control syringe respirometers




containing 0.5 to 0.75 cc of sediment were included in each experiment,




       28.  For N. arenaceodentata, routine rates of oxygen consump-




tion were measured using a 10-cc syringe (filled to 8 cc) containing




no sediment.  Rather, worms were offered 5-mm (outside diameter)




glass tubes inside which they would begin to form a mucus tube.




This glass tube could then be handled without any apparent effect




on the worms.  After 2 hr, the oxygen concentration of the seawater




was determined by injecting a 1-cc sample onto the oxygen probe.




With this species, the syringe was not refilled as the remaining




seawater (7 cc) was used to determine ammonia excretion rates.




       29.  Stender dishes containing approximately 25 ml of sedi-




ment were used to test the validity of oxygen consumption rates of
                                23

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  N. incisa obtained from the syringe respirometers.  A 35- by 50-mm

  stender dish with a hole drilled in the side was fitted with a

  Radiometer oxygen probe (Figure 5).  Declining oxygen tension in

  the dish was monitored using a strip chart recorder.  The water

  within the dish was stirred using a magnetic stir bar held to the

  top of the respirometer by a water-driven magnetic stirrer.  Control

  runs were made with the sediment alone to determine the amount of

  oxygen depletion due solely to the biological oxygen demand of the

  sediment.  The result of this test indicated that there was no

  significant difference in measuring respiration rate in syringes

  than  in  the larger  sediment-filled stender dishes (syringe:  1.59 +

  0.61  yl  02/mg/hr;   stender dish:   1.33 +  0.54 yl  02/mg/hr).  This

  experiment was repeated 6 times, with one stender dish respirometer

  and five syringe  respirometers per experimental run.



                              RESPIROMETERS
 TO 0
METER
RECORDER
                    CHAMBER
                                                        SYRINGE
                                  MAGNETIC STIRRER
                                — STIR BAR
                                        SEDIMENT
     Figure 5.  Schematic of respirometers (syringe and stender  dish)
                used to determine oxygen consumption rates of N. incisa
                juveniles
                                   24

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       30.  In all experiments oxygen consumption rates  are  reported




as weight-specific rates.  When reported,  the  energy expended  during




routine metabolism was considered to be the average oxygen consumption




rate (microliters per hour) during the experimental period times




the energy equivalent for oxygen (4.80 cal/ml; Elliot and Davison




1975).






Excretion




       31.  Following their use in the respiration tests, individual




NL_ incisa were placed in 6 ml of filtered seawater without sediment




for approximately 3 hr to determine ammonia excretion rates.  Follow-




ing this period of time, the dry weight of each worm was taken.




For N^ arenaceodentata,  the water for  ammonia determination was




taken  directly from the  syringe respirometers.




       32.  Ammonia concentrations were determined according  to  the




technique of  Bower and Holm-Hansen  (1980).  Calories lost per unit




time  as  excreted  ammonia were  calculated  by multiplying the excretion




rate  of  an  individual worm (micrograms of NH4N per hour) during  the




experimental  period by the heat  of  formation  of  ammonia (0.62 cal/mg;




Elliot and  Davison  1975).   Atomic ratios  of oxygen consumed to




ammonia  excreted (0:N ratios) were  calculated following the method




of  Corner and Cowey  (1968).






                         Burrowing Activity






        33.   Quantitative estimates  could not  be  made of the feeding




 rate for Nephtys incisa.  Since this species  is  thought to ingest
                                 25

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 sediment  in  its search  for  food  (i.e., burrow), it was felt that




 making quantitative measurements on burrowing activity could provide




 a  qualitative  estimate  of feeding  rates.  Burrowing activity of N_._




 incisa juveniles in the solid phase assay treatments was estimated




 using narrow,  glass-walled  aquaria.  The aquaria were 14 cm long by




 1  cm wide and  15 cm high (Figure 6; Davis 1979).  Individual aquaria




 were filled with homogenized mixtures of the appropriate sediment




 and allowed to equilibrate  in a flowing seawater bath.  Following




 this equilibration period (1 to 2  hr), two juvenile worms were




 randomly  placed on the sediment surface.  At the end of the 10-day




 exposure  period, plastic sheets were then laid against the glass




 walls of  the aquarium and the visible burrows traced.  Total burrow




 lengths were calculated from these tracings using a Numonics 1224




 electronic digitizer (Numonics Corp., Lansdale, N.J.).  In the




 particulate phase assays, a tracing was made of those burrows (for




 both N_^ incisa and N_._ arenaceodentata) visible on the side of the




 exposure  dish.




       34.  As the narrow aquaria  restrict worm movements to essen-




 tially two dimensions, data collected from these aquaria do not




 necessarily reflect the normal patterns and depths that might be




expected  in the field from the sizes of worms used.  They do, however,




 provide a relative index of the worm activity in the various treatment




conditions.
                                26

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                  16cm
                                   14cm
Figure 6.  Schematic of the narrow, glass-walled aquarium used to
           determine burrowing activity and maximum burrow depth
                        Statistical Analysis


        35.  In order to satisfy the objectives of this study, it was

 important to establish the variation expected from an individual

 worm when making physiological measurements as well as the degree

 of variation expected between individuals.  To establish this,

 repeated rate determinations (respiration or excretion rate) were

 made on a series of individual N_._ incisa.

        36.  Sources of variation in physiological rate measurements

 can include:  (a) temporal changes in the rate within an organism,

 (b) differences in rates between organisms, and  (c) the effects  of

 a particular treatment on the physiological rate.  Of prime  importance
                                 27

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in effects studies is an ability to detect significant changes in

the measured parameter (i.e., physiological rate) due to the treat-

ment.  Within-organism and between-organism variation, therefore,

must be quantified so that appropriate sample sizes needed to detect

a treatment effect can be determined.  The between-worm and within-

worm variation for the physiological measure was then determined

using the VARACOMP procedure (Goodnight 1979).

       37.  Based on these variations, sample size was then calcu-

lated using a general sample size formula (Snedecor and Cochran

1980).  The following sample size determination formula was used to

determine the minimum sample size needed:


                            n = C x (2S)/D                      (4)
where

    n = sample size

    C » constant which is a function of a type I and type II error

    S = estimate of variability from the pilot study

    D = percentage deviation from the mean that is to be
        detected (Snedecor and Cochran  1980)


       38.  Where appropriate within an assay (solid phase or par-

ticulate phase), data from a particular treatment from each repeated

experiment were combined for analysis.  The decision to combine

these data was based on the fact that within each assay there were

no significant differences in worm size or respiration or excretion

rates at the start of the experiment.  It can therefore be concluded

that there were no major physiological differences in the worms

from each experiment at the beginning of the 10-day assay.
                                28

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       39.  A one-way analysis of variance was computed to determine




the effects of the various treatments on the physiolgical function




of both polychaetes (Snedecor and Cochran 1980).  If significant




differences (at P = 0.05) were found, a Tukey and Kramer pairwise




comparison was used to determine where the differences existed




(Snedecor and Cochran 1980).  Data on burrow length and maximum




burrow depth were rank transformed prior to one-way analysis of




variance  (Conover and Iman 1981).
                                 29

-------
                          PART III:  RESULTS






                         Sources  of  Variation






 Respiration rate




        40.   In this  study,  the greatest  variance  found in  the




 respiration rates  of N.  incisa juveniles was  the  between-worm compo-




 nent  (Table 2).  Assuming that a 50 percent change in respiration




 rate  is biologically significant and  is  the percent change at which




 treatment effects  should be detectable,  a  sample  size of 10 was




 found to be sufficient based  on  a level  of confidence set at 90




 percent, using the between-worm  variation  as  the  estimate of varia-




 bility.  If a  25 percent change  in  respiration rate had been chosen




 as  the desired level, a  sample size of 22 would be needed to detect



 significant differences.






 Excretion rate




       41.   Using  the approach outlined  in paragraph 36, sources of




 variation were determined for  ammonia excretion rates in N. incisa.




 As with respiration  rates,  the between-worm component was the greatest




 source of variation  (Table  2).   Because of the low variation found




 in the between- and within-worm  variation for ammonia excretion




rates, only  3 worms would be needed to detect significant differences




at the 90 percent  confidence level.  However, ten worms were used




in order to provide a complementary set of measurements to the




respiration rate determinations.
                                30

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                                   Table 2


       Sources of variation in respiration and ammonia excretion rate

         of N. "incisa juveniles determined during initial experiments



                            Respiration Rate*  N**   Excretion Rate* H**
Within Worm Variance
(Repeated measurements)
Between Worm Variance
0.166
0.340
25       0.00000391   25
25       0.00000624   25
 * Variance.
** N= Number of Determinations.
                            Treatment  of  Data


   Nephtys incisa

          42.   Within an assay,  no significant differences were found

   in worm size and respiration  and excretion rates  of  N.  incisa at  the

   start of the experiment.   Therefore,data from each individual exper-

   iment (within an assay) were  combined  for analysis.   Summary data

   from each individual experiment can be found in Appendix A.


   Neanthes arenaceodentata

          43.   Unlike N. incisa, there were significant differences in

   worm size at the beginning of each  experiment within an assay.  For

   this reason, data from the various  experiments within the particular

   assay were  analyzed separately.
                                   31

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                            Consumption






       44.  Food consumption rates of N. arenaceodentata juveniles




exposed to  100-percent reference sediment for 10 days were signif-




icantly higher than those exposed to 100-percent BRH sediment (Table




3).  Feeding rates of those worms exposed to the 50:50 mixture of




reference and BRH sediments were not significantly different from




those of the control group although they were significantly higher




than those worms exposed to 100-percent BRH.




       45.  Juvenile N. arenaceodentata exposed in the particulate




phase assays exhibited similar food ingestion patterns to those




found in the solid phase assays.  In experiment I, consumption




rates of worms from the reference/reference (particulate phase/solid




phase) and reference/BRH treatments were significantly higher than




those rates for worms exposed in either the BRH/reference or BRH/BRH




regimes (Table 3).  In experiment II, there were significantly




higher food consumption rates in juveniles from the reference/refer-




ence and BRH/reference treatments compared to those from the BRH/BRH




exposure (Table 3).  There was no difference, however, between refer-




ence/BRH and BRH/BRH ingestion rates as well as no significant




difference between reference/BRH and both reference/reference and




BRH/reference treatments (Table 3).
                                32

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                                  Table 3


          Food consumption rates of N. arenaceodentata juveniles
exposed to various sediment conditions

Experiment Treatment
I 100% REF*
50% REF/ 50% BRH
100% BRH
I REF/REF
REF/ BRH
BRH/ REF
BRH/ BRH
II REF/REF
REF/BRH
BRH/ REF
BRH/ BRH
Consumption Rates**
mg/24 hr
Solid Phase Assay
4.49 + 0.68
4.34 + 0.72
3.52 + 0.85
Particulate Phase Assay
4.49 + 0.96
4.63 + 1.58
3.31 + 1.23
3.38 + 1.23
6.65 + 0.92
6.35 + 1.14
6.62 + 1.36
5.54 + 1.00
Nt Gptt
5 A
5 A
5 B
10 A
10 A
10 B
10 B
10 A
10 A,B
10 A
10 B
 * Ref =• Reference Sediment
** Means + S.D.
 t N - Number of Determinations
tt Gp - Grouping Letter.  Means having the same Gp are not
   significantly different at p - 0.05.
                                    33

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                             Production






Nephtys incisa



       46.  Exposure to 100-percent BRH sediment for 10 days had a




significant effect on the growth of juvenile Nephtys incisa (Table




4).  Worms maintained in 100-percent reference and 50-percent refer-




ence/50-percent BRH sediment conditions gained an average dry weight




of 24 and 9 percent, respectively, during the 10-day exposure,




while those worms in the 100-percent BRH treatment lost an average




of 16 percent of their dry weight during the same period.




       47.  Juvenile N^ incisa exposed in the particulate phase




assays exhibited similar changes in dry weight (Table 4).  In both




experiments conducted with these conditions, worms from the reference/




reference treatments (particulate phase/solid phase) were significantly




heavier at the end of the 10-day exposure than those exposed to



BRH/BRH.  Worms from the BRH/reference treatments were not signifi-




cantly different from the reference/reference worms.
                                 34

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                                Table 4


        Changes in dry weight of N.  incisa juveniles exposed to

                      various sediment conditions
                                                              Change In
                                                                Weight
                                                          Gpt   mg   %
                               Dry Weight**
                        Initial        Final
Treatment                 mg             mg


                             Solid Phase Assay
        100% REF*          1.411+0.246   1.743+0.204   A   +.33 ;  +24

        50% REF/50% BRH    1.411 + 0.246   1.543 + 0.389   A   +.13 ;  + 9

        100% BRH           1.411+0.246   1.189+0.105   B   -.22;  -16



                                Particulate Phase Assay
REF/REF
REF /BRH
BRH/REF
BRH/BRH
3
3
3
3
.277
.277
.277
.277
+
+
+
+
0.762
0.762
0.762
0.762
4
3
3
3
.173
.742
.850
.228
+ 0
+ 0
+ 0
+ 0
.535
.755
.723
.578
A
A
A
B
+.95
+ .47
+.57
-.05
; +29
; +14
; +18
; -2
 * Ref - Reference Sediment
** Means +1 S.D.
 t Gp - Grouping letter.  Means having the same Gp are not significantly
   different at p - 0.05
                                   35

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Neanthes arenaceodentata




       48.  Exposure to 100-percent BRH sediment did not significantly




alter growth of N_. arenaceodentata during the 10-rday exposure period




(Table 5) although the trend was for the mean dry weight of the 100-




percent BRH-exposed worms to be lower than for the worms exposed in




the reference sediment.




       49.  For experiment I of the particulate phase assay, there




was no significant difference in worm dry weights at the end of a




10-day exposure to the various sediment combinations.  In experiment




II, however, growth of those N. arenaceodentata Juveniles exposed




to the BRH/BRH conditions was significantly lower than exhibited by




worms from the other conditions (Table 5).
                                 36

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                               Table  5
     Changes in dry weight  of  N.  arenaceodentanta juveniles exposed
to various sediment

Experiment Treatment




Dry V
Initial
mg
Solid Pt)
I 100% REF*
50% REF/50% BRH
100% BRH
4.
4.
4.
16 +
16 +
16 +
1
1
1
.73
.73
.73
Particulate
I REF/REF
REF /BRH
BRH/REF
BRH/BRH
II REF/REF
REF /BRH
BRH/REF
BRH/BRH
3.
3.
3.
3.
4.
4.
4.
4.
52 +
52 +
52 +
52 +
43 +
43 +
43 +
43 +
2
2
2
2
0
0
0
0
.09
.09
.09
.09
.92
.92
.92
.92
conditions




/eight**
Final
mg Gp*
iase Assay
6.44
5.03
5.16
Phase
5.41
5.98
5.87
5.35
6.51
6.27
6.26
4.64
+ 1
+ 1
+ 1
.69
.21
.08
A
A
A



Change In
Weight
mg %
+2
+0
+1
.28
.87
.00
+55
+21
+24
Assay
+ 1
+ 1
+ 1
+ 1
+ 1
+ 1
+ 1
+ 0
.80
.47
.95
.58
.91
.93
.53
.88
A
A
A
A
A
A
A
B
+1
+2
+2
+1
+2
+1
+1
+0
.89
.46
.35
.83
.08
.84
.83
.21
+54
+70
+67
+52
+47
+42
+41
+5
 * REF - Reference Sediment.
** Means +1 S.D.
 t Gp - Grouping letter. Means having the same Gp are not
   significantly different at p - 0.05.
                                   37

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                         Maintenance Costs






Respiratory expenditures




       50.  Nephtys incisa.  Weight-specific respiration rates of




N^ incisa juveniles exposed to 100-percent reference sediment were




significantly higher than those of juvenile worms exposed to 100-




percent BRH sediment (Table 6).  Worms from the 50-percent reference/




50-percent BRH treatment exhibited oxygen consumption rates inter-




mediate to those from the 100-percent reference and 100-percent BRH




conditions.




       51.  A similar pattern was found for those worms from the




particulate phase experiments.  In this assay, N. incisa juveniles




from the reference/reference treatment had significantly higher




respiration rates than those worms from the BRH/BRH exposure (Table




6).  Respiration rates of worms from the BRH/reference treatment




were also significantly higher than those of the BRH/BRH-treated




worms but were not statistically different from the reference/refer-




ence individuals.  Worms from the reference/BRH treatment exhibited




oxygen consumption rates similar to those found in the BRH/BRH-




treated worms (Table 6).




     52.  Neanthes arenaceodentata.  No significant differences in




weight-specific respiration rate were found in N. arenaceodentata




exposed to the various sediment treatments (Table 7).
                                38

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                               Table 6


       Weight-specific respiration rate of  N.  incisa juveniles

                exposed to various sediment conditions
   Treatment
Respiration Rate**
  yd 07/mg/hr
N1
Gp
                                                       TT
Solid Phase Assay
100% REF*
50% REF/ 50% BRH
100% BRH

REF/REF
REF/BRH
BRH/REF
BRH/BRH
1.65 + 0.77
1.36 + 0.63
1.10 + 0.73
Particulate
0.75 + 0.20
0.62 + 0.25
0.86 + 0.25
0.66 + 0.28
62 A
56 A,B
44 B
Phase Assay
42 A
42 B
42 A
44 B
 * REF » Reference Sediment.
** Means + 1 S.D.
 tN= Number of determinations.
tt Gp - Grouping Letter. Means having the same Gp are not
    significantly different.
                                   39

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                                 Table  7
     Weight-specific respiration rate of N. arenaceodentata juveniles
exposed

Experiment Treatment
I 100% REF*
50% REF/ 50%
100% BRH
I REF/REF
REF/ BRH
BRH/REF
BRH/ BRH
II REF/REF
REF/ BRH
BRH/REF
BRH/BRH
to various sediment conditions

Respiration Rate**
11 «. 0,/mg/hr NT
Solid Phase Assay
1.23+0.17 4
BRH 1.18+0.27 5
0.94+0.19 7
Particulate Phase Assay
2.30+0.61 10
2.23 + 0.65 10
2.05 + 0.70 10
2.57 + 0.74 10
1.74 + 0.41 10
1.93+0.50 10
1.81 + 0.31 10
2.16 + 0.37 9

GP"
A
A
A
A
A
A
A
A
A
A
A
 * REF - Reference Sediment.
** Means + 1 S.D.
 t N - Number of Determinatons.
ft Gp » Grouping letter.  Means having the same Gp are not
   significantly different.
                                     40

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 Ammonia excretion rates and 0;N ratios




       53.  Nephtys incisa.  As with weight-specific respiration




rates, weight-specific ammonia excretion rates of N. incisa juveniles




exposed to 100-percent BRH and BRH/BRH conditions were significantly




lower than those rates of worms exposed to 100-percent reference




and reference/reference (Tables 8).  The other treatments were




intermediate between these two values.




       54.  No significant differences were found in 0:N ratios




between the various treatments in either the solid phase or the




particulate phase assay (Table 8).  In all cases, 0:N ratios were




relatively high (greater than 50) indicating that lipids and carbo-




hydrates are being used as part of the substrate for energy produc-




tion.




       55.  Neanthes arenaceodentata.  For two of the three experi-




ments conducted with N_._ arenaceodentata  (solid phase experiment and




experiment I, particulate  phase), no  significant differences in




weight-specific ammonia excretion rates  were  found  (Table  9).   In




the other particulate  phase  experiment,  however, a  significant




increase in ammonia excretion  rate was detected  in  those worms




exposed to either  the  BRH/BRH  or BRH/reference treatment.




       56.  As with N_-_ incisa  no significant  differences were  found




in 0:N ratios between  the  various  experiments in either assay.   The




0:N ratios in all  treatments ranged  between  84 and  229  (Table  9).
                                 41

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                                 Table 8
   Weight-specific ammonia excretion rate and OtN ratios of N. incisa

Treatment
100% REF*
50% REF/ 50%
100% BRH
REF/ REF
REF/ BRH
BRH/ REF
BRH/ BRH
juveniles exposed to various sediment

Excretion Rate**
v$ NH4N/mg/hr Nt Gptt
Solid Phase Assay
0.011 + 0.003 62 A
BRH 0.010 + 0.002 56 A,B
0.008 + 0.004 44 B
Particulate Phase Assay
0.012 + 0.003 42 A
0.010 + 0.002 42 B
0.013 + 0.004 42 A
0.010 + 0.004 44 B
conditions

0:N
Ratio**
147 + 69
121 + 56
98 + 60
67 + 18
55 + 22
77 + 22
59 + 25

A
A
A
A
A
A
A
 * REF - Reference Sediment.
** Means + 1 S.D.
 t N « Number of determinations, all three experiments combined.
Tt Gp - Grouping letter.  Means with the same Gp are not
   significantly different.
                                     42

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                                  Table 9


Weight-specific ammonia excretion rate and OtN ratios of N. arenaceodentata
juveniles
exposed to various sediment conditions

Experiment Treatment
I 100% REF*
50% REF/ 50%
100% BRH
I REF/ REF
REF/ BRH
BRH/ REF
BRH/ BRH
II REF/REF
REF/BRH
BRH/REF
BRH/BRH
Excertion Rate**
ug NH4N/mg/hr Nt
Solid Phase Assay
0.0045 + 0.001 4
BRH 0.0034 + 0.001 5
0.0050 + 0.001 7
Particulate Phase Assay
0.010 + 0.007 10
0.011 + 0.004 9
0.015 + 0.007 8
0.013 + 0.007 7
0.008 + 0.002 9
0.008 + 0.004 10
0.013 + 0.003 10
0.014 + 0.004 9
0:N
Gptt Ratio** Gptt
A 110 + 21 A
A 105 + 17 A
A 84 + 25 A
A 205 + 55 A
A 199 + 58 A
A 183 + 62 A
A 229 + 66 A
B 155 + 37 A
B 172 + 45 A
A 161 + 32 A
A 193 + 42 A
  * REF • Reference sediment.
 ** Means +  1 S.D.
  t N - Number of determinations.
 tt Gp - Grouping letter.  Means with  the same  Gp  are  not
    significantly different.
                                       43

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                  Partitioning of Energy Resouces






Nephtys incisa




       57.  The preceding physiological data were used to calculate




the efficiency at which available energy (P+R+E) was partitioned




between growth and maintenance (Respiratory energy expenditure and




Excretory energy losses).  In both the solid phase and particulate




phase experiments, maintenance costs were higher in the reference




treatments (100-percent reference;  reference/reference) than they




were in the Black Rock Harbor treatments (100-percent BRH; BRH/BRH)




(Table 10).  Estimated maintenance costs for the worms from the




other treatments were generally intermediate between these two




extremes.




       58.  Energy losses through ammonia excretion were not impor-




tant when compared to other energy costs involved in maintenance




(i.e., respiratory expenditures; Table 10).  However small, there




were significant differences in energy lost when the excretion




rates of worms maintained in 100-percent reference and reference/




reference were compared to 100-percent BRH and BRH/BRH.

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                                   Table 10


        Cummulative energy values for production and maintenance costs

                            of N. incisa juveniles


                                     Respiratory              Excretory
                     Growth**       Energy Expenditure**     Energy Loss**
Treatment           J^	Gptt   _jt	Gptt          jt	Gptt


                               Solid Phase Assay


100% REF*         4.85 + 0.97  A   12.6 + 3.8   A        0.035 + 0.009    A


50% REF/50% BRH   2.01 + 0.48  B    8.9 + 3.6   A,B      0.029 + 0.008   A,B


100% BRH         -2.30+0.36  C    5.6+2.1   B        0.020+0.006      B


                             Particulate Phase Assay


REF/REF           9.77 + 1.86  A   14.1 + 4.7   A        0.077 + 0.018      A

REF/BRH           4.62 + 0.92  B   11.2 + 3.0   B        0.064 + 0.018      B

BRH/REF           6.44 + 1.42  A   15.8 + 5.0   A        0.084 + 0.027      A

BRH/BRH          -1.73+0.52  C    9.8+3.1   B        0.054+0.012      B
 * REF » Reference Sediment.
** Means +  1 S.D.
 t J » Joules.
tt Gp » Grouping letter.  Means having  the  same  Gp are  not
   significantly different at p -  0.05.
                                      45

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       59.  Comparison of net growth efficiencies of  N.  incisa

juveniles from the different treatments showed that worms maintained

in 100-percent reference sediment were most efficient (+27  percent)

at converting available energy to new tissue (Table 11;  Figure  7).

Worms maintained in 100-percent BRH were the least efficient  (-24

percent) and in fact lost weight during the exposure  period.  In

the 50-percent reference/50-percent BRH treatment, worms converted

energy to tissue with an efficiency of +17 percent, which was not

statistically different from the efficiency for worms maintained in

the 100-percent reference condition.

            60-i
            40-
         I  20-
         UJ
         S+
         u_
         LU
              0-
         o
         (E
         UJ
            20-
            40-
            60J
                       0%            50%
                              % BRH SEDIMENT
                                                   100%
Figure 7.
             Net growth efficiency of  N.  incisa juveniles  exposed
             to  various mixtures  of reference  and  BRH sediment  for
             10  days.

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                                    Table 11


              Net Growth Efficiency of N. incisa juveniles exposed

                         to various sediment conditions
                                        Net Growth
             Treatment                  Efficiency**
                              Solid Phase Assay


             100% REF*                   +27 + 9         A

             50% REF/ 50% BRH             +17 + 9         A

             100% BRH                    -24 +30        B


                        Particulate Phase Assay

              REF/REF                    +39 +10        A

              REF/BRH                    +32 +16        A

              BRH/REF                    +27 +7         A

              BRH/ BRH                    -28 +40        B
 * REF - Reference Sediment.
** Means +1 S.D.
 t Gp - Grouping letter.  Means having the same  Gp  are  not
   significantly different at p - 0.05.
                                       47

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       60.  A similar pattern of net growth efficiency was found in



the particulate phase assays.  N. incisa juveniles maintained under



reference/reference conditions exhibited the highest net growth



efficiency (+39 percent) while those in the BRH/BRH treatment had a



negative net growth efficiency (-28 percent) (Table 11; Figure 8).



Net growth efficiencies of worms kept in reference/BRH and BRH/refer-



ence exposures were +32 and +27 percent, respectively.  These values



were not significantly different from those of the worms in the



reference/reference conditions.




              60-
           o

           g  20

           o
           -
           lt.
           UJ
               0-
          O
          cc.
          UJ
              40-
              60J
                      con/con  con/BRH  BRH/con  BRH/BRH


                           SEDIMENT  COMBINATION


                            (particulate/ sediment)



 Figure 8.  Net growth efficiency of  N^ incisa juveniles exposed to

            various combinations of sediment and suspended particles

            for 10 days
                                48

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Neanthes arenaceodentata




       61.  The appropriate data were also used to calculate cummula-




tive energy values and net growth efficiency for N. arenaceodentata.




As can be seen in Table 12, a great amount of variance was connected




with estimating tissue production during the 10-day experimental




period.  As with Nephtys incisa, energy losses through ammonia




excretion were not important when compared to energy costs involved




in other maintenance processes  (respiratory energy expenditure;




Table 12).




       62.  The large variation in production in turn leads to a




large variation in calculated net growth efficiencies (Table 13).




The mean net growth efficiencies of the worms maintained in the




reference sediment, however, are in some agreement with those found




for N. incisa.  In experiment II of the particulate phase assay,




the pattern of net growth efficiency is similar to that of N^




incisa.  In this experiment, the net growth efficiencies of worms




exposed to the BRH/BRH treatment was significantly lower than it




was for worms in the other treatments.
                                 49

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                                Table  12


         Cummulative energy values for  production and maintenance

         costs of N. arenaceodentata juveniles exposed to various

                          sediment conditions
                                         Respiratory          Excretory
                           Growth**      Energy Expenditure** Energy Loss**
Experiment  Treatment      .P	Gp"     jt	Gptt       jt	Gp^
                                   Solid Phase Assay

  I        100% REF*       22+20   A     38+9    A     0.057 + 0.016  A

           50% REF/50% BRH 14+19   A     30 + 11   A,B   0.028 + 0.001  B

           100% BRH        13+14   A     23+3   B     0.041 + 0.008  A


                                  Particulate Phase Assay
I REF/REF
REF /BRH
BRH/REF
BRH/BRH
II REF/REF
REF/BRH
BRH/REF
BRH/BRH
39
45
49
37
36
38
38
3
+ 38
+ 27
+ 40
+ 32
+ 33
+ 39
+ 31
± 14
A
A
A
A
A
A
A
B
57 +
61 +
54 +
62 +
52 +
56 +
53 +
47 +
13
8
13
7
6
10
6
9
A
A
A
A
A
A
A
A
0
0
0
0
0
0
0
0
.081
.108
.140
.126
.080
.083
.140
.113
+
+
+
+
+
+
+
+
0.040
0.027
0.052
.069
0.021
0.028
0.030
0.029
A
A
A
A
A
A
A
A
 * REF - Reference Sediment.
** Means +1 S.D.
 t J  - Joules.
tt Gp - Grouping letter.  Means with the same Gp are not significantly
        different.
                                   50

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                                  Table 13


       Net growth efficiency of N. arenaceodentata juveniles exposed

                       to various sediment conditions
                                           Net Growth
     Experiment     Treatment               Efficiency** Gpt
                               Solid Phase Assay

                     100% REF*             +29 +34       A

                     50Z REP/50Z BRH       +18+42       A

                     100% BRH              +27 + 32       A
                         Particulate Phase Assay
I



II



REF/REF
REF/BRH
BRH/REF
BRH/BRH
REF/REF
REF/BRH
BRH/REF
BRH/BRH
+30 + 27
+39 + 16
+41 + 20
+31 + 23
+35 + 19
+34 + 24
+35 + 20
+ 3 + 23
A
A
A
A
A
A
A
B
 * REF « Reference Sediment.
** Means + 1 S.D.
 t Gp • Grouping letter.  Means having the same Gp are not
   significantly different at p - 0.05.
                                       51

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                          Scope for Growth






       63.  In order to calculate scope for growth (SFG) values,  the




energy content of the food offered to N. arenaceodentata had to be




determined.  In addition, the effeciency at which ingested food was




assimilated had to be estimated.  The prawn flake used as a food




source in these experiments was found to have an energy value of




18.53 + .471 J/mg while the assimilation efficiency was estimated to




be between 20 and 27 percent.




       64.  In all experiments, the SFG following a 10-day exposure




to BRH sediment (100-percent BRH or BRH/BRH treatments) was signifi-




cantly reduced (Table 14).  The SFG values from these treatments




represent a 27- to 65-percent reduction in the growth potential




compared to SFG values for worms from the reference sediment condi-




tions (100-percent reference and reference/reference).
                                 52

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                               Table 14


           Scope for growth for N. arenaceodentata juveniles

                      exposed to various sediments
                                               Scope for Growth**
Experiment            Treatment                    J/Day    Gpt
                             Solid Phase Assay


                    100% REF*                 12.1 +  2.5     A

                    50% REF/50% BRH           10.8 +  1.4   A,B

                    100% BRH                    8.8 +  2.5     B


                       Particulate Phase Assay


II
REF/REF
REF/BRH
BRH/REF
BRH/BRH
I REF/REF
REF/BRH
BRH/REF
BRH/BRH
7.2 + 2.1
6.8 + 3.1
2.6 + 2.1
2.5 + 2.9
17.8 + 3.1
16.8 + 4.0
17.6 + 4.2
10.1 + 2.9
A
A
B
B
A
A
A
B
 * REF - Reference  Sediment.
** Means +  1  S.D.
 t Gp - Grouping letter.  Means  having the  same Gp are not
    significantly different at p - 0.05.
                                    53

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                        Burrowing Activity


Nephtys incisa

       65.  Both burrow length and maximum depth of burrowing in the

100-percent reference treatment were significantly different  from

similar measurements taken from the 100-percent BRH exposure  (Table

15).  The total length of visible burrows was approximately three

times greater in the 100-percent reference condition than it  was in

100-percent BRH (Figure 9).  The depth to which the worms burrowed

was three times greater in 100-percent reference sediment (30.4  mm)

when compared to the maximum burrow depth in 100-percent BRH  (9.5

mm) .

                   SOUTH REFERENCE SEDIMENT
                       Burrow Length  = 42.2 ±9.9  cm
               Maximum Burrow Depth  = 30.417.3  mm
                  BLACK ROCK HARBOR  SEDIMENT
Figure 9.
           Burrow Length = 14.2 + 4.7 cm
    Maximum Burrow Depth =  9.5 ±2.3 mm

Drawing of burrows created by N. incisa juveniles  exposed
to reference  and BRH sediments for 10 days
                                54

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                                Table 15


 Burrow length and maximum burrow depth of N. inclsa juveniles exposed

                     to various sediment conditions
    Treatment
Burrow Length**   Percent in  Maximum Burrow Depth**
    cm       Gptt  Sediment           mm	Gptt

100% REF*
50% REF/ 50% BRH
100% BRH

REF/REF
REF/ BRH
BRH/ REF
BRH/ BRH
Solid Phase Assay
42.2 + 9.9 A - 30.4 + 7.3
N.D.t - N.D.
14.2 + 4.7 B - 9.5 + 2.3
Particulate Phase Assay
43.1 + 8.9 A 58.7 29.3 + 2.0
21.8 + 1.1 B 40.8 17.0 + 2.4
36.9+6.2 A 77.0 31.2+6.1
9.3 + 0.8 C 31.3 16.8 + 2.5

A

B

A
B
A
B
 * REF - Reference Sediment.
** Means +1 S.D.
 t N.D. - No data.
tt Gp - Grouping letter.  Means with the same Gp are not significantly
   different at p - 0.05.
                                    55

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         66.   Results of measurements of burrowing activity  in the

  particulate phase experiments were similar to  the  preceding results,

  although somewhat complicated by the fact that  suspended particles

  settled out during the course of the experiment to form a  layer of

  sediment over the bedded sediment.  Nephtys incisa from the reference/

  reference condition produced significantly more burrows and burrowed

  to  a  significantly greater depth than did those worms maintained in

  BRH/BRH (Table 15;  Figure 10).
                        PARTICULATE/SOLID  PHASE
           control/control
      burrow length =43.1 ±8.9
      BRH/control
burrow length = 36.9 ±6.2
           control/BRH
      burrow length = 21.8± I.I
      BRH/BRH
burrow  length = 9.310.8
Figure 10.  Drawing of burrows created by N_._ incisa juveniles  exposed
            to various combinations of reference and BRH sediment
            for 10 days
                                  56

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       67.  Burrowing activity in the other two particulate phase




treatments depended on whether the reference sediment was in the




upper particulate layer or was present as the original,  bedded




sediment.  In the BRH/reference treatment, most of the burrowing




activity was concentrated in the original sediment (77 percent of




activity) and the depth of burrowing was not significantly different




from that found in the reference/reference treatment (Table 15;




Figure 10).  In the reference/BRH exposure, on the other hand, most




of the burrowing activity (59 percent) was found in the particulate




layer which formed during the 10-day exposure.  Maximum burrow




depth in this treatment was only 17 mm, which is very similar to




the maximum burrow depth found in the BRH/BRH treatment.






Neanthes arenaceodentata




       68.  Neanthes arenaceodentata juveniles exhibited significantly




greater burrowing activity (both burrow length and maximum depth of




burrowing) in those treatments that had reference sediment as the




bedded sediment than in those worms which had BRH as the bedded




sediment (Table 16; Figure 11).




       69.  In all treatments, more than half of the burrowing activity




occurred in the upper particulate layer which formed during the




10-day exposure, regardless of the type of sediment used in the




suspended particulate phase.
                                57

-------
                         PARTICULATE/SOLID PHASE
          control/control
    burrow length = 39.7 ±9.6
                                                       BRH/control
 burrow length = 4 1.3 ± 9.8
          control/BRH
         BRH/BRH
    burrow length = 14.9 ±6.8
burrow length - 20.2 ±1.0
Figure  11.  Drawing of burrows created  by N. arenaceodentata  juveniles
             exposed to various combinations of reference and  BRH
             sediment for 10  days
                                     58

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                                 Table 16
       Burrow length and maximum burrow depth of N. arenaceodentata
juveniles exposed to various sediment conditions

Treatment

REF/REF*
REF/BRH
BRH/REF
BRH/BRH
Burrow Length** Percent in Maximum Burrow Depth**
cm Gpt Sediment mm Gpt
Particulate Phase Assay
39.7 + 9.6 A 49 26.8+11.3 A
14.9 + 6.8 B 24 17.0+ 4.1 B
41.3 + 9.8 A 32 34.2 + 4.0 A
20.2 + 1.0 B 37 17.0 + 9.2 B
 * REF • Reference Sediment.
** Means + 1 S.D.
 t Gp » Grouping letter.  Means having the same Gp  letter are not
   significantly different at p -  0.05.
                                    59

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                        PART IV: DISCUSSION






       70.  Nephtys incisa is a nonselective deposit feeder and is




typically found in soft sediments.  It does not build permanent




tubes, but rather burrows indiscriminately, ingesting sediment and




associated microorganisms as a food source (Davis 1979).  This




species is considered a member of the equilibrium assemblage in




Long Island Sound (Rhoads and Germane 1982), and is usually associated




with sediments that are oxygenated to depths of up to 10 cm.  The




physical effects of the errant burrowing behavior of N^ incisa on




local sediments are the provision of vertical particle mixing and




the enhancement of pore water exchange (Rhoads and Germane* 1982).




Much of the burrowing activity occurs at the redox boundary, a zone




typically high in microorganism productivity (Yingst and Rhoads




1980).




       71.  The data presented here for the reference sediment




treatments indicate that N^ incisa juveniles perform much the same




organism-sediment roles as do the adult stages.  Burrowing activity




during the 10-day experimental period was extensive, with burrows




penetrating up to 30 mm in depth.  Respiratory pumping also appeared




to be effective in pore water exchange, as the burrow tubes had an




observable 'halo' of light sediment which is an indicator of oxidized




sediment (Aller and Yingst 1978).




       72.  The physiological measurements determined with Nephtys




incisa juveniles were reproducible and repeatable.  (Review Appendix




A for data from each individual experiment.) Within an experiment,
                                 60

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each response measured (growth,  respiration and ammonia excretion




rate, and burrowing activity) exhibited standard deviations  about




the mean value that were less than 28 percent.  This is an indication




that the response measurements are reproducible, providing the measure-




ment technique is followed properly.  To adequately measure growth,




for example, care must be taken at the beginning of the experiment




to size the worms.  Failure to control the variance in initial  dry




weight will reduce the chances (statistical) of being able to detect




growth differences among the various treatments at the end of a




10-day experiment.




       73.  Almost all of the biological endpoints evaluated with




Nephtys incisa followed a dose-response to additions of BRH sediment.




These responses were repeatable from one experiment to another.  The




only exception to this was the calculated 0:N ratios which did not




exhibit a  response across the treatments used.




       74.  Net growth efficiencies for N. incisa  Juveniles main-




tained in  control sediments  are within the range of efficiencies




reported for  polychaetes under a  variety of conditions.   For instance,




Carey (1962)  estimated population production  and respiration values




for  N._ incisa from a Long Island  Sound study  site. Using data




from his Table 9, a mean net growth efficiency  of  36  percent is




calculated (population production/population  production + respiration),




Net  growth efficiency  for omnivorous  polychaetes  (both indiscriminate




sediment ingesters, as well  as  surface detrital feeders)  has been




found to be generally  between 14  and  40 percent (Kay  and  Brafield




 1973; Tenore  and  Gopalan  1974;  Neuhoff 1979).
                                 61

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        75.  The effects of exposure to BRH sediment are twofold.




First,  N. incisa juveniles greatly reduce their burrowing activity




when exposed to whole BRH sediment.  This was graphically demonstrated




in Figures 9 and 10, where worms maintained in BRH sediment for 10




days demonstrated almost no burrowing activity and no depth penetra-




tion.   Similar results were found in the lugworm, Arenlcola cristata,




exposed to kepone-contaminated sediments (Rubinstein 1979).




        76.  In the suspended particulate assays, where a choice




between sediments was offered as settling of the particulate dose




occurred, N. incisa appeared to avoid BRH sediment.  Comparison of




the reference/BRH treatment to the BRH/reference treatment illustrates




this point.  Almost all of the burrowing activity in the reference/BRH




treatment was in the upper settled layer.  This activity occurred




only after the sediment layer was formed.  Worms from the BRH/refer-




ence exposure exhibited an opposite activity level.  In this treat-




ment, there was extensive reworking of the lower, reference sediment,




while there was very little activity in the upper, sedimented BRH




material.  The only burrows present in this upper layer were a few,




fairly vertical ones to the surface (when compared to reference/




reference and reference/BRH).  These burrows probably provide the




worms with a supply of oxygenated water to meet their respiratory




requirements.




       77.  The dramatic decrease in burrowing activity of N^ incisa




juveniles that occurred during exposure to BRH indicates that the




worms are probably curtailing physiological functions.  If the




primary energy source of this species is ingested sediment, then
                                 62

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curtailing burrowing activity has the effect of  starving  the  indi-




vidual.  This is, in essence, the second effect  BRH sediment  has on




the species.  This effect is brought about by the failure of  the




worms to consume a sufficient amount of energy for normal physio-




logical and biochemical processes.  Nephtys incisa exposed to BRH




sediments generally exhibited lowered maintenance costs,  most




probably in response to the reduced energy intake and reduced




burrowing activity.  Despite these lowered costs, individuals in




the BRH sediment had to catabolize some tissue in order to provide




the energy requirements of routine metabolism.  Catabolism of tissue




to meet energy requirements is a common phenomenon in organisms




that are below their required maintenance ration (Pandian 1975).




       78.  In those treatments where there was a choice between BRH




and reference sediments, however, there was an increase in burrowing




activity (with a majority of the activity occurring in the reference




sediment; Fig. 10).  The increase in activity noted in the BRH/




reference and reference/BRH sediments is apparently sufficient to




provide enough energy for both maintenance  costs and growth.  Worms




under  these sediment conditions had  lower,  but not significantly




different, net growth efficiencies  than did worms maintained in the




reference sediment  (Table  11).




       79.  In contrast  to N^ incisa, N^_ arenaceodentata  is  primarily




a surficial deposit feeder that  builds  somewhat  permanent mucus




tubes  near  the sediment  surface.  Unlike N^ incisa, N^ arenaceodentata




will leave  their  tubes to search  the sediment surface  for food.




Despite  these differences  in feeding habits,  N^ arenaceodentata
                                 63

-------
 juveniles exhibited burrowing patterns similar to  those of  N^ incisa.




 Burrowing activity  was  reduced in  participate  phase  assays  in which




 BRH sediment was the bedded sediment.   In addition,  depth of  burrowing




 was also  shallower  in those treatments where BRH sediment existed




 as the bedded sediment.




        80.  The physiological responses of N.  arenaceodentata to




 exposure  to  BRH sediment,  however, were not as dramatic as  they




 were with N. incisa.  In only one  of the three experiments  conducted




 with ^ arenaceodentata was there  a significant  difference  in dry




 weight at the end of  the 10-day exposure.   In  addition,  no  significant




 differences  in  respiration rates were  found in any of the various




 experiments.  Food  consumption rates,  however, were  found to  be




 affected,  with  lower  rates found in those  worms  exposed  to  BRH




 sediment.  The  general lack of physiological response by N_._




 arenaceodentata  to  BRH sediment may have been  due to several  factors.




        81.   One  is  simply  that exposure to BRH sediment  does  not




 cause  any  change in the growth rate or  physiolgical rate  of this




 species.   Unlike N^ incisa,  which appears  to curtail burrow/feeding




 activity in BRH sediment,  N.  arenaceodentata feeds on detrital




 material found on the sediment surface  and thus  can avoid ingesting




 BRH sediment under  some conditions.  The conclusion that BRH sediment




 does not have a  physiological  effect on N_._ arenaceodentata, however,




is not probable since food consumption rates were affected by exposure




 to BRH sediment.




       82.   It is also possible that the mucus tube formed by this




species may reduce its contact with surrounding sediments.  This,
                                64

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in turn, would reduce its exposure to contaminants that are found




in BRH sediment.  Although not specifically tested for in this




study, it would be interesting to determine what degree of protection




from sediment contaminants is offered by mucus tubes.




       83.  Another explanation, however, for the lack of any measur-




able growth response to BRH sediment is that the variance in worm




sizes at the beginning of an experiment was so great that any changes




in growth rate resulting from the treatment exposure could not be




detected after the 10-day exposure period.  With N_._ incisa the




coefficient of variation in dry weight at the beginning of an exper-




iment was 24 percent, while with N. arenaceodentata the coefficient




of variation in worm size was quite large in the first two experiments




(42 and 59 percent, respectively).  This degree of variation made




it difficult at best to measure growth in as short a time period




(10 days) as used in this study.  In the third experiment (particulate




phase, experiment III), on the other hand, where significantly




lowered growth was detected in those worms from the BRH/BRH condition,




the coefficient of variation for dry weight at the start of the




experiment was only 21 percent.  While it is not known whether the




relationship between low variation in worm size at the beginning of




an experiment and the ability to detect changes in growth after 10




days is causal or casual, more time will be spent in future experiments




with N. arenaceodentata in selecting worms for experimentation.




       84.  The net growth efficiencies calculated in this study for




N. arenaceodentata were within the range of values reported in the




literature for detrital feeding polychaetes (Kay and Brafield 1973;
                                65

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 Neuhoff 1979).   The large  variation  found  about  the mean net growth




 efficiency  in all  of these experiments was probably largely due to




 poor  control in  worm size  at  the  beginning of  the  experiments.  It




 is  interesting to  note  that the net  growth efficiencies of both N.




 arenaceodentata  and N.  incisa in  reference sediment conditions were




 similar.




        85.  Insight into differences in mode of  exposure that may




 exist for these  two species can be seen by examining the net growth




 efficiencies from  the BRH  exposures.  For  Nj,_ incisa, exposure to




 BRH sediment greatly reduced  burrowing activity  which directly




 affected the amount of  energy (food)  ingested.   The impact of this




 is  apparent in the  negative net growth efficiency  for !*._ incisa




 from the BRH treatments.




        86.  Although burrowing activity is reduced in BRH sediment




with N. arenaceodentata, this fact does not necessarily lead to




 reduced energy injestion rates as with N^_  incisa.  Since N.




arenaceodentata is  a surficial detrital feeder,  rates of food




ingestion are probably not  related to burrowing  activity, at least




not as stongly as it does with sediment injesters such as N. incisa.




Net growth efficiency for KL^  areanaceodentata in BRH/BRH treatment




(experiment II,  particulate phase assay) was lower than for those




individuals in the  reference/reference condition, but the efficiency




remained positive,  suggesting that feeding did continue in BRH




sediment although at some reduced level.




       87.   Unlike  net growth efficiency which is an integrative




measure of  past physiological condition (i.e., a measure of the
                                 66

-------
efficiency at which an organism grew during the experimental  period),




scope for growth is an index by which the  current  physiological




condition of an organism is evaluated (i.e., is a  measure of  the




potential for growth).  It does not provide a time course estimate




of the cummulative effects a particular condition  has had on  an




organism as does net growth efficiency.  Rather, SFG evaluates




current physiological condition—the current physiological state




obviously being a product of those conditions to which the organism




was exposed to in the recent past.  Combined, these two estimators




of organismal health (net growth efficiency and SFG) offer insights




into what the effects of a particular set of conditions have been.




       88.  In the present study, the SFG values indicate that BRH




sediment  (100-percent BRH, BRH/BRH treatments) reduces the amount




of energy available for growth in N. arenaceodentata juveniles




(Table 14).  This is particularly interesting in  light of the fact




that the net growth efficiency values for two of  the experiments




did not  indicate any effect of BRH sediment  on  the growth of  this




species.  SFG values of N_._ arenaceodentata  followed a dose-reponse




to additions of BRH sediment in  all  experiments,  while net growth




efficiency did not.  The reason  for  this is  that  SFG values are not




as sensitive to differences in  size  as  is  the  measure of net  growth




efficiency.  Changes in size over  the  experimental period are not




taken into  account in  SFG  determinations whereas  these changes  in




growth are an important factor  in  calculating  net growth




efficiency.
                                 67

-------
        89.  The SFG values presented in this study are probably an




overestimate of what may be the actual energy available for growth.




The SFG estimates for worms from the reference sediment exposure




are at  least twice as high as the amount of growth that was recorded




during  the  10-day experimental period.  This was determined by




multiplying the mean SFG value (which is calculated for a 24-hr




period) by  the length of the experiment (Table 14) and comparing




this to the average energy equivalent for the amount of tissue




produced during the 10-day exposure (Table 12).  This is not an




entirely correct procedure for comparing actual growth with data




which allows for the prediction of growth (in this case, SFG) and




is probably partly responsible for the overestimation of the growth




potential in this particular case.  Scope for growth determinations




should instead be made at several times during the experimental




period  in which growth will actually be measured to better track




potential for growth.  By determining SFG only at the end of the




experimental period, all estimates for the potential for growth are




for individuals larger than existed during most of the experiment.




As the potential for growth probably increases with size (at least




within the juvenile stage), SFG values from worms at the end of the




10-day experimental period should be greater than those from worms




from the beginning of the experiment and would thus tend to over-




estimate previous potential for growth.




       90.  Another reason to believe that the SFG values presented




here are an overestimate of growth potential is the fact that no




attempt was made to calculate the amount of energy loss associated
                                 68

-------
with mucus production.  As noted earlier,  N.  arenaceodentata builds




tubes of mucus and sediment and these tubes appear to be con-




tinually rebuilt.  (Personal observation of the Neanthes cultures




suggest that mucus production is virtually constant.) As mucopoly-




saccharides are high in energy content, the production of these




compounds would be at a considerable cost in terms of energy.  For




example, it is estimated that approximately 30 percent of assimilated




energy is partitioned as the cost of mucus production in some inverte-




brate species which produce copious quantities of mucus  (Pandian




1975).  Future bioenergetics research with this species  will take




into account  the  amount of assimilated  energy  that is used  in mucus




production.




        91.  The  effects of BRH sediment are  probably due to at  least




one of  two  factors.   Either the worms  (especially N. incisa) were




responding  to a  difference  in  particle  size, or  they were reacting




to  contaminants  within  the  BRH sediment.   While  the  first possibility




cannot  be completely  discounted, Carey  (1962)  found  no  correlation




between the occurrence  of N^ incisa populations  in Long Island




Sound  and particle-size distribution despite significant differences




in  the  granulometry among the  various sites.




        92.   The second possibility appears to be the most likely




factor  affecting N^ incisa.  Black Rock Harbor sediment is a multi-




 contaminated sediment that contains a wide variety of anthropogenic




 chemicals.   Qualitative analysis of BRH sediments indicates the




 presence of high concentrations of polychlorinated  biphenyls,




 polynuclear aromatic hydrocarbons, and heavy metals including  Cu,
                                  69

-------
Pb, Cd, and Cr (Rogerson et al. 1984).  Despite this large spectrum




of contaminants, BRH sediment is not acutely toxic to N. incisa




juveniles.




       93.  The exposure conditions presented in this report do not




represent realistic exposure regimes for N_^ incisa in Long Island




Sound.  Rather, they were employed to produce a sublethal response.




Current research is aimed at establishing the exposure regimes that




!N^ incisa would be exposed to near the Black Rock Harbor site.




Once these limits are established, long-term laboratory studies




will be conducted with environmental conditions that more closely




approximate field conditions.
                                70

-------
                         PART V:   CONCLUSIONS






        94.  The results of this  study indicate that  the  principles of




bioenergetics can be applied to study the effect of sediment  disposed




on polychaetes.  With Nephtys incisa, the physiological measurements




were found to be reproducible within a particular treatment.   The




physiological measurements were also found to be repeatable.   In all




instances, measurement values from a particular treatment in one




experiment were not significantly different from the same treatment




in repeated experiments.  In most cases, the physiological measures




followed a dose-response to additions of BRH sediment.  The only




exception were the 0:N ratios which did not show a response to exposure




to BRH sediment.




        95.  The findings with Neanthes arenaceodentata were not as




dramatic as they were for N^ incisa.  In only one of the three




experiments was there a dose-response to additions of BRH sediment.
                                 71

-------
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Davis, W.R.  1979.  "The Burrowing, Feeding and Respiratory Activities
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Gilfillan, E.S.  1980.  "The Use of Scope-for-growth Measurements
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Gilfillan, E.S.,  and Hanson, S.A.  1975.  "Effects of Paralytic
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Mclntyrne, A.D.,  and Pearce, J.B.  (eds), 1980.  "Biological Effects
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McKinney, C.L., Jr.  1982.  "Interrelationships Between Energy
Metabolism, Growth Dynamics, and Reproduction During the Life Cycle
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Neuhoff, H.-G.  1979.  "Influence of Temperature and Salinity on
Food Conversion and Growth of Different Nereis Species (Polychaeta,
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Pandian, T.J.  1975.  "Mechanisms of Heterotrophy," Marine Ecology,
Vol.  II.  Physiological Mechanisms, Part 1, 0.  Kinne, ed.  Wiley,
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Pesch, C.E., and Hoffman, G.L.  1983.  "Interlaboratory Comparison
of a 28-day Toxicity Test With the Polychaete Neanthes arenaceodentata,
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Rhoads, D.C., and Germane, J.D.  1982.   "Characterization of Organism-
sediment Relations Using Sediment Profile Imaging: An Efficient
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Rogerson, P., Schimmel, S., and Hoffman, G.  1984.  "Chemical and
Biological Characterization of Black Rock Harbor Dredged Material,"
Technical Report D-84-, prepared by U.S. Environmental Protection
Agency, Narragansett, R.I., for the U.S. Army Engineer Waterways
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Rubinstein, N.  1979.  "A Benthic Bioassay Using Time-lapsed Photog-
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of Lugworms (Polychaeta:  Arenicolida)," Marine Pollution;  Functional
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Vernberg, eds.. Academic Press, New York, pp 341-351.
                                 74

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Sanders, H.L.  1956.  "Oceanography of Long Island Sound.   X.   The
Biology of Bottom Marine Communities," Bulletin Binghamton Oceanograghy
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Schauer, P.S., and Pesch, C.E.  In Prep.  "Influence of Diet on
Growth, Survival and Tube Production of Laboratory Cultured Polychaete
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Tenore, K.R., and Gopalan, U.K.  1974.  "Feeding Efficiencies of
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Warren, C.E., and Davis, G.E.  1967.  "Laboratory. Studies on the
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Yingst, J.Y., and Rhoads, D.C.  1980.  "The Role of Bioturbation in
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of South Carolina Press, Columbia, pp 407-421.
                                 75

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APPENDIX A:  SOLID PHASE AND PARTICULATE PHASE DATA SHEETS
                            Al

-------
                     LABORATORY WORM DATA SHEET
                             COE/ERLN FVP
 STUDY PLAN:  3

 EXPERIMENT DESCRIPTION: SOLID
                                   INVESTIOATOR: JOHNS/GU T JAHR

                                        DATE OF TEST:     821014
                                      SPECIES: NEPHTYS  INC ISA
                      ** EXPERIMENTAL CONDITIONS **
 TEMPERATURE:  21. OO DEGREES CENTIGRADE
 SALINITY:  28. OO    PARTS PER THOUSAND
 EXPOSURE DURATION: 1O DAYS
 PHOTOPERIOD:  13 HOURS
 FLOW RATE:   SO MLS/MIN
 NUMBER OF ANIMALS/REPLICATE:
 ANIMAL'S LIFE STAGE:  JUVENILE
 SIZE:  1.39 +/- 0.21 MG DRY WT
 CONTROLS:  1OO PERCENT REF
 FOOD:  PRAWN FLAKE SUSPENSION
 ANIMAL SOURCE:  SOUTH REFERENCE, LONG ISLAND SOUND
 COLLECTION TEMPERATURE:  19. 3 C
 ACCLIMATION:  20 C.
              SEDIMENT SCUHCE: BARREL OR COLLECTION/JAR  NUMBER
 SOLID REFERENCE:   11/SO                    SOLID BRH:  LL/23
 SUSPENDED REFERENCE:                       SUSPENDED  BRH:
                                         RANGE: 2O.30  - 21. SO
                                         RANGE: 28. OO  - 29. OO
                                  VOLUME ADDITIONS/DAY:
                               13  NUMBER OF REPLICATES/TREATMENT:
                                        COLLECTION  SALINITY:  29
SAMPLE: EXPOSURE CONCENIRATIONS!
                                   GROWTH
                                   JOULES
NUMBER
                                  !RESPIRATORY!EXCRETORY  !NET GROWTH
                                  !  ENERGY    !   ENERGY  !EFFICIENCY
NOMINAL    !MEASURED!       !      !EXPENDITURE!    LOSS    !IN PERCENT
           !          !       !      i  JOULES    !   JOULES  !       !
           !          !MEAN +/-STD  !MEAN+/-STD  1MEAN+/-STD1MEAN+/-STD
400020 100 PERCENT REF!

400021 30:30 REF:BRH   !
                       !
400022 100 PERCENT BRH!
                                !  . 87  !  . O7  i 6. 1  !  1. 8  i  0. O4!O. 01 !  17   !  10
                                i       ;:::;:       i
                                ! -. 34  !  . 11  1 3. 3  !  2. 1  !  0. O3!0. Ol!  -3   !  14
                                !       !!!!!!       i
                                !-3. 33  !  . 34  ! 6. 3  !  2. 8  !  0. 02!0. 01!  -6O  !  20
                                        A3

-------
                      LABORATORY WORM DATA SHEET
                              COE/ERLN FVP
 STUDY FLAN:  3

 EXPERIMENT DESCRIPTION:  SOLID
                                  INVESTIGATOR:  JOHNS/CUTJAHR

                                        DATE OF TEST:      821203
                                       SPECIES:  NEPHTYS INC ISA
                       **  EXPERIMENTAL CONDITIONS »»
TEMPERATURE: 2O. 30  DEGREES CENTIGRADE
SALINITY: 30. 00     PARTS  PER  THOUSAND
EXPOSURE DURATION:  10  DAYS
PHOTOPERIOD: 13  HOURS
FLOW RATE:   50 KLS/MIN
NUMBER OF ANIMALS/REPLICATE:
ANIMAL'S LIFE STAGE: JUVENILE
SIZE:  1.4O +/- O. 33 MG DRY WI
CONTROLS: 1OO PERCENT  REF
FOOD:  PRAWN  FLAKE SUSPENSION
ANIMAL SOURCE:  SOUTH REFERENCE, LONG  ISLAND  SOUND
COLLECTION TEMPERATURE: 6. 7 C
ACCLIMATION: 2O  C.
             SEDIMENT  SOURCE:  BARREL  OR  COLLECTION/JAR NUMBER
SOLID REFERENCE: 11/50                    SOLID BRH:  LL/25
SUSPENDED REFERENCE:                       SUSPENDED BRH:
                                          RANGE:  2O.OO - 21. 5O
                                          RANGE:  28. 5O - 31. OO
                                   VOLUME ADDITIONS/DAY:
                                15  NUMBER OF REPLICATES/TREATMENT:
                                        COLLECTION SALINITY:  29
SAMPLE! EXPOSURE CONCENTRATIONS!
NUMBER!
       !
       !
          NOMINAL
            GROWTH    !RESPIRATORY{EXCRETORY  ! NET GROWTH
        !    JOULES    !  ENERGY    !  ENERGY   !EFFICIENCY
MEASURED!        !      !EXPENDITURE!   LOSS    !IN PERCENT
        !        i      i  JOULES    i  JOULES   i       !
        !MEAN +/-STD  SMEAN+/-STD  !MEAN+/-STD!MEAN-t-/-STD
400023!100 PERCENT REF
       I
       I

40OO23!5O:5O REF:BRH
       I
       f

40OO27J100 PERCENT BRH
                                5  . 87!  . 47  i  13. 5!  3. 7 !  0. O4!0. 01!  30   !   8
                                      !!!!!!       !
                                4. 13  ! 1.2   !  11.31  2.9 !  0. O3:O.Oi:  26   !   7
                                      I      J      I      «      «     t       I

                               -O. 04  !  .0   !   5.3!  1.2 !  0. O2!0. 01!  -1   !   4
                                      !!!!!!       i
                                        A4

-------
                    LABORATORY WORM  DATA  SHEET
                            COE/ERLN FVP
STUDY PLAN: 3
                                  INVESTIGATOR:  JOHNS /GUTJAHR
EXPERIMENT DESCRIPTION: SOLID
                                        DATE  OF  TEST:
                                                          821203
                                     SPECIES:  NEPHTYS INC ISA
                     »•* EXPERIMENTAL CONDITIONS  **
                                        RANGE:  SO.OO -  21. 30
                                        RANGE:  28.90 -  31. OO
TEMPERATURE: 20.50 DEGREES CENTIGRADE
SALINITY: 30. OO    PARTS PER THOUSAND
EXPOSURE DURATION: 1O DAYS
PHOTOPERIOD: 13 HOURS
FLOU RATE:    SO I1LS/MIN
NUMBER OF ANIMALS/REPLICATE:
ANIMAL'S LIFE STAGE:  JUVENILE
SIZE: 1.43 +/- 0.73 MG DRY WT
CONTROLS: 1OO PERCENT REF
FOOD: PRAWN  FLAKE SUSPENSION
ANIMAL SOURCE:  SOUTH REFERENCE
COLLECTION TEMPERATURE. 6. 7 C
ACCLIMATION: 20 C.
             SEDIMENT SOURCE: BARREL OR  COLLECTION/JAR  NUMBER
SOLID REFERENCE:  11/50                     SOLID BRH:  LL/23
SUSPENDED REFERENCE:                        SUSPENDED BRH:
                                 VOLUME ADDITIONS/DAY:
                               13  NUMBER QF  REPLICATES/TREATMENT:
                                LONG  ISLAND  SOUND
                                      COLLECTION SALINITY:  29
SAMPLE! EXPOSURE CONCENTRATIONS!
! ! !
NUMBER! NOMINAL ! MEASURED i
! ! !
! ! !
GROV
JOUl
MEAN +i
400036! 100 PERCENT REF! ! 4.04
! ! i
400O37!5O:30 REF: BRH ! ! O. 65
! ! !
400O38J1OO PERCENT BRH! !-l. 85
! ! !
•ITH ! RESPIRATORY! EXCRETORY
.ES ! ENERGY ! ENERGY
{EXPENDITURE! LOSS
! JOULES ! JOULES
'-STD SMEAN+/-STD !MEAN+/-STD
. 73
. 16
. 17
11. 8! 2. 5 ! 0. 04!0. 01
i i !
6. 2! 1.3 i O. O310. Ol
III
3. 3! 2. 3 ! 0. 0210. 01
! ! !
NET GROWTH
EFFICIENCY
IN PERCENT
!
MEAN+/-STD
23 i 5
!
1O ! 2
1
-24 ! 14
!
                                       A5

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                    LABORATORY WORM DATA SHEET
                             COE/ERLN FVP

STUDY PLAN: 3                     INVESTIGATOR:  JOHNS/CUTJAHR

EXPERIMENT DESCRIPTION:  SUSPENDED          DATE OF TEST:      830902

                                      SPECIES:  NEPHTYS INC ISA
                     **  EXPERIMENTAL CONDITIONS *»
TEMPERATURE: 21.20  DEGREES CENTIGRADE
SALINITY: 30. SO     PARTS PER THOUSAND
EXPOSURE DURATION:  10  DAYS
PHOTOPERIOD: 13  HOURS
FLOW RATE:   35 MLS/MIN
NUMBER OF ANIMALS/REPLICATE:
ANIMAL'S LIFE STAGE: JUVENILE
SIZE: 3. 13 +/- 0.60 MO DRY WT
CONTROLS: 2OO MG/L  REF/REF
FOOD: PRAWN  FLAKE SUSPENSION
ANIMAL SOURCE:  SOUTH REFERENCE/  LONG ISLAND SOUND
COLLECTION TEMPERATURE:  2O. 3 C
ACCLIMATION: 20  C.
             SEDIMENT  SOURCE: BARREL OR COLLECTION/ JAR NUMBER
SOLID REFERENCE:  1 1 1/19                    SOLID ERH:  EE/3, 8
SUSPENDED REFERENCE: 111/19,21,22          SUSPENDED BRH: EE/7, 11,12
                                         RANGE:  20. 30 - 22. SO
                                         RANGE:  30.OO - 31. OO
                                  VOLUME ADDITIONS/DAY:  67
                               IS   NUMBER CF REPLICATES/TREATMENT:
                                        COLLECTION SALINITY:  25 8
SAMPLE! EXPOSURE CONCEf
NUMBER! NOMINAL
40O1301200MG/L REF/REF
1
400131 I200MG/L BRH/REF
I
1
40Q132S200MG/L REF/BRH
400133 S200MS/L BRH/BRH
v'TRATIONS
MEASURED
211+87
171+53
211+87
171+33
GRO'v
JOUt
MEAN +/
6. 09
4. 35
3. OO
-2. 30
^TH
-ES
'-STD
1.22
. 96
1. 1O
. 78
RESPIRATORY
ENERGY
EXPENDITURE
JOULES
MEAN+/-STD
14. 1
13. 7
10. 2
9. 0
4. 7
3. 2
4. 7
3. 3
EXCRETORY
ENERGY
LOSS
JOULES
MEAN+/-STD
O. O8!O. 01
1
O. O8!0. O2
O. O3!0. 01
1
0. 0410. 01
NET GROWTH
EFFICIENCY
IN PERCENT
MEAN+/-STD
35 ! 5
1
1
24 ! 3
36 ! 22
-78 ! 48
                                       A6

-------
                    LABORATORY  WORM DATA SHEET
                             COE/ERLN FVP
STUDY PLAN:  3
                                  INVESTIGATOR:  JOHNS/OUFJAHR
EXPERIMENT DESCRIPTIO:4: SUSPENDED
                                            DATE OF TEST:
830920
                                      SPECIES:  NEPHTYS INC ISA
                     ** EXPERIMENTAL  CONDITIONS •**
                                         RANGE:  19. SO - 22. GO
                                         RANGE:  30.00 - 31. 8O
TEMPERATURE: 2O. 60 DEGRESS  CENTIGRADE
SALINITY: 3O.70    PARTS  PER  THOUSAND
EXPOSURE DURATION: 10  DAYS
PHOTOPERIOD: 13 HOURS
FLOW RATE:  32 fS-S/HIN           VOLUilE ADDITIONS/DAY: 61
NUM3ER OF ANIMALS/REPLICATE:   15  NUMBER OF REPLICATES/TREATMENT:    1
ANIMAL'S LIFE STAGE: JUVENILE
SIZE: 3.43 +/- 0.92 KG DRY  Wf
CONTROLS: 2CO KG/L REF/REF
FOOD: PRAWN FLAKE SUSPENSION
ANIMAL SOURCE:  SOUTH REFERENCE,  LONG ISLAND SOUND
COLLECTION TEMPERATURE: 21.6  C          COLLECTION SALINITY: 29.2
ACCLIMATION: 2O C.
             SEDIMENT  SOUHCE:  BARREL OR COLLECTION/JAR NUMBER
SOLID REFERENCE:  111/23                    SOLID BRH: EE/17
SUSPENDED REFERENCE: HI/6, 7, 36             SUSPENDED BRH: EE/8. 10, 14
SAMPLE! EXPOSURE CONCENTRATIONS! GROWTH
! i JOULES
NUMBER ! NOM I NAL ! MEASURED :
1
1
4OG134I20OKG/L REK/REF
I
1
400133 J20CKS/L BRH/REF
!
4O013612OCKG/L REF/BRH
j
40O13712OOMO/L BRH/BRH
!
(MEAN +/
193+73 !13. 38
!
226+48 i 8 48
t
I
193+73 ! 5. 11
j
226+48 i 1. 41


r-STD
. 94

1. 36

. 92

RESPIRATORY
ENERGY
EXPENDITURE
JOULES
MEAN+/-STD
17. 8! 4.4
t
18. 3! 3. 8
t
1
12. 2! 2. 8
1
.27 ! 10. 6! 2. 3
EXCRETORY
ENERGY
LOSS
JOULES
MEAN+/-STD
O. 0710. 01
1
1
0. 07 !0. 03
1
1
O. 07 !0. 02
j
0. 0510. 01
NET GROWTH
EFFICIENCY
IN PERCENT

MEAN+/-
44

33

29


•STD
7

7

4

12 ! 2
                                        A7

-------
                     LABORATORY WORM DATA SHEET
                             COE/ERLN FVP
STUDY PLAN: 3

EXPERIMCNT DESCRIPTION:  SOLID
   INVESTIGATOR: JOHNS/GUTJAHR

         DATE OF TEST:     830516
                                      SPECIES:  NEANTHES ARENACEODENTATA
                      **  EXPERIMENTAL CONDITIONS **
TEMPERATURE: 21. OO  DEGREES  CENTIGRADE
SALINITY: 28. OO     PARTS PER  THOUSAND
EXPOSURE DURATION:  1O  DAYS
PHOTOPERIOO: 13 HOURS
FLOW RATE:   100 HLS/MIN
NUMBER OF ANIMALS/REPLICATE:
ANIMAL'S LIFE STAGE: JUVENILE
SIZE: 4. 16 + /-  1.73 MS DRY  WT
CONTROLS: 100 PERCENT  REF
FOOD: PRAWN  FLAKE SUSPENSION
ANIMAL SOURCE:  CULTURE
COLLECTION TEMPERATURE:
ACCLIMATION: 20 C.
             SEDIMENT  SOURCE:
SOLID REFERENCE:  I I/SO
SUSPENDED REFERENCE:
          RANGE: 20. 30 - 21. 30
          RANGE: 28. 00 - 29. 00
    VOLUME ADDITIONS/DAY  67
13  NUMBER OF REPLICATES/TREATMENT:
   AGE:   41 DAYS
         COLLECTION SALINITY:

BARREL OR COLLECTION/JAR NUMBER
            SOLID BRH: LL/23
            SUSPENDED BRH:
SAMPLE! EXPOSURE CONCENTRATIONS
! i
NUMBER! NOMINAL {MEASURED
!
'•
4OOO23S10O PERCENT REF
!
400029! 50: 50 REF: BRH
j
400030 110O PERCENT BRH







! !
GROWTH
JOULES
I
1
t
MEAN -»-/-STD
22 ! 20
!
14 ! 19
!
13 ! 14
1
RESPIRATORY
ENERGY
EXPENDITURE
JOULES
MEAN+/-STD
38 ! 9
!
30 ! 11
1
23 i 3
EXCRETORY !NET GROWTH
ENERGY {EFFICIENCY
LOSS ! IN PERCENT
JOULES i !
MEAN+/-STD ! MEAN+/-STD
0. 06 { 0. O2 ! 29 ! 34
t t <
1 1 I
0.03! 0.02! 18 ! 42
! ! !
0. 04 !0. 01! 27 ! 32
! ! ! ! i
                                       A8

-------
                    LABORATORY  WORM  DATA SHEET
                             COE/ERLN FVP
STUDY PLAN: 3
                                  INVESTIGATOR:  JOHNS/GUTJAHR
EXPERIMENT DESCRIPTION: SUSPENDED
                                              DATE CF TEST:
                                                                830*722
                                      SPECIES:  NEANTHES ARENACEOCENTATA
                     *+ EXPERIMENTAL  CONDITIONS **
                                         RANGE:  19, 80 - 22. OO
                                         RANGE:  3O. OO - 31. 80
TEMPERATURE: 2O. 6O DECREES CENTIGRADE
SALINITY: 30. 70    PARTS PER  THOUSAND
EXPOSURE DURATION: 10 DAVS
PHOTOPERIOD: 13 HOURS
FLOW RATE:   33 MLS/MIM            VOLUME ADDITIQNS/'DAY  63
NUMBER OF ANIMALS/REPLICATE:   13   NUMBER CF REPLICATES/TREATMENT:
ANIMAL'S LIFE STAGE:  JUVtNILE    AGE:   42 DAYS
SIZE: 3. 52 +/- 2.09  MG DRY WT
CONTROLS: 20O MG/L REF/REF
FOOD: PRAWN FLAKE SUSPENSION
ANIMAL SOURCE:  CULTURE
COLLECTION TEMPEKATURE:                  COLLECTION SALINITY:
ACCLIMATION: 20 C.
             SEDIMENT SOURCE:  BARREL  OR COLLECTION/JAR NUMBER
                                           SOLID BRH:  EE/17,18
SOLID REFERENCE: 111/23,26
SUSPENDED REFERENCE:  111tb, 7,36
                                           SUSPENDED BRH:  EE/8. 1O, 14
SAMPLE !
!
NUMBER !
1

4O0133!
t
4O0139!
1
4O0140!
i
400141!
EXPOSURE CONCENTRATIONS!

NOMI


20CMG/L

20CMO/L

20CM3/L

20CKO/L

MAL


RE»=YREF

BRH/REF

REF/BRH


1
GROWTH J RESPIRATORY
JOULES ! ENERGY
MEASURED i


199

222

199

3RH/BRHS222
1
1
1
+ 73!
1
+ 44!
1
1
+ 73!
1
+ 44!

MEAN
39

49

43

37
I ! EXPENDITURE
! ! JOULES
+/-STD !MEAN-t-/-STD
: 38
1
5 40
•*
i 27
37

34

61
f 1
t 1
13

13

3

! 32 5 62 ! 7
EXCRETORY 1NET
ENERGY !
LOSS i
JOULES !
KEAN+/-STD!
0. 08 !O. O4!
i ;
O. 14!0. O3!
! i
O. 1110.03!
t I
0. 13J0.07!
GROWTH
EFFICIENCY
IN PERCENT

5
MEAN-f/-STD
3O

41

39

31
! 27
*
; 20
j
i 16
1
! 23
                                      A9

-------
                     LABORATORY WORM DATA SHEET
                             CCE/ERLN FVP

STUDY PLAN:  3                     INVESTIGATOR:  JOHNS/CUTJAHR

EXPERIMENT  DESCRIPTION:  SUSPENDED            DATE OF TEST:
                                                                 830816
                                         SPECIES: NEANTHES ARENACEODENTATA
                      -*•» EXPERIMENTAL CONDITIONS **
                                         RANGE: 20. SO - 21. 50
                                         RANGE: 30.00 - 31. 00
TtKPERATUSE:  21. OO DEGREES CENTIGRADE
SALINITY: 30. 00     PARTS PER THOUSAND
EXPOSURE DURATION:  10 DAYS
PHOTOPERIOD:  13  HOURS
FLOW RATE:    33 HLS/MIN           VOLUME ADDITIONS/DAY  67
NUMBER OF ANIMALS/REPLICATE:   13  NUMBER OF REPLICATES/TREATMENT:
ANIMAL'S LIFE STAGE:  JUVtNILE    AGE:   37 DAYS
SIZE:  4. 43 +/- 0. 092   MG DRY WT
CONTROLS: 200 MG/L REF/REF
FOOD:  PRAWN  FLAKE SUSPENSION
ANIMAL SOURCE: CULTURE
COLLECTION TEMPERATURE:                 COLLECTION SALINITY:
ACCLIMATION: 20  C.
             SEDIMENT SOURCE:  BARREL OR COLLECTION/JAR NUMBER
                                           SOLID BRH:  EE/1.2
                                           SUSPENDED BRH:  EE/l-3
SOLID REFERENCE:  111/13,14
SUSPENDED REFERENCE:  111/13-17
SAMPLE: EXPOSURE CONCENTRATIONS
NUMBER: NOMINAL [MEASURED
40O126I20CMG/L RE*-/REF!217 + 86
1 1
400127S20CMG/L RE*-/BRH!190 + 61
400123!20CKS/L BRH/REFI217 + 86
t i
40O129S2OCMC/L BRH/BRH!190 + 61
GRO;
JOUl
MEAN -»•
36
38
3
4TH
.ES
'-STD
33
39
31
14
RESPIRATORY
ENERGY
EXPENDITURE
JOULES
MEAN* /-STD
52
36
33
47
6
10
6
9
EXCRETORY !NET GROWTH
ENERGY ! EFFICIENCY
LOSS ! IN PERCENT
JOULES ! !
MEAN+/-STD ! MEAN-t-/-STD
0.08! 0.02! 33 ! 19
i ! !
O. 08 JO. 03! 34 ! 24
ft i
O. 14!0. 03! 33 ! 20
1 f 1
0. 11 SO. 03! 3 ! 23
                                       A10

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