$EPA NRMRL

\ NATIONAL RISK MANAGEMENT RESEARCH LABORATORY
www.epa.gov/nrmrl GROUND WATER AND ECOSYSTEMS RESTORATION RESEARCH

Quantifying and Modeling the Risk of Disturbance to
Ecosystems Caused by Invasive Species

Introduction to the Problem

Invasive species are biological pollutants that threaten ecosystem health.
Identifying the mechanisms of invasion and developing predictive models
of invasion will be critical to developing risk management strategies for
limiting the economic and environmental damage caused by invaders.

Background

Eastern redcedar (Juniperus virginianct), a prolific evergreen tree and its
associated vertebrate seed dispersers, will be used as a model invader-
disperser system. Limiting factors of invasion (growth and seed dispersal;
see Figure 1) will be studied through manipulative field experiments
conducted at Oklahoma State University's (OS Us) Cross Timbers
Experimental Range Research Site. Researchers from OSU will
collaborate to complete research tasks.

Objectives

•	Investigate the mechanisms of invasion by Juniperus virginianct

•	Quantify the susceptibility of ecosystems to invasion

•	Develop predictive models of invasion probability based on
empirical data

Approach

The effects of local plant diversity and soil characteristics on
J. virginianct survival and growth (which are measures of invasion ability)
will be examined in a field experiment. J. virginianct seedlings will be
planted in three distinct ecosystem types: old field, deciduous forest, juniper
forest (see Figure 2). Seedling growth and survival will be analyzed in
relation to adjacent plant species and soil type and chemistry.

J. virginianct dispersal limits will be studied by monitoring seed consumers
(see Figure 3) and their rates of consumption at J. virginianct trees, fenced
to exclude small mammals and deer. A greenhouse experiment will be
conducted to determine the effects of both nitrate nitrogen and ammonium
nitrogen on seedling growth. Models of ecosystem susceptibility to
J. virginianct invasion will be developed using empirical data collected in
this study and projecting onto existing geographical data bases on
vegetation type, soil type, animal distributions, and nitrogen deposition
in the United Sates.

The Circle of Invasion

Juniper
Invasion

Seedling
Growth

Seed
Dispersal

Figure 1: The Circle of Invasion

Figure 2: Plant diversity and soil effects on growth

Figure 3: Seed Dispersers: Cedar Waxwing

The National Risk Management Research Laboratory's mission is to advance scientific and engineering
solutions that enable EPA and others to effectively manage current and future environmental risks.
NRMRL possesses unique strengths and capabilities and is dedicated to providing credible
technological information and scientific solutions that support national priorities
and protect human health and the environment.


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Accomplishments to Date (February 2003)

•	Completed all field work on plant diversity, soil characteristics, and seed consumption

•	Completed lab analysis of soils

•	Presented papers and posters at three national meetings

•	Greenhouse study is under way

Near-Future Tasks

•	Continue greenhouse experiment on nitrogen effects on growth

•	Complete reports for field and lab work portions of study

•	Develop predictive models of invasion

Goals

•	Arrive at a better understanding of the mechanisms of ecosystem invisibility

•	Develop unique field techniques

•	Identify vulnerable stages of invader life history

•	Develop predictive models of invisibility applicable to other species and useful to land managers
responsible for addressing invasive species problems

Investigators	Collaborators

Paul Mayer	Oklahoma State University

Ground Water and Ecosystem Restoration Division	U.S. Geological Survey

Ada, Oklahoma 74820
580-436-8647

The National Risk Management Research Laboratory's mission is to advance scientific and engineering
solutions that enable EPA and others to effectively manage current and future environmental risks.
NRMRL possesses unique strengths and capabilities and is dedicated to providing credible
technological information and scientific solutions that support national priorities
and protect human health and the environment.


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