United States       Prevention, Pesticides     EPA712-C-96-338
          Environmental Protection    and Toxic Substances     February 1996
          Agency        (7101)
&EPA    Microbial Pesticide
          Test Guidelines
          OPPTS 885.5200
          Expression in a
          Terrestrial Environment

-------
                           INTRODUCTION
     This guideline is one  of a  series  of test  guidelines that have been
developed by the Office of Prevention, Pesticides and Toxic Substances,
United States Environmental  Protection Agency for use  in the testing of
pesticides and toxic substances, and the  development of test data that must
be submitted to the Agency  for review under Federal regulations.

     The Office of Prevention, Pesticides and Toxic Substances (OPPTS)
has  developed this guideline through  a process of harmonization that
blended the testing  guidance  and requirements that  existed in the Office
of Pollution Prevention and  Toxics  (OPPT) and appeared in Title  40,
Chapter I,  Subchapter R of the Code of Federal Regulations  (CFR),  the
Office of Pesticide Programs (OPP) which appeared in publications of the
National Technical  Information Service (NTIS) and the guidelines pub-
lished by the Organization  for Economic Cooperation and Development
(OECD).

     The purpose of harmonizing these  guidelines  into a single set of
OPPTS guidelines is to minimize  variations among the testing procedures
that must be performed to meet the data  requirements of the U. S. Environ-
mental Protection Agency  under  the Toxic  Substances  Control Act  (15
U.S.C. 2601) and the Federal Insecticide, Fungicide and Rodenticide Act
(7U.S.C. I36,etseq.).

     Final  Guideline Release: This guideline  is available from the U.S.
Government Printing Office, Washington, DC 20402 on The Federal Bul-
letin   Board.   By  modem  dial   202-512-1387,  telnet   and   ftp:
fedbbs.access.gpo.gov    (IP     162.140.64.19),    internet:     http://
fedbbs.access.gpo.gov, or call 202-512-0132 for disks  or paper copies.
This guideline is also available electronically in ASCII and PDF (portable
document format) from the EPA Public Access Gopher  (gopher.epa.gov)
under the heading "Environmental Test  Methods and Guidelines."

-------
OPPTS 885.5200   Expression in a terrestrial environment.
     (a) Scope—(1) Applicability. This guideline is intended to meet test-
ing requirements of the  Federal  Insecticide, Fungicide,  and Rodenticide
Act (FIFRA) (7 U.S.C. 136, et seq.}.

     (2) Background. The source material used in developing this har-
monized OPPTS test guideline is OPP guideline 155A-10.

     (b) Test  standards.  This   guideline is to be  used  with OPPTS
885.5000.  (1)  Tests shall be conducted in a greenhouse  environment to
determine whether the microbial pest control abent (MPCA) is able to sur-
vive, persist, and replicate in a terrestrial environment consisting of soil
and vegetation representative of the proposed use site. The following pa-
rameters should be varied to determine their effect on the survival and
growth of the MPCA population:

     (i) Temperature.

     (ii) Humidity.

     (iii) Precipitation (amount, frequency,  pH).

     (iv) Sunlight.

     (v) pH (soil and foliar surfaces).

     (vi) Nutrients (soil, vegetation).

     (2) The values selected for each parameter should be selected to ap-
proximate the conditions expected at the intended use site.

     (3) Laboratory  studies  designed to determine  the microbial  agent's
growth requirements (e.g., temperature, humidity, pH,  sunlight and nutri-
ents)  may  supplement the  greenhouse  study described  in  paragraph
(d)(l)(i) of OPPTS 885.5000. Laboratory studies may demonstrate that the
MPCA will be unable to survive and the Agency will  consider studies
on a case-by-case basis to meet the intent of testing in lieu of the green-
house study.

     (c) Test substance.  A typical end-use product  or the technical grade
of the active ingredient shall be tested.

     (d) Test duration. Data to establish a population  decline curve shall
be collected at intervals until two half-life  determinations have been made
or until data establish that the microbial agent population is  able to main-
tain itself in the terrestrial environment  at or above the level present imme-
diately after test initiation.

     (e) Reporting and evaluation of  data. The reporting and  evaluation
provisions are the same as those set forth in OPPTS 885.5000.

-------
     (f) Tier progression. If results of this study indicate that the MPCA
is able to persist in the terrestrial environment such that susceptible nontar-
get organisms tested in Tier I are likely to be exposed, then the appropriate
testing in Tier III (OPPTS 885.4550, 885.4600, or 885.4850) is required
as specified in 40 CFR  158.740.

     (g) References. The following references contain information for de-
veloping acceptable protocols:

     (1) Anthony, D.W.et al. Field test with Nosema algerae Vavra and
Undeen (Microsporida, Nosematidae) against Anopheles albimanus Wiede-
mann in Panama. Miscellaneous Publications of the Entomological Society
of America 11:17-28 (1978).

     (2) Armstrong, J.L et al. A microcosm method to assess survival of
recombinant bacteria associated  with plants and herbivorus insects. Cur-
rent Microbiology: in press (1987).

     (3) Cunningham, J.C. Persistence of the nuclear polyhedrosis virus
of the eastern hemlock looper on balsam  foliage. Insect Pathology Re-
search Institute,  Sault  Ste. Marie, Ontario, Canada. Bimonthly Research
Notes 26:24-25 (1970).

     (4) Elgee, E. Persistence of a virus of the white-marked tussock moth
on balsam fir foliage. Maritime Forest Research Centre, Fredericton, New
Brunswick, Canada. Bimonthly Research Notes 31:33-34 (1975).

     (5) Grison, P.  et  al. Microbial  pesticides  and environment. Annals
of Zoological Ecology of Animals 8:133-160 (1976).

     (6) Harcourt,  D.J. Persistence of  a granulosis virus of Pieris rapae
in soil. Journal of Invertebrate Pathology 11:142-143 (1968).

     (7)  Hukuhara,  T.  and  H.  Namura.  Distribution  of a  nuclear
polyhedrosis virus of the fall webworm Hyphantria cunea in soil. Journal
of Invertebrate Pathology 19:308-316 (1972).

     (8) Ignoffo, C.M.  et al. Stability of conidia of an  entomopathogenic
fungus Nomuraea rileyi in and on soil.  Environmental Entomology 7:724-
727 (1978).

     (9) Ignoffo, C.M.  et al. Vertical movement of conidia of Nomuraea
rileyi through sand and loam soils. Environmental Entomology 7:270-272
(1977).

     (10) Jaques, R.P. The persistence  of a nuclear polyhedrosis virus in
the habitat of the host insect Trichoplusia  ni 11. Polyhedra in soil. Cana-
dian Entomology 99:820-829 (1967).

-------
     (11)  Jaques,  R.P. Leaching of the nuclear polyhedrosis virus of
Trichoplusia ni from soil. Journal of Invertebrate Pathology  13:256-263
(1969).

     (12)  Jaques,  R.P.  Occurrence and  accumulation  of  viruses of
Trichoplusia ni in  treated field plots. Journal of Invertebrate Pathology
23:140-152(1974).

     (13) Jaques, R.P. Occurrence and accumulation of the granulosis virus
of Pieris  rapae  in  treated field plots. Journal of Invertebrate Pathology
23:351-359(1974).

     (14)  Kerr,  A.   Soil   microbiological  studies  on  Agrobacterium
radiobacter and biological control of crown gall. Soil Science 118:158-
172 (1974).

     (15) Ladd, Jr. T.L. and P.J. McCabe. Persistence of spores of Bacillus
popilliae,  the causal organism of Type A milky disease of Japanese beetle
larvae in New Jersey soils. Journal of Economic Entomology  60:493-495
(1967).

     (16) Liang, L.N. et al. Fate in model ecosystems of microbial species
of potential use in genetic engineering. Applied Environmental Microbi-
ology 44:708-714 (1982).

     (17) Lingg, A.J. and J.J. McMahon.  Survival of lyophilized Bacillus
popilliae in  soil. Applied Microbiology 17:718-720 (1969).

     (18) Milner, R.J.  and Lutton, G.G. Metarrhizium anisoplia:  Survival
of Conidia in the soil. Proceedings of the First International  Colloquium
on Invertebrate Pathology. Queen's  University  at Kingston, Canada, pp.
428-429 (1976).

     (19) Morris, O.N. A method of visualizing and assessing deposits of
aerially sprayed  insect  microbes.  Journal  of Invertebrate  Pathology
22:115-121(1973).

     (20) Narayanan, K.  et al. Preliminary observations  on the persistence
of nuclear polyhedrosis virus of Spodoptera litura F. Madras Agricultural
Journal 64:487-488 (1977).

     (21) Pinnock,  D.E.  et al. Effect of tree species on the coverage and
field persistence of Bacillus  thuringiensis spores. Insect biological control.
Journal of Invertebrate Pathology 25:209-214 (1975).

     (22)  Roone,  R.E.  and  R.A.   Dacust.  Survival  of  the   nuclear
polyhedrosis virus of Heliothis armigera on crops and in soil in Botswana.
Journal of Invertebrate Pathology 27:7-12 (1976).

-------
     (23) Swift, M.J. Microbial succession  during the decay of organic
matter. In: Burns, R.G.; Slater, J.H. (eds.) Experimental microbial ecology.
Oxford: Blackwell, 164-177 (1982).

     (24) Thomas, E.D. et al.  The effect of soil pH of cabbage looper
nuclear polyhedrosis virus  in soil.  Journal of Invertebrate Pathology
21:21-25 (1973).

     (25) Vankova, J. and M.  Svestka, M. The  field  persistence  and effi-
cacy of Bacillus thuringiensis  formulations. Biological  control  of forest
pests. Anz Schadlingskd Pflanzenschutz 49:33-38.  (English Summary)
(1976).

     (26) Walter, M.V. et al. A method to evaluate survival of genetically
engineered microorganisms in soil extracts. Current Microbiology: in press
(1987).

     (27) Walter, M.V.  et  al.  Measuring genetic stability in  bacteria  of
potential use in genetic engineering. Applied Environmental Microbiology
53:105-109(1987).

     (28)  Wojciechlowska,   M.   et   al.   Duration  of   activity   of
entomopathogenic microorganisms introduced into the soil. Polish Eco-
logical Studies 3:141-148 (1977).

     (29) Young, S.Y. Pre- and posttreatment assessment of virus levels.
Pages 139-142 In selected papers from EPA-USDA Working Symposium.
M.  Summers, R. Engler, L. Falcon, and  P. Vail (eds.) American Society
of Microbiology, Washington, DC (1975).

-------