ROOTED AQUATIC PLANTS
IN THE
UPPER POTOMAC RIVER BASIN
DIVISION OF FIELD INVESTIGATIONS, CINCINNATI
OFFICE OF ENFORCEMENT AND STANDARDS COMPLIANCE
WATER QUALITY OFFICE
ENVIRONMENTAL PROTECTION AGENCY
CINCINNATI, OHIO

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ROOTED AQUATIC PLANTS IN THE UPPER POTOMAC RIVER BASIN
by
Delbert B. Hicks
Division of Field Investigations, Cincinnati
Office of Enforcement and Standards Compliance
Water Quality Office
Environmental Protection Agency
Cincinnati, Ohio
1971

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TABLE OF CONTENTS
CONCLUSIONS 		1
INTRODUCTION		2
STUDY AREA		3
SAMPLING STATION AND METHODS		h
AQUATIC PLANT GROWTH FACTORS 		5
STUDY RESULTS		6
SIGNIFICANCE OF ROOTED AQUATIC
PLANTS AS RESERVOIRS OF NUTRIENTS'		9
ACKNOWLEDGEMENTS		 .	10
LITERATURE CITED	11
TABLES
Table 1. Location of Aquatic Plant Sampling Stations,
Upper Potomac River Basin. ... 	 . .	Follow
Page 9
Table 2. Standing Crop of Root Aquatic Plants in the
Upper Potcmac River Basin and Weight of
Phosphorus and Organic Nitrogen in Storage . . .
Table 3. Phosphorus and Nitrogen Content of Rooted
Aquatic Plants from the Upper Potcmac
River Basin 	

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CONCLUSIONS
1.	Rooted aquatic plants in the Upper Potomac Basin axe not a
significant factor in the nutrient budget of the Lover
Potomac River.
2.	In August 1969 the standing crop of rooted aquatic plants
in the Upper Potomac River Basin contained 400 pounds of
phosphorus (P) and	pounds of nitrogen (N). These
quantities are equivalent to approximately 40 percent of
the phosphorus and nitrogen in wastewaters discharged to
the Upper Potomac River in a single day.
5- Standing crop estimates were made for growths in Antietam
Creek, Conococheague Creek and the Potomac River. Other
basin streams contained too few growths of rooted plants to
merit sampling for standing crop estimates.
1

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INTRODUCTION
Conferees of the Potomac River-Washington Metropolitan Area
Enforcement Conference recommended that the Interstate Commission
on the Potomac River Basin, the States of Maryland, Virginia,
Pennsylvania and West Virginia, the District of Columbia and the
Federal Water Pollution Control Administration make joint studies
of the entire Potomac Basin to determine detrimental effects on the
Potomac River and estuary. Concern developed for the effects of
rooted aquatic plants or pond-weeds on the nutrient regimen cf the
Lower Potomac River and estuary. These plants accumulate and store
nutrients during the spring and summer growing season. Upon death
and decay of these plants in late summer and autumn, stored nutrients
would be released for transport to the lower river and estuary where
nuisance algal growths occur. The Regional Director, Middle Atlantic
Region, Federal Water Pollution Control Administration, requested the
Division of Field Investigations, Cincinnati, to conduct a study of
plants in the Upper Potomac River Basin to:
1.	Estimate the quantity of phosphorus (p) and organic
nitrogen (N) accumulated in rooted aquatic plant
growth.
2.	Assess the significance of these stored nutrients
on the nutrient budget of the Potomac River Basin.
2

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STUDY AREA
The Potomac River portion included in the study ex-
tended from Great Falls, Maryland, upstream 220 miles to the
confluence of the Savage River, Maryland. Major tributary
streams investigated were the Monocacy River, Antietam Creek,
Conococheague Creek, South Branch Potomac River, Cacapon River,
and Shenandoah River.
The field survey was conducted August 11 to 19, 19^9>
a period of the year when aquatic plant6 growths would be
expected to reach maximum abundance.
5

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SAMPLING STATION AND METHODS
When possible, sampling stations were located near highway
bridge crossings. Bridges provided good vantage points to observe
the overall streambed coverage by rooted aquatic plants. For
purpose of study, rooted aquatic plants were defined as vascular
plants attached to the stream bottom by a root system and submerged
except for floating portions of the plants and the macro-alga Chara.
The quantity of rooted aquatic plants was determined when
plant growths covered more than five percent of the stream bottom.
The percent coverage and quantity of plants for the interstation
reaches were estimated by extrapolation. Fifty sampling stations
were established in the study area (Figure 1 and Table l).
Plant samples were taken with an open-ended, steel cylinder,
10 inches in diameter and 36 inches in length. The sampler was
forced into the 6tream bottom with a rotary motion. Entrapped
vegetation was removed, damp dried, segregated into species and
weighed. Sampling was conducted at 20-foot intervals along a 100-
foot transect of the plant growths.
Complete plants representative of the kinds found in the
quantitative samples were collected, damp-dried, weighed, frozen
on dry ice, and returned to the laboratory. Each specimen was
homogenized, freeze-dried, and analyzed for phosphorus (P) and
organic nitrogen (N) content.
4

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POTOMAC RIVER BASIN

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AQCJATIC PLAM* GROWTH FACTORS
The establishment and growth of rooted aquatic plants is
governed by physical factors such as the availability of suit-
able substratej current velocities, water depth and turbidity.
Increased current velocity can cause the bottom to become
physically unstable for root attachment by scouring away the
finer particles in which plants root. Increases in depth or
turbidity reduces available sunlight for plant germination and
photosynthesis. If adverse conditions persist into the growing
season, lush growths may fail to develop in areas that in previous
years may have supported abundant growths.
Phosphorus and nitrogen sure two major nutrients essential
for growth of rooted aquatic plants. Plants assimilate these
nutrients and place them in temporary storage. The quantities of
nutrients held in storage are dependent upon the standing crop of
plants. With the annual die-down of plants in the fall, stored
nutrients are slowly released to the stream as plants undergo de-
composition. The released nutrients are then available for recycling
by other primary producers.
5

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STUDY RESULTS
UPPER POTOMAC RIVER
The main stem of the Upper Potomac River upstream from
the Great Falls supported a standing crop of rooted aquatic
plants estimated at TOO tons and contained 280 pounds of phos-
phorus (P) and 3660 pounds of nitrogen (N) (Table 2). Sixty-
six percent of the standing crop was contained in an 18 mile
reach extending from the confluence of Antietam Creek (Station
17) upstream to where Conococheague Creek enters the Potcmac
River. The remaining portion of the standing crop was in a
35-mile section of the river extending from the mouth of the
South Branch Potomac River upstream to Dawsen, Maryland.
ANTIETAM CREEK
Antietam Creek, Maryland had an estimated 5k tons of
rooted aquatic plants with a potential yield of 3*+ pounds phos-
phorus and 220 pounds nitrogen (Table 2). Fifty-three percent
of the plants were found in two 600-foot sections of Antietam
Creek: 1. at Burnside Bridge (Station 13) near Sharpsberg and
2. at Antietam (Station 17) near the mouth of the stream. These
two very dense patches of plants were atypical for the stream.
6

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7
At these points, the stream was shallow and uniform in depth,
lacked shading by upper-story stream-side vegetation and
supported one species of rooted aquatic vegetation.
Thirty-six percent of the aquatic weeds were contained
in a 15-mile reach extending downstream from the mouth of the
West Branch of Antietam Creek. The West Branch (Station 13A)
supported 11 percent of the standing crop in a one-mile reach
of stream flowing through a meadow. The East Branch of Antietam
Creek (Station lk) supported too few plants for determining
standing crop estimates.
CONOCOCHEAGUE CREEK
Conococheague Creek supported an estimated 128 tons of
rooted aquatic plants that contained 80 pounds of phosphorus (p)
and 520 pounds of nitrogen (N) (Table 2). The plants were dis-
tributed in an eight-mile section of stream located at Station 19-
MONOCACY RIVER, SOUTH BRANCH POTOMAC RIVER,
SHENANDOAH RIVER, AND CACAPON RIVER
These streams and their tributaries contained insufficient
growths of rooted aquatic plants from which standing crop estimates
could be determined. Aquatic plant coverage was less than five
percent. Streams had high flows which were considered unseasonable

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8
for the year because of heavy summer rains. The streams and
their tributaries were very turbid with visibility reduced to
less them 12 inches in depth. High flows and increased tur-
bidity were apparent factors limiting the abundance of rooted
aquatic plants in these streams.

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SIGNIFICANCE OF ROOTED AQUATIC
PLANTS AS RESERVOIRS OF NUTRIENTS
Rooted aquatic plants in the Potomac River Basin are not
a significant factor affecting nutrient loading of the Lower
Potomac River.
The August standing crop of rooted aquatics in the Potomac
River upstream from the Great Falls was estimated to be 880 tons
and contained 400 pounds phosphorus (P) and M*00 pounds nitrogen
(N). These nutrients are equivalent to approximately UO percent
of the average total pounds of phosphorus and nitrogen in waste-
waters discharged to the Upper Potomac River system in a single
day w.
In past years, more extensive growths of rooted aquatic
(2 3 1<.)
plants in the Upper Basin have been reportedv ' ' ' . Even under
conditions of a superabundance of rooted aquatic plants, the
quantity of phosphorus and nitrogen in storage remains insignificant
when compared to the known nutrient discharges. Assuming that
stream beds of the Upper Potomac River and its major tributaries
were 100 percent covered with plants at densities equal to the
maximum observed in this study, total phosphorus and nitrogen in
storage would be less than .04 percent and 2.0 percent, respectively,
of that discharged annually to the Lower Potomac River as measured
at the Great Falls.
9

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Table 1. Location of Aquatic Plant
Sampling Stations,
Upper Potomac River Basin
Station
No.	Stream	Location
11
Monocacy River
Md. Rt. 28 Bridge
10
Monocacy River
Lily Pons Bridge, Gland Road, Md.
9
Monocacy River
Md. Rt. 80 Bridge east of
Buckeystown, Md.
8
Monocacy River
Gas House Pike Road Bridge
near Frederick, Md.
7
Monocacy River
Md. Rt. 26 Bridge at Ceresville, Md.
6
Monocacy River
Md. Rt. 76 Bridge (Le Gore Bridge)
5
Monocacy River
Mumma Ford Bridge, Md.
k
Monocacy River
West of Harvey, Md.
5
Marsh Creek
Greenmount, Pa.
2
Rock Creek
Pa. Rt. 13k Bridge at Barlow, Pa.
IT
Antietam Creek
Antietam, Md.
13
Antietam Creek
Burnside Bridge, Antietam Nat. Park, Md.
16
Antietam Creek
Roxbury Road Bridge, Md.
15
Antietam Creek
Md. 60 Bridge
Ik
£. Branch Antietam Creek
Pa. Rt. 16 Bridge, east of Waynesboro,
Pa.
1JA
W. Branch Antietam Creek
Pa. Rt. 516, northwest of Waynesboro, Pa.
18
Conococheague Creek
Md. Rt. 68 Bridge, near mouth
19
Conococheague Creek
Conococheague, Md.
20
Conococheague Creek
Broadfording Road Bridge, Md-
21
Conococheague Creek
Pa. Rt. l6 Bridge west of Greencastle,
Pa.

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Table 1 Cont.
Station
No.	Stream	Location
26	Potomac River	Md. Rt. 135 at mouth Savage R., Md.
27	Potomac River	Md. Rt. 9 near Pinto, Md.
29	Potomac River	Toll Bridge at Old Town, Md.
2k	Potomac River	W. Va. Rt. 29 Bridge at Paw Paw,
W. Va.
52	Potomac River	W. Va. Rt. 48 Bridge at
Sheperdstown, W. Va.
50	Potomac River	Harpers Ferry, W. Va.
53	Potomac River	Md. Rt. 15 Bridge at Pt. Rocks, Md.
57	Potomac River	Great Falls, Cheaapeake and Ohio
Canal Park, Md.
30	S. Branch, Potomac R.	French Sta. W. Va. near mouth
31	S. Branch Potomac R.	W. Va. Rt. 28 Bridge downstream
Romney, W. Va.
32	S. Branch Potomac R.	W. Va. Rt. 28 Bridge upstream
Romney, W. Va.
33	S. Branch Potomac R.	U.S. 220 Bridge downstream
Moorefield, W. Va.
35	S. Branch Potomac R.	U.S. 220 Bridge downstream
Petersburg, W. Va.
36	S. Branch Potomac R.	1 mile downstream Petersburg, W. Va.
37	S. Branch Potomac R.	U.S. 220 Bridge upstream
Petersburg, W. Va.
39	s- Branch Potomac R.	U.S. 33 Bridge E. of Franklin, W. Va.
38	N. Fork of S. Branch	U. S. 35 Bridge at Judy Gap, W. Va.
Potomac River

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Table 1 Cont.
Station
No •	Stream
JU	S. Fork of S. Branch,
Potomac River
UO	S. Fork of S. Branch,
Potomac River
22	Cacapon River
23	Cacapon River
k9	Shenandoah River
U8	Shenandoah River
k6	S. Fork Shenandoah River
bk	S. Fork Shenandoah River
i+3	South River of S. Fork
Shenandoah River
k2	Middle River of S. Fork
Shenandoah River
kl	North River of S. Fork
Shenandoah River
kj	North Fork Shenandoah
River
1*5	North Fork Shenandoah
River
Location
U.S. 220 Bridge at Moorefield,
W. Va.
U. S. 33 Bridge near Brandyvine,
W.Va.
W.Va. Rt. 9 Bridge near mouth
W.Va. Rt. 9> 2nd Bridge upstream
from mouth
Harpers Ferry National Park, W.Va.
Va. Rt. 7 Bridge east of Berryville,
Va.
Va. Rt. 211 Bridge west of Luray, Va.
Va. Rt. 33 Bridge east of Elkton, Va.
Va. Rt. 256 at Grottoes, Va.
Va. Rt. 256 east of Weyers Cave,
Va
U. S. 11 Bridge south Mt. Crawford,
Va.
Va. Rt. 55 Bridge
Va. Rt. U2 Bridge at Timberville, Va.

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Table 2. Standing Crop of Root Aquatic Plants
In the Upper Potomac River Basin and
Weight of Phosphorus and Organic Nitrogen in Storage






Tons2
Weeds
per Strean
Mi.
Plants Contained]
Stream
Sta.
No.
Mi. of
Stream1
Aver.
Stream
Width,
Ft.
$ Weed
Coverage
lbs2
Weeds
per
S^.Ft.
lbs P
per
Streasi
Mi.
lbs N
per
Stream
Mi.
.
Potomac River
52
18
550
20
.09
26
10
102
ti
27
18
105
20
•09
5
2
13
If
29
17
185
30
.06
9
1+
1*0
Antietam Ck.
17
0.1
125
80
•51
15
8
ko
ti
15
0.1
110
80
• 51
13
7
35
M
16
17
75
< 5
-
-
-
-
fl
15
15
•F*
vn
15
.07
1
1
7
II
13A
1
15
30
• 50
6

36
II
ll*
-
20
< 5
-
-
-
-
Conococheague Ck.
18
fc-5
150
< 5
-
-
-
-
It
19
8
200
30
.10
16
10
55
II
20
11+
125
< 5
-
-
-
-
fl
21
-
100
< 5
-
-
-
—
es of stream estimated as representative of data.
Stet weight
^Nutrient data based on phosphorus and nitrogen analyses
of plant types present at sampling station (Table 3)•

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Table 3¦ Phosphorus and Nitrogen Content
of Rooted Aquatic Plants from the Upper Potcmac River Basin
pTof~TotaI Wet Wt- Dry Vt.
Station Kinds of Plants	Weeds at Nutrient Nutrient $ P	$ P	$N	% H
No- from Sq. Ft. Sample Station Sample(gm) SampLe(gjn) Dry Vt. Vet Vt. Dry Vt. Vet .
13
Heterantherla dubi
100
28
1-59
0.44
0.025
2.4
0.13
13A
Elodea canadensis
94
30
4.28
0.25
0.035
2.1
0.30

Potamogeton crispus
6
31
2.98
0.29
0.028
2.2
0.21
15
E. canadensis
67
42
3-78
O.56
0-050
3-0
0.27

H. dubi
13
12
1.22
0.48
0.049
2.7
0.27
19
H. dubi
69
25
2.10
0.36
0.030
1-9
0.16

P. americamifl
16
13
1.57
0-30
0.036
2.0
0.24

E. canadensis
15
18
1.83
O.36
0.030
1-7
0.17
28
Najas flexilis
46
14
1.76
O.17
0.021
1.1
0.14

H. dubi
29
17
1.30
0.14
0.010
1.8
0.14

Chara sp.
10
5
0.93
0.12
0.022
0.8
0.15

E. canadensis
5
6
1.11
0.10
0.018
0.0
0.14
29
P. illinoens is
59
16
1.87
0.24
0.028
2.2
0.25

P. diverBifollus
41
22
3-03
0.12
0.016
1.4
0.19
52
Vallisneria sp.
74
24
1.86
0.19
0.015
2.2
0.17

H. dubi
11
13
1.03
0.30
0.024
2.6
O.25

P. illinoens is
8
24
2.87
0.22
0.027
2.2
0.24

N. Ruadalupensis
4
14
1.77
0.28
O.O36
2.6
0-33

E. canadensis
3
7
1.30
0.14
0.026
1.8
0.33

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ACKNOWLEDGEMENTS
James LaBuy, Aquatic Biologist, Middle Atlantic Region,
EPA, Charlottesville, Virginia, assisted with field investi-
gations .
Chemists, Division of Field Investigations, Cincinnati,
EPA, Cincinnati, Ohio, conducted plant nutrient analyses.
10

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LITERATURE CITED
1.	Javorski, N. A., D. W. Lear, and J. A. Aalto. A Technical
Assessment of Current Water Quality Conditions and Factors
Affecting Water Quality in the Upper Potomac Estuary. Tech.
Rept. No. 3} Chesapeake Technical Support Laboratory, Middle
Atlantic Region, PVPCA, Dept. of Interior. 19^9 •
2.	DeRose, C. R. The Monocacy River, Physical, Chemical and
Bacteriological Water Quality, Rept. No. 1, State of Maryland,
Dept. of Water Resources, Division of Water Quality Investi-
gations. 1966.
3.	LaBuy, J. Biological Survey of the Monocacy River and
Tributaries. CB-SRBP Working Docunent No. 2J, Middle
Atlantic Region, FWPCA, Dept. of Interior. 1968.
LaBuy, J. Biological Survey Antietam Creek and Some Tribu-
taries. CB-SRBP Working Document No. 22, Middle Atlantic
Region, JVPCA, Dept. of Interior. 1968.
11

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Rooted aquatic plants in
the upper Potomac River
basin
EJDD PT 00027

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