United States      Prevention, Pesticides     EPA712-C-98-351
          Environmental Protection    and Toxic Substances     January 1998
          Agency        (7101)
&EPA    Fate, Transport and
          Transformation Test
          Guidelines
          OPPTS 835.3160
          Biodegradability in Sea
          Water

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                           INTRODUCTION
     This guideline is one  of a  series  of test  guidelines that have been
developed by the Office of Prevention, Pesticides and Toxic Substances,
United States Environmental  Protection Agency for use  in the testing of
pesticides and toxic substances, and the  development of test data that must
be submitted to the Agency  for review under Federal regulations.

     The Office of Prevention, Pesticides and Toxic Substances (OPPTS)
has  developed this guideline through  a process of harmonization that
blended the testing  guidance  and requirements that  existed in the Office
of Pollution Prevention and  Toxics  (OPPT) and appeared in Title  40,
Chapter I,  Subchapter R of the Code of Federal Regulations  (CFR),  the
Office of Pesticide Programs (OPP) which appeared in publications of the
National Technical  Information Service (NTIS) and the guidelines pub-
lished by the Organization  for Economic Cooperation and Development
(OECD).

     The purpose of harmonizing these  guidelines  into a single set of
OPPTS guidelines is to minimize  variations among the testing procedures
that must be performed to meet the data  requirements of the U. S. Environ-
mental Protection Agency  under  the Toxic  Substances  Control Act  (15
U.S.C. 2601) and the Federal Insecticide, Fungicide and Rodenticide Act
(7U.S.C. I36,etseq.).

     Final  Guideline Release: This guideline  is available from the U.S.
Government Printing Office, Washington, DC 20402 on The Federal Bul-
letin   Board.   By  modem  dial   202-512-1387,  telnet   and   ftp:
fedbbs.access.gpo.gov  (IP 162.140.64.19), or  call 202-512-0132 for disks
or paper copies.  This  guideline is also available electronically in ASCII
and PDF (portable document format) from EPA's World Wide Web  site
(http://www.epa.gov/epahome/research.htm) under the heading "Environ-
mental Test Methods and Guidelines."

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OPPTS 835.3160  Biodegradability in sea water.
     (a) Scope—(1) Applicability. This guideline is intended to meet test-
ing  requirements  of  both  the  Federal   Insecticide,  Fungicide,  and
Rodenticide Act (FIFRA) (7 U.S.C. 136, et seq.) and the Toxic Substances
Control Act (TSCA) (15 U.S.C. 2601).

     (2) Background.  The source material  used in developing this har-
monized OPPTS test guideline is OECD guideline  306, Biodegradability
in Sea Water.

     (b) General introduction.  (1) When the original Organization  for
Economic Cooperation and Development (OECD)  Test Guidelines were
developed, it was  not  known to what extent results  from the screening
tests for ready biodegradability using  fresh water and sewage effluent or
activated sludge as inoculum  could be applied to the marine environment.
Variable results on this point have been reported (e.g., see paragraph (e)(l)
of this guideline).

     (2) Many industrial waste waters, containing a variety of chemicals,
reach the sea either by direct discharge or via estuaries and rivers in which
the residence times are low compared with the period necessary for com-
plete  biodegradation of many of the  chemicals present. Because  of the
growing awareness of the  need to protect the marine environment against
increasing loads of chemicals and the need to estimate the probable con-
centration of chemicals in the sea, test methods for biodegradability in
sea water have been developed.

     (3) The methods  described here use natural  sea water  both  as  the
aqueous phase  and as  the source  of microorganisms. In an  endeavor to
conform with the methods for ready biodegradability in fresh water,  the
use of ultrafiltered and centrifuged sea water was investigated, as was the
use of marine sediments as inocula. These investigations were unsuccess-
ful. The test medium therefore is natural sea water pre-treated to remove
coarse particles.

     (4) This guideline consists  of two test methods: the Shake  Flask
Method and the Closed  Bottle Method.  In  order  to assess  ultimate
biodegradability with the  Shake  Flask  Method, relatively high concentra-
tions  of the test substance must be used because  of the poor sensitivity
of the dissolved organic  carbon  (DOC) analytical  method. This  in turn
necessitates the addition to the sea water of mineral nutrients (N and P),
the low concentrations of which would otherwise limit the removal  of
DOC. It is also  necessary to add  the nutrients in the Closed Bottle Method
because of the concentration of the added test substance.

     (5) Hence, the methods are  not tests for ready  biodegradability since
no  inoculum is added  in  addition to the microorganisms already present
in the sea water. Neither do  the tests  simulate the marine environment

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since nutrients are added and the concentration of test substance is very
much higher than would be present in the sea.

     (c) Application. (1) The methods in this guideline are intended for
application when the pattern of use and disposal of the chemical substance
in question indicates a route  to  the sea.  If the result is positive (>70%
DOC removal; >60% ThOD—theoretical oxygen demand), it may be con-
cluded that there is a potential for biodegradation in the marine environ-
ment. However, a negative result  does not preclude such potential but indi-
cates that further study is necessary, for example using as low a concentra-
tion of the test substance as possible.

     (2) In either case, if a more definitive value  for the rate  or degree
of biodegradation in sea water at a particular  site  is required, other more
complex and sophisticated, and hence more costly, methods would have
to be applied.  For  example, a simulation test could be applied using a
concentration  of test substance nearer to  the likely environmental con-
centration,  or non-fortified, non-pretreated sea water taken from the loca-
tion of interest could be used and primary biodegradation followed by  spe-
cific chemical analysis. For ultimate biodegradability, 14C-labeled test  sub-
stance would be necessary to  allow  rates of disappearance of soluble or-
ganic 14C  and production of 14CC>2 to be measured at environmentally
realistic concentrations of that  substance.

     (d) Choice of methods.  The selection of test method depends on a
number of factors. The following table 1 is given  to help the investigator
select a test method. Whereas  chemicals having water solubility  below the
equivalent  of about 5 mg C/L cannot be tested  using the Shake Flask
Method, in principle they may be tested using the Closed  Bottle Method.

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   Table 1.—Advantages and Disadvantages of the Shake Flask and Closed Bottle Methods
       Method
        Advantages
        Disadvantages
Shake flask
simple apparatus except for C
analyzer
60 d duration is not a problem
no interference from
nitrification
can be adapted for volatile chemi-
cals
  needs C analyzer
  uses 5-40 mg DOC/L; could
  be inhibitory
  DOC determination is difficut
  at low concentrations in
  sea water (chloride effect)

  DOC sometimes high in
  sea water
Closed bottle
simple apparatus
simple end determination
uses low concentration of test
compound (2 mg/L), thus less
chance of inhibition
easily adapted for volatile
chemicals
- can be difficut to maintain
  airtightness of bottles
- wall growth of bacteria can
  lead to false values
- blank 02 uptake values can be
  high,  especially after 28 days;
  can be overcome by aging
  the sea water
- possible interference from 02
  uptake by nitrification
          (1) Shake flask method—(i) Introduction. (A) This method is a sea
     water variant of the Modified OECD Screening Test (see paragraph (e)(2)
     of this  guideline).  It was finalized as  a result of a ring test organized for
     the Europeon Economic Community (EEC) by the Danish Water Quality
     Institute (see paragraph (e)(3) of this guideline).

          (B) In common with the accompanying sea water Closed Bottle Meth-
     od, the results  from this test are not to be taken as indicators of ready
     biodegradability, but are to be used specifically for obtaining information
     about the biodegadability of chemicals in marine environments.

          (ii) Principle of  the  method. A predetermined amount of the  test
     substance is dissolved in the test medium  to yield a concentration of 5-
     40 mg  DOC/L. If the  limits of sensitivity  of organic carbon  analyses are
     improved, the use of lower concentrations of test substance may be advan-
     tageous, particularly for inhibitory substances. The solution of the test sub-
     stance  in  the test  medium  is incubated with agitation in the dark or in
     diffuse light under aerobic conditions at a fixed temperature (controlled
     to +  1°C) which will normally be within the  range  15-20°C. In cases
     where the objective  of the  study is to simulate environmental  situations,
     tests  may  be  performed at temperatures  outside this  range. The rec-
     ommended maximum  test duration is 60  days. Degradation is followed
     by DOC measurements (ultimate degradation) and, in some cases, by spe-
     cific analysis (primary  degradation).

          (iii)  Information on  the test substance. (A) To  detemine whether
     the test may be applied to  a particular substance, certain properties of the

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substance must be  known.  The organic carbon content of the substance
must be  established, its volatility must be such that  significant losses do
not occur during  the course of the  test, and its solubility in water should
be greater than the equivalent of 25-40 mg C/L. Further, the test substance
should not significantly sorb to glass surfaces. Information  on the purity
or the relative proportions of major components of the  test substance will
be useful in interpreting the results obtained, especially  when the result
lies close to the "pass" level.

    (B)  Information on the toxicity of the test substance to bacteria, for
example  as measured  in short-term respiration rate  tests (see paragraph
(e)(4) of this guideline), may  be useful  when  selecting  appropriate test
concentrations, and may be essential for the correct interpretation of low
biodegradation values.  However, such information is not always sufficient
for interpreting results obtained in  the biodegradation test, and the proce-
dure described in paragraph (d)(l)(vi)(F)(3) of this guideline  may be more
suitable.

    (iv)  Reference substance. (A) Suitable reference substances  shall be
used to check the microbial activity of the sea water sample. Sodium ben-
zoate, sodium acetate and aniline are examples of chemicals that may be
used for  this purpose.  If reference substances  are not  degraded within  a
reasonably short time, it is recommended that the test be repeated using
another sea water sample.

    (B)  In the EEC ring test (see  paragraph (e)(3) of this guideline), the
lag phase (TL) and time to achieve 50 per cent degradation (tso) excluding
the lag phase were  1 to 4 days and 1 to 7 days, respectively,  for sodium
benzoate. For aniline the TL ranged from  0 to 10 days and the t50 from
1 to 10 days.

    (v) Reproducibility and sensitivity of the method. The reproducibil-
ity of the  method was established in the  ring test (see paragraph (e)(3)
of this guideline). The lowest  concentration of test  substance for which
this method can be used with DOC analysis is largely  determined by the
detection limit in  the organic  carbon analysis  (about 0.5 mg  C/L,  at
present) and the concentration of dissolved organic carbon in the sea water
used (usually on  the order of 3-5  mg/L for water from the  open ocean).
The background concentration  of DOC should not exceed about  20% of
the total  DOC concentration after addition of test substance. If this is not
feasible,  the background concentration of DOC may sometimes be  reduced
by aging the sea water prior to  testing. If the method  is used with specific
chemical analysis only (by which  primary degradation is measured),  the
investigator must document, by supplying  additional information,  whether
ultimate  degradability  can  be expected.  This additional information may
consist   of  the  results   from  other   tests   for   ready  or   inherent
biodegradability.

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     (vi) Description of the method — (A) Apparatus. Normal laboratory
apparatus and:

     (7) Shaking machine accommodating 0.5- to 2-L  Erlenmeyer flasks,
either with automatic temperature control or used in a constant temperature
room at 15-20°C controlled to ±1°C.

     (2) Narrow neck, 0.5- to 2-L Erlenmeyer flasks.

     (3) Membrane filtration apparatus, or centrifuge.

     (4) Membrane filters, 0.2-0.45(j,m.

     (5) Carbon analyzer.

     (6) Equipment for specific analysis (optional).

     (B) Sea water.  (7) Collect a sample of sea  water in a thoroughly
cleaned container and  transport it to  the laboratory, preferably within
1-2  days of collection.  During transport do not allow  the temperature of
the sample to exceed significantly  the temperature to be used in the test.
Identify the sampling location precisely  and describe it in terms  of its
pollutional and nutrient status. Especially  for coastal  waters, include in
this characterization  a heterotrophic microbial colony count and the  deter-
mination of the concentrations of dissolved nitrate, ammonium and phos-
phate.

     (2) Provide the  following information for the sea water sample itself:

     (/) Date of collection.

     (//) Depth of collection.

     (///) Appearance of sample — turbid, etc.

     (/v) Temperature at the time of collection.

     (v) Salinity.

     (v/) DOC.

     (v//) Delay between collection  and use in the test.
        If the DOC content of the sea water sample is found to be high
(see paragraph (d)(l)(v) of this guideline), it is recommended that the sea-
water be aged for approximately  one week prior to use.  This  is accom-
plished by storing the sample under aerobic conditions at the test tempera-
ture and in the dark or in diffuse light. If necessary, maintain aerobic con-
ditions by aerating gently.  During aging, the content of easily degradable
organic material is reduced. In the ring test (see paragraph (e)(3) of this
guideline), no difference was revealed between the degradation potential
of aged and freshly collected sea water samples. Prior to use, pretreat the

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sea water to remove coarse particles, e.g., by  filtering through a nylon
or coarse paper filter (not membrane or GF-C filters),  or by ceatrifuging
gently. The procedure used must be reported. Pretreat after aging, if used.

     (C)  Stock solutions for mineral nutrients. (1) Prepare the following
stock solutions using analytical grade reagents:

     (/) Potassium dihydrogen orthophosphate, KH2PO2. . . 8.50 g

Dipotassium hydrogen orthophosphate, K2HPO4. .. 21.75 g

Disodium hydrogen orthophosphate dihydrate,

     Na2HPO4.2H2O. . . 33.30 g

Ammonium chloride, NfUCl. . . 0.50 g

Dissolve and make up to 1 L with distilled water.

     (//) Calcium chloride, CaCl2. . . 27.50 g

26Dissolve and make up to 1 L with distilled water.

     (Hi) Magnesium sulfate heptahydrate, MgSO4.7H2O . .  . 22.50 g

Dissolve and make up to 1 L with distilled water.

     (iv)  Iron (III) chloride hexahydrate, FeCl3.6H2O. . .  0.25 g

Dissolve and make up to 1 L with distilled water.

     (2) Precipitation in solution (see paragraph (d)(l)(vi)(C)(7) (iv) of this
guideline) may be prevented by adding one drop of concentrated HC1  or
0.4 g ethylenediaminetetraacetic acid (EOTA,  disodium salt) per L. If a
precipitate forms in a stock solution, replace it with freshly made solution.

     (D)  Preparation of test medium. Add  1  mL  of each of the  stock
solutions    described    in    paragraphs   (d)(l)(vi)(C)(7)(/)   through
(d)(l)(vi)(C)(7)(/v) of this guideline per L of pretreated sea water.

     (E) Inoculum. Do not add any inoculum in addition to the microorga-
nisms already present in the sea water. Optionally, determine the number
of colony-forming heterotrophs  in the sea water test medium (and pref-
erably also in the original  sea water  samples)  using  a suitable method,
such as plate  counts with marine agar. This is particularly desirable for
samples from  coastal or polluted sites. Check the heterotrophic microbial
activity in the sea water by using a reference compound.

     (F)  Preparation of flasks. (1) Ensure  that all  glassware is  scru-
pulously clean (e.g., using alcoholic hydrochloric  acid), rinsed and  dried

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before use in order to avoid contamination with residues from previous
tests.  Flasks must also be cleaned before they are used for the  first time.

    (2)  Evaluate test substances in duplicate  flasks simultaneously, to-
gether with a single flask for the reference compound.  Carry out a blank
test, in duplicate, with neither test nor reference substance, for  the  deter-
mination of analytical blanks.  Dissolve the test substance  in the test me-
dium—it may be conveniently added via a concentrated stock solution—
to give the desired starting  concentration of 5-40 mg DOC/L. Test the
reference substance at a starting concentration corresponding  to 20  mg
DOC/L.  If stock  solutions of test and/or reference substances  are used,
ensure that the  salinity  of the  sea water  medium  is not  greatly altered.

    (3) If toxic effects can be expected  or cannot be ruled out, it may
be advisable to  include an inhibition experiment, in duplicate, in the  test
design. Add the test and reference substances to the same vessel, the con-
centration  of the reference compound being normally the  same as in the
control test (i.e., 20 mg DOC/L) in order to allow comparison.

    (4) Dispense adequate amounts of test solutions into the Erlenmeyer
flasks  (up  to  about half the flask volume is  a convenient  amount)  and
subsequently provide each flask with a loose cover (e.g., aluminium foil)
that makes gas  exchange between the  flask and the surrounding air pos-
sible.  (Cotton wool plugs are unsuitable if DOC analysis  is used).  Place
the vessels on the shaker and  shake  continuously at a gentle  rate (e.g.,
100 rpm) throughout the test. Control the temperature (15-20°C and within
±2°C), and shield the vessels from light in order to  avoid growth of algae.
Ensure that the air is free of toxic materials.

    (G) Physical-chemical  control test (optional). If abiotic degradation
or loss mechanisms are suspected, such as  hydrolysis (a  problem with spe-
cific analysis  only), volatilization, or sorption,  it is advisable to perform
a physical-chemical control. This can be done by adding mercury (II) chlo-
ride (HgCL:)1 (50-100 mg/L) to  vessels with  test substance in order to
inhibit microbial activity.  A  significant  decrease  in  DOC or  specific
compound concentration in the physical-chemical control test indicates abi-
otic removal mechanisms. (If mercury chloride is  used, attention  should
be paid to interferences or catalyst poisoning in DOC analysis).

    (H) Number of flasks. In a  typical experiment, the following  flasks
are used:

    (7) Flasks 1 & 2—containing test substance.

    (2) Flasks 3 & 4—containing sea water only (blank).
    iMercury (II) chloride (HgCb) is a very toxic substance that should be handled
with suitable precautions. Aqueous wastes containing this chemical should be disposed
of appropriately; they should not be discharged down the drain.

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     (3) Flask 5—containing reference substance.

     (4) Flask 6—containing test  and reference substance (toxicity con-
trol)—optional.

     (5) Flask 7—containing test substance and sterilizing agent (abiotic
sterile control)—optional.

     (I) DOC analysis. In the course of the test, withdraw samples at suit-
able  intervals   for   DOC   analysis   as   described   in   paragraphs
(d)(l)(vi)(I)(7)(/)  through (d)(l)(vi)(I)(7)(jc) of this guideline. Always take
samples at the start of the test (day 0) and  at day  60.  A minimum of
five samples in total are required to describe the time-course of degrada-
tion. No fixed time schedule for  sampling can be stated since  the  rate
of biodegradation varies.  Carry out the  DOC determination in duplicate
on each sample.

     (7) Determination of organic carbon in sea water. (/) For the deter-
mination of organic carbon of a water sample, the organic compounds in
the sample are oxidized to carbon dioxide using one of the following three
techniques:

     (a) Wet oxidation by persulfate/UV-irradiation.

     (b) Wet oxidation by persulfate/elevated temperature (116-130°C).

     (c) Combustion.

     (//) Evolved  CO2 is then quantified employing infrared spectrometry
or titrimetry. Alternatively, CO2 is reduced to methane, which is quantified
using a flame ionization detector (FID).

     (///) The persulfate/UV-method is commonly used for the analysis of
"clean" water with a low content of particulate  matter. The  persulfate/
elevated temperature  and combustion methods  can  be  applied  to most
kinds of water samples, the former being most suitable for samples with
low levels of nonvolatile organic carbon  (NVOC), and  the combustion
method being applicable to samples with NVOC content well above 1 mg
C/L.

     (2) Interferences.  (/) All three methods depend on eliminating or
compensating for inorganic carbon (1C) present in the sample. Purging of
CO2 from the  acidified sample is the  most  frequently used method to
eliminate  1C, although this also  results in a loss of volatile organic com-
pounds  (see  paragraph (e)(5) of this guideline). Complete elimination or
compensation of  1C must  be ensured for each sample matrix,  and it may
be necessary to determine volatile organic  carbon  (VOC) separately from
NVOC.

                                  8

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     (//) High chloride concentrations  result in  decreased  oxidation effi-
ciency using the persulfate/UV-method (see paragraph (e)(6) of this guide-
line). Application of an oxidation reagent modified by the addition of mer-
cury (II) nitrate may remove this interference. It is recommended that the
maximum tolerable sample volume be used to evaluate each type of chlo-
ride-containing sample. High salt concentrations in samples  analyzed using
the combustion method can cause salt coating of the catalyst and excessive
corrosion of the combustion tube.  Precautions should be taken according
to the manufacturer's manual.

     (///) Highly turbid samples  as well as samples containing particulate
matter may be incompletely oxidized by the persulfate/UV-method.
        An example of a suitable method. (/) Nonvolatile organic carbon
is determined by oxidation with persulfate/UV-irradiation and subsequent
quantification of evolved CO2  employing non-dispersive infrared spec-
trometry.

     (//) The  oxidation reagent is modified in accordance with the sugges-
tions given in paragraph (e)(6) of this guideline, as described in the manu-
facturer's manual:

     (a) 8.2 g of HgCl2 and 9.6  g of Hg(NO3)2.H2O are  dissolved in sev-
eral hundred mL of low-carbon concentration reagent water.
     (b) 20 g of K2S2O8 are dissolved in the mercuric salt solution.

     (c) 5 mL of concentrated HNOs  are added to the mixture.

     (d) The reagent is diluted to 1000 mL.

     (Hi) The interference from chloride is removed using a 40-(iL sample
volume for 10% chloride and 200-(iL sample volume for 1.9% chloride.
Samples with high chloride concentrations and/or larger sample volumes
can be analyzed using this method provided that build-up of chloride in
the oxidation vessel is prevented. Determination of volatile organic carbon
can subsequently be performed, if relevant, for the sample type in question.

     (iv) Automated systems have also been described in the  literature (see
paragraph (e)(7) of this guideline).

     (J) Sampling. (1) The required  volume of the samples depends upon
the analytical method (specific  analysis), the carbon analyzer used, and
the procedure (membrane filtration or centrifugation) selected for sample
treatment before carbon determination (see paragraphs (d)(l)(vi)(J)(3) and
(d)(l)(vi)(J)(/) of this guideline). Before sampling ensure that the test me-
dium is mixed well and that any material adhering to the wall of the flask
is dissolved or suspended.

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     (2) Membrane filter or centrifuge immediately after sampling. If nec-
essary,  store the filtered or centrifuged samples  at 2-4°C for up to 48
hours or below -18°C for longer periods. If it is known that the substance
will remain unaffected by  acidification, acidify to pH  2  before  storing.
        Membrane filters (0.2-0.45 (im) are suitable if it is ensured that
they neither  release carbon nor  sorb  the substance  in the filtration step.
Polycarbonate filters are generally suitable.  Some membrane filters are im-
pregnated with surfactants for hydrophilization and may release consider-
able quantities of dissolved  carbon.  Prepare such  filters by boiling  in
deionized water for three consecutive periods, each of one hour. After boil-
ing,  store the filters in  deionized water. Discard  the first 20  mL of the
filtrate.

     (4) Centrifugation of the samples may  be an acceptable alternative
to membrane filtration. Centrifuge at  40,000  m.s-2 (-4000 g) for 15 min-
utes, preferably in a refrigerated centrifuge.

    Note: The differentiation of Total Organic Carbon (TOC) from DOC  by
centrifugation at very low concentrations does not seem to work,  since either
not all  bacteria are removed, or carbon as part of the  bacterial plasma is redis-
solved. At higher test concentrations  (> 10  mg C/L) the centrifugation error
seems to be comparatively small.
     (K) Frequency of sampling.  (7) If analyses are  performed  imme-
diately after  sampling, determine the  next sampling time  by considering
the result of the analytical determination.

     (2) If samples are  preserved (see  paragraph (d)(l)(vi)(J)(2) of this
guideline) for analysis at a later time,  take more samples than the required
minimum number of five. Analyze the  last samples first,  and by a step-
wise "backwards "selection of appropriate  samples for analysis, it is pos-
sible to obtain a  good description of the biodegradation curve  with a rel-
atively small number  of analytical  determinations. If no  degradation has
taken place by the end of the test, no further samples need to be analyzed,
and in this situation, the  "backwards" strategy may save considerable ana-
lytical costs.

     (3) If a  plateau on the degradation curve is observed before the 60th
day, end the test. If degradation has obviously started by day 60, but has
not reached a plateau, extend the experiment for a further period.
                                  10

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    (vii) Data and reporting—(A) Treatment of results. (7) Record the
results on the data sheet below,  and calculate the biodegradation values
for both test and reference substances from the equation:
1-
                                -C
                               t   H
                                          xlOO
  where:

  Dt = degradation in percentage DOC or specific compound removal at
    time t;

  C0 = starting concentration of DOC or specific compound in the test
    medium;

  Ct = concentration of DOC or specific compound in the test medium
    at time t;
        =  starting concentration of DOC or specific compound in the
    blank;

        = concentration of DOC  or  specific  compound in  the blank at
    time t.
                                11

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 BIODEGRADATION IN SEA WATER—SHAKE FLASK METHOD
                         DATA SHEET
LABORATORY:	
DATE AT START OF TEST:	
TEST SUBSTANCE:	
    Name:	
    Stock solution concentration (mg/L as chemical):	
    Initial concentration in medium, t0 (mg/L as chemical):.
                                      thus (mg DOC/L):
1. SEA WATER:
    Source:	
    Date of collection:	
    Depth of collection:.
    Appearance at time of collection (e.g., turbid, etc.):.
    Salinity at collection (%):	
    Temperature at collection (°C):	
    DOC "x"hours after collection (mg/L):.
    Pretreatment prior to testing (e.g., filtration, sedimentation, aging,
        etc.):	
    Microbial colony count of original sample (colonies/mL):	
    At start of test (colonies/mL):	
    Other characteristics:	
                               12

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2. CARBON DETERMINATIONS:
    Carbon analyzer:

Test: nutrient-fortified
sea water with test
substance
Blank: nutrient-for-
tified as waterout
test substance
Flask no.
1
2
1
2

ai
a2
mean, Ca(t)
bi
b2
mean, Cb(t>
Cl
C2
mean, CC(t)
di
d2
mean, Cd(t)
mean (Cc(t) + Cdm)/2
DOC after n days (mg/L)
0













ni













n2













n3













nx













                            13

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3. EVAULATION OF RAW DATA
Flask No.
1
2
Mean (*)
Calculation of results
D j Qt) ~ Cbl(D x 10Q
"i A r - r
'-o uW(o)
D j CKO ~ cww „ 100
^2 ' p _ ,-,
uo cW(o)
„ _ D, + D2
' 2
% Degradation after n days
0
0
0
0
n,



n,



n.



n,



    * Dj and D2 should not be averaged if there is a considerable difference.

    Note:  Similar formats may be used when degradation is followed by specific analysis and for the
         reference compound and toxicity controls.
4. ABIOTIC DEGRADATION (optional)

DOC cone. (mg/L) in sterile control
Time (days)
0
Cs(0)
t
C,(.)
                                      C  —C
                    % abiotic degradation = -^	S^L x 100
                                         Cs(o}
     (2) State degradation as the percentage DOC  removal (ultimate deg-
radation)  or  specific compound removal (primary degradation) at time t.
Calculate the DOC  concentrations to the nearest 0.1 mg/L, and round up
the means of the Dt  values to the nearest  whole per cent.

     (3) Illustrate  the time course of degradation graphically in a diagram
as shown in the figure in paragraph (d)(l)(vii)(C)(3)  of this guideline. If
there are  sufficient  data, calculate from  the curve the lag phase (TL)  and
the time to reach  50 per cent removal from the end of the lag phase (tso).

     (B) Test report. (7) The test report shall contain the following  infor-
mation:

     (/) Test substance, (a) Physical nature and, where relevant, physical/
chemical properties.

     (b) Identification of the substance.

                                   14

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     (//) Test conditions, (a) Location and description of the sampling site;
pollutional and nutrient status (colony counts, and nitrate, ammonium and
phosphate levels if appropriate).

     (b) Characteristics of the sample (date of sampling, depth, appearance,
temperature,  salinity, DOC (optional), delay between  collection  and use
in the test.

     (c) Method used (if any) for aging the sea water.

     (d) Method used for pretreatment (filtration/sedimentation) of the sea
water.

     (e) Method used for DOC determination.

     (f) Method used for specific analysis (optional).

     (g) Method used for determining the number of heterotrophs in the
sea water (plate count method or alternative procedure) (optional).

     (h) Other methods  (optional) used to characterise the sea water (ATP
measurements,  etc.).

     (Hi) Results,  (a) Analytical data  reported on a  data sheet as shown
in paragraph (d)(l)(vii)(A)(7) of this guideline.

     (b) The  course of degradation represented graphically  in  a  diagram
showing the lag phase (TL), slope,  and time  (starting from the end of the
lag phase)  to reach 50% removal (tso). The lag phase may be estimated
graphically as  shown in the figure in paragraph (d)(l)(vii)(C)(3) of this
guideline or conveniently taken as  the time needed for 10% degradation.

     (c) Percentage degradation measured  after 60 days, or at end of test.

     (iv) Discussion of results.

     (C) Validity  and interpretation  of results. (1) The results obtained
with the reference compounds (e.g., sodium benzoate, sodium acetate or
aniline) should be  comparable to results obtained in the ring test (see para-
graphs (d)(l)(iv) and (e)(3) of this  guideline. If results obtained with ref-
erence compounds are atypical, the test should be repeated using another
sea water  sample.  Although results  of inhibition tests may not always be
straightforward to interpret because of the  contribution  of DOC by the
test material, a significant reduction of the total DOC removal  rate, com-
pared with that of the control, is a positive sign of toxic effects.

     (2) Owing to  the relatively high test concentrations used as compared
with most natural  systems and consequently  an unfavorable ratio between
the concentrations  of test substances and other carbon sources, the method
is to be regarded as a preliminary test that can be used to indicate  whether
or not a substance is easily biodegradable  in sea water. Accordingly a

                                  15

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low result does not necessarily mean that the test substance is  not bio-
degradable in marine environments, but indicates that more work will be
necessary to establish this.

    (3) An  example of graphical presentation of data and estimation of
the values of TL (length of lag phase) and t50 (time, starting at tL, needed
to reach 50% removal) is given in the following Figure 1.

  FIGURE i.—TYPICAL BIODEGRADATION CURVE FOR THE SHAKE FLASK
                              METHOD
        100-
         20-
     (2) Closed bottle method—(i) Introduction. (A) This method is a
sea water variant of the Closed Bottle Test (see paragraph (e)(8) of this
guideline) and was finalized as a result of a ring test organised for the
EEC by the Danish Water Quality Institute (see paragraph (e)(3) of this
guideline).

     (B) In common with the accompanying sea water Shake Flask Meth-
od, results  of  this test  are not to  be  taken as  indications  of ready
biodegradability, but are to be used specifically for obtaining information
about the biodegradability of chemicals in marine environments.

     (ii) Principle of the method.  (A) A  predetermined amount  of the
test substance is dissolved in the test medium at a typical concentration
of 2 to 10 mg  of test substance/L. One or more concentrations may be

                                 16

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used. The solution is kept in a filled closed bottle in the dark in a constant
temperature bath  or enclosure controlled to ±1°C within  the range  15-
20°C. In those cases where the objective of the study is to simulate envi-
ronmental situations, tests  may be performed at temperatures outside  this
range provided that the temperature is properly controlled.  Degradation
is followed by oxygen analyses over a 28-day period.

    (B)  The ring test showed that if the test  was extended beyond 28
days no useful information could be gathered, in most cases, due to severe
interferences. The blank biological oxygen  demand (BOD)  values were
excessively high probably due to wall growth, caused by lack of agitation,
and to nitrification. Thus, the recommended duration is 28  days, but if
the blank BOD value remains within the 30  per cent limit  (see paragraph
(d)(2)(vi)(B)(/) of this guideline) the test could be prolonged.

    (iii) Information on  the test  substance.  (A) To determine whether
the test may be applied to a particular substance, certain properties of the
substance must be known.  The empirical formula is required so that the
theoretical oxygen  demand  (ThOD) may  be calculated as described in
paragraphs (d)(2)(iii)(A)(7)(/) through  (vf) of this guideline; otherwise the
chemical oxygen  demand (COD) of the substance must be determined to
serve  as  a surrogate for the ThOD. The use of COD is less  satisfactory
since some chemicals are not fully oxidized in the COD test.

    (7) Calulation of the theoretical oxygen demand.  (/)  The ThOD
of the  substance CcHhClciNnNanaOoPpSs of molecular weight MW is  cal-
culated as follows:
                   16 2c + —(h- cl- 3n) + 3s+-p + -na - 0
                                             2    2
                                     MW
    (if) This calculation implies that C is mineralized to CO2, H to H2O,
P to P2Os and Na to Na2O. Halogen is eliminated as hydrogen halide
and nitrogen as ammonia.
                                 17

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Example:
              Glucose C6H1206, MW = 180
                     16(2 x 6 + - x 12 - 6)

              ThOD =	= 1.07 mg O2 / mg glucose
                            180
     (///) Molecular weights of salts other than those of the alkali metals

are calculated on the assumption that the salts have been hydrolyzed.


     (iv) Sulfur is assumed to be oxidized to the state of +6.


Example:



       Sodium dodecylbenzenesulfonate C18H29S03Na, MW = 348



                             29     1
                       16(36 + — + 3 + --3)
                ThOD= - - - - - = 2.34 mg 0? / mg substance
                               348                  2

     (v)  In the case of nitrogen-containing  substances the nitrogen may

be  eliminated as ammonia,  nitrite,  or nitrate, and each corresponds to a

different theoretical oxygen demand.


     For nitrite,
                  , .L     1 „    ^   o    3     5     1       1
                  16  2c + — (h-cl) + 3s + — n + — p +   na - o
                           2               222
      ThODNO? =— ^ - - - - - - - - - ^
               2                        MW

     For nitrate,
                   16

       ThODxjn  =
 2c +   (h-cl) + 3s + —n + —p +   na - o
L     2              222.
                                  18

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     (v/) Suppose full nitrate formation had been observed by analysis in
the case of a secondary amine: (C 12^.25)2 NH, MW = 353. Then
                   16(48 + 51/2 + 5/2)
        ThODNO  =	= 3.44 mg O2 /mg substance
               3          353
     (B) The solubility of the substance should be at least 2 mg/L, though
in principle less soluble compounds could be tested (e.g., after sonication)
as could volatile compounds. Information on the purity or the relative pro-
portions of major components of the test substance will be useful in inter-
preting the  results obtained, especially when the result lies close to the
"pass" level.

     (C) Information on the toxicity of the substance to bacteria, for exam-
ple as measured in  short-term respiration tests (see paragraph (e)(4)  of
this guideline), may be useful when selecting  appropriate  test concentra-
tions, and may be essential for the correct interpretation of low biodegrada-
tion values. However, such  information is not  always sufficient for inter-
preting results obtained in the biodegradation test, and the procedure de-
scribed in paragraph (d)(2)(vi)(F)(7) of this guideline may be  more suit-
able.

     (iv)  Reference substance. (A) Suitable reference substances shall be
used to check the microbial activity of the sea water sample. Aniline, so-
dium acetate and sodium benzoate are examples  of  substances that may
be used for this purpose. Degradation of these compounds must reach at
least 60 per cent with  respect to their ThOD within a reasonably short
time span; otherwise it  is recommended that  the test  be  repeated using
another sea water sample.

     (B) In the EEC ring test (see paragraph (e)(3) of this guideline), the
lag phase  (TL) and the time to achieve 50 per cent degradation (tso) exclud-
ing the lag phase were 0 to 2 days  and  1 to 4 days, respectively, for
sodium benzoate. For aniline the TL and tso values were 0 to 7 and 2 to
12 days, respectively.

     (v) Reproducibility. The reproducibility  of the method was estab-
lished in the EEC ring test (see paragraph (e)(3) of this guideline).

     (vi)  Description of the method—(A) Apparatus.  Normal laboratory
equipment and:

     (7) 250- to 300-mL BOD bottles  with glass stoppers or narrow neck
250-mL bottles with glass stoppers.

     (2) Several 2-, 3- and 4-L bottles for the preparation of the experiment
and for the filling of BOD bottles.

                                  19

-------
     (3) Waterbath or constant temperature room for keeping the bottles
at constant temperature (+ 1°C)) with the exclusion of light.

     (4) Equipment for analysis of dissolved oxygen.

     (5) Membrane filters, 0.2 to 0.45 (im (optional).

     (6) Equipment for specific analysis (optional).

     (B) Sea Water. (7) Collect a sea water sample  and provide informa-
tion  on the sampling  location and sample  as described in paragraphs
(d)(l)(vi)(B)(7) and (d)(l)(vi)(B)(2) of this guideline.

     (2) If the  DOC content of the sample is found to be high or if it
is thought that the blank BOD after 28 days  may be more than 30% of
that  of the reference substance, it is recommended that the sea water be
aged for about a week  prior to use. Follow the procedures  described in
paragraph (d)(l)(vi)(B)(3) of this guideline.

     (C) Stock solutions for mineral  nutrients. Prepare stock solutions
as described in paragraph (d)(l)(vi)(C) of this guideline.

     (D) Preparation of test  medium. Add  1  mL  of  each  of the stock
solutions  described in paragraph  (d)(l)(iv)(C) of this guideline per L of
sea water. Saturate the  test medium with air at the test temperature  by
aerating with clean compressed air for approximately 20  minutes. Deter-
mine the concentration of dissolved oxygen for control purposes. The satu-
rated concentration of dissolved oxygen as a function of salinity and tem-
perature may be read from the nomogram in the following Figure 2.
                                 20

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FIGURE 2.—NOMOGRAM OF SATURATION CONCENTRATION OF OXYGEN
           vs. TEMPERATURE AT VARIOUS SALINITIES
                          21

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     (E) Inoculum. See paragraph (d)(l)(vi)(E) of this guideline.

     (F) Preparation of test bottles. (7) Perform all necessary manipula-
tions including aging and pretreatment of the sea water at the chosen test
temperature in the range of 15 to 20°C, ensuring cleanliness but not steril-
ity of all glassware.

     (2) Prepare groups of BOD bottles for the  determination of the BOD
of the test and reference substances in simultaneous experimental series.
Perform all analyses on duplicate bottles (i.e., duplicate blanks, duplicate
reference  substance bottles and duplicate test substance bottles). A mini-
mum of five  sampling times in total are  required to describe the time
course of degradation.  For oxygen analyses,  five sampling times require
a total of3x2x5 = 30 bottles (blank, reference substance and test sub-
stance), and thus about 10 L of test medium.

     (3) Prepare separate solutions of test and reference substances in large
bottles of sufficient volume (see paragraph (d)(2)(vi)(A)(2) of this guide-
line) by  first  adding test  and  reference substances either  directly  or by
adding a concentrated stock solution to the  partly filled large  bottles. Add
further test medium to give the final desired concentrations. If stock solu-
tions of test and/or reference substances are used, ensure that the salinity
of the sea water medium is not significantly  altered.

     (4) Select concentrations of test  and reference substances by taking
into  account:

     (/) The solubility of dissolved oxygen in sea water at the prevailing
test temperature and salinity (see the nomogram in paragraph (d)(2)(vi)(D)
of this guideline).

     (//) The blank BOD of the  sea water.

     (///) The expected biodegradability of the test substance.

     (5) At 15°C and 20°C and 32 parts per thousand salinity  (i.e.,  that
of ocean water),  the solubility of dissolved oxygen  is about 8.1 and 7.4
mg/1, respectively. The  oxygen consumption of the  seawater  itself (blank
respiration) may be 2 mg O2/L  or more if the sea water is not aged. There-
fore, in order  to ensure that a significant amount of oxygen remains after
oxidation of  the  test  substance,  use  a  starting  concentration of  test
compound of approximately 2 to 3 mg/L (depending  on  the ThOD) for
the compounds that are  expected to be completely degraded under the con-
ditions of the test, such as reference  substances. Use  higher initial con-
centrations for less degradable  substances, up to about  10 mg/L, provided
that toxic effects  do not occur.  It can be advantageous to run parallel tests
with low (about 2 mg/L) and high (about 10 mg/L) concentrations of test
substance.

                                  22

-------
     (6} An oxygen blank must be determined at each time point using
parallel bottles containing neither test nor reference substance.

     (7) If inhibitory effects are to be determined, prepare the following
series of solutions in separate large bottles (see paragraph (d)(2)(vi)(A)(2)
of this guideline):

     (/) 2 mg/L of an easily degradable compound, e.g., any of the ref-
erence substances listed in paragraph (d)(2)(iv)(A) of this guideline.

     (//) x mg/L of test substance (x is usually 2).

     (///) 2 mg/L of the easily degradable compound plus  x mg/L  of test
substance.

     (G) Physical-chemical control test (optional). If the option of using
specific analyses is used, a physical-chemical  control test  may be per-
formed to  determine whether the test  substance  is removed by  abiotic
mechanisms, such as hydrolysis or sorption.  A  physical-chemical control
test may be performed by adding mercury (II) chloride (HgCL:)2  (50  to
100 mg/L) to duplicate flasks with test substance in order to inhibit  micro-
bial  activity. A significant decrease in specific compound concentration
in the course of the test indicates abiotic  removal.

     (H) Number of BOD bottles. In a typical experiment the following
bottles are used:

     (1) At least 10 containing test substance.

     (2) At least 10 containing nutrient-fortified sea water only.

     (3) At least 10 containing reference substance.

     (4) 6  bottles  containing  test and reference substances (toxicity con-
trol)—optional.

     (vii) Procedure. (A) After preparation, immediately siphon each solu-
tion from the lower quarter (not from the bottom) of the appropriate large
bottle, to fill the respective  group  of BOD bottles.  Immediately analyze
the  time zero  samples for dissolved oxygen (see paragraph (d)(2)(vii)(D)
of this guideline), or preserve them for later chemical analysis by precipita-
tion with MnCl2 (manganese (II) chloride) and NaOH (sodium hydroxide).

     (B) Incubate the remaining BOD bottles at the test temperature (15-
20°C) in the dark, and remove bottles for analysis at appropriate time inter-
vals. No fixed time schedule can be stated since the rate of biodegradation
    2Mercury (II) chloride (HgCb) is a very toxic substance that should be handled
with suitable precautions. Aqueous wastes containing this chemical should be disposed
of appropriately; they should not be discharged down the drain.

                                  23

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varies. Analyze for dissolved oxygen (see paragraph (d)(2)(vii)(D) of this
guideline).

     (C) Membrane filter (0.2-0.45 (im) or centrifuge for 15 minutes sam-
ples for specific analyses (optional). Store these samples for up to 48 hours
at 2-4°C, or for longer periods at -18°C, if they are not analyzed imme-
diately. If it is known that the test substance will remain unaffected by
acidification, acidify to pH 2 before storing.

     (D) Determine the concentration of dissolved oxygen using a chemi-
cal or electrochemical method that is recognized nationally or internation-
ally.

     (viii) Data  and reporting—(A) Treatment of results. (7) Record
analytical results on the data sheet below.
                                 24

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     BIODEGRADATION IN SEA WATER—CLOSED BOTTLE
                           METHOD

                         DATA SHEET
LABORATORY:	

DATE AT START OF TEST:	

TEST SUBSTANCE:	

    Name:	

    Stock solution concentration (mg/L):	
    Initial concentration in medium, t0 (mg/L):

    ThOD or COD (mg Ch/mg test substance):

1. SEA WATER:

    Source:	

    Date of collection:	
    Depth of collection:.
    Appearance at time of collection (e.g., turbid, etc.):_

    Salinity at collection (%):	

    Temperature at collection (°C):	
    DOC "x"hours after collection (mg/L):.
    Pretreatment prior to testing (e.g., filtration, sedimentation, aging,

        etc.):	

    Microbial colony count of original sample (colonies/mL):	

    At start of test (colonies/mL):	

    Other characteristics:	

    Test medium:

    Temperature after aeration (°C):	
                               25

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              concentration after aeration and standing before start of test

               (mg 02/L):	
2. DO DETERMINATION:

     Method: Winkler/electrode

Test: nutrient-for-
tified sea water
with test sub-
stance
Blank: nutrient-for-
tified sea water
but without test
substance
Flask no.
1
2
Mean blank
1
2
Mean test

3i
a2
rrit = (ai + a2)/2
Ci
C2
mb = (Ci + c2)/2
mg O2 after n days
0






HI






n2






n3






n4






    Note: Similar format may be used for reference compound and toxicity controls.

3. DO DEPLETION: % DEGRADATION (%D):

(mb - mt)o>
%D = (mb - mt)«) x 100/test substance (mg/l) x ThOD
DO depletion after n days
ni


n2


n3


n4


       This assumes that nib(0) = mt(0), where
       o) = blank value at day 0,
    mt(o) = test substance value at day 0.
    If mb(o) does not equal mt(0), use (mt(0) - mt(x)) - (mb(0) - mb(X)), where
    mb(x) = blank value at day x,
    mt(x) = test substance value at day x.
    (2) Calculate the BOD as the difference in the oxygen depletion between
the blank and a solution of test substance under the conditions of the test. Divide
the net oxygen depletion by the concentration (w/v) of the substance in order
to express the BOD as mg BOD/mg test substance. Extent of degradation is
defined as the ratio of the BOD to either the ThOD or the COD,  but preferably
the former, and is expressed as a percentage (see paragraph (d)(2)(viii)(A)(3)
of this guideline).
                                        26

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     (3) Calculate the extent of biodegradation for each sampling time, for both
test and reference substances, using one of the following equations:
                                     mg O, / mg tested substance
                     % biodegradation =	x 100
                                   mg ThOD / mg tested substance
                                    mg O, / mg tested substance
                     % biodegradation =	x 100
                                   mg COD / mg tested substance
         where:

         ThOD =  theoretical  oxygen demand (for method of calculation,  see
           paragraph (d)(2)(iii)(A) of this guideline).

         COD = chemical oxygen demand, determined experimentally.

           NOTE: Sometimes the two methods of calculation (percentage of the ThOD
       and percentage of the COD)  do not give the same results.  It is preferable to
       use ThOD because some chemicals are not fully oxidized in the COD test.
           (4) Illustrate the time course of degradation graphically by means of
       a diagram;  see  the example in paragraph  (d)(2)(viii)(C)(<5)  of this guide-
       line. If there are  sufficient data,  calculate  the lag phase  (TL) and the time
       from the end of the  lag phase that is required  to reach  50% degradation
       (tso).

           (5) If specific analysis is used (optional),  express  the  percentage of
       primary degradation as the percentage of specific substance removed with-
       in the test period, corrected for analytical blanks.

           (B) Test report. (7) The test report must contain the following infor-
       mation:

           (/) Test substance:

           (a) Physical nature and, where relevant, physical/chemical properties.

           (b) Identification of the substance.

           (//) Test conditions:

           (a) Location and description of the  sampling site: pollutional and  nu-
       trient status  (colony counts, and  nitrate, ammonium  and phosphate levels
       if appropriate).

                                         27

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     (b) Characteristics of the sample: date of sampling, depth, appearance,
temperature,  salinity, DOC (optional),  delay between collection and use
in the test.

     (c) Method used (if any) for aging the sea water.

     (d) Method used for pretreatment (filtration/sedimentation) of the sea
water.

     (e) Method used for COD determination (if performed).

     (f) Method used for oxygen measurements.

     (g) Method of dispersion for substances that are poorly soluble under
the test conditions.

     (h) Method used for determining the number of heterotrophs in the
sea water (plate count method or alternative procedure).

     (/) Method used for determining DOC in sea water (optional).

     (/) Method used for specific analysis (optional).

     (k) Other  optional  methods used to characterize the sea water (ATP
measurements,  etc.).

     (Hi) Results:

     (a)  Analytical data  reported  on  a  data  sheet  (see  paragraph
(d)(2)(viii)(A)(7) of this guideline).

     (b) The  time  course of degradation represented graphically in a dia-
gram showing  the lag phase, (ti_),  slope, and time starting from the  end
of the lag phase to reach 50% of the final oxygen uptake caused by oxida-
tion  of the test compound (tso). The lag phase may be  estimated graphi-
cally as shown  in Figure 3 in paragraph (d)(2)(viii)(C)(<5) of this guideline,
or conveniently in taken as the time needed for 10% degradation.

     (c) % degradation measured after 28 days.

     (iv) Discussion of results.

     (C) Validity and interpretation of results. (1) The blank respiration
should not exceed 30% of the oxygen in the test bottle. If it is not possible
to meet this criterion using  freshly  collected sea water, the seawater must
be aged (stabilized) before use.

     (2) The possibility that nitrogen-containing compounds may affect the
results should be considered.

     (3) Results obtained  with the  reference substances sodium benzoate
and  aniline should  be comparable to the results obtained  in the ring test

                                  28

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(see paragraphs (d)(2)(iv)(B) and (e)(3) of this guideline). If results  ob-
tained with reference compounds are atypical, the test should be repeated
using another sea water sample.

     (4) The test substance can be considered to be inhibitory to bacteria
at the concentration used if the BOD of the mixture of reference and  test
substances is less than the sum of the BOD of the separate  solutions of
the  two substances.

     (5) Owing to the relatively high test concentrations as compared with
most natural systems,  and consequently an unfavorable ratio between the
concentrations  of test  substance and other carbon sources, the method is
regarded as a preliminary test that can be used to indicate whether or not
a substance is  easily biodegradable in sea water. Accordingly, a low result
does not necessarily mean that the test substance is not biodegradable in
marine  environments,  but indicates  that more work will be necessary to
establish this.

     (6) An example of graphical presentation of data and estimation of
the  values of TL (length of lag phase) and tso (time, starting at TL, needed
to reach 50%  of the final oxygen uptake caused by oxidation of the  test
substance), is given in the following Figure 3:
                                  29

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 FIGURE 3.—TYPICAL BIODEGRADATON CURVE FOR THE CLOSED BOTTLE
                             METHOD
       100
        80
        60
        40
        20
            oxygen consumption
            % of final
                                                      days
            "tog phase"
    (e) References. The following references should be consulted for ad-
ditional background material on this test guideline.

    (1) de Kreuk J.F. and Hanstveit A.O. 1981. Determination of the
biodegradability of the organic fraction of chemical wastes. Chemosphere
10 (6): 561-573.

    (2) OECD, Paris. 1992. Test Guideline 301 E.

    (3) Nyholm N. and Kristensen P. 1987. Screening Test Methods for
Assessment of Biodegradability of Chemical Substances in Seawater. Final
Report of the ring test programme 1984-1985, March 1987, Commission
of the European Communities.

    (4) OECD, Paris. 1984. Test Guideline 209.

    (5) International Standards Organization (ISO). 1986. Water quality—
determination of total organic  carbon. Draft International  Standard ISO/
DIS 8245, January 16.

    (6) American Public Health Association. 1985. Standard Methods for
the Examination of Water and Wastewater, 16th ed.
                                30

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    (7) Schreurs W. 1978. An automated colorimetric method for the de-
termination of dissolved organic carbon in seawater by UV destruction.
Hydrobiological Bulletin 12: 137-142.

    (8) OECD, Paris. 1992. Test Guideline 301 D.
                                31

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