United States Environmental Protection Agency	Office of Research and Development

National Exposure Research Laboratory
Research Abstract

Government Performance Results Act Goal: Clean and Safe Water

Significant Research Findings:

Occurrence of CCL-Related Pathogens
in Source and Drinking Waters

Scientific Problem The Safe Drinking Water Act requires EPA to establish a list of
and Policy Issues	unregulated microbiological and chemical contaminants to aid in

priority setting for the Agency's drinking water regulation development
program. The list, referred to as the Contaminant Candidate List (CCL),
identifies chemical and microbiological contaminants the EPA has
selected as potential regulatory candidates from the universe of
contaminants which may possibly pose a significant public health risk to
consumers if present in drinking water. There are three bacterial
species listed on the CCL (Aeromonas, Mycobacterium avium complex
and Helicobacter pylori) that was published in the Federal Register on
March 2, 1998. More information is needed on the occurrence of these
species in drinking water in the U.S. Currently, suitable methods for
obtaining occurrence data are available only for Aeromonas and
Mycobacterium avium complex (MAC) organisms.

MAC organisms are members of the nontuberculous mycobacteria
(NTM) group in the genus Mycobacterium. Although it was generally
thought that Mycobacterium tuberculosis was the only clinically
significant species, changes in attitudes toward species of mycobacteria
other than M. tuberculosis came from reports in the 1950's and 1960's
that acid-fast bacilli had been cultured from pathological material under
circumstances that led microbiologists to believe that some may be
clinically significant. Searches for mycobacteria in the environment
were renewed. Among the many sources of what became known as
NTM water was not at first identified as important. Recently, however,
there has been increasing evidence that water may be one of the vehicles
by which these organisms infect or colonize the human body. In recent
years NTM have emerged as a major cause of opportunist infection in
individuals with impaired immunity such as AIDS patients, the elderly,
or those undergoing immunosuppressive drugs or chemotherapy.

Studies have shown that NTM can survive, grow, and colonize drinking
water supply systems. Without evidence of person-to-person

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transmission, there is a strong possibility that humans may be infected
from environmental sources.

Because NTM disease in immunocompromised hosts is primarily
disseminated to many organs, questions have been raised concerning the
portal of entry of these mycobacteria. Before the discovery of AIDS and
in the absence of evidence of person-to-person spread of NTM,
pulmonary infection was thought to be due to aerosol inhalation.
However, in AIDS patients and other immunocompromised hosts,
infection can occur via the gastrointestinal tract, lungs, or both. The
environment is considered a likely source. Thus, a wide range of
sources, exposures, and modes of transmission need to be investigated.
Having data available showing the occurrence of these important
emerging opportunistic pathogens greatly aids in the assessment of risks
to susceptible hosts from exposure to drinking water and other
environmental samples that may contain these organisms.

Aeromonas are Gram negative, facultatively anaerobic, rod-shaped
bacteria in the family Aeromonadaceae that can be found in all surface
water environments. Many of the recognized species or biotypes have
been implicated as causing human disease, although only three, A
hydrophila, A. caviae and A. veronii/sobria are considered to be major
human pathogens. Primary effects are gastrointestinal disease resulting
in diarrhea. However, these bacteria can also cause serious wound
infections and bacteremia. Although they are generally susceptible to
chlorine disinfection, the presence of Aeromonas in source waters
indicates that these bacteria could be present in municipal drinking
water. Past published studies have confirmed this possibility, reporting
the isolation of Aeromonas bacteria in drinking water from distribution
systems. It is also possible that these bacteria could evade disinfection
and reproduce as a part of the biofilm material inside distribution
system pipes.

Research Approach To aid the Agency in assessing the risks associated with contact with

drinking water, the objectives of this research are as follows:

•	To determine the potential for human exposure by measuring the
occurrence of MAC and other NTM organisms in drinking water
supply system samples and other environmental samples.

•	To determine the potential for human exposure by measuring the
occurrence of Aeromonas bacteria in drinking water samples.

Mycobacterium: A total of 139 samples were collected from five

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different types of sources. Sources included public drinking water
utilities, cisterns, bottled waters, drinking water treatment samples, and
ice samples. Drinking water samples were collected from
geographically dispersed sites (21 states). Samples were collected from
cold taps, warm taps, or showers. The temperatures of the samples were
recorded, and total and free chlorine analyses were performed. The age
of the sample site (residence or hospital), the type of drinking water
treatment, and the source water information was provided with the
samples. Noncarbonated bottled water samples were collected from
multiple bottlers. Ice samples were collected in sterile sample bottles
and allowed to melt at room temperature before analyses. Samples were
analyzed for NTM by a membrane filtration method using 7H10 agar
after treatment of the sample with cetylpyridinium chloride to reduce
background organism levels. The plates were incubated for eight weeks
at 37° C and the NTM isolates were identified by a polymerase chain
reaction (PCR) -mediated sequence analysis of the gene encoding the
16S rRNA. Heterotrophic plate counts were performed by the R2A
membrane filtration method.

Aeromonas: Utilities were chosen to participate in the survey with
regard to known characterisitics of their drinking water distribution
systems. Characteristics of particular interest were low free chlorine
residuals in the distribution systems, the use of surface waters as the
source water, long residence times for water in the system, high
assimilable organic carbon values and the use of chloramine or chlorine
dioxide disinfection. Usually 4-6 samples were collected by utility
personnel and shipped cold overnight for filtration and Aeromonas
analysis using ampicillin-dextrin agar (ADA). In addition, turbidity
levels, the presence of total coliforms, E. coli and heterotrophic plate
counts were also determined. Water temperature was measured at the
tap by the utility. Presumptive Aeromonas colonies on ADA were
confirmed by testing for cytochrome c oxidase and the ability to ferment
trehalose. Confirmed isolates were archived for future analyses,
including identification to the species level and presence of virulence
factors.

Results and

Implications

Mycobacterium: Drinking water samples were collected from 42
drinking water supply systems from 21 states. NTM were isolated from
16 (38%) of these systems. In all there were eight different NTM
species recovered from surface water drinking water supplies. The most
frequently occurring isolate wasM mucogenicum followed by M
intracellulare, and M. gastri/kansasii. MAC organisms were found in
9% of the drinking water supplies examined. NTM or MAC organisms
were not found predominately in hot water versus cold water.

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Heterotrophic bacterial levels, chlorine concentration, or the age of the
sampling site were not found to correlate with the occurrence of NTM.

A number of outbreaks of nosocomial (i.e. hospital-acquired) disease
caused by rapidly growing NTM have been reported. In this study M
mucogenicum was isolated from 41% of the NTM-positive samples.
The most frequent disease associated with M. mucogenicum are
catheter sepsis and posttraumatic skin infections. M. mucogenicum was
the most frequently occurring NTM in this study and was isolated from
all geographic areas. This indicates that the organism may be ubiquitous
in drinking water and may pose a potential health risk.

In summary, NTM were isolated from 38% of the drinking water
supplies examined from a wide geographic area and 33% of all
samples. Not only were MAC organisms isolated from 19% of the NTM
- positive samples, but other clinically significant mycobacterial
opportunist pathogens, such asM kansasii andM mucogenicum, were
also present. These organisms were isolated from well-operated, well-
maintained drinking water utilities with heterotrophic plate count (HPC)
levels of 500 colony forming units (CFU) /ml or less and chlorine
residuals of greater than 0.5 mg/L. This study, along with other studies,
indicates that exposure to drinking water containing NTM could pose a
health risk to immunocompromised hosts and, to a lesser degree, to
immunocompetent individuals.

Aeromonas: To date 16 utilities have participated by providing water
samples for analysis, totaling over 900 samples. Of these, 1.8% have
contained Aeromonas bacteria, ranging in density from 1 colony forming
unit (CFU)/500ml to >300 CFU/500ml. The occurrence of these
bacteria were not randomly distributed among the samples. One utility
accounted for half of the positive samples, and one tap at this utility
accounted for 30% of the total overall positive samples.

Research
Collaboration and
Publications

This research supplies critical information to the EPA Office of Water
and will be used in determining whether the MAC group of organisms
and Aeromonas bacteria, included on the current Contaminant Candidate
List, should be regulated as contaminants in drinking water. This
Annual Performance Measure (APM 46) supports FY01 Annual
Performance Goal 007, "Reduce uncertainties and improve methods to
assess and control risks of microbes, especially CCL."

The NTM study was conducted by the EPA's National Exposure
Research Laboratory. Publications and presentations from this research
include:

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Covert, T. C., Rodgers, M. R., and Reyes, A. L. "Nontuberculous mycobacteria."
Presented at the Annual Water Quality and Measurement Conference, Dallas,
Texas. September 10, 1996.

Covert, T. C., Rodgers, M. R., Reyes, A. L. and Stelma, G. N. "Occurrence of
nontuberculous mycobacteria in environmental samples." Presented at the
Annual Meeting of the American Society for Microbiology, Miami Beach,
Florida. May 4-8, 1997.

Kralovic, S. M., Cannon, C. G., Covert, T. C., Pennington, J. L., Cushion, M. T.,
Hagland, L. A., Perlada, D. A., Reyes, A. L., Rodgers, M. R, and Rhodes J. C.
"Increased isolation of Mycobacterium fortuitum from patients associated with
a common nursing unit." Presented at the Annual Meeting of the American
Society for Microbiology, Miami Beach, Florida. May 4-8, 1997.

Covert, T. C. "Mycobacterium avium - intracellulare." Presented at the AWWA
Water Quality Technology Conference. Emerging Waterborne Pathogens
Seminar (ST8). San Diego, California. November 4, 1998.

Covert, T. C., Rodgers, M. R., Reyes, A. L. and Stelma, G. N. "Occurrence of
nontuberculous mycobacteria in environmental samples." Appl. Environ.
Microbiol. 65:2492-2496, 1999.

The Aeromonas occurrence project is being conducted as a collaboration between
the National Exposure Research Laboratory (ORD) and the Technical Service
Center (OGWDW). Publications and presentations from this research include:

Rodgers, M., Shadix, L. and Feige, M. "Occurrence ofAeromonas bacteria in
potable waters." Presented at the Water Quality Technology Conference, Salt
Lake City, Utah. November 5-9, 2000.

Rodgers, M., Shadix, L.and Feige ,M. "The use of Ampicillin-dextrin agar for the
recovery of Aeromonas bacteria from drinking water. " (In preparation).

Future Research	Scientists at the EPA's National Exposure Research Laboratory are now

in the process of developing improved cultural and rapid molecular
methods for detection of NTM in drinking water.

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Questions and inquires concerning the NTM study can be directed to:
Terry C. Covert

US EPA, Office of Research and Development
National Exposure Research Laboratory
Cincinnati, Ohio 45268

Phone: 513/569 - 7318
E-mail: covert.terry@epa.gov

Questions and inquires concerning the Aeromonas study can be directed
to:

Mark Rodgers

US EPA, Office of Research and Development
National Exposure Research Laboratory
Cincinnati, Ohio 45268

Phone: 513/569 -7225
E-mail: rodgers.mark@epa.gov

Contacts for

Additional

Information

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