United States Environmental Protection Agency	Office of Research and Development

National Exposure Research Laboratory
Research Abstract

Government Performance Results Act Goal: Clean and Safe Water

Significant Research Findings:

Detecting Emerging and CCL-Related Pathogens

Scientific Problem and This research is designed to provide improved methods for detecting
Policy Issues	an(j enumerating emerging and Contaminant Candidate List (CCL^-

related microbial pathogens that are potentially transmitted through
drinking water. The CCL was developed in response to the 1996
amendments to the Safe Drinking Water Act (SDWA), which required
that EPA periodically identify new drinking water contaminants for
potential regulation. Emerging pathogens are contaminants that are
likely to appear on future CCL lists. The methods developed under
this project area are needed to obtain occurrence and exposure data
that are required for adequate characterization of potential public
health risks. They can also be used to evaluate the effectiveness of -
various drinking water treatment options or for routine compliance
monitoring. The development of policies, guidance and regulations
that ensure the availability of safe, pathogen-free drinking water are
thus dependent on the data obtained by these improved methods.

Research Approach The objective for this project area is to develop and evaluate practical,

sensitive, and economical technologies for the detection and
measurement of specific emerging and CCL-related microbes.

Meeting this objective required the development of multiple
approaches, including: 1) new cell lines and cell culture methods to
identify pathogenic human enteric viruses, protozoa, and bacteria; 2) -
genomic and immuno-based methods for rapid detection of pathogenic
microbes, especially for those organisms which cannot be cultured or
which are difficult to culture; 3) scanning and transmission electron
microscopy to monitor and identify microbial contaminants and to
confirm cell culture and immunoassay results; 4) fluorescence based
confocal microscopy techniques to image and identify the ultrastruc-
ture of living pathogenic agents; and 5) source water quality indicator
systems that accurately predict the presence and source of human fecal
contamination in environmental waters.

Results and	This Annual Performance Measure (APM 45) supports FY01 Annual

Implications	Performance Goal 007 which states: Reduce Uncertainties and

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Improve Methods Associated with the Assessment and Control of
Risks Posed by Exposure to Microbial Contaminants in Drinking
Water with a Focus on the Emerging Pathogens on the CCL."

Significant findings are as follows:

A rapid molecular method has been developed to detect and identify
caliciviruses in environmental and drinking water. Recently, this
method was used to detect and identify a genogroup 2 calicivirus in
well water following a waterborne outbreak. Caliciviruses have been
the responsible etiological agent in many waterborne disease outbreaks
and probably account for the majority of outbreaks where the
etiological agents have not been identified. The Centers for Disease
Control estimate that caliciviruses are responsible for greater than 90
percent of all non-bacterial gastroenteritis in the U.S. and worldwide.

A rapid molecular method was developed that will identify all known
hepatitis E viral strains in drinking and source waters. The hepatitis E
virus is a waterborne emerging pathogen that has caused many
outbreaks of illness in the developing world, including Mexico. Thus
far, an outbreak of this virus has not been reported in the U.S.,
however, several sporadic cases have occurred, and there is concern
that it will be an emerging infectious disease in this country. An
important bearing on this disease is the finding that a variant of the
virus is common in hogs in the U.S. It is thought that hogs may serve
as a reservoir for human infection, inasmuch as hepatitis E viral
isolates from two of the U.S. human cases were more closely related to
the U.S. hog hepatitis E virus than to human strains from other parts of
the world.

Three new chapters describing virus detection technology have been
appended to the U.S. EPA Manual of Methods for Virology. Chapter
14 describes the most widely used viral method for recovering human
enteric viruses from water matrices. The method takes advantage of
positively charged filters to concentrate viruses from water. Chapter 15
describes a quantal method for assaying culturable waterborne human
enteric viruses. The assay differs from the plaque assay in that it is
based on a direct microscopic viewing of cells for virus-induced
cytopathic effects and has a greater sensitivity. The methods described
in Chapters 14 and 15 primarily detect infectious enteroviruses and
reoviruses. The enterovirus group includes the coxsackieviruses and
echoviruses, two groups that are included on the CCL. Chapter 16
describes procedures for the detection of coliphage in water matrices.
Two quantitative procedures and one qualitative presence-absence

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procedure are presented. The procedures described have an important
consideration as a general indicator of water quality. Currently, hard
copies of these chapters are being distributed. In the near future
electronic text of these publications will be available in the EPA
Microbiology Home page (www.epa.gov/microbes/).

A method has been developed by which a microsporidia species can be
identified from processed water samples. The method has been
published and is the first report of a flourescent in situ hybridization
assay that utilizes a species-specific fluorescent-labeled oligonuclotide
probe. Microsporidia are obligate intracellular protozoan parasites that
are on the CCL.

Research collaborations are ongoing with several governmental
organizations on basic work to develop research tools and to provide
organisms for several of the emerging parasites. These collaborators
include the World Health Organization, the Centers for Disease
Control, the Department of Agriculture (Interagency Agreement #
DW12938843), the U.S. Geological Survey (Interagency Agreements #
DW14938174 and DW14939295), the University of Cincinnati
(Assistance Agreement # CR826758) and the Department of Defense
(Interagency Agreement #DW97937961). Other collaborative projects
are with Kansas State University which has undertaken the
development of a cell culture model for parasitic protozoan
propagation, and with the Veteran's Administration Palo Alto Health
Care System (Interagency Agreement # DW36938176) and the Johns
Hopkins School of Public Health which are developing molecular
testing methods for emerging and CCL list parasites. Innovative
methods of isolating etiological agents of emerging diseases from large
volume water samples are being investigated under a collaborative
project with the Marshfield Medical Research and Education
Foundation. A examination of calicivirus strains in latrine and well
water samples is being conducted in Kenya, Africa in conjunction with
the University of Surrey in the United Kingdom (Assistance
Agreement # R82860301).

Examples of recent publications from this study include:

Bennett, J.W., Gauci, M.R., LeMoenic, S., Schaefer, III, F.W., and Lindquist,
H.D.A. "A comparison of enumeration techniques for Cryptosporidium par\>um
oocysts." J. Parasitol. 85:1165-1168, 1999.

Fout, G.S., Dahling D.R, and Saffennan, R.S. "Collecting and processing of water-
borne viruses by positive charged filtrationand organic flocculation." In USEPA
Manual of Methods for Virology, EPA-600/4-84-013 (N14), U. S. Enviromnental

National Exposure Research Laboratory — October 2001

Research
Collaboration and
Publications


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Protection Agency, Cincinnati, OH. (April 2001).

Fout, G.S., Dahling D.R, and Safferman, R.S. " Total culturable virus quantal assay."
InUSEPAManual of Methods for Virology, EPA-600/4-84-013 (N15), U.
S. Environmental Protection Agency, Cincinnati, OH (April 2001).

Grimm, A.C. and Fout, G.S. "Development of a molecular method to identify
hepatitis E virus in environmental water." J. Virol. Meth. (submitted).

Hester, J.D., Lindquist, H.D.A., Bobst, A.M., and Schaefer III, F.W. "Fluorescent in
situ detection of Encephalitozoon hellem spores with a 6-
carboxyfluorescein-labeled ribosomal RNA-targeted oligonucleotide
probe." J. Eukaryot. Microbiol. 47:299-308, 2000.

Lindquist H.D.A., Bennett, J.W., Ware, M., Stetler, R.E., Gauci, M., and Schaefer
III, F.W. "Testing methods for detection of Cryptosporidium spp. in water
samples." Supplement to the Southeast Asian Journal of Tropical Medicine
and Public Health (In press).

Lindquist, H.D.A., Ware, M., Stetler, R.E., Wymer, L., and Schaefer, III, F.W. "A
comparison of four fluorescent antibody based methods for purifying,
detecting and confirming Cryptosporidium parvum in surface waters." J.
Parasitol. (In press).

Pandian, A, Berg, G., Dahling, D.R., Cashdollar, J.C., Wymer, L., and Fout G. S.
and L. Wymer. "Comparison of a new beef extract preparation with beef
extract V for eluting and reconcentrating poliovirus 1 adsorbed on 1MDS
filters." J. Virol. Meth. (Cleared for publication).

Simmons III, O.D., Sobsey, M.D., Heaney, C.D., Schaefer III, F.W., and Francy,
D.S. 2001. "Concentration and detection of Cryptosporidium oocysts in
surface water samples by method 1622 using ultrafiltration and capsule
filtration."^/?/?/. Environ. Microbiol. 67:1123-1127, 2001.

Schaefer, III, F.W. 2001. "Detection of protozoans and helminths in source and

finished drinking water." InManual of Environmental Microbiology, 2nd ed.
(Cleared for publication).

Williams, F.P., Stetler, R.E., and Safferman, R.S. "Procedures for detecting coli-
phages." InUSEPAManual of Methods for Virology, EPA-600/4-84-013
(N16), U. S. Environmental Protection Agency, Cincinnati, OH (June
2001).

Future Research	Future research will continue to focus on reducing deficiencies in the

current technology needed to process and detect emerging and CCL-
related pathogens. These studies are of importance to the Office of
Water, Agency risk assessors, the scientific community and industry
spokes groups as they will enhance the ability to measure the
occurrence and exposure of the public to waters contaminated with
pathogenic agents and thereby reduce the public health uncertainties.

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This research will include the introduction of new technologies, such
as combining cell culture and molecular methods in order to identify
and characterize the listed pathogenic agents. At the same time,
studies will be continuing on the development of faster and simpler
indicator methods for evaluating microbial water quality using
innovative cultural, immunological, genomic and biochemical
approaches. In addition, future research will also be supporting the
development of procedures that will differentiate viable from non-
viable and infectious from non-infectious microbial pathogens. Other
methods will be developed that will aid in identifying the sources of
microbial pathogens such as human versus animal contamination.

Questions and inquiries can be directed to:

G. Shay Fout, Ph.D. or Robert S. Safferman, Ph.D.

US EPA, Office of Research and Development
National Exposure Research Laboratory
Cincinnati, OH 45268-1320

Phone 513/569-7387 or 513/569-7334

E-mail: fout.shav@epa.gov or safferman.robert@epa.gov

Contacts for

Additional

Information

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